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<document id="C951FDABD70C7B27DFFB39BB7FB94EF5" ID-DOI="10.11646/zootaxa.4028.4.10" ID-GBIF-Dataset="6b873979-bec1-4751-906d-569e43d2d7fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="241732" ID-ZooBank="6A395098-2EC3-41B5-9E4D-8BEAA8C2FDC3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461288304186" checkinUser="plazi" docAuthor="Ortega-Salas, Héctor &amp; González-Soriano, Enrique" docDate="2015" docId="038A48709D256A50FF7BFF3AFF73FA36" docLanguage="en" docName="zt04028p594.pdf" docOrigin="Zootaxa 4028 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Libellula coahuiltecana Ortega-Salas &amp; González-Soriano, 2015, sp. nov." docType="treatment" docVersion="7" lastPageNumber="593" masterDocId="FFB330089D246A54FFECFFADFFAAFFB9" masterDocTitle="A new specIes of Libellula LInnaeus, 1758, from the Cuatro CIénegas basIn, CoahuIla, MéxIco (AnIsoptera: LIbellulIdae)" masterLastPageNumber="594" masterPageNumber="589" pageNumber="590" updateTime="1698615638711" updateUser="plazi">
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<mods:title id="7A3B04913035BC4F8F5F457D7B0A1D04">A new specIes of Libellula LInnaeus, 1758, from the Cuatro CIénegas basIn, CoahuIla, MéxIco (AnIsoptera: LIbellulIdae)</mods:title>
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<mods:name id="1D30535BC44512070D909A473A916E98" type="personal">
<mods:role id="B9879BC2FDCF14EF7F52EEA896BCD1EA">
<mods:roleTerm id="8720CFD005BA3C53CCC2AE551B2CC05C">Author</mods:roleTerm>
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<mods:namePart id="ADE9E94CA614C00ED0CD813545E6E02C">Ortega-Salas, Héctor</mods:namePart>
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<mods:roleTerm id="1FEEBAB8E6A29A49F4366D435E2C36E4">Author</mods:roleTerm>
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<mods:namePart id="08579690464A1EE7C9AD55E25342963A">González-Soriano, Enrique</mods:namePart>
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<mods:typeOfResource id="9DDAB0A43CB7829B0F9F88278B03323F">text</mods:typeOfResource>
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<mods:title id="A6ED2B620550D0F46DF2CEB2DD9C0A06">Zootaxa</mods:title>
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<mods:part id="E58784147A4DE99ABF0BDA3C1918F868">
<mods:date id="57CDAF24FE51A93E689901E8F68FA4E4">2015</mods:date>
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<treatment id="038A48709D256A50FF7BFF3AFF73FA36" ID-DOI="http://doi.org/10.5281/zenodo.6112961" ID-GBIF-Taxon="119641785" ID-Zenodo-Dep="6112961" LSID="urn:lsid:plazi:treatment:038A48709D256A50FF7BFF3AFF73FA36" httpUri="http://treatment.plazi.org/id/038A48709D256A50FF7BFF3AFF73FA36" lastPageId="4" lastPageNumber="593" pageId="1" pageNumber="590">
<subSubSection id="C339AAED9D256A55FF7BFF3AFE8AFF68" pageId="1" pageNumber="590" type="nomenclature">
<paragraph id="8B9CF9669D256A55FF7BFF3AFC61FF0B" blockId="1.[151,971,151,209]" box="[151,971,151,178]" pageId="1" pageNumber="590">
<heading id="D0D44E0A9D256A55FF7BFF3AFC61FF0B" bold="true" box="[151,971,151,178]" fontSize="11" level="1" pageId="1" pageNumber="590" reason="1">
<emphasis id="B95725749D256A55FF7BFF3AFC61FF0B" bold="true" box="[151,971,151,178]" pageId="1" pageNumber="590">
<taxonomicName id="4C2382E59D256A55FF7BFF3AFE1CFF08" ID-CoL="9X38P" box="[151,438,151,177]" class="Insecta" family="Libellulidae" genus="Libellula" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="590" phylum="Arthropoda" rank="species" species="coahuiltecana" status="sp. nov.">
<emphasis id="B95725749D256A55FF7BFF3AFE1CFF08" bold="true" box="[151,438,151,177]" italics="true" pageId="1" pageNumber="590">Libellula coahuiltecana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264980F9D256A55FE51FF35FDB1FF0B" box="[445,539,152,178]" pageId="1" pageNumber="590" rank="species">sp. nov.</taxonomicNameLabel>
Ortega-Salas &amp; González-Soriano
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9CF9669D256A55FF7BFF17FE8AFF68" blockId="1.[151,971,151,209]" box="[151,288,186,209]" pageId="1" pageNumber="590">
(
