688 lines
87 KiB
XML
688 lines
87 KiB
XML
<document id="A9C3C6E3C7272C219BA4918B8B90A6E3" ID-DOI="10.11646/zootaxa.5293.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="7960232" ID-ZooBank="F68B3A5C-E100-4F7A-A1A1-FC9933403700" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1684798263703" checkinUser="plazi" docAuthor="Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder & Lira, André Felipe De Araújo" docDate="2023" docId="0389886AE943FFB61399109DFD2D2110" docLanguage="en" docName="zootaxa.5293.2.6.pdf" docOrigin="Zootaxa 5293 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5293.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Progarypus smaugi Bedoya-Roqueme & Tizo-Pedroso & Barbier & Lira 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="327" masterDocId="FFB0F012E94BFFBD130E1314FF832309" masterDocTitle="Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil" masterLastPageNumber="332" masterPageNumber="317" pageNumber="325" updateTime="1684800287596" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="10DC6A40F5BC303982C9AB7360F23175">
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<mods:title id="FFE2B00434A3DBE56ADDC9B47DA96740">Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil</mods:title>
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<mods:namePart id="EF4A26F42DD02D8DFF3B57206312562D">Bedoya-Roqueme, Edwin</mods:namePart>
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<mods:affiliation id="2D1ADFE89A09091492FC7B5C0DBA0232">Programa de Pós-graduação em Recursos Naturais do Cerrado, Laboratório de Ecologia Comportamental de Aracnídeos, Universidade Estadual de Goiás, Anápolis, Brazil.</mods:affiliation>
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<mods:namePart id="8B3CEBB98C82C9DD43965EA9E253DB8E">Tizo-Pedroso, Everton</mods:namePart>
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<mods:affiliation id="2B7EC81ACAE094641CC3DD2E87FA0BC3">Laboratório de Ecologia Comportamental de Aracnídeos, Programa de Pós-graduação em Recursos Naturais do Cerrado, Centro de Ensino e Aprendizagem em Rede, Universidade Estadual de Goiás, Anápolis, Brazil.</mods:affiliation>
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<mods:namePart id="F7725729717BE84EBDFD41FF21F52268">Barbier, Eder</mods:namePart>
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<mods:affiliation id="5941481696F92DB4211620F3ED20E30D">Programa de Pós-graduação em Biologia Animal, Departamento de Zoologia, Universidade Federal de Pernambuco, Recife, Brazil. barbier. eder @ gmail. com; http: // orcid. org / 0000 - 0001 - 5068 - 7048</mods:affiliation>
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<mods:nameIdentifier id="67DE9CBDB9ABAF5F817E8A2A778A7848" type="email">barbier.eder@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FA37732AC1A15B072A84A6E96BBCA5E1">Lira, André Felipe De Araújo</mods:namePart>
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<mods:affiliation id="2222D30DA0236A86CC678FE4FB014B06">Centro de Educação e Saúde, Universidade Federal de Campina Grande, Cuité, Brazil.</mods:affiliation>
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<mods:title id="FFEBC8DCB8CDD86293410A77E9C7A04C">Zootaxa</mods:title>
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<mods:date id="B625D7C6C07F99885F24F7AE68512861">2023</mods:date>
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<mods:number id="C9799FD1EE2F7412A25D615D1A254F3D">2023-05-22</mods:number>
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<treatment id="0389886AE943FFB61399109DFD2D2110" ID-DOI="http://doi.org/10.5281/zenodo.7960274" ID-Zenodo-Dep="7960274" LSID="urn:lsid:plazi:treatment:0389886AE943FFB61399109DFD2D2110" httpUri="http://treatment.plazi.org/id/0389886AE943FFB61399109DFD2D2110" lastPageId="11" lastPageNumber="327" pageId="8" pageNumber="325">
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<taxonomicName id="4C2042FFE943FFB51399109DFE0020AB" authority="Bedoya-Roqueme & Tizo-Pedroso & Barbier & Lira, 2023" authorityName="Bedoya-Roqueme & Tizo-Pedroso & Barbier & Lira" authorityYear="2023" box="[151,387,904,931]" class="Arachnida" family="Hesperolpiidae" genus="Progarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="8" pageNumber="325" phylum="Arthropoda" rank="species" species="smaugi" status="sp. nov.">
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<emphasis id="B954E56EE943FFB51399109DFE0020AB" bold="true" box="[151,387,904,931]" italics="true" pageId="8" pageNumber="325">Progarypus smaugi</emphasis>
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<taxonomicNameLabel id="A2675815E943FFB51284109DFE6B20AA" box="[394,488,905,931]" pageId="8" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="131B25F9E943FFB5139910B8FE8220CE" box="[151,257,940,967]" captionStart="FIGURES 1–4" captionStartId="5.[152,270,1492,1516]" captionTargetBox="[170,1417,558,1466]" captionTargetId="figure-182@5.[170,1417,558,1466]" captionTargetPageId="5" captionText="FIGURES 1–4. Habitus: 1–2, Geogarypus gollumi sp. nov.; 1, female holotype, dorsal view; 2, male paratype, dorsal view. Habitus: 3–4, Progarypus smaugi sp. nov.; 3, female paratype, dorsal view; 4, male holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7960234" httpUri="https://zenodo.org/record/7960234/files/figure.png" pageId="8" pageNumber="325">
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Figs. 3
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<emphasis id="B954E56EE943FFB513ED10B9FF7220CE" bold="true" box="[227,241,941,967]" pageId="8" pageNumber="325">–</emphasis>
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4
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,
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<figureCitation id="131B25F9E943FFB5120310B8FED020CE" box="[269,339,940,967]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="8" pageNumber="325">
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15
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<emphasis id="B954E56EE943FFB5122710B9FEB420CE" bold="true" box="[297,311,941,967]" pageId="8" pageNumber="325">–</emphasis>
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24
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<emphasis id="B954E56EE943FFB513991705FEFC2722" bold="true" box="[151,383,1041,1067]" pageId="8" pageNumber="325">Material examined.</emphasis>
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, E
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. Barbier leg. (
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, E
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. Barbier leg. (
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||
<collectionCode id="ED31A1B9E943FFB51644174EFA1B277D" box="[1354,1432,1114,1140]" pageId="8" pageNumber="325">LECA</collectionCode>
|
||
; Ps-017);
|
||
<specimenCount id="9D26F2F5E943FFB513F0176AFEED2791" box="[254,366,1150,1176]" pageId="8" pageNumber="325" type="female">4 females</specimenCount>
|
||
,
|
||
<collectingRegion id="49E4F79EE943FFB51275176AFD892791" box="[379,522,1150,1176]" country="Brazil" name="Pernambuco" pageId="8" pageNumber="325">Pernambuco</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BFBA306E943FFB51118176AFD352791" box="[534,694,1150,1176]" pageId="8" pageNumber="325">Meu Rei cave</collectingMunicipality>
|
||
,
|
||
<geoCoordinate id="EE145FBBE943FFB511CD176AFCDC2791" box="[707,863,1150,1176]" degrees="08" direction="south" minutes="29" orientation="latitude" pageId="8" pageNumber="325" precision="1" seconds="14.1" value="-8.48725">08°29’14.1”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE145FBBE943FFB51062176AFB932791" box="[876,1040,1150,1176]" degrees="37" direction="west" minutes="16" orientation="longitude" pageId="8" pageNumber="325" precision="1" seconds="48.8" value="-37.280224">37°16’48.8”W</geoCoordinate>
|
||
,
|
||
<elevation id="000DDE4FE943FFB51713176AFB202791" box="[1053,1187,1150,1176]" metricMagnitude="2" metricUnit="m" metricValue="7.77" pageId="8" pageNumber="325" unit="m" value="777.0">
|
||
