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<document id="0904CD32C42394E3E69628CA7ED26712" ID-DOI="10.11646/zootaxa.5293.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="7960232" ID-ZooBank="F68B3A5C-E100-4F7A-A1A1-FC9933403700" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1684798263703" checkinUser="plazi" docAuthor="Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder &amp; Lira, André Felipe De Araújo" docDate="2023" docId="0389886AE949FFBF13991174FBA22562" docLanguage="en" docName="zootaxa.5293.2.6.pdf" docOrigin="Zootaxa 5293 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5293.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudochthonius biseriatus Mahnert 2001" docType="treatment" docVersion="2" lastPageNumber="319" masterDocId="FFB0F012E94BFFBD130E1314FF832309" masterDocTitle="Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil" masterLastPageNumber="332" masterPageNumber="317" pageNumber="319" updateTime="1684800287596" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3C8D3DAD05813444652CF2E3FA784829">Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil</mods:title>
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<mods:namePart id="E77506C4BE21B66890F3C80560CE0F7A">Bedoya-Roqueme, Edwin</mods:namePart>
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<mods:namePart id="8EBF7A8055FE7556EBF748FBFE602B64">Tizo-Pedroso, Everton</mods:namePart>
<mods:affiliation id="EF7ABA9354C845582B5447E0006A1107">Laboratório de Ecologia Comportamental de Aracnídeos, Programa de Pós-graduação em Recursos Naturais do Cerrado, Centro de Ensino e Aprendizagem em Rede, Universidade Estadual de Goiás, Anápolis, Brazil.</mods:affiliation>
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<mods:namePart id="1E4EF0E0D917FFE32A5942C3F5CE4D5C">Barbier, Eder</mods:namePart>
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<mods:affiliation id="067E743E7395BEE1F35FD9D2640CA619">Programa de Pós-graduação em Biologia Animal, Departamento de Zoologia, Universidade Federal de Pernambuco, Recife, Brazil. barbier. eder @ gmail. com; http: // orcid. org / 0000 - 0001 - 5068 - 7048</mods:affiliation>
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<treatment id="0389886AE949FFBF13991174FBA22562" ID-DOI="http://doi.org/10.5281/zenodo.7960252" ID-Zenodo-Dep="7960252" LSID="urn:lsid:plazi:treatment:0389886AE949FFBF13991174FBA22562" httpUri="http://treatment.plazi.org/id/0389886AE949FFBF13991174FBA22562" lastPageNumber="319" pageId="2" pageNumber="319">
<subSubSection id="C33A6AF7E949FFBF13991174FD272173" box="[151,676,608,635]" pageId="2" pageNumber="319" type="nomenclature">
<paragraph id="8B9F397CE949FFBF13991174FD272173" blockId="2.[151,676,608,635]" box="[151,676,608,635]" pageId="2" pageNumber="319">
<heading id="D0D78E10E949FFBF13991174FD272173" bold="true" box="[151,676,608,635]" fontSize="11" level="1" pageId="2" pageNumber="319" reason="1">
<taxonomicName id="4C2042FFE949FFBF13991174FD272173" ID-CoL="4NTPD" authority="Mahnert, 2001" authorityName="Mahnert" authorityYear="2001" box="[151,676,608,635]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="319" phylum="Arthropoda" rank="species" species="biseriatus">
<emphasis id="B954E56EE949FFBF13991174FD272173" bold="true" box="[151,676,608,635]" pageId="2" pageNumber="319">
<emphasis id="B954E56EE949FFBF13991174FE672172" bold="true" box="[151,484,608,635]" italics="true" pageId="2" pageNumber="319">Pseudochthonius biseriatus</emphasis>