<figureCitation id="1318E5E39D256A55FF72FF17FF48FF68" box="[158,226,186,209]" captionStart="FIGURE 1" captionStartId="1.[151,244,1872,1894]" captionTargetBox="[151,1436,991,1848]" captionTargetId="figure@1.[151,1436,991,1848]" captionTargetPageId="1" captionText="FIGURE 1. Libellula coahuiltecana sp. nov. Male from Cuatro Ciénegas de Carranza, Coahuila, Mexico." httpUri="https://zenodo.org/record/241733/files/figure.png" pageId="1" pageNumber="590">Figs 1</figureCitation>
,
<figureCitation id="1318E5E39D256A55FF03FF17FF56FF68" box="[239,252,186,209]" captionStart="FIGURE 2" captionStartId="2.[151,244,1951,1973]" captionTargetBox="[229,1305,209,1922]" captionTargetId="figure@2.[178,1409,191,1930]" captionTargetPageId="2" captionText="FIGURE 2. Libellula coahuiltecana. Male paratype. Left wings; FIGURE 3. Libellula coahuiltecana. Male paratype. Secondary genitalia; FIGURE 4. Libellula coahuiltecana. Male paratype. Vesica spermalis; FIGURE 5. Libellula coahuiltecana. Male paratype. Cercus; FIGURE 6. Libellula coahuiltecana. Female paratype. Vulvar lamina" httpUri="https://zenodo.org/record/241734/files/figure.png" pageId="1" pageNumber="590">2</figureCitation>
6)
</paragraph>
</subSubSection>
<subSubSection id="C339AAED9D256A55FF7BFEADFAC6FE80" pageId="1" pageNumber="590" type="etymology">
<paragraph id="8B9CF9669D256A55FF7BFEADFAC6FE80" blockId="1.[151,1436,256,937]" pageId="1" pageNumber="590">
<emphasis id="B95725749D256A55FF7BFEADFEBDFEAE" bold="true" box="[151,279,256,279]" pageId="1" pageNumber="590">Etymology.</emphasis>
The name is to be considered a noun in apposition and refers to the Coahuiltecans, an indigenous group who inhabited the territory of the present day States of Coahuila and Nuevo León in
<collectingCountry id="F334B9F69D256A55FC04FE8FFB90FE80" box="[1000,1082,290,313]" name="Mexico" pageId="1" pageNumber="590">México</collectingCountry>
and southern Texas in
<collectingCountry id="F334B9F69D256A55FAC3FE8FFAC2FE80" box="[1327,1384,290,313]" name="United States of America" pageId="1" pageNumber="590">USA</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C339AAED9D256A50FF2BFEE8FD83FBA8" lastPageId="4" lastPageNumber="593" pageId="1" pageNumber="590" type="materials_examined">
<paragraph id="8B9CF9669D256A55FF2BFEE8FDF0FDC8" blockId="1.[151,1436,256,937]" pageId="1" pageNumber="590">
<emphasis id="B95725749D256A55FF2BFEE8FEC7FEE5" bold="true" box="[199,365,325,348]" pageId="1" pageNumber="590">
<typeStatus id="549847C49D256A55FF2BFEE8FF55FEE5" box="[199,255,325,348]" pageId="1" pageNumber="590">Type</typeStatus>
material.
</emphasis>
Total:
<specimenCount id="9D2532EF9D256A55FE55FEE8FE71FEE5" box="[441,475,325,348]" pageId="1" pageNumber="590" type="generic">9♂</specimenCount>
1♀.
<typeStatus id="549847C49D256A55FDFDFEE8FDDEFEE5" box="[529,628,325,348]" pageId="1" pageNumber="590" type="holotype">Holotype</typeStatus>
♂:
<collectingCountry id="F334B9F69D256A55FD72FEE8FD58FEE5" box="[670,754,325,348]" name="Mexico" pageId="1" pageNumber="590">México</collectingCountry>
, Coahuila State, Cuatro Ciénegas de Carranza, Rancho Orozco, pond Tío Candido (
<geoCoordinate id="EE179FA19D256A55FE9BFEC5FD84FEC6" box="[375,558,360,383]" direction="north" orientation="latitude" pageId="1" pageNumber="590" precision="1" value="26.869705">26°5210.94 N</geoCoordinate>
,
<geoCoordinate id="EE179FA19D256A55FDD2FEC5FD52FEC6" box="[574,760,360,383]" direction="west" orientation="longitude" pageId="1" pageNumber="590" precision="1" value="-102.078255">102°441.72 W</geoCoordinate>
,
<quantity id="4CDB54839D256A55FCEBFEC5FCF9FEC6" box="[775,851,360,383]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="1" pageNumber="590" unit="m" value="720.0">720 m</quantity>
),
<date id="FF9DDFA69D256A55FC87FEC5FC5FFEC6" box="[875,1013,360,383]" pageId="1" pageNumber="590" value="2013-07-05">5 July 2013</date>
, leg. H. Ortega-Salas;
<typeStatus id="549847C49D256A55FB16FEC5FACEFEC6" box="[1274,1380,360,383]" pageId="1" pageNumber="590" type="paratype">paratypes</typeStatus>
: 1♀,