<quantity id="4CD89499E943FFB51713176AFBE62791" box="[1053,1125,1150,1176]" metricMagnitude="2" metricUnit="m" metricValue="7.77" pageId="8" pageNumber="325" unit="m" value="777.0">777 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FF9E1FBCE943FFB517BE176AFAA02791" box="[1200,1315,1150,1176]" pageId="8" pageNumber="325" value="2016-01-19">
|
||
<collectingDate id="EFDAE654E943FFB517BE176AFAA02791" box="[1200,1315,1150,1176]" pageId="8" pageNumber="325" value="2016-01-19">19-I-2016</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26D55CAAE943FFB51621176AFED727B5" pageId="8" pageNumber="325">
|
||
<collectingCounty id="62FE41F0E943FFB51621176AFEB627B5" pageId="8" pageNumber="325">Catimbau National Park</collectingCounty>
|
||
, E
|
||
</collectorName>
|
||
. Barbier leg. (
|
||
<collectionCode id="ED31A1B9E943FFB512FC17B6FDC327B5" box="[498,576,1186,1212]" pageId="8" pageNumber="325">LECA</collectionCode>
|
||
; Ps-018)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6AF7E943FFB713C917D1FEFD2030" lastPageId="10" lastPageNumber="327" pageId="8" pageNumber="325" type="diagnosis">
|
||
<paragraph id="8B9F397CE943FFB513C917D1FEB92679" blockId="8.[151,1437,1041,2041]" pageId="8" pageNumber="325">
|
||
<emphasis id="B954E56EE943FFB513C917D1FEC127D6" bold="true" box="[199,322,1221,1247]" pageId="8" pageNumber="325">Diagnosis.</emphasis>
|
||
Distribution of the trichobothria on the fixed finger chelal, being trichobothrium
|
||
<emphasis id="B954E56EE943FFB517E717D3FA8027E9" box="[1257,1283,1223,1248]" italics="true" pageId="8" pageNumber="325">ist</emphasis>
|
||
distal de
|
||
<emphasis id="B954E56EE943FFB5167917D3FA1E27E9" box="[1399,1437,1223,1248]" italics="true" pageId="8" pageNumber="325">est.</emphasis>
|
||
Trichobothrium
|
||
<emphasis id="B954E56EE943FFB5124017FFFEDE260D" box="[334,349,1259,1284]" italics="true" pageId="8" pageNumber="325">it</emphasis>
|
||
proximal to
|
||
<emphasis id="B954E56EE943FFB512FA17FFFD91260D" box="[500,530,1259,1284]" italics="true" pageId="8" pageNumber="325">est</emphasis>
|
||
;
|
||
<emphasis id="B954E56EE943FFB5112B17FEFDB9260D" box="[549,570,1258,1284]" italics="true" pageId="8" pageNumber="325">ib</emphasis>
|
||
near the level of
|
||
<emphasis id="B954E56EE943FFB5100417FEFCB2260D" box="[778,817,1258,1284]" italics="true" pageId="8" pageNumber="325">isb.</emphasis>
|
||
Nodus ramosus on movable finger chelal proximal to trichobothria
|
||
<emphasis id="B954E56EE943FFB51221161BFEB52621" box="[303,310,1295,1320]" italics="true" pageId="8" pageNumber="325">t</emphasis>
|
||
; the robust pedipalps, male femur
|
||
<geoCoordinate id="EE145FBBE943FFB511C0161AFC882621" box="[718,779,1294,1320]" degrees="0.604" direction="north" orientation="latitude" pageId="8" pageNumber="325" precision="55" value="0.604">0.604</geoCoordinate>
|
||
/
|
||
<quantity id="4CD89499E943FFB5101A161AFC092621" box="[788,906,1294,1320]" metricMagnitude="-4" metricUnit="m" metricValue="2.29" pageId="8" pageNumber="325" unit="mm" value="0.229">
|
||
<geoCoordinate id="EE145FBBE943FFB5101A161AFCD22621" box="[788,849,1294,1320]" degrees="0.229" direction="east" orientation="longitude" pageId="8" pageNumber="325" precision="55" value="0.229">0.229</geoCoordinate>
|
||
mm
|
||
</quantity>
|
||
, 2.6× (female,
|
||
<geoCoordinate id="EE145FBBE943FFB51732161AFBFA2620" box="[1084,1145,1294,1321]" degrees="0.681" direction="north" orientation="latitude" pageId="8" pageNumber="325" precision="55" value="0.681">0.681</geoCoordinate>
|
||
/
|
||
<quantity id="4CD89499E943FFB5178F161AFB7B2621" box="[1153,1272,1294,1321]" metricMagnitude="-4" metricUnit="m" metricValue="2.51" pageId="8" pageNumber="325" unit="mm" value="0.251">
|
||
<geoCoordinate id="EE145FBBE943FFB5178F161AFB3D2620" box="[1153,1214,1294,1321]" degrees="0.251" direction="east" orientation="longitude" pageId="8" pageNumber="325" precision="55" value="0.251">0.251</geoCoordinate>
|
||
mm
|
||
</quantity>
|
||
, 2.7×), patella
|
||
<geoCoordinate id="EE145FBBE943FFB513991626FF572644" box="[151,212,1330,1357]" degrees="0.485" direction="north" orientation="latitude" pageId="8" pageNumber="325" precision="55" value="0.485">0.485</geoCoordinate>
|
||
/
|
||
<quantity id="4CD89499E943FFB513D21626FED32645" box="[220,336,1330,1357]" metricMagnitude="-4" metricUnit="m" metricValue="2.09" pageId="8" pageNumber="325" unit="mm" value="0.209">
|
||
<geoCoordinate id="EE145FBBE943FFB513D21626FE9A2644" box="[220,281,1330,1357]" degrees="0.209" direction="east" orientation="longitude" pageId="8" pageNumber="325" precision="55" value="0.209">0.209</geoCoordinate>
|
||
mm
|
||
</quantity>
|
||
, 2.3× (female,
|
||
<geoCoordinate id="EE145FBBE943FFB512F41626FDB42644" box="[506,567,1330,1357]" degrees="0.552" direction="north" orientation="latitude" pageId="8" pageNumber="325" precision="55" value="0.552">0.552</geoCoordinate>
|
||
/
|
||
<quantity id="4CD89499E943FFB511311626FD302645" box="[575,691,1330,1357]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="325" unit="mm" value="0.25">
|
||
<geoCoordinate id="EE145FBBE943FFB511311626FDFF2644" box="[575,636,1330,1357]" degrees="0.250" direction="east" orientation="longitude" pageId="8" pageNumber="325" precision="55" value="0.25">0.250</geoCoordinate>
|
||
mm
|
||
</quantity>
|
||
, 2.5×). Fixed finger with 39 pointed teeth, movable finger with 30 flattened teeth.
|
||
</paragraph>
|
||
<paragraph id="8B9F397CE943FFB513C9166DFB8526D5" blockId="8.[151,1437,1041,2041]" pageId="8" pageNumber="325">
|
||
<emphasis id="B954E56EE943FFB513C9166DFE95269A" bold="true" box="[199,278,1401,1427]" pageId="8" pageNumber="325">Adults</emphasis>
|
||
. Carapace (
|
||
<figureCitation id="131B25F9E943FFB512A8166EFD94269D" box="[422,535,1402,1428]" captionStart="FIGURES 1–4" captionStartId="5.[152,270,1492,1516]" captionTargetBox="[170,1417,558,1466]" captionTargetId="figure-182@5.[170,1417,558,1466]" captionTargetPageId="5" captionText="FIGURES 1–4. Habitus: 1–2, Geogarypus gollumi sp. nov.; 1, female holotype, dorsal view; 2, male paratype, dorsal view. Habitus: 3–4, Progarypus smaugi sp. nov.; 3, female paratype, dorsal view; 4, male holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7960234" httpUri="https://zenodo.org/record/7960234/files/figure.png" pageId="8" pageNumber="325">Figs. 3–4</figureCitation>
|
||
), and pedipalps reddish-brown, carapace in both male and female with pale posterior area, more evident in males than in females, all legs light yellow, all tergites divided (except XI), anterior tergites light-brown colors, chela pedipalpal darker reddish-brown (
|
||
<figureCitation id="131B25F9E943FFB5108316D6FC7926D5" box="[909,1018,1474,1500]" captionStart="FIGURES 1–4" captionStartId="5.[152,270,1492,1516]" captionTargetBox="[170,1417,558,1466]" captionTargetId="figure-182@5.[170,1417,558,1466]" captionTargetPageId="5" captionText="FIGURES 1–4. Habitus: 1–2, Geogarypus gollumi sp. nov.; 1, female holotype, dorsal view; 2, male paratype, dorsal view. Habitus: 3–4, Progarypus smaugi sp. nov.; 3, female paratype, dorsal view; 4, male holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7960234" httpUri="https://zenodo.org/record/7960234/files/figure.png" pageId="8" pageNumber="325">Figs. 3–4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F397CE943FFB513C916F2FB242541" blockId="8.[151,1437,1041,2041]" pageId="8" pageNumber="325">
|
||
Chelicera with five setae on hand, fixed finger with one apical tooth, and four subapical teeth, movable finger with tooth-like subapical lobe (
|
||
<figureCitation id="131B25F9E943FFB512FB151EFDC4252D" box="[501,583,1546,1572]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="8" pageNumber="325">Fig. 16</figureCitation>
|
||
), rallum with three blade, first and second long toothed (
|
||
<figureCitation id="131B25F9E943FFB517C8151EFA9B252D" box="[1222,1304,1546,1572]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="8" pageNumber="325">Fig. 17</figureCitation>
|
||
), gale long, with three apical branches (
|
||
<figureCitation id="131B25F9E943FFB512C2153AFD9D2541" box="[460,542,1582,1608]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="8" pageNumber="325">Fig. 18</figureCitation>
|
||
), serrula exterior with 18 blades (female, with 20 blades).