<bibRefCitation id="EFB1448DE949FFBF12E21175FD272173" author="Mahnert, V." box="[492,676,608,635]" pageId="2" pageNumber="319" pagination="95 - 148" refId="ref14699" refString="Mahnert, V. (2001) Cave-dwelling pseudoscorpions (Arachnida, Pseudoscorpiones) from Brazil. Revue Suisse de Zoologie, 108, 95 - 148. https: // doi. org / 10.5962 / bhl. part. 79622" type="journal article" year="2001">Mahnert, 2001</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="C33A6AF7E949FFBF139911BCFB9F21EE" pageId="2" pageNumber="319" type="materials_examined">
<paragraph id="8B9F397CE949FFBF139911BCFB9F21EE" blockId="2.[151,1437,680,1643]" pageId="2" pageNumber="319">
<emphasis id="B954E56EE949FFBF139911BCFE0021CB" bold="true" box="[151,387,680,706]" pageId="2" pageNumber="319">Material examined.</emphasis>
<materialsCitation id="3B483321E949FFBF128511BDFB9A21EE" collectingDate="2019-05-09" collectionCode="LECA" collectorName="Catimbau National Park, E &amp; E. Barbier" country="Brazil" elevation="556" latitude="-8.570583" location="Furna do Morcego cave" longLatPrecision="1" longitude="-37.38211" municipality="Catimbau National Park" pageId="2" pageNumber="319" specimenCount="2" specimenCount-male="2" stateProvince="Pernambuco">
<specimenCount id="9D26F2F5E949FFBF128511BDFE6521CB" box="[395,486,680,707]" pageId="2" pageNumber="319" type="male">2 males</specimenCount>
,
<collectingRegion id="49E4F79EE949FFBF12FA11BCFD0021CB" box="[500,643,680,706]" country="Brazil" name="Pernambuco" pageId="2" pageNumber="319">Pernambuco</collectingRegion>
,
<collectingCountry id="F33779ECE949FFBF119E11BCFD5A21CB" box="[656,729,680,706]" name="Brazil" pageId="2" pageNumber="319">Brazil</collectingCountry>
:
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,
<geoCoordinate id="EE145FBBE949FFBF170B11BDFB1721CA" box="[1029,1172,681,707]" degrees="8" direction="south" minutes="34" orientation="latitude" pageId="2" pageNumber="319" precision="1" seconds="14.1" value="-8.570583">8°3414.1”S</geoCoordinate>
;
<geoCoordinate id="EE145FBBE949FFBF17AC11BCFAC521CA" box="[1186,1350,680,707]" degrees="37" direction="west" minutes="22" orientation="longitude" pageId="2" pageNumber="319" precision="1" seconds="55.6" value="-37.38211">37°2255.6”W</geoCoordinate>
,
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<quantity id="4CD89499E949FFBF165D11BCFA1F21CA" box="[1363,1436,680,707]" metricMagnitude="2" metricUnit="m" metricValue="5.56" pageId="2" pageNumber="319" unit="m" value="556.0">556 m</quantity>
a.s.l.
</elevation>
,
<date id="FF9E1FBCE949FFBF13D711D9FED621EE" box="[217,341,717,743]" pageId="2" pageNumber="319" value="2019-05-09">
<collectingDate id="EFDAE654E949FFBF13D711D9FED621EE" box="[217,341,717,743]" pageId="2" pageNumber="319" value="2019-05-09">09-V-2019</collectingDate>
</date>
;
<collectorName id="26D55CAAE949FFBF126E11D8FD1221EE" box="[352,657,716,743]" pageId="2" pageNumber="319">
<collectingMunicipality id="6BFBA306E949FFBF126E11D8FDF121EF" box="[352,626,716,742]" pageId="2" pageNumber="319">Catimbau National Park</collectingMunicipality>
, E
</collectorName>
. Barbier leg. (
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; Ps-SCR008)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A6AF7E949FFBF13C911E4FD5B277A" pageId="2" pageNumber="319" type="diagnosis">
<paragraph id="8B9F397CE949FFBF13C911E4FD5B277A" blockId="2.[151,1437,680,1643]" pageId="2" pageNumber="319">
<emphasis id="B954E56EE949FFBF13C911E4FEC12003" bold="true" box="[199,322,752,778]" pageId="2" pageNumber="319">Diagnosis.</emphasis>
According to
<bibRefCitation id="EFB1448DE949FFBF12E911E4FD1A2002" author="Mahnert, V." box="[487,665,752,779]" pageId="2" pageNumber="319" pagination="95 - 148" refId="ref14699" refString="Mahnert, V. (2001) Cave-dwelling pseudoscorpions (Arachnida, Pseudoscorpiones) from Brazil. Revue Suisse de Zoologie, 108, 95 - 148. https: // doi. org / 10.5962 / bhl. part. 79622" type="journal article" year="2001">Mahnert (2001)</bibRefCitation>