<collectingCountry id="F334B9F69D256A55FF7BFE27FF41FE18" box="[151,235,394,417]" name="Mexico" pageId="1" pageNumber="590">México</collectingCountry>
, Coahuila State, Cuatro Ciénegas de Carranza. Rancho PRONATURA, pond Quintero (
<geoCoordinate id="EE179FA19D256A55FB32FE27FA32FE18" box="[1246,1432,394,417]" direction="north" orientation="latitude" pageId="1" pageNumber="590" precision="1" value="26.80566">26°4820.38 N</geoCoordinate>
,
<geoCoordinate id="EE179FA19D256A55FF7BFE00FEE8FE7D" box="[151,322,429,452]" direction="west" orientation="longitude" pageId="1" pageNumber="590" precision="1" value="-102.017006">102°11.22 W</geoCoordinate>
,
<quantity id="4CDB54839D256A55FEBDFE00FE33FE7D" box="[337,409,429,452]" metricMagnitude="2" metricUnit="m" metricValue="7.18" pageId="1" pageNumber="590" unit="m" value="718.0">718 m</quantity>
)
<date id="FF9DDFA69D256A55FE40FE00FDE6FE7D" box="[428,588,429,452]" pageId="1" pageNumber="590" value="2013-04-26">26 April 2013</date>
, leg. M. Trujano-Ortega;
<specimenCount id="9D2532EF9D256A55FC81FE00FC3AFE7D" box="[877,912,429,452]" pageId="1" pageNumber="590" type="generic">2♂</specimenCount>
, same data as
<typeStatus id="549847C49D256A55FBD5FE00FB3CFE7D" box="[1081,1174,429,452]" pageId="1" pageNumber="590" type="holotype">holotype</typeStatus>
(H. Ortega-Salas);
<specimenCount id="9D2532EF9D256A55FA9FFE00FA3DFE7D" box="[1395,1431,429,452]" pageId="1" pageNumber="590" type="generic">3♂</specimenCount>
,
<collectingCountry id="F334B9F69D256A55FF7BFE7DFF41FE5E" box="[151,235,464,487]" name="Mexico" pageId="1" pageNumber="590">México</collectingCountry>
, Coahuila State, Cuatro Ciénegas de Carranza. Rancho PRONATURA, mesquite forest (
<geoCoordinate id="EE179FA19D256A55FB33FE7DFA32FE5E" box="[1247,1432,464,487]" direction="north" orientation="latitude" pageId="1" pageNumber="590" precision="1" value="26.80963">26°4834.67 N</geoCoordinate>
,
<geoCoordinate id="EE179FA19D256A55FF7BFE5FFEE7FDB0" box="[151,333,498,521]" direction="west" orientation="longitude" pageId="1" pageNumber="590" precision="1" value="-102.01951">102°110.21 W</geoCoordinate>
,
<quantity id="4CDB54839D256A55FEBBFE5FFE34FDB0" box="[343,414,498,521]" metricMagnitude="2" metricUnit="m" metricValue="7.27" pageId="1" pageNumber="590" unit="m" value="727.0">727 m</quantity>
),
<date id="FF9DDFA69D256A55FE5EFE5FFD9BFDB0" box="[434,561,498,521]" pageId="1" pageNumber="590" value="2013-07-08">8 July 2013</date>
, leg. H. Ortega-Salas;
<specimenCount id="9D2532EF9D256A55FCC8FE5FFCE2FDB0" box="[804,840,498,521]" pageId="1" pageNumber="590" type="generic">2♂</specimenCount>
,
<collectingCountry id="F334B9F69D256A55FCBFFE5FFC0DFDB0" box="[851,935,498,521]" name="Mexico" pageId="1" pageNumber="590">México</collectingCountry>
, Coahuila State, Cuatro Ciénegas de Carranza. Rancho PRONATURA, Poza Azul 1 (26°4938.4, 102°0127.8),
<date id="FF9DDFA69D256A55FC91FDB8FBAAFD95" box="[893,1024,533,556]" pageId="1" pageNumber="590" value="2013-07-08">8 July 2013</date>
, leg. H. Ortega-Salas;
<specimenCount id="9D2532EF9D256A55FB16FDB8FA8CFD95" box="[1274,1318,533,556]" pageId="1" pageNumber="590" type="generic">1 ♂</specimenCount>
,,
<collectingCountry id="F334B9F69D256A55FAAFFDB8FA3DFD95" box="[1347,1431,533,556]" name="Mexico" pageId="1" pageNumber="590">Mexico</collectingCountry>
, Coahuila state, Cuatro Ciénegas de Carranza. Rancho Orozco, pond Tío Candido (
<geoCoordinate id="EE179FA19D256A55FBF3FD95FB79FDF6" box="[1055,1235,568,591]" direction="north" orientation="latitude" pageId="1" pageNumber="590" precision="1" value="26.891033">26°5327.72 N</geoCoordinate>
,
<geoCoordinate id="EE179FA19D256A55FB0DFD95FA32FDF6" box="[1249,1432,568,591]" direction="west" orientation="longitude" pageId="1" pageNumber="590" precision="1" value="-102.08634">102°510.82 W</geoCoordinate>
,
<quantity id="4CDB54839D256A55FF7BFDF7FF77FDC8" box="[151,221,602,625]" metricMagnitude="2" metricUnit="m" metricValue="7.22" pageId="1" pageNumber="590" unit="m" value="722.0">722 m</quantity>
),
<date id="FF9DDFA69D256A55FF1CFDF7FEC5FDC8" box="[240,367,602,625]" pageId="1" pageNumber="590" value="2013-07-09">9 July 2013</date>
, leg. H. Ortega-Salas.