|
||
</paragraph>
|
||
<paragraph id="8B9F397CE943FFB513C91546FB602461" blockId="8.[151,1437,1041,2041]" pageId="8" pageNumber="325">
|
||
Pedipalps robust (
|
||
<figureCitation id="131B25F9E943FFB5129B1546FDA32565" box="[405,544,1618,1644]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="8" pageNumber="325">Figs. 19–20</figureCitation>
|
||
), in both male and female strongly granulated, trochanter 1.3× (female, 1.2×) as long as broad, femur 2.6× (female 2.7×) as long as broad, chela with pedicel 3.2× (female 3.4×), chela without pedicel 3.0× (female 3.2×), inner margin of hand in female rounded (
|
||
<figureCitation id="131B25F9E943FFB510B8158EFB8E25BD" box="[950,1037,1690,1716]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="8" pageNumber="325">Fig. 20</figureCitation>
|
||
). Movable finger chelal
|
||
<quantity id="4CD89499E943FFB51626158EFA1E25BD" box="[1320,1437,1690,1717]" metricMagnitude="-4" metricUnit="m" metricValue="5.18" pageId="8" pageNumber="325" unit="mm" value="0.518">0.518 mm</quantity>
|
||
(female
|
||
<quantity id="4CD89499E943FFB513F915AAFEF325D1" box="[247,368,1726,1752]" metricMagnitude="-4" metricUnit="m" metricValue="6.41" pageId="8" pageNumber="325" unit="mm" value="0.641">0.641 mm</quantity>
|
||
), fixed finger with 39 pointed teeth (female with 37 pointed teeth), movable finger with 30 flattened teeth (female with 32 flattened teeth) (
|
||
<figureCitation id="131B25F9E943FFB511A915F6FD7925F5" box="[679,762,1762,1788]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="8" pageNumber="325">Fig. 21</figureCitation>
|
||
). Trichobothrium
|
||
<emphasis id="B954E56EE943FFB510CD15F7FC5E25F5" box="[963,989,1763,1788]" italics="true" pageId="8" pageNumber="325">ist</emphasis>
|
||
distal de
|
||
<emphasis id="B954E56EE943FFB5174915F7FBE525F5" box="[1095,1126,1763,1788]" italics="true" pageId="8" pageNumber="325">est</emphasis>
|
||
; trichobothrium
|
||
<emphasis id="B954E56EE943FFB5162C15F7FAB125F5" box="[1314,1330,1763,1788]" italics="true" pageId="8" pageNumber="325">it</emphasis>
|
||
proximal to
|
||
<emphasis id="B954E56EE943FFB513BA1413FF502429" box="[180,211,1799,1824]" italics="true" pageId="8" pageNumber="325">est</emphasis>
|
||
;
|
||
<emphasis id="B954E56EE943FFB513EF1412FF742429" box="[225,247,1798,1824]" italics="true" pageId="8" pageNumber="325">ib</emphasis>
|
||
near the level of
|
||
<emphasis id="B954E56EE943FFB512B61412FE5A2429" box="[440,473,1798,1824]" italics="true" pageId="8" pageNumber="325">isb</emphasis>
|
||
; trichobothrium
|
||
<emphasis id="B954E56EE943FFB511981413FD2A2429" box="[662,681,1799,1824]" italics="true" pageId="8" pageNumber="325">st</emphasis>
|
||
proximal to the middle of the finger,
|
||
<emphasis id="B954E56EE943FFB517451412FBE72429" box="[1099,1124,1798,1824]" italics="true" pageId="8" pageNumber="325">sb</emphasis>
|
||
closer to
|
||
<emphasis id="B954E56EE943FFB517DF1412FB5C2429" box="[1233,1247,1798,1824]" italics="true" pageId="8" pageNumber="325">b</emphasis>
|
||
than to
|
||
<emphasis id="B954E56EE943FFB516371413FACF2429" box="[1337,1356,1799,1824]" italics="true" pageId="8" pageNumber="325">st</emphasis>
|
||
, nodus ramosus on fixed finger distal of
|
||
<emphasis id="B954E56EE943FFB51102143FFDA8244D" box="[524,555,1835,1860]" italics="true" pageId="8" pageNumber="325">est</emphasis>
|
||
, and in the movable finger chelal proximal to trichobothrium
|
||
<emphasis id="B954E56EE943FFB517ED143FFB68244D" box="[1251,1259,1835,1860]" italics="true" pageId="8" pageNumber="325">t</emphasis>
|
||
(
|
||
<figureCitation id="131B25F9E943FFB517F5143EFACE244D" box="[1275,1357,1834,1860]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="8" pageNumber="325">Fig. 21</figureCitation>
|
||
), well-developed venom apparatus in both on the fixed finger and on the movable finger chelal (
|
||
<figureCitation id="131B25F9E943FFB5178C145AFB562461" box="[1154,1237,1870,1896]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="8" pageNumber="325">Fig. 21</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F397CE943FFB513C91466FC3024F0" blockId="8.[151,1437,1041,2041]" pageId="8" pageNumber="325">
|
||
Carapace triangular (
|
||
<figureCitation id="131B25F9E943FFB512B81466FD8A2484" box="[438,521,1906,1933]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="8" pageNumber="325">Fig. 15</figureCitation>
|
||
), squamose sculpturing, posteriorly reticulate, cucullus shorts, with an indistinct longitudinal furrow, with medial furrow and indistinct posterior furrow (
|
||
<figureCitation id="131B25F9E943FFB510B91482FB8B24B8" box="[951,1032,1942,1969]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="8" pageNumber="325">Fig. 15</figureCitation>
|
||
), four large eyes, all setae very short and acuminate, with approx. 38 setae (female approx. 36 setae), four setae on the anterior margin, one seta preocular on each side, with six to twelve setae on the posterior margin (
|
||
<figureCitation id="131B25F9E943FFB5105C14CAFC2524F0" box="[850,934,2014,2041]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="8" pageNumber="325">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF5F69F4E942FFB413961432FD1724A2" ID-DOI="http://doi.org/10.5281/zenodo.7960240" ID-Zenodo-Dep="7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="9" pageNumber="326" startId="9.[152,270,1830,1854]" targetBox="[187,1367,192,1797]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8B9F397CE942FFB413961432FD1724A2" blockId="9.[151,1437,1830,1963]" pageId="9" pageNumber="326">
|
||
<emphasis id="B954E56EE942FFB413961432FED92437" bold="true" box="[152,346,1830,1854]" pageId="9" pageNumber="326">FIGURES 15–24.</emphasis>
|
||
<taxonomicName id="4C2042FFE942FFB4126E1433FDA52437" authority="Bedoya-Roqueme & Tizo-Pedroso & Barbier & Lira, 2023" authorityName="Bedoya-Roqueme & Tizo-Pedroso & Barbier & Lira" authorityYear="2023" box="[352,550,1831,1854]" class="Arachnida" family="Hesperolpiidae" genus="Progarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="326" phylum="Arthropoda" rank="species" species="smaugi" status="sp. nov.">
|
||
<emphasis id="B954E56EE942FFB4126E1433FDA52437" box="[352,550,1831,1854]" italics="true" pageId="9" pageNumber="326">Progarypus smaugi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954E56EE942FFB411221433FDFE2436" bold="true" box="[556,637,1831,1855]" pageId="9" pageNumber="326">
|
||
<taxonomicNameLabel id="A2675815E942FFB411221433FDFE2436" box="[556,637,1831,1855]" pageId="9" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F397CE941FFB713C91383FB3023F0" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="327">Coxal region: female and male with an apical lobe of coxa pedipalpal with two setae marginal, and three discal setae, approx. 26 setae (female approx. 28), coxa I ca. 22 setae (female ca. 26 setae), Coxa II ca. 24 setae (female ca. 28), Coxa III ca. 30 setae (female ca. 32 setae), Coxa IV ca. 34 setae (female ca. 36 setae).</paragraph>
|
||
<paragraph id="8B9F397CE941FFB713C91217FBEA22FC" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="327">
|
||
Opisthosoma: all tergites are divided (except XI); reticulate sculpture, all setae acuminate, pleural membrane strongly striated. Tergal chaetotaxy of
|
||
<typeStatus id="549B87DEE941FFB711451233FD2E2248" box="[587,685,295,321]" pageId="10" pageNumber="327">holotype</typeStatus>
|
||
male, tergites I–XI: 6: 8: 8: 8: 6: 6: 4: 4: 6: 6: 4 [4ST]: 2, females
|
||
<typeStatus id="549B87DEE941FFB71399125FFE86226C" box="[151,261,331,357]" pageId="10" pageNumber="327">paratypes</typeStatus>
|
||
, tergites I–XI: 6: 6–8: 6–8: 8: 6: 6: 6–4: 6–4: 6: 6: 4 [4ST]: 2, males
|
||
<typeStatus id="549B87DEE941FFB7170E125FFBED226C" box="[1024,1134,331,357]" pageId="10" pageNumber="327">paratypes</typeStatus>
|
||
, tergites I–XI: 6: 6: 8: 8: 6: 6–8: 6–8: 6–8: 6: 6–8: 4 [4ST]: 2, tergite XI with four setae tactile. Sternal chaetotaxy of the male
|
||
<typeStatus id="549B87DEE941FFB717DE127BFAB62280" box="[1232,1333,367,393]" pageId="10" pageNumber="327">holotype</typeStatus>
|
||
, sternites II–XII: 8(4): (1)6(1): (1)6(1): 8: 8: 8: 8: 8: 10: 10: 2, females
|
||
<typeStatus id="549B87DEE941FFB710581287FC4722A4" box="[854,964,403,429]" pageId="10" pageNumber="327">paratypes</typeStatus>
|
||
, sternites II–XII: 6: (1)6(1): (1)6(1): 6–8: 6–8: 8: 8–10: 8–10: 10–12: 10: 2, males
|
||
<typeStatus id="549B87DEE941FFB7115812A3FD4722D8" box="[598,708,439,465]" pageId="10" pageNumber="327">paratypes</typeStatus>
|
||
, sternites II–XII: 8(4): (1)6(1): (1)6(1): 6: 6–8: 8–10: 8–10: 8–10: 10–12: 10: 2, sternites XI with two setae tactile, pleural membrane III–X with one seta.
|
||
</paragraph>
|
||
<paragraph id="8B9F397CE941FFB713C912EBFE4E218C" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="327">
|