<taxonomicName id="4C2042FFE949FFBF11AC11E4FC5B2003" authorityName="Mahnert" authorityYear="2001" box="[674,984,752,778]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="319" phylum="Arthropoda" rank="species" species="biseriatus">
<emphasis id="B954E56EE949FFBF11AC11E4FC5B2003" box="[674,984,752,778]" italics="true" pageId="2" pageNumber="319">Pseudochthonius biseriatus</emphasis>
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, it can be easily distinguished from the other species of the genus by the carapace reticule, as long as broad, absent eyes, anterior margin with a prominent dentate epistome, 16 strong setae, one preocular seta present on each side (
<bibRefCitation id="EFB1448DE949FFBF10D9102CFBF9205A" author="Mahnert, V." box="[983,1146,824,851]" pageId="2" pageNumber="319" pagination="95 - 148" refId="ref14699" refString="Mahnert, V. (2001) Cave-dwelling pseudoscorpions (Arachnida, Pseudoscorpiones) from Brazil. Revue Suisse de Zoologie, 108, 95 - 148. https: // doi. org / 10.5962 / bhl. part. 79622" type="journal article" year="2001">Mahnert 2001</bibRefCitation>
;
<figureCitation id="131B25F9E949FFBF178B102CFB55205A" box="[1157,1238,824,851]" captionStart="FIGURES 514" captionStartId="6.[152,270,1830,1854]" captionTargetBox="[188,1405,212,1797]" captionTargetId="figure-23@6.[170,1417,181,1806]" captionTargetPageId="6" captionText="FIGURES 514. Geogarypus gollumi sp. nov. female holotype, dorsal view, unless stated otherwise: 5, carapace, dorsal view. 6, left chelicera, dorsal view. 7, left galea, lateral view. 8, left rallum, lateral view. 9, male paratype, right pedipalp aspect. 10, female holotype, right pedipalp aspect. 11, right chela, lateral view (arrows indicating aspect of the teeth). 12, genital sternites, ventral view. 1314, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960238" httpUri="https://zenodo.org/record/7960238/files/figure.png" pageId="2" pageNumber="319">Fig. 13</figureCitation>
); Chelicerae with five setae, fixed finger with 911 acute teeth, movable finger with 810 teeth, absent spinneret tooth-like in male, serrula exterior with 20 blades, rallum nine setae, first only shorts dentate (see
<bibRefCitation id="EFB1448DE949FFBF17281094FB492092" author="Mahnert, V." box="[1062,1226,896,923]" pageId="2" pageNumber="319" pagination="95 - 148" refId="ref14699" refString="Mahnert, V. (2001) Cave-dwelling pseudoscorpions (Arachnida, Pseudoscorpiones) from Brazil. Revue Suisse de Zoologie, 108, 95 - 148. https: // doi. org / 10.5962 / bhl. part. 79622" type="journal article" year="2001">Mahnert 2001</bibRefCitation>
;
<figureCitation id="131B25F9E949FFBF17D61094FAAF2092" box="[1240,1324,896,923]" captionStart="FIGURES 514" captionStartId="6.[152,270,1830,1854]" captionTargetBox="[188,1405,212,1797]" captionTargetId="figure-23@6.[170,1417,181,1806]" captionTargetPageId="6" captionText="FIGURES 514. Geogarypus gollumi sp. nov. female holotype, dorsal view, unless stated otherwise: 5, carapace, dorsal view. 6, left chelicera, dorsal view. 7, left galea, lateral view. 8, left rallum, lateral view. 9, male paratype, right pedipalp aspect. 10, female holotype, right pedipalp aspect. 11, right chela, lateral view (arrows indicating aspect of the teeth). 12, genital sternites, ventral view. 1314, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960238" httpUri="https://zenodo.org/record/7960238/files/figure.png" pageId="2" pageNumber="319">Fig. 14</figureCitation>