</paragraph>
<paragraph id="8B9CF9669D256A55FF2BFDD0FEB6FD0E" blockId="1.[151,1436,256,937]" pageId="1" pageNumber="590">
<typeStatus id="549847C49D256A55FF2BFDD0FF56FD2D" box="[199,252,637,660]" pageId="1" pageNumber="590">Type</typeStatus>
specimens are deposited in the Colección Nacional de Insectos (
<collectionCode id="ED3261A39D256A55FC03FDD0FB9FFD2D" box="[1007,1077,637,660]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="1" pageNumber="590">CNIN</collectionCode>
), Instituto de Biología,
<collectionCode id="ED3261A39D256A55FAA6FDD0FA31FD2D" box="[1354,1435,637,660]" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="1" pageNumber="590">UNAM</collectionCode>
(
<collectionCode id="ED3261A39D256A55FF72FD0DFEBAFD0E" box="[158,272,672,695]" httpUri="http://grbio.org/cool/9wfd-7x2v" name="Instituto de BiIología, Universidad Nacional Autónoma de México" pageId="1" pageNumber="590">IBUNAM</collectionCode>
).
</paragraph>
<paragraph id="8B9CF9669D256A55FF2BFD6FFF74FCA6" blockId="1.[151,1436,256,937]" pageId="1" pageNumber="590">
<emphasis id="B95725749D256A55FF2BFD6FFE13FD60" bold="true" box="[199,441,706,729]" pageId="1" pageNumber="590">
<typeStatus id="549847C49D256A55FF2BFD6FFE85FD60" box="[199,303,706,729]" pageId="1" pageNumber="590" type="holotype">Holotype</typeStatus>
male. Head
</emphasis>
. Labium cream; labrum reddish, pale in basal 0.2; anteclypeus reddish brown; postclypeus and frons red; vertex red, black basally; antenna black; occipital triangle brown; eyes (in life) reddish above, blue-gray below.
</paragraph>
<paragraph id="8B9CF9669D256A55FF2BFC87FDC0FC10" blockId="1.[151,1436,256,937]" pageId="1" pageNumber="590">
<emphasis id="B95725749D256A55FF2BFC87FEBEFCF8" box="[199,276,810,833]" italics="true" pageId="1" pageNumber="590">Thorax</emphasis>
. Prothorax: anterior and middle lobes pale reddish, basally black; hind lobe pale. Pterothorax: mesepisternum red, mesepimeron reddish brown, remainder of thorax pale brown laterally, humeral and metapleural sutures washed with dark brown at upper 0.25. Legs pale brown; dashes of brown at distal end of femora, tibiae darker especially at inner surfaces, armature black.
</paragraph>
<caption id="DF5CA9EE9D256A55FF7BF8FDFB38F8DF" box="[151,1170,1872,1894]" httpUri="https://zenodo.org/record/241733/files/figure.png" pageId="1" pageNumber="590" targetBox="[151,1436,991,1848]" targetPageId="1">
<paragraph id="8B9CF9669D256A55FF7BF8FDFB38F8DF" blockId="1.[151,1170,1872,1894]" box="[151,1170,1872,1894]" pageId="1" pageNumber="590">
<emphasis id="B95725749D256A55FF7BF8FDFEA6F8DF" bold="true" box="[151,268,1872,1894]" pageId="1" pageNumber="590">FIGURE 1.</emphasis>
<taxonomicName id="4C2382E59D256A55FEFEF8FDFE5CF8DF" box="[274,502,1872,1894]" class="Insecta" family="Libellulidae" genus="Libellula" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="590" phylum="Arthropoda" rank="species" species="coahuiltecana" status="sp. nov.">
<emphasis id="B95725749D256A55FEFEF8FDFE5CF8DF" box="[274,502,1872,1894]" italics="true" pageId="1" pageNumber="590">Libellula coahuiltecana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264980F9D256A55FE10F8FDFDE3F8DF" box="[508,585,1872,1894]" pageId="1" pageNumber="590" rank="species">
<emphasis id="B95725749D256A55FE10F8FDFDE3F8DF" bold="true" box="[508,585,1872,1894]" pageId="1" pageNumber="590">sp. nov.</emphasis>
</taxonomicNameLabel>
Male from Cuatro Ciénegas de Carranza, Coahuila, Mexico.