||
The genital region of the male
|
||
<typeStatus id="549B87DEE941FFB7111112EBFD022110" box="[543,641,511,537]" pageId="10" pageNumber="327">holotype</typeStatus>
|
||
with eighth setae, and four discal setae on the anterior operculum (
|
||
<figureCitation id="131B25F9E941FFB7166112EBFF352134" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="10" pageNumber="327">Fig. 22</figureCitation>
|
||
), the posterior operculum with six marginal setae (
|
||
<figureCitation id="131B25F9E941FFB711F91137FCC82134" box="[759,843,547,573]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="10" pageNumber="327">Fig. 22</figureCitation>
|
||
), internal genital structures, indistinctly visible (
|
||
<figureCitation id="131B25F9E941FFB716611137FF342168" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="10" pageNumber="327">Fig. 22</figureCitation>
|
||
). Genital region of the female
|
||
<typeStatus id="549B87DEE941FFB7111F1153FDF22168" box="[529,625,583,609]" pageId="10" pageNumber="327">paratype</typeStatus>
|
||
with six central setae on the anterior operculum, and six marginal setae on the posterior operculum.
|
||
</paragraph>
|
||
<paragraph id="8B9F397CE941FFB713C9119BFEFD2030" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="327">
|
||
Legs: all legs light-yellow (
|
||
<figureCitation id="131B25F9E941FFB7110D119BFD0C21A0" box="[515,655,655,681]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="10" pageNumber="327">Figs. 23–24</figureCitation>
|
||
), Leg I (
|
||
<figureCitation id="131B25F9E941FFB711FD119BFCCA21A0" box="[755,841,655,681]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="10" pageNumber="327">Fig. 23</figureCitation>
|
||
): trochanter 1.5× (female 1.7×), femur 2.8× (female 2.7×), patella 2.4× (female 2.6×), tibia 2.4× (female 2.6×), metatarsus 3.2× (female 3.3×), tarsus 3.3× (female 4.8×), Leg IV (
|
||
<figureCitation id="131B25F9E941FFB7124F11C3FE1521F8" box="[321,406,727,753]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="10" pageNumber="327">Fig. 24</figureCitation>
|
||
): trochanter 1.1× (female 1.2×), femur+patella 2.6× (female 2.7×), tibia 4.0× (female 4.1×), metatarsus 3.1× (female 3.2×), tarsus 3.7× (female 3.8×), subterminal setae smooth, arolium undivided and longer than claws (
|
||
<figureCitation id="131B25F9E941FFB71213100BFEF12030" box="[285,370,799,825]" captionStart="FIGURES 15–24" captionStartId="9.[152,270,1830,1854]" captionTargetBox="[187,1367,192,1797]" captionTargetId="figure-19@9.[170,1417,181,1806]" captionTargetPageId="9" captionText="FIGURES 15–24. Progarypus smaugi sp. nov. male holotype, dorsal view, unless stated otherwise: 15, carapace, dorsal view. 16, left chelicera, dorsal view. 17, left galea, lateral view. 18, left rallum, lateral view. 19, male holotype, right pedipalp aspect. 20, female paratype, right pedipalp aspect. 21, right chela, lateral view. 22, genital sternites, male genitalia, ventral view. 23–24, left leg I, lateral view, and left leg IV, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7960240" httpUri="https://zenodo.org/record/7960240/files/figure.png" pageId="10" pageNumber="327">Fig. 24</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6AF7E941FFB713C91057FC37265C" pageId="10" pageNumber="327" type="description">
|
||
<paragraph id="8B9F397CE941FFB713C91057FD8F2750" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="327">
|
||
<emphasis id="B954E56EE941FFB713C91057FE3F2055" bold="true" box="[199,444,834,861]" pageId="10" pageNumber="327">Measurements (mm)</emphasis>
|
||
:Male
|
||
<typeStatus id="549B87DEE941FFB7110B1057FDE42054" box="[517,615,835,861]" pageId="10" pageNumber="327">holotype</typeStatus>
|
||
(
|
||
<typeStatus id="549B87DEE941FFB7117C1057FD552054" box="[626,726,835,861]" pageId="10" pageNumber="327">Paratype</typeStatus>
|
||
in parentheses):body length 1.796(1.813); carapace
|
||
<geoCoordinate id="EE145FBBE941FFB716181057FAD62054" box="[1302,1365,835,861]" degrees="0.609" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.609">0.609</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB716531057FA1F2054" box="[1373,1436,835,861]" degrees="0.486" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.486">0.486</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB713911073FF622088" box="[159,225,871,897]" degrees="0.612" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.612">0.612</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB713E41073FEAF2088" box="[234,300,871,897]" degrees="0.489" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.489">0.489</geoCoordinate>
|
||
); Chelicerae; movable finger. pedipalps trochanter:
|
||
<geoCoordinate id="EE145FBBE941FFB7108B1072FC472088" box="[901,964,870,897]" degrees="0.255" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.255">0.255</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB710C31072FB882088" box="[973,1035,870,897]" degrees="0.191" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.191">0.191</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB717101072FBE32088" box="[1054,1120,870,897]" degrees="0.257" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.257">0.257</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB717671072FB292088" box="[1129,1194,870,897]" degrees="0.193" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.193">0.193</geoCoordinate>
|
||
); femur:
|
||
<geoCoordinate id="EE145FBBE941FFB716181073FAD62088" box="[1302,1365,871,897]" degrees="0.604" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.604">0.604</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB716501073FA1F2088" box="[1374,1436,871,897]" degrees="0.229" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.229">0.229</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB71391109FFF6320AC" box="[159,224,907,933]" degrees="0.607" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.607">0.607</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB713E7109FFEA820AC" box="[233,299,907,933]" degrees="0.230" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.23">0.230</geoCoordinate>
|
||
); patella
|
||
<geoCoordinate id="EE145FBBE941FFB71292109EFE5920AC" box="[412,474,906,933]" degrees="0.485" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.485">0.485</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB712EC109EFDA220AC" box="[482,545,906,933]" degrees="0.209" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.209">0.209</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB71138109FFDF420AC" box="[566,631,907,933]" degrees="0.487" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.487">0.487</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB7118E109FFD4120AC" box="[640,706,907,933]" degrees="0.210" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.21">0.210</geoCoordinate>
|
||
); chela with pedicel
|
||
<geoCoordinate id="EE145FBBE941FFB710B1109FFC7D20AC" box="[959,1022,907,933]" degrees="0.929" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.929">0.929</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB71708109FFBC620AC" box="[1030,1093,907,933]" degrees="0.289" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.289">0.289</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB71754109FFB1920AC" box="[1114,1178,907,933]" degrees="0.931" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.931">0.931</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB717AD109FFB6520AC" box="[1187,1254,907,933]" degrees="0.290" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.29">0.290</geoCoordinate>
|
||
); chela without pedicel
|
||
<geoCoordinate id="EE145FBBE941FFB713FA10BBFEB120C0" box="[244,306,943,969]" degrees="0.873" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.873">0.873</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB7123410BBFEFA20C0" box="[314,377,943,969]" degrees="0.289" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.289">0.289</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB7128310BAFE4D20C0" box="[397,462,942,969]" degrees="0.853" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.853">0.853</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB712D910BAFD9A20C0" box="[471,537,942,969]" degrees="0.290" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.29">0.290</geoCoordinate>
|
||
), movable finger 0.578 (0.581). Leg I: trochanter
|
||
<geoCoordinate id="EE145FBBE941FFB7176510BBFB2920C0" box="[1131,1194,943,969]" degrees="0.136" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.136">0.136</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB717BD10BBFB7120C0" box="[1203,1266,943,969]" degrees="0.129" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.129">0.129</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB7160B10BBFAC420C0" box="[1285,1351,943,969]" degrees="0.137" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.137">0.137</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB7165E10BBFA1320C0" box="[1360,1424,943,969]" degrees="0.131" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.131">0.131</geoCoordinate>