); Tergites whitish-yellow, tergites IIIIV with round medial spots, chaetotaxy: 2: 2: 4: 4-6: 6: 6: 6: 6: 4: 4(2ST): 2; Pedipalps, femur 6.46.6×, patella 2.22.5×, hand 2.32.4×, chela 7.58.0×, finger 2.3×, fixed finger with 3741 pointed teeth arranged like offset saw teeth and therefore giving the impression of heterodonty, trichobothrium
<emphasis id="B954E56EE949FFBF17E810F9FA83270F" box="[1254,1280,1005,1030]" italics="true" pageId="2" pageNumber="319">ist</emphasis>
much near to
<emphasis id="B954E56EE949FFBF13991704FF3F2723" box="[151,188,1040,1066]" italics="true" pageId="2" pageNumber="319">esb</emphasis>
than to
<emphasis id="B954E56EE949FFBF12131705FEBF2723" box="[285,316,1041,1066]" italics="true" pageId="2" pageNumber="319">est</emphasis>
, and one sensillum distal to
<emphasis id="B954E56EE949FFBF11861704FD222723" box="[648,673,1040,1066]" italics="true" pageId="2" pageNumber="319">sb</emphasis>
on movable finger (see
<bibRefCitation id="EFB1448DE949FFBF10B41704FBDE2722" author="Mahnert, V." box="[954,1117,1040,1067]" pageId="2" pageNumber="319" pagination="95 - 148" refId="ref14699" refString="Mahnert, V. (2001) Cave-dwelling pseudoscorpions (Arachnida, Pseudoscorpiones) from Brazil. Revue Suisse de Zoologie, 108, 95 - 148. https: // doi. org / 10.5962 / bhl. part. 79622" type="journal article" year="2001">Mahnert 2001</bibRefCitation>
;
<figureCitation id="131B25F9E949FFBF17651704FB762722" box="[1131,1269,1040,1067]" captionStart="FIGURES 1524" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 1524. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 2324, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="2" pageNumber="319">Figs. 1516</figureCitation>
); Leg I: femur 6.16.2×, 4.24.5×, tibia 4.95.3×, tarsus 10.411.3×; Leg IV: femur+patella 3.13.4×, tibia 5.05.2×, metatarsus 3.53.8×, tarsus 10.611.9× (one basal tactile seta).
</paragraph>
</subSubSection>
<subSubSection id="C33A6AF7E949FFBF13C91769FE452666" pageId="2" pageNumber="319" type="discussion">
<paragraph id="8B9F397CE949FFBF13C91769FE452666" blockId="2.[151,1437,680,1643]" pageId="2" pageNumber="319">
<emphasis id="B954E56EE949FFBF13C91769FEB8279E" bold="true" box="[199,315,1149,1175]" pageId="2" pageNumber="319">Remarks.</emphasis>
Measurements and proportion of pedipalps, distribution of trichobothria, and some other taxonomic characters correspond to the description of
<taxonomicName id="4C2042FFE949FFBF118B17B5FC9327B3" authorityName="Mahnert" authorityYear="2001" box="[645,784,1184,1210]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="319" phylum="Arthropoda" rank="species" species="biseriatus">
<emphasis id="B954E56EE949FFBF118B17B5FC9327B3" box="[645,784,1184,1210]" italics="true" pageId="2" pageNumber="319">P. biseriatus</emphasis>
</taxonomicName>
from
<collectingRegion id="49E4F79EE949FFBF105617B4FC7027B3" box="[856,1011,1184,1210]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="319">Minas Gerais</collectingRegion>
,
<collectingCountry id="F33779ECE949FFBF170E17B4FBC727B3" box="[1024,1092,1184,1210]" name="Brazil" pageId="2" pageNumber="319">Brazil</collectingCountry>
(
<bibRefCitation id="EFB1448DE949FFBF175A17B4FB7A27B2" author="Mahnert, V." box="[1108,1273,1184,1211]" pageId="2" pageNumber="319" pagination="95 - 148" refId="ref14699" refString="Mahnert, V. (2001) Cave-dwelling pseudoscorpions (Arachnida, Pseudoscorpiones) from Brazil. Revue Suisse de Zoologie, 108, 95 - 148. https: // doi. org / 10.5962 / bhl. part. 79622" type="journal article" year="2001">Mahnert 2001</bibRefCitation>
;