</paragraph>
</caption>
<caption id="DF5CA9EE9D266A56FF7BF832FC2FF849" httpUri="https://zenodo.org/record/241734/files/figure.png" pageId="2" pageNumber="591" targetBox="[229,1305,209,1922]" targetPageId="2">
<paragraph id="8B9CF9669D266A56FF7BF832FC2FF849" blockId="2.[151,1437,1951,2032]" pageId="2" pageNumber="591">
<emphasis id="B95725749D266A56FF7BF832FEA5F80C" bold="true" box="[151,271,1951,1973]" pageId="2" pageNumber="591">FIGURE 2.</emphasis>
<taxonomicName id="4C2382E59D266A56FEF4F832FE55F80C" box="[280,511,1951,1973]" class="Insecta" family="Libellulidae" genus="Libellula" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="591" phylum="Arthropoda" rank="species" species="coahuiltecana">
<emphasis id="B95725749D266A56FEF4F832FE55F80C" box="[280,511,1951,1973]" italics="true" pageId="2" pageNumber="591">Libellula coahuiltecana</emphasis>
</taxonomicName>
. Male paratype. Left wings;
<emphasis id="B95725749D266A56FCC8F832FC36F80C" bold="true" box="[804,924,1951,1973]" pageId="2" pageNumber="591">FIGURE 3.</emphasis>
<taxonomicName id="4C2382E59D266A56FC49F832FB26F80C" box="[933,1164,1951,1973]" class="Insecta" family="Libellulidae" genus="Libellula" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="591" phylum="Arthropoda" rank="species" species="coahuiltecana">
<emphasis id="B95725749D266A56FC49F832FB26F80C" box="[933,1164,1951,1973]" italics="true" pageId="2" pageNumber="591">Libellula coahuiltecana</emphasis>
</taxonomicName>
. Male paratype. Secondary genitalia;
<emphasis id="B95725749D266A56FF1AF811FEC1F86B" bold="true" box="[246,363,1980,2002]" pageId="2" pageNumber="591">FIGURE 4.</emphasis>
<taxonomicName id="4C2382E59D266A56FE9CF811FDFFF86B" box="[368,597,1980,2002]" class="Insecta" family="Libellulidae" genus="Libellula" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="591" phylum="Arthropoda" rank="species" species="coahuiltecana">
<emphasis id="B95725749D266A56FE9CF811FDFFF86B" box="[368,597,1980,2002]" italics="true" pageId="2" pageNumber="591">Libellula coahuiltecana</emphasis>
</taxonomicName>
. Male paratype. Vesica spermalis;
<emphasis id="B95725749D266A56FC4EF811FBBDF86B" bold="true" box="[930,1047,1980,2002]" pageId="2" pageNumber="591">FIGURE 5.</emphasis>
<taxonomicName id="4C2382E59D266A56FBF0F811FAABF86B" box="[1052,1281,1980,2002]" class="Insecta" family="Libellulidae" genus="Libellula" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="591" phylum="Arthropoda" rank="species" species="coahuiltecana">
<emphasis id="B95725749D266A56FBF0F811FAABF86B" box="[1052,1281,1980,2002]" italics="true" pageId="2" pageNumber="591">Libellula coahuiltecana</emphasis>
</taxonomicName>
. Male paratype. Cercus;
<emphasis id="B95725749D266A56FF0AF877FEF1F849" bold="true" box="[230,347,2010,2032]" pageId="2" pageNumber="591">FIGURE 6.</emphasis>
<taxonomicName id="4C2382E59D266A56FE8DF877FDEFF849" box="[353,581,2010,2032]" class="Insecta" family="Libellulidae" genus="Libellula" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="591" phylum="Arthropoda" rank="species" species="coahuiltecana">
<emphasis id="B95725749D266A56FE8DF877FDEFF849" box="[353,581,2010,2032]" italics="true" pageId="2" pageNumber="591">Libellula coahuiltecana</emphasis>
</taxonomicName>
. Female paratype. Vulvar lamina
</paragraph>
</caption>
<caption id="DF5CA9EE9D276A57FF7BF832FB11F86C" httpUri="https://zenodo.org/record/241735/files/figure.png" pageId="3" pageNumber="592" targetBox="[278,1347,222,1921]" targetPageId="3">
<paragraph id="8B9CF9669D276A57FF7BF832FB11F86C" blockId="3.[151,1436,1951,2005]" pageId="3" pageNumber="592">
<emphasis id="B95725749D276A57FF7BF832FEA7F80C" bold="true" box="[151,269,1951,1973]" pageId="3" pageNumber="592">FIGURE 7.</emphasis>
<taxonomicName id="4C2382E59D276A57FEFFF832FE64F80C" box="[275,462,1951,1973]" class="Insecta" family="Libellulidae" genus="Libellula" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="592" phylum="Arthropoda" rank="species" species="needhami">
<emphasis id="B95725749D276A57FEFFF832FE64F80C" box="[275,462,1951,1973]" italics="true" pageId="3" pageNumber="592">Libellula needhami</emphasis>
</taxonomicName>
. Left wings;
<emphasis id="B95725749D276A57FDBCF832FD6CF80C" bold="true" box="[592,710,1951,1973]" pageId="3" pageNumber="592">FIGURE 8.</emphasis>