|
||
), femur
|
||
<geoCoordinate id="EE145FBBE941FFB713EA10C7FEA020E4" box="[228,291,979,1005]" degrees="0.220" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.22">0.220</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB7122210C7FEE920E4" box="[300,362,979,1005]" degrees="0.077" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.077">0.077</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB7127210C6FE3E20E4" box="[380,445,978,1005]" degrees="0.225" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.225">0.225</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB712C810C6FD8B20E4" box="[454,520,978,1005]" degrees="0.079" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.079">0.079</geoCoordinate>
|
||
), patella
|
||
<geoCoordinate id="EE145FBBE941FFB7117D10C7FD3020E4" box="[627,691,979,1005]" degrees="0.149" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.149">0.149</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB711B210C7FD7920E4" box="[700,762,979,1005]" degrees="0.061" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.061">0.061</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB7100510C6FCCF20E4" box="[779,844,978,1005]" degrees="0.151" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.151">0.151</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB7105B10C6FC1420E4" box="[853,919,978,1005]" degrees="0.063" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.063">0.063</geoCoordinate>
|
||
), tibia
|
||
<geoCoordinate id="EE145FBBE941FFB710E410C6FBAA20E4" box="[1002,1065,978,1005]" degrees="0.159" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.159">0.159</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB7173F10C6FBEC20E4" box="[1073,1135,978,1005]" degrees="0.065" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.065">0.065</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB7178C10C7FB4020E4" box="[1154,1219,979,1005]" degrees="0.161" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.161">0.161</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB717C510C7FA8E20E4" box="[1227,1293,979,1005]" degrees="0.067" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.067">0.067</geoCoordinate>
|
||
); metatarsus
|
||
<geoCoordinate id="EE145FBBE941FFB7139910E3FF552718" box="[151,214,1015,1041]" degrees="0.131" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.131">0.131</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB713D010E3FE9E2718" box="[222,285,1015,1041]" degrees="0.041" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.041">0.041</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB7123B10E3FEF42718" box="[309,375,1015,1041]" degrees="0.133" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.133">0.133</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB7128E10E3FE412718" box="[384,450,1015,1041]" degrees="0.043" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.043">0.043</geoCoordinate>
|
||
); tarsus
|
||
<geoCoordinate id="EE145FBBE941FFB7112110E3FDED2718" box="[559,622,1015,1041]" degrees="0.134" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.134">0.134</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB7117910E3FD362718" box="[631,693,1015,1041]" degrees="0.043" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.043">0.043</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB711C310E2FC8C2718" box="[717,783,1014,1041]" degrees="0.136" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.136">0.136</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB7101610E2FCDA2718" box="[792,857,1014,1041]" degrees="0.045" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.045">0.045</geoCoordinate>
|
||
). Leg IV: trochanter
|
||
<geoCoordinate id="EE145FBBE941FFB7176810E3FB262718" box="[1126,1189,1015,1041]" degrees="0.231" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.231">0.231</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB717A310E3FB6F2718" box="[1197,1260,1015,1041]" degrees="0.131" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.131">0.131</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB7160A10E2FAC52718" box="[1284,1350,1014,1041]" degrees="0.235" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.235">0.235</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB7164110E2FA122718" box="[1359,1425,1014,1041]" degrees="0.133" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.133">0.133</geoCoordinate>
|
||
); femur+patella:
|
||
<geoCoordinate id="EE145FBBE941FFB7124A170FFE01273C" box="[324,386,1051,1077]" degrees="0.843" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.843">0.843</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB71284170FFE4A273C" box="[394,457,1051,1077]" degrees="0.180" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.18">0.180</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB712D7170EFD98273C" box="[473,539,1050,1077]" degrees="0.845" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.845">0.845</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB7112A170EFDE5273C" box="[548,614,1050,1077]" degrees="0.183" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.183">0.183</geoCoordinate>
|
||
); tibia:
|
||
<geoCoordinate id="EE145FBBE941FFB711B4170FFD7A273C" box="[698,761,1051,1077]" degrees="0.361" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.361">0.361</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB7100F170FFCC3273C" box="[769,832,1051,1077]" degrees="0.091" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.091">0.091</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB71041170EFC12273C" box="[847,913,1050,1077]" degrees="0.365" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.365">0.365</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB71094170EFC5F273C" box="[922,988,1050,1077]" degrees="0.093" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.093">0.093</geoCoordinate>
|
||
); metatarsus
|
||
<geoCoordinate id="EE145FBBE941FFB71761170FFB2C273C" box="[1135,1199,1051,1077]" degrees="0.219" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.219">0.219</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB717B9170FFB76273C" box="[1207,1269,1051,1077]" degrees="0.071" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.071">0.071</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB7160A170FFAC7273C" box="[1284,1348,1051,1077]" degrees="0.221" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.221">0.221</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB71643170FFA13273C" box="[1357,1424,1051,1077]" degrees="0.070" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.07">0.070</geoCoordinate>
|
||
); tarsus
|
||
<geoCoordinate id="EE145FBBE941FFB713EE172AFEA32750" box="[224,288,1086,1113]" degrees="0.189" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.189">0.189</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB71226172AFEE52750" box="[296,358,1086,1113]" degrees="0.051" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.051">0.051</geoCoordinate>
|
||
(
|
||
<geoCoordinate id="EE145FBBE941FFB7127A172AFE352750" box="[372,438,1086,1113]" degrees="0.187" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.187">0.187</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB712B1172AFD822750" box="[447,513,1086,1113]" degrees="0.053" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.053">0.053</geoCoordinate>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F397CE941FFB713C91777FC37265C" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="327">
|
||
<materialsCitation id="3B483321E941FFB713C91777FC33265C" collectionCode="IV" collectorName="Body & Carapace & Chelicerae & Pedipalps & Leg I & Leg" latitude="0.641" longLatPrecision="78" longitude="-0.643" pageId="10" pageNumber="327" specimenCount="1" typeStatus="paratype">
|
||
Female
|
||
<typeStatus id="549B87DEE941FFB7122F1777FE122774" box="[289,401,1123,1149]" pageId="10" pageNumber="327">paratypes</typeStatus>
|
||
:
|
||
<collectorName id="26D55CAAE941FFB712951777FE5B2774" box="[411,472,1123,1149]" pageId="10" pageNumber="327">Body</collectorName>
|
||
length
|
||
<geoCoordinate id="EE145FBBE941FFB711231776FDEF2774" box="[557,620,1122,1149]" degrees="2.241" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="2.241">2.241</geoCoordinate>
|
||
<geoCoordinate id="EE145FBBE941FFB711621776FD3E2774" box="[620,701,1122,1149]" degrees="2.250" direction="west" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="-2.25">–2.250</geoCoordinate>
|
||
.