<figureCitation id="131B25F9E949FFBF160817B4FA1327B3" box="[1286,1424,1184,1210]" captionStart-0="FIGURES 514" captionStart-1="FIGURES 1524" captionStartId-0="6.[152,270,1830,1854]" captionStartId-1="9.[152,270,1830,1854]" captionTargetBox-0="[188,1405,212,1797]" captionTargetBox-1="[187,1367,192,1797]" captionTargetId-0="figure-23@6.[170,1417,181,1806]" captionTargetId-1="figure-19@9.[170,1417,181,1806]" captionTargetPageId-0="6" captionTargetPageId-1="9" captionText-0="FIGURES 514. Geogarypus gollumi sp. nov. female holotype, dorsal view, unless stated otherwise: 5, carapace, dorsal view. 6, left chelicera, dorsal view. 7, left galea, lateral view. 8, left rallum, lateral view. 9, male paratype, right pedipalp aspect. 10, female holotype, right pedipalp aspect. 11, right chela, lateral view (arrows indicating aspect of the teeth). 12, genital sternites, ventral view. 1314, left leg I, lateral view, and left leg IV, lateral view." captionText-1="FIGURES 1524. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 2324, left leg I, lateral view, and left leg IV, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.7960238" figureDoi-1="http://doi.org/10.5281/zenodo.7960240" httpUri-0="https://zenodo.org/record/7960238/files/figure.png" httpUri-1="https://zenodo.org/record/7960240/files/figure.png" pageId="2" pageNumber="319">Figs. 1316</figureCitation>
). The specimens of the cave from
<collectingRegion id="49E4F79EE949FFBF110317D0FD1A27D7" box="[525,665,1220,1246]" country="Brazil" name="Pernambuco" pageId="2" pageNumber="319">Pernambuco</collectingRegion>
(
<collectingCountry id="F33779ECE949FFBF11A417D0FD7227D7" box="[682,753,1220,1246]" name="Brazil" pageId="2" pageNumber="319">Brazil</collectingCountry>
) slightly differ from the other specimens in the body length (
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<geoCoordinate id="EE145FBBE949FFBF13EE17FCFEB3260A" box="[224,304,1256,1283]" degrees="1.656" direction="west" orientation="longitude" pageId="2" pageNumber="319" precision="55" value="-1.656">1.656</geoCoordinate>
), smaller than those described by
<bibRefCitation id="EFB1448DE949FFBF11BC17FCFCE6260A" author="Mahnert, V." box="[690,869,1256,1283]" pageId="2" pageNumber="319" pagination="95 - 148" refId="ref14699" refString="Mahnert, V. (2001) Cave-dwelling pseudoscorpions (Arachnida, Pseudoscorpiones) from Brazil. Revue Suisse de Zoologie, 108, 95 - 148. https: // doi. org / 10.5962 / bhl. part. 79622" type="journal article" year="2001">Mahnert (2001)</bibRefCitation>
, similarly in the number of teeth on the fingers of the chela, 35 teeth on the fixed finger and 31 on the movable finger, however, these differences can be attributed to intraspecific variations by the geographical distribution of the species, since all the diagnostic characters correspond to the species
<taxonomicName id="4C2042FFE949FFBF123B1641FE3C2667" authorityName="Mahnert" authorityYear="2001" box="[309,447,1364,1390]" class="Arachnida" family="Chthoniidae" genus="Pseudochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="319" phylum="Arthropoda" rank="species" species="biseriatus">
<emphasis id="B954E56EE949FFBF123B1641FE3C2667" box="[309,447,1364,1390]" italics="true" pageId="2" pageNumber="319">P. biseriatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A6AF7E949FFBF13C9166DFD8C254E" pageId="2" pageNumber="319" type="description">
<paragraph id="8B9F397CE949FFBF13C9166DFD8C254E" blockId="2.[151,1437,680,1643]" pageId="2" pageNumber="319">
<emphasis id="B954E56EE949FFBF13C9166DFE4F269B" bold="true" box="[199,460,1400,1427]" pageId="2" pageNumber="319">Measurements (mm).</emphasis>
Body length:
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<geoCoordinate id="EE145FBBE949FFBF11B0166CFC8D269A" box="[702,782,1400,1427]" degrees="1.656" direction="west" orientation="longitude" pageId="2" pageNumber="319" precision="55" value="-1.656">1.656</geoCoordinate>