<taxonomicName id="4C2382E59D276A57FD20F832FC2CF80C" box="[716,902,1951,1973]" class="Insecta" family="Libellulidae" genus="Libellula" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="592" phylum="Arthropoda" rank="species" species="needhami">
<emphasis id="B95725749D276A57FD20F832FC2CF80C" box="[716,902,1951,1973]" italics="true" pageId="3" pageNumber="592">Libellula needhami</emphasis>
</taxonomicName>
. Secondary genitalia;
<emphasis id="B95725749D276A57FBB3F832FB7FF80C" bold="true" box="[1119,1237,1951,1973]" pageId="3" pageNumber="592">FIGURE 9.</emphasis>
<taxonomicName id="4C2382E59D276A57FB30F832FA3CF80C" box="[1244,1430,1951,1973]" class="Insecta" family="Libellulidae" genus="Libellula" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="592" phylum="Arthropoda" rank="species" species="needhami">
<emphasis id="B95725749D276A57FB30F832FA3CF80C" box="[1244,1430,1951,1973]" italics="true" pageId="3" pageNumber="592">Libellula needhami</emphasis>
</taxonomicName>
. Vesica spermalis;
<emphasis id="B95725749D276A57FEA8F812FE6FF86C" bold="true" box="[324,453,1983,2005]" pageId="3" pageNumber="592">FIGURE 10.</emphasis>
<taxonomicName id="4C2382E59D276A57FE27F812FD2FF86C" box="[459,645,1983,2005]" class="Insecta" family="Libellulidae" genus="Libellula" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="592" phylum="Arthropoda" rank="species" species="needhami">
<emphasis id="B95725749D276A57FE27F812FD2FF86C" box="[459,645,1983,2005]" italics="true" pageId="3" pageNumber="592">Libellula needhami</emphasis>
</taxonomicName>
. Cercus;
<emphasis id="B95725749D276A57FD0CF812FCCAF86C" bold="true" box="[736,864,1983,2005]" pageId="3" pageNumber="592">FIGURE 11.</emphasis>
<taxonomicName id="4C2382E59D276A57FC8AF812FBB5F86C" box="[870,1055,1983,2005]" class="Insecta" family="Libellulidae" genus="Libellula" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="592" phylum="Arthropoda" rank="species" species="needhami">
<emphasis id="B95725749D276A57FC8AF812FBB5F86C" box="[870,1055,1983,2005]" italics="true" pageId="3" pageNumber="592">Libellula needhami</emphasis>
</taxonomicName>
. Vulvar lamina.
</paragraph>
</caption>
<paragraph id="8B9CF9669D206A50FF2BFF35FD02FE18" blockId="4.[151,1436,152,1423]" pageId="4" pageNumber="593">
Wings: Mostly hyaline with orange-red markings as follows. In FW a basal spot extending from base to subtriangle, in HW the same spot extending from base to triangle; spot at nodus reaching bridge below; costal and subcostal areas and tips of wings suffused with same color (
<figureCitation id="1318E5E39D206A50FDD3FF70FD2AFF4D" box="[575,640,221,244]" captionStart="FIGURE 2" captionStartId="2.[151,244,1951,1973]" captionTargetBox="[229,1305,209,1922]" captionTargetId="figure@2.[178,1409,191,1930]" captionTargetPageId="2" captionText="FIGURE 2. Libellula coahuiltecana. Male paratype. Left wings; FIGURE 3. Libellula coahuiltecana. Male paratype. Secondary genitalia; FIGURE 4. Libellula coahuiltecana. Male paratype. Vesica spermalis; FIGURE 5. Libellula coahuiltecana. Male paratype. Cercus; FIGURE 6. Libellula coahuiltecana. Female paratype. Vulvar lamina" httpUri="https://zenodo.org/record/241734/files/figure.png" pageId="4" pageNumber="593">Fig. 2</figureCitation>
). One cubito-anal Cx in all wings; Ax in FW 14, in HW 12 (L) and 10 (R); Px in Fw 8 (L) and 9 (R), in HW 10 (L) and 9 (R); FW triangle with 4 (L) and 3 (R) Cx, HW triangle with a single Cx; FW subtriangle with 5 (L) and 6 (R) cells; FW discoidal field starting with 4 cell rows for a space of one cell length, then decreasing to 3 rows at commencement of Rspl, at hind margin increasing to 12 (LFW) and 11 (RFW) rows, 3 paranal cells in HW; FW bridge Cx 2, HW bridge Cx 1; FW Pt orange,
<quantity id="4CDB54839D206A50FCDAFEC5FC39FEC6" box="[822,915,360,383]" metricMagnitude="-3" metricUnit="m" metricValue="4.62" pageId="4" pageNumber="593" unit="mm" value="4.62">4.62 mm</quantity>
long, with 3 (R) or (4) Cx immediately below, in HW
<quantity id="4CDB54839D206A50FF26FE27FE83FE18" box="[202,297,394,417]" metricMagnitude="-3" metricUnit="m" metricValue="4.81" pageId="4" pageNumber="593" unit="mm" value="4.81">4.81 mm</quantity>
long with 3 Cx immediately below.