|
||
<collectorName id="26D55CAAE941FFB711C91777FCAC2774" box="[711,815,1123,1149]" pageId="10" pageNumber="327">Carapace</collectorName>
|
||
0.691–0.693/0.552–0.553;
|
||
<collectorName id="26D55CAAE941FFB7176B1777FB632774" box="[1125,1248,1123,1149]" pageId="10" pageNumber="327">Chelicerae</collectorName>
|
||
; movable finger.
|
||
<collectorName id="26D55CAAE941FFB713991793FE8B27A8" box="[151,264,1159,1185]" pageId="10" pageNumber="327">Pedipalps</collectorName>
|
||
:trochanter 0.226–0.229/0.210–0.213; femur 0.681–0.683/0.251–0.253; patella 0.550–0.553/0.220–0.223; chela with pedicel1.091–1.093/0.321–0.323; chela without pedicel 1.031–1.033/0.321–0.323; movable finger
|
||
<geoCoordinate id="EE145FBBE941FFB7164117BFFA0E27CC" box="[1359,1421,1195,1221]" degrees="0.641" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.641">0.641</geoCoordinate>
|
||
<geoCoordinate id="EE145FBBE941FFB7168017BFFF5927E0" degrees="0.643" direction="west" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="-0.643">– 0.643</geoCoordinate>
|
||
.
|
||
<collectorName id="26D55CAAE941FFB713E617DBFEA927E0" box="[232,298,1231,1257]" pageId="10" pageNumber="327">Leg I</collectorName>
|
||
: trochanter 0.050–0.060/0.136–0.137; femur 0.263–0.265/0.080–0.090; patella 0.167–0.169/0.102– 0.103; tibia 0.215–0.217/0.080–0.090; metatarsus 0.203–0.206/0.060–0.070; tarsus 0.195–0.197/0.041–0.043.
|
||
<collectorName id="26D55CAAE941FFB7167F17E7FA1E2604" box="[1393,1437,1267,1293]" pageId="10" pageNumber="327">Leg</collectorName>
|
||
<collectionCode id="ED31A1B9E941FFB713991603FF342638" box="[151,183,1303,1329]" pageId="10" pageNumber="327">IV</collectionCode>
|
||
: trochanter 0.230–0.231/0.130–0.133; femur+patella 0.803–0.805/0.251–0.253; tibia 0.532–0.533/0.110–0.112; metatarsus 0.316–0.319/0.060–0.070; tarsus 0.249–0.251/0.040–0.050
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6AF7E941FFB713C9164BFAE72694" pageId="10" pageNumber="327" type="etymology">
|
||
<paragraph id="8B9F397CE941FFB713C9164BFAE72694" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="327">
|
||
<emphasis id="B954E56EE941FFB713C9164BFECD2670" bold="true" box="[199,334,1375,1401]" pageId="10" pageNumber="327">Etymology.</emphasis>
|
||
The specific epithet (
|
||
<taxonomicName id="4C2042FFE941FFB7114C164BFD102671" authorityName="Bedoya-Roqueme & Tizo-Pedroso & Barbier & Lira" authorityYear="2023" box="[578,659,1375,1400]" class="Arachnida" family="Hesperolpiidae" genus="Progarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="327" phylum="Arthropoda" rank="species" species="smaugi">
|
||
<emphasis id="B954E56EE941FFB7114C164BFD102671" box="[578,659,1375,1400]" italics="true" pageId="10" pageNumber="327">smaugi</emphasis>
|
||
</taxonomicName>
|
||
) derives its name from the mythical (fictional) dragon, the last of the great dragons remaining in Middle Earth, in the fantasy literary book “The Hobbit” written by J.R.R. Tolkien.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6AF7E941FFB713C916B2FB1626C8" box="[199,1173,1446,1473]" pageId="10" pageNumber="327" type="distribution">
|
||
<paragraph id="8B9F397CE941FFB713C916B2FB1626C8" blockId="10.[151,1437,151,2013]" box="[199,1173,1446,1473]" pageId="10" pageNumber="327">
|
||
<emphasis id="B954E56EE941FFB713C916B2FEE226C9" bold="true" box="[199,353,1446,1472]" pageId="10" pageNumber="327">Distribution.</emphasis>
|
||
Only known from the
|
||
<typeStatus id="549B87DEE941FFB7116E16B3FD1326C8" box="[608,656,1447,1473]" pageId="10" pageNumber="327">type</typeStatus>
|
||
locality (
|
||
<collectingCountry id="F33779ECE941FFB711F516B3FCC626C8" box="[763,837,1447,1473]" name="Brazil" pageId="10" pageNumber="327">Brazil</collectingCountry>
|
||
:
|
||
<collectingRegion id="49E4F79EE941FFB7104116B3FC5D26C8" box="[847,990,1447,1473]" country="Brazil" name="Pernambuco" pageId="10" pageNumber="327">Pernambuco</collectingRegion>
|
||
, Meu Rei cave).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6AF7E941FFB613C916DFFD2D2110" lastPageId="11" lastPageNumber="328" pageId="10" pageNumber="327" type="discussion">
|
||
<paragraph id="8B9F397CE941FFB613C916DFFD2D2110" blockId="10.[151,1437,151,2013]" lastBlockId="11.[151,1436,151,537]" lastPageId="11" lastPageNumber="328" pageId="10" pageNumber="327">
|
||
<emphasis id="B954E56EE941FFB713C916DFFEB826EC" bold="true" box="[199,315,1483,1509]" pageId="10" pageNumber="327">Remarks.</emphasis>
|
||
According to the descriptions by
|
||
<bibRefCitation id="EFB1448DE941FFB711BC16DFFCC726EC" author="Beier, M." box="[690,836,1483,1509]" pageId="10" pageNumber="327" pagination="49 - 56" refId="ref13486" refString="Beier, M. (1931 a) Zur Kenntnis der Chthoniiden (Pseudoskorpione). Zoologischer Anzeiger, 93, 49 - 56." type="journal article" year="1931">Beier (1931a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB1448DE941FFB7104116DFFC1426EC" author="Beier, M." box="[847,919,1483,1509]" pageId="10" pageNumber="327" pagination="1 - 294" refId="ref13581" refString="Beier, M. (1932 b) Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich, 58, i - xxi + 1 - 294." type="journal article" year="1932">1932b</bibRefCitation>
|
||
, 1959, 1964) and the taxonomic key presented by
|
||
<bibRefCitation id="EFB1448DE941FFB713B516FBFEEC2500" author="Mahnert, V." box="[187,367,1519,1545]" pageId="10" pageNumber="327" pagination="95 - 148" refId="ref14699" refString="Mahnert, V. (2001) Cave-dwelling pseudoscorpions (Arachnida, Pseudoscorpiones) from Brazil. Revue Suisse de Zoologie, 108, 95 - 148. https: // doi. org / 10.5962 / bhl. part. 79622" type="journal article" year="2001">Mahnert (2001)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C2042FFE941FFB7127216FBFDDB2501" authority="Bedoya-Roqueme & Tizo-Pedroso & Barbier & Lira, 2023" authorityName="Bedoya-Roqueme & Tizo-Pedroso & Barbier & Lira" authorityYear="2023" box="[380,600,1519,1544]" class="Arachnida" family="Hesperolpiidae" genus="Progarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="327" phylum="Arthropoda" rank="species" species="smaugi" status="sp. nov.">
|
||
<emphasis id="B954E56EE941FFB7127216FBFDDB2501" box="[380,600,1519,1544]" italics="true" pageId="10" pageNumber="327">Progarypus smaugi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954E56EE941FFB7116E16FBFD392500" bold="true" box="[608,698,1519,1545]" pageId="10" pageNumber="327">
|
||
<taxonomicNameLabel id="A2675815E941FFB7116E16FBFD392500" box="[608,698,1519,1545]" pageId="10" pageNumber="327" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from other species of
|
||
<taxonomicName id="4C2042FFE941FFB7170516FBFA9A2500" authority="Beier, 1931" authorityName="Beier" authorityYear="1931" box="[1035,1305,1519,1545]" class="Arachnida" family="Hesperolpiidae" genus="Progarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="327" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954E56EE941FFB7170516FBFB0D2501" box="[1035,1166,1519,1544]" italics="true" pageId="10" pageNumber="327">Progarypus</emphasis>
|
||
Beier, 1931
|
||
</taxonomicName>
|
||