. Carapace: 0.5230.526/0.4380.456. Pedipalps: femur: 0.4260.459/0.1140.116, patella 0.2370.241/0.1010.103, chela: 0.7920.796/0.1570.159, movable finger:
<geoCoordinate id="EE145FBBE949FFBF139916D4FF5426D2" box="[151,215,1472,1499]" degrees="0.513" direction="north" orientation="latitude" pageId="2" pageNumber="319" precision="55" value="0.513">0.513</geoCoordinate>
/
<geoCoordinate id="EE145FBBE949FFBF13EE16D4FEA226D2" box="[224,289,1472,1499]" degrees="0.030" direction="east" orientation="longitude" pageId="2" pageNumber="319" precision="55" value="0.03">0.030</geoCoordinate>
. Chelicera:
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/
<geoCoordinate id="EE145FBBE949FFBF12F316D5FDBE26D2" box="[509,573,1473,1499]" degrees="0.090" direction="east" orientation="longitude" pageId="2" pageNumber="319" precision="55" value="0.09">0.090</geoCoordinate>
, movable finger length: 0.125. Leg I: femur: 0.4610.463/0.0700.080, patella: 0.2730.281/0.0600.080, tibia: 0.2510.273/0.0500.070, tarsus: 0.4770.479/0.0500.060. Leg IV: femur+patella: 0.6890.690/0.2130.232, tibia: 0.4410443/
<geoCoordinate id="EE145FBBE949FFBF106C151DFC21252A" box="[866,930,1545,1571]" degrees="0.070" direction="north" orientation="latitude" pageId="2" pageNumber="319" precision="55" value="0.07">0.070</geoCoordinate>
<geoCoordinate id="EE145FBBE949FFBF10AC151DFC72252A" box="[930,1009,1545,1571]" degrees="0.090" direction="west" orientation="longitude" pageId="2" pageNumber="319" precision="55" value="-0.09">0.090</geoCoordinate>
, metatarsus: 0.250.27/
<geoCoordinate id="EE145FBBE949FFBF1604151CFAC9252A" box="[1290,1354,1544,1571]" degrees="0.070" direction="north" orientation="latitude" pageId="2" pageNumber="319" precision="55" value="0.07">0.070</geoCoordinate>
<geoCoordinate id="EE145FBBE949FFBF1644151CFA1B252A" box="[1354,1432,1544,1571]" degrees="0.090" direction="west" orientation="longitude" pageId="2" pageNumber="319" precision="55" value="-0.09">0.090</geoCoordinate>
, tarsus: 0.4930.495/0.0500.070.
</paragraph>
</subSubSection>
<subSubSection id="C33A6AF7E949FFBF13C91544FBA22562" box="[199,1057,1616,1643]" pageId="2" pageNumber="319" type="distribution">
<paragraph id="8B9F397CE949FFBF13C91544FBA22562" blockId="2.[151,1437,680,1643]" box="[199,1057,1616,1643]" pageId="2" pageNumber="319">
<emphasis id="B954E56EE949FFBF13C91544FEE02563" bold="true" box="[199,355,1616,1642]" pageId="2" pageNumber="319">Distribution:</emphasis>
<collectingCountry id="F33779ECE949FFBF12641544FE2D2563" box="[362,430,1616,1642]" name="Brazil" pageId="2" pageNumber="319">Brazil</collectingCountry>
(
<bibRefCitation id="EFB1448DE949FFBF12B31545FDD02562" author="Harvey, M. S." box="[445,595,1617,1643]" pageId="2" pageNumber="319" refId="ref14336" refString="Harvey, M. S. (2013) Pseudoscorpions of the World. Version 3.0. Western Australian Museum, Perth. Available from: http: // www. museum. wa. gov. au / catalogues / pseudoscorpions (accessed 20 February 2023)" type="url" year="2013">Harvey 2013</bibRefCitation>
;
<bibRefCitation id="EFB1448DE949FFBF11501544FB962562" author="World Pseudoscorpiones Catalog" box="[606,1045,1616,1643]" pageId="2" pageNumber="319" refId="ref15487" refString="World Pseudoscorpiones Catalog (2022) World Pseudoscorpiones Catalog. Natural History Museum Bern. Available from: http: // wac. nmbe. ch (accessed 20 February 2022)" type="url" year="2022">
World
<taxonomicName id="4C2042FFE949FFBF11A41544FCF02563" authorityName="de Geer" authorityYear="1778" box="[682,883,1616,1642]" class="Arachnida" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="319" phylum="Arthropoda" rank="order">Pseudoscorpiones</taxonomicName>
Catalog 2022
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>