</paragraph>
<paragraph id="8B9CF9669D206A50FF2BFE00FD77FD0E" blockId="4.[151,1436,152,1423]" pageId="4" pageNumber="593">
<emphasis id="B95725749D206A50FF2BFE00FE86FE7D" box="[199,300,429,452]" italics="true" pageId="4" pageNumber="593">Abdomen</emphasis>
. Abdomen orange. Dorsum of S1 and narrow latero-basal stripe on S10 dark brown. Latero-apical spots of same color on S8 and S9. In ventral view abdomen gradually narrowing toward apex. S4 about as long as its basal width (ratio width/length= 0.97) in ventral view. In lateral view anterior lamina lower than hamule and genital lobe, genital lobe high and rounded; hamule bifurcate, inner branch narrow, vertically raised, ending in an externally directed recurved black tooth; outer branch low, rounded (Fig. 3). Vesica spermalis: segment 4 highly sclerotized with lateral lobes long, tubular, directed caudally, with tips converging medially, median lobe with two very thin wirelike cornua directed ventrally (Fig 4). Cercus with a row of seven denticles along its lower margin, tip slightly upturned in lateral view (Fig. 5); cercus length 2.73; epiproct length 2.36.
</paragraph>
<paragraph id="8B9CF9669D206A50FF2BFD6FFF79FD45" blockId="4.[151,1436,152,1423]" pageId="4" pageNumber="593">Dimensions: Total length 47.89; abdomen length 31.7; FW length 36.58; HW length 34.89; HW maximum width 10.52</paragraph>
<paragraph id="8B9CF9669D206A50FF2BFCA5FA3AFCA6" blockId="4.[151,1436,152,1423]" box="[199,1424,776,799]" pageId="4" pageNumber="593">
Variation in male
<typeStatus id="549847C49D206A50FE69FCA5FE41FCA6" box="[389,491,776,799]" pageId="4" pageNumber="593" type="paratype">paratypes</typeStatus>
(n=8): Little variation observed in male
<typeStatus id="549847C49D206A50FC72FCA5FBAEFCA6" box="[926,1028,776,799]" pageId="4" pageNumber="593" type="paratype">paratypes</typeStatus>
except for wing venation as follows:
</paragraph>
<paragraph id="8B9CF9669D206A50FF2BFC87FDADFC10" blockId="4.[151,1436,152,1423]" pageId="4" pageNumber="593">
Ax in FW 1317, in Hw 912; Px in FW 811, in Hw 911; triangle with 23 Cx in FW; FW subtriangle with 36 cells; FW discoidal field starting with 34 rows of cells then increasing to 1013 at hind margin, HW starting with 34 rows increasing to 1114 at hind margin; bridge Cx,
<quantity id="4CDB54839D206A50FD27FCDDFCA4FC3E" box="[715,782,880,903]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="4" pageNumber="593" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">13 in</quantity>
Fw and
<quantity id="4CDB54839D206A50FC87FCDDFC05FC3E" box="[875,943,880,903]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="4" pageNumber="593" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">14 in</quantity>
Hw; Pt
<quantity id="4CDB54839D206A50FBEBFCDDFB27FC3E" box="[1031,1165,880,903]" metricMagnitude="-1" metricUnit="m" metricValue="1.1557000000000002" metricValueMax="1.2471400000000001" metricValueMin="1.0642600000000002" pageId="4" pageNumber="593" unit="in" value="4.55" valueMax="4.91" valueMin="4.19">4.194.91 in</quantity>
Fw,
<quantity id="4CDB54839D206A50FB2DFCDDFAE2FC3E" box="[1217,1352,880,903]" metricMagnitude="-1" metricUnit="m" metricValue="1.19761" metricValueMax="1.28016" metricValueMin="1.11506" pageId="4" pageNumber="593" unit="in" value="4.715" valueMax="5.04" valueMin="4.39">4.395.04 in</quantity>
Hw, 2 4 Cx behind Pt in FW,
<quantity id="4CDB54839D206A50FE67FC3FFE64FC10" box="[395,462,914,937]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="4" pageNumber="593" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">34 in</quantity>
HW.
</paragraph>
<paragraph id="8B9CF9669D206A50FF2BFC18FBD3FC75" blockId="4.[151,1436,152,1423]" box="[199,1145,949,972]" pageId="4" pageNumber="593">S4 width-length ratio 0.871.08 (1.07); cerci 2.582.88 (2.77); epiproct 2.172.36 (2.26)</paragraph>
<paragraph id="8B9CF9669D206A50FF2BFC75FD83FBA8" blockId="4.[151,1436,152,1423]" pageId="4" pageNumber="593">Dimensions: Total length 44.849.6 (48.3); abdomen 29.233.3 (31.9); FW 34.738.2 (37.0); HW 33.236.3 (35.4); HW maximum width 9.711.7 (11.0).</paragraph>
</subSubSection>
<subSubSection id="C339AAED9D206A50FF2BFBB0FF73FA36" pageId="4" pageNumber="593" type="description">
<paragraph id="8B9CF9669D206A50FF2BFBB0FC8CFBEE" blockId="4.[151,1436,152,1423]" pageId="4" pageNumber="593">
<emphasis id="B95725749D206A50FF2BFBB0FE26FB8D" bold="true" box="[199,396,1053,1076]" pageId="4" pageNumber="593">
Female
<typeStatus id="549847C49D206A50FECAFBB0FE26FB8D" box="[294,396,1053,1076]" pageId="4" pageNumber="593" type="paratype">paratype</typeStatus>
</emphasis>
(young specimen): Head: Labrum cream, rest of the head yellowish brown, rear of vertex, antennae and adjacent areas dark brown, occiput light brown.
</paragraph>
<paragraph id="8B9CF9669D206A50FF2BFBCFFC23FB25" blockId="4.[151,1436,152,1423]" pageId="4" pageNumber="593">
<emphasis id="B95725749D206A50FF2BFBCFFEBEFBC0" box="[199,276,1122,1145]" italics="true" pageId="4" pageNumber="593">Thorax</emphasis>
: Entirely tan but mesepisternum suffused with light brown; legs yellow, apex of femora and remainder of leg dark brown, armature black; legs yellow at base to light brown at tarsi;
</paragraph>
<paragraph id="8B9CF9669D206A50FF2BFB05FD77FABD" blockId="4.[151,1436,152,1423]" pageId="4" pageNumber="593">
Wings with orange patches as in male although lighter; Ax in FW 14, 9 in HW; Px in FW 8, in HW 9 (left) and 8 (right); FW subtriangle with 5 (left) and 4 (right) cells; bridge Cx
<quantity id="4CDB54839D206A50FC8EFB67FC26FB58" box="[866,908,1226,1249]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="593" unit="in" value="2.0">2 in</quantity>
FW, 1 (left) and 3 (right) in HW; Pt
<quantity id="4CDB54839D206A50FAF1FB67FAC2FB58" box="[1309,1384,1226,1249]" metricMagnitude="-1" metricUnit="m" metricValue="1.2319" pageId="4" pageNumber="593" unit="in" value="4.85">4.85 in</quantity>
FW,
<quantity id="4CDB54839D206A50FF7BFB40FF4BFABD" box="[151,225,1261,1284]" metricMagnitude="-1" metricUnit="m" metricValue="1.2573" pageId="4" pageNumber="593" unit="in" value="4.95">4.95 in</quantity>
HW; 2 Cx behind pterostigma in FW,
<quantity id="4CDB54839D206A50FD91FB40FD0FFABD" box="[637,677,1261,1284]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="593" unit="in" value="3.0">3 in</quantity>
HW.
</paragraph>
<paragraph id="8B9CF9669D206A50FF2BFABDFAEEFAF0" blockId="4.[151,1436,152,1423]" pageId="4" pageNumber="593">
<emphasis id="B95725749D206A50FF2BFABDFE86FA9E" box="[199,300,1296,1319]" italics="true" pageId="4" pageNumber="593">Abdomen</emphasis>
: pale brown, S49 with a narrow middorsal black stripe (diffuse on S4 and S89); S8 flanked with narrow ventrolateral flaps. Cercus length 1.48. Vulvar lamina &quot;V&quot; shaped with two nipplelike structures at base (Fig. 6).
</paragraph>
<paragraph id="8B9CF9669D206A50FF2BFAF8FF73FA36" blockId="4.[151,1436,152,1423]" pageId="4" pageNumber="593">Dimensions: Total length 44.72; abdomen length 28.71; FW length 36.78; HW length 34.45; HW maximum length 11.67.</paragraph>
</subSubSection>
</treatment>
</document>