as follows: it differs from the
|
||
<typeStatus id="549B87DEE941FFB712601507FE1D2524" box="[366,414,1555,1581]" pageId="10" pageNumber="327">type</typeStatus>
|
||
species of the genus
|
||
<taxonomicName id="4C2042FFE941FFB711971507FBBC2524" authority="(Balzan, 1887)" baseAuthorityName="Balzan" baseAuthorityYear="1887" box="[665,1087,1555,1581]" class="Arachnida" family="Hesperolpiidae" genus="Progarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="327" phylum="Arthropoda" rank="species" species="ramicola">
|
||
<emphasis id="B954E56EE941FFB711971507FC092524" box="[665,906,1555,1581]" italics="true" pageId="10" pageNumber="327">Progarypus ramicola</emphasis>
|
||
(Balzan, 1887)
|
||
</taxonomicName>
|
||
, from
|
||
<collectingCountry id="F33779ECE941FFB717831507FB752524" box="[1165,1270,1555,1581]" name="Paraguay" pageId="10" pageNumber="327">Paraguay</collectingCountry>
|
||
, for the femur pedipalpal without pedicel, clearly separated, somewhat concave, femur pedipalps
|
||
<geoCoordinate id="EE145FBBE941FFB717621523FB2F2558" box="[1132,1196,1591,1617]" degrees="0.720" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.72">0.720</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB717BB1523FB762558" box="[1205,1269,1591,1617]" degrees="0.160" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.16">0.160</geoCoordinate>
|
||
, patella 0.690 (0.540)/0.201, rallum of chelicera with three denticulate blades. It differs from
|
||
<taxonomicName id="4C2042FFE941FFB71707154FFAF9257C" authority="Beier, 1964" authorityName="Beier" authorityYear="1964" box="[1033,1402,1627,1653]" class="Arachnida" family="Hesperolpiidae" genus="Progarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="327" phylum="Arthropoda" rank="species" species="longipes">
|
||
<emphasis id="B954E56EE941FFB71707154FFB71257C" box="[1033,1266,1627,1653]" italics="true" pageId="10" pageNumber="327">Progarypus longipes</emphasis>
|
||
Beier, 1964
|
||
</taxonomicName>
|
||
by each tergite with six to eight very short marginal setae; pedipalps relatively very thinned, femur pedipalpal on the apical third with an extended tactile seta, scaly sculpture in the medial area and granular in the rest. Fixed finger chelal with 45 densely spaced teeth, finger movable chelal with 35 pointed teeth in apical third, but completely flat at the base, and
|
||
<taxonomicName id="4C2042FFE941FFB7127015FFFC97240C" authority="Beier, 1964" authorityName="Beier" authorityYear="1964" box="[382,788,1771,1797]" class="Arachnida" family="Hesperolpiidae" genus="Progarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="327" phylum="Arthropoda" rank="species" species="marginatus">
|
||
<emphasis id="B954E56EE941FFB7127015FFFD0A240D" box="[382,649,1771,1796]" italics="true" pageId="10" pageNumber="327">Progarypus marginatus</emphasis>
|
||
Beier, 1964
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F33779ECE941FFB7105515FFFC1B240C" box="[859,920,1771,1797]" name="Chile" pageId="10" pageNumber="327">Chile</collectingCountry>
|
||
by the
|
||
<emphasis id="B954E56EE941FFB710E115FFFBE1240C" box="[1007,1122,1771,1797]" italics="true" pageId="10" pageNumber="327">eb -esb -isb</emphasis>
|
||
trichobothria at the base of the fingers,
|
||
<emphasis id="B954E56EE941FFB71210141BFEBB2421" box="[286,312,1807,1832]" italics="true" pageId="10" pageNumber="327">ist</emphasis>
|
||
is closer to
|
||
<emphasis id="B954E56EE941FFB712CC141BFE512421" box="[450,466,1807,1832]" italics="true" pageId="10" pageNumber="327">it</emphasis>
|
||
than to
|
||
<emphasis id="B954E56EE941FFB71121141BFDC62420" box="[559,581,1807,1833]" italics="true" pageId="10" pageNumber="327">ib</emphasis>
|
||
; male femur pedipalpal
|
||
<geoCoordinate id="EE145FBBE941FFB71054141AFC182420" box="[858,923,1806,1833]" degrees="0.950" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.95">0.950</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB710AA141AFC602420" box="[932,995,1806,1833]" degrees="0.201" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.201">0.201</geoCoordinate>
|
||
, 4.7×, patella
|
||
<geoCoordinate id="EE145FBBE941FFB7178C141BFB422420" box="[1154,1217,1807,1833]" degrees="0.801" direction="north" orientation="latitude" pageId="10" pageNumber="327" precision="55" value="0.801">0.801</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE145FBBE941FFB717C4141BFA882420" box="[1226,1291,1807,1833]" degrees="0.240" direction="east" orientation="longitude" pageId="10" pageNumber="327" precision="55" value="0.24">0.240</geoCoordinate>
|
||
, 3.3×. Differ from
|
||
<taxonomicName id="4C2042FFE941FFB713D81427FDAD2444" authority="Beier, 1931" authorityName="Beier" authorityYear="1931" box="[214,558,1843,1869]" class="Arachnida" family="Hesperolpiidae" genus="Progarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="327" phylum="Arthropoda" rank="species" species="novus">
|
||
<emphasis id="B954E56EE941FFB713D81427FE202445" box="[214,419,1843,1868]" italics="true" pageId="10" pageNumber="327">Progarypus novus</emphasis>
|
||
Beier, 1931
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F33779ECE941FFB7117B1427FD3A2444" box="[629,697,1843,1869]" name="Brazil" pageId="10" pageNumber="327">Brazil</collectingCountry>
|
||
by the carapace 1.5× as long as wide, gradually narrowing, with truncated anterior margin, and approximately net-like sculpturing. Pedipalps slender, femur 4.0×, as long as wide, sessile, patella 3.5×, chelal fingers slightly shorter than hand without pedicel
|
||
<quantity id="4CD89499E941FFB710F1146EFBE4249C" box="[1023,1127,1914,1941]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="10" pageNumber="327" unit="mm" value="0.54">0.54 mm</quantity>
|
||
. It differs from
|
||
<taxonomicName id="4C2042FFE941FFB71617146FFE0424B0" authority="Beier, 1959" authorityName="Beier" authorityYear="1959" class="Arachnida" family="Hesperolpiidae" genus="Progarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="327" phylum="Arthropoda" rank="species" species="peruanus">
|
||
<emphasis id="B954E56EE941FFB71617146FFF7C24B1" italics="true" pageId="10" pageNumber="327">Progarypus peruanus</emphasis>
|
||
Beier, 1959
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F33779ECE941FFB712C5148BFE7D24B0" box="[459,510,1951,1977]" name="Peru" pageId="10" pageNumber="327">Peru</collectingCountry>
|
||
by the carapace as long as wide at the base, conically narrowed at anterior margin, densely granulated. Pedipalps almost as long as the body, femur without pedicel 4.3×, patella 3×, fingers clearly longer than hand chela with pedicel and as long as femur, tarsus IV shorter than tarsus I. It differs from
|
||
<taxonomicName id="4C2042FFE940FFB616171383FE4523DC" authority="(Balzan, 1887)" baseAuthorityName="Balzan" baseAuthorityYear="1887" class="Arachnida" family="Hesperolpiidae" genus="Progarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="328" phylum="Arthropoda" rank="species" species="oxydactylus">
|
||
<emphasis id="B954E56EE940FFB616171383FE9823DC" italics="true" pageId="11" pageNumber="328">Progarypus oxydactylus</emphasis>
|
||
(Balzan, 1887)
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F33779ECE940FFB6110913AFFDEC23DC" box="[519,623,187,213]" name="Paraguay" pageId="11" pageNumber="328">Paraguay</collectingCountry>
|
||
by the carapace clearly longer than wide, finely granulated, and reticulate at the base, pedipalps rather long and thin. Femur 4.1× times as long as wide, weakly pedicelled, finely granulated, and striated, patella 3.2× as long as wide, notably shorter than the femur. Similarly, it differs from the species
|
||
<taxonomicName id="4C2042FFE940FFB613991233FDA42248" authority="Mahnert, 2001" authorityName="Mahnert" authorityYear="2001" box="[151,551,295,321]" class="Arachnida" family="Hesperolpiidae" genus="Progarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="328" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B954E56EE940FFB613991233FEFB2248" box="[151,376,295,321]" italics="true" pageId="11" pageNumber="328">Progarypus gracilis</emphasis>
|
||
<bibRefCitation id="EFB1448DE940FFB612711233FDA42248" author="Mahnert, V." box="[383,551,295,321]" pageId="11" pageNumber="328" pagination="95 - 148" refId="ref14699" refString="Mahnert, V. (2001) Cave-dwelling pseudoscorpions (Arachnida, Pseudoscorpiones) from Brazil. Revue Suisse de Zoologie, 108, 95 - 148. https: // doi. org / 10.5962 / bhl. part. 79622" type="journal article" year="2001">Mahnert, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2042FFE940FFB6113C1233FC252248" authority="Mahnert, 2001" authorityName="Mahnert" authorityYear="2001" box="[562,934,295,321]" class="Arachnida" family="Hesperolpiidae" genus="Progarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="328" phylum="Arthropoda" rank="species" species="liliae">
|
||
<emphasis id="B954E56EE940FFB6113C1233FD742248" box="[562,759,295,321]" italics="true" pageId="11" pageNumber="328">Progarypus liliae</emphasis>
|
||
<bibRefCitation id="EFB1448DE940FFB611F01233FC252248" author="Mahnert, V." box="[766,934,295,321]" pageId="11" pageNumber="328" pagination="95 - 148" refId="ref14699" refString="Mahnert, V. (2001) Cave-dwelling pseudoscorpions (Arachnida, Pseudoscorpiones) from Brazil. Revue Suisse de Zoologie, 108, 95 - 148. https: // doi. org / 10.5962 / bhl. part. 79622" type="journal article" year="2001">Mahnert, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C2042FFE940FFB610EF1233FADD2248" authority="Mahnert, 2001" authorityName="Mahnert" authorityYear="2001" box="[993,1374,295,321]" class="Arachnida" family="Hesperolpiidae" genus="Progarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="328" phylum="Arthropoda" rank="species" species="setifer">
|
||
<emphasis id="B954E56EE940FFB610EF1233FB312248" box="[993,1202,295,321]" italics="true" pageId="11" pageNumber="328">Progarypus setifer</emphasis>
|
||
<bibRefCitation id="EFB1448DE940FFB617B71233FADD2248" author="Mahnert, V." box="[1209,1374,295,321]" pageId="11" pageNumber="328" pagination="95 - 148" refId="ref14699" refString="Mahnert, V. (2001) Cave-dwelling pseudoscorpions (Arachnida, Pseudoscorpiones) from Brazil. Revue Suisse de Zoologie, 108, 95 - 148. https: // doi. org / 10.5962 / bhl. part. 79622" type="journal article" year="2001">Mahnert, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F33779ECE940FFB61399125FFF58226C" box="[151,219,331,357]" name="Brazil" pageId="11" pageNumber="328">Brazil</collectingCountry>
|
||
by the length of the femur pedipalps 0.7-1.0-
|
||
<quantity id="4CD89499E940FFB611D1125FFCC5226C" box="[735,838,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" pageId="11" pageNumber="328" unit="mm" value="1.21">1.21 mm</quantity>
|
||
, by the distribution of the trichobothria
|
||
<emphasis id="B954E56EE940FFB61605125FFAA9226D" box="[1291,1322,331,356]" italics="true" pageId="11" pageNumber="328">est</emphasis>
|
||
,
|
||
<emphasis id="B954E56EE940FFB61637125FFAD0226D" box="[1337,1363,331,356]" italics="true" pageId="11" pageNumber="328">ist</emphasis>
|
||
,
|
||
<emphasis id="B954E56EE940FFB6166D125FFAF0226D" box="[1379,1395,331,356]" italics="true" pageId="11" pageNumber="328">it</emphasis>
|
||
,
|
||
<emphasis id="B954E56EE940FFB6168F125FFA16226D" box="[1409,1429,331,356]" italics="true" pageId="11" pageNumber="328">et</emphasis>
|
||
, the length of the fingers of the pedipalpal hand. The new species resembles
|
||
<taxonomicName id="4C2042FFE940FFB610EF127BFA1B2280" authority="Mahnert, 2001" authorityName="Mahnert" authorityYear="2001" box="[993,1432,367,393]" class="Arachnida" family="Hesperolpiidae" genus="Progarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="328" phylum="Arthropoda" rank="species" species="nigrimanus">
|
||
<emphasis id="B954E56EE940FFB610EF127BFB682281" box="[993,1259,367,392]" italics="true" pageId="11" pageNumber="328">Progarypus nigrimanus</emphasis>
|
||
<bibRefCitation id="EFB1448DE940FFB617FF127BFA1B2280" author="Mahnert, V." box="[1265,1432,367,393]" pageId="11" pageNumber="328" pagination="95 - 148" refId="ref14699" refString="Mahnert, V. (2001) Cave-dwelling pseudoscorpions (Arachnida, Pseudoscorpiones) from Brazil. Revue Suisse de Zoologie, 108, 95 - 148. https: // doi. org / 10.5962 / bhl. part. 79622" type="journal article" year="2001">Mahnert, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
, from
|
||
<collectingCountry id="F33779ECE940FFB613D81287FE9922A4" box="[214,282,403,429]" name="Brazil" pageId="11" pageNumber="328">Brazil</collectingCountry>
|
||
however, it differs by the femur pedipalpal length 0.7–1.0 mm; at least one of trichobothrium
|
||
<emphasis id="B954E56EE940FFB616431287FAEF22A5" box="[1357,1388,403,428]" italics="true" pageId="11" pageNumber="328">est</emphasis>
|
||
,
|
||
<emphasis id="B954E56EE940FFB616751287FA1622A5" box="[1403,1429,403,428]" italics="true" pageId="11" pageNumber="328">ist</emphasis>
|
||
,
|
||
<emphasis id="B954E56EE940FFB6139912A3FF2422D9" box="[151,167,439,464]" italics="true" pageId="11" pageNumber="328">it</emphasis>
|
||
,
|
||
<emphasis id="B954E56EE940FFB613BB12A3FF4A22D9" box="[181,201,439,464]" italics="true" pageId="11" pageNumber="328">et</emphasis>
|
||
proximal to
|
||
<emphasis id="B954E56EE940FFB6125512A3FEED22D9" box="[347,366,439,464]" italics="true" pageId="11" pageNumber="328">st</emphasis>
|
||
, finger relatively shorter, at most 1.3×, pedipalps bicolored, chelal hand darker than femur and patella, femur pedipalpal 4.7–5.3× (length
|
||
<quantity id="4CD89499E940FFB6118D12CFFD6E22FC" box="[643,749,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="11" pageNumber="328" unit="mm" value="0.86">0.86 mm</quantity>
|
||
), patella 3.8–3.9× (length
|
||
<quantity id="4CD89499E940FFB6171312CFFB3922FC" box="[1053,1210,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="7.85" metricValueMax="7.9" metricValueMin="7.8" pageId="11" pageNumber="328" unit="mm" value="0.785" valueMax="0.79" valueMin="0.78">0.78–0.79mm</quantity>
|
||
) longer than broad, carapace anteriorly with tessellated sculpturing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |