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<document id="E5BD83849BBCE5A713C787DBE07D232E" ID-DOI="10.11646/zootaxa.5293.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="7960232" ID-ZooBank="F68B3A5C-E100-4F7A-A1A1-FC9933403700" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1684798263703" checkinUser="plazi" docAuthor="Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder &amp; Lira, André Felipe De Araújo" docDate="2023" docId="0389886AE943FFB513991383FA1F200B" docLanguage="en" docName="zootaxa.5293.2.6.pdf" docOrigin="Zootaxa 5293 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5293.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geogarypus Chamberlin 1930" docType="key" docVersion="3" lastPageNumber="325" masterDocId="FFB0F012E94BFFBD130E1314FF832309" masterDocTitle="Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil" masterLastPageNumber="332" masterPageNumber="317" pageNumber="325" updateTime="1684871452790" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="D613E712AEB51029FF2AAC4B65479656">Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil</mods:title>
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<mods:namePart id="FDE3D818B971CB331D2243488027950E">Bedoya-Roqueme, Edwin</mods:namePart>
<mods:affiliation id="C5184503BBB5C668D14229D6BC90C6F0">Programa de Pós-graduação em Recursos Naturais do Cerrado, Laboratório de Ecologia Comportamental de Aracnídeos, Universidade Estadual de Goiás, Anápolis, Brazil.</mods:affiliation>
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<mods:namePart id="D0DC93CB44577C4562F57EAF961E5532">Tizo-Pedroso, Everton</mods:namePart>
<mods:affiliation id="CA5A45F556E7B0D41F7B889E727E68F7">Laboratório de Ecologia Comportamental de Aracnídeos, Programa de Pós-graduação em Recursos Naturais do Cerrado, Centro de Ensino e Aprendizagem em Rede, Universidade Estadual de Goiás, Anápolis, Brazil.</mods:affiliation>
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<mods:namePart id="101D130AFCD2DDFB2575F0ED073D39C9">Barbier, Eder</mods:namePart>
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<mods:affiliation id="609D52CE3F4D5AA1B05CF1ED28ED2500">Programa de Pós-graduação em Biologia Animal, Departamento de Zoologia, Universidade Federal de Pernambuco, Recife, Brazil. barbier. eder @ gmail. com; http: // orcid. org / 0000 - 0001 - 5068 - 7048</mods:affiliation>
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<mods:namePart id="3B1756A535599501D55728FA3B931671">Lira, André Felipe De Araújo</mods:namePart>
<mods:affiliation id="9A4186C1B5EDAF06A1B910A3A4451D15">Centro de Educação e Saúde, Universidade Federal de Campina Grande, Cuité, Brazil.</mods:affiliation>
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Key to
<taxonomicName id="4C2042FFE943FFB513FF1383FDE923B8" ID-CoL="4MX9" ID-ENA="535072" authority="Chamberlin, 1930" authorityName="Chamberlin" authorityYear="1930" box="[241,618,151,177]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="8" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="B954E56EE943FFB513FF1383FE0123B8" bold="true" box="[241,386,151,177]" italics="true" pageId="8" pageNumber="325">Geogarypus</emphasis>
Chamberlin, 1930
</taxonomicName>
species reported from
<collectingCountry id="F33779ECE943FFB510861383FC5623B8" box="[904,981,151,177]" name="Brazil" pageId="8" pageNumber="325">Brazil</collectingCountry>
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1 Chela of the pedipalp with eight trichobothria; trichobothrium
<emphasis id="B954E56EE943FFB5103513C8FCD423F8" box="[827,855,220,241]" italics="true" pageId="8" pageNumber="325">isb</emphasis>
present........................................ (2
<emphasis id="B954E56EE943FFB5169A13C8FA1F23FB" bold="true" box="[1428,1436,220,242]" pageId="8" pageNumber="325">)</emphasis>
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<paragraph id="8B9F397CE943FFB5139913EEFA1F2225" blockId="8.[151,1437,220,770]" pageId="8" pageNumber="325">
<keyLead id="30D19A49E943FFB5139913EEFA1F2225" pageId="8" pageNumber="325">
- Chela of the pedipalp with seven trichobothria; trichobothrium
<emphasis id="B954E56EE943FFB5105F13EDFCEE2207" box="[849,877,249,270]" italics="true" pageId="8" pageNumber="325">isb</emphasis>
absent; femur pedipalpal 3.5× (male, 3.5×); patella 2.3× (male, 2.2×); leg I and IV diplotarsate.......................................................
<taxonomicName id="4C2042FFE943FFB517ED1203FACA2225" authority="Bedoya-Roqueme &amp; Tizo-Pedroso &amp; Barbier &amp; Lira, 2023" authorityName="Bedoya-Roqueme &amp; Tizo-Pedroso &amp; Barbier &amp; Lira" authorityYear="2023" box="[1251,1353,279,300]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="8" pageNumber="325" phylum="Arthropoda" rank="species" species="gollumi" status="sp. nov.">
<emphasis id="B954E56EE943FFB517ED1203FB772225" box="[1251,1268,279,300]" italics="true" pageId="8" pageNumber="325">G</emphasis>
.
<emphasis id="B954E56EE943FFB5160E1203FACA2225" box="[1280,1353,279,300]" italics="true" pageId="8" pageNumber="325">gollumi</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="A2675815E943FFB5165E1203FA1F2225" box="[1360,1436,279,300]" pageId="8" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
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<keyLead id="30D19A49E943FFB513991220FA1F226E" pageId="8" pageNumber="325">2 Femur of pedipalp more than 3.0× as long as wide, chela of pedipalp 1.63.2×; movable finger more than 4048 teeth, fixed finger with 2732 teeth............................................................................... (3)</keyLead>
</paragraph>
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<keyLead id="30D19A49E943FFB51399127BFA1F22AB" pageId="8" pageNumber="325">
- Femur of pedipalp 3.0× longer than wide; pedipalp chela 1.8× (see Mello-Leit„o, 1937; fig.1); movable finger with 38 teeth, fixed finger with 32 teeth...........................................................
<taxonomicName id="4C2042FFE943FFB517AC1298FA0A22AB" authority="Mello-Leit" authorityName="Mello-Leit" box="[1186,1417,396,418]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="8" pageNumber="325" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B954E56EE943FFB517AC1298FB3022A8" box="[1186,1203,396,417]" italics="true" pageId="8" pageNumber="325">G</emphasis>
.
<emphasis id="B954E56EE943FFB517CE1298FA9B22A8" box="[1216,1304,396,417]" italics="true" pageId="8" pageNumber="325">formosus</emphasis>
Mello-Leit
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„o
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<paragraph id="8B9F397CE943FFB5139912BEFA1F22D5" blockId="8.[151,1437,220,770]" pageId="8" pageNumber="325">
<keyLead id="30D19A49E943FFB5139912BEFA1F22D5" pageId="8" pageNumber="325">3 Femur of pedipalp 3.13.5× as long as wide; patella 2.12.4× longer than wide; fingers 1.11.2× longer than hand with pedicel............................................................................................. (4)</keyLead>
</paragraph>
<paragraph id="8B9F397CE943FFB5139912F1FA1F213D" blockId="8.[151,1437,220,770]" pageId="8" pageNumber="325">
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- Femur of pedipalp more than 3.5× as long as wide; patella thinner, 2.4× longer than wide (see
<bibRefCitation id="EFB1448DE943FFB5178E12F0FA8122F0" author="Balzan, L." box="[1152,1282,484,506]" pageId="8" pageNumber="325" pagination="401 - 454" refId="ref13379" refString="Balzan, L. (1890). Revisione dei Pseudoscorpioni del Bacino dei Fiumi Parana e Paraguay nell'America meridionale. Annali del Museo Civico di Storia Naturale di Genova, 9 (2 a), 401 - 454." type="journal article" year="1890">Balzan, 1890</bibRefCitation>
; fig. 22); finger shorter (
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) than the hand with pedicel (
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) and longer than the chela (
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)......................................................................................................
<taxonomicName id="4C2042FFE943FFB5178A110BFA1F213D" authority="(Balzan, 1887)" baseAuthorityName="Balzan" baseAuthorityYear="1887" box="[1156,1436,543,565]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="8" pageNumber="325" phylum="Arthropoda" rank="species" species="cuyabanus">
<emphasis id="B954E56EE943FFB5178A110BFB16213D" box="[1156,1173,543,564]" italics="true" pageId="8" pageNumber="325">G</emphasis>
.
<emphasis id="B954E56EE943FFB517AF110BFA8B213D" box="[1185,1288,543,564]" italics="true" pageId="8" pageNumber="325">cuyabanus</emphasis>
(Balzan, 1887)
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4 Femur of the pedipalp 3.03.3×, patella 2.42.5× as long as wide (see
<bibRefCitation id="EFB1448DE943FFB5108C1128FB912158" author="Mahnert, V." box="[898,1042,572,594]" pageId="8" pageNumber="325" pagination="719 - 810" refId="ref14603" refString="Mahnert, V. (1979) Pseudoskorpione (Arachnida) aus dem Amazonas-Gebiet (Brasilien). Revue Suisse de Zoologie, 86, 719 - 810. https: // doi. org / 10.5962 / bhl. part. 82338" type="journal article" year="1979">Mahnert, 1979</bibRefCitation>
; fig. 87); fingers (0.470.48 (0.490.55)) longer than hand (0.400.42 (0.450.49)) with pedicel; fixed finger with 4148 teeth (distal 12 small, half teeth arranged in two rows), movable finger with 2731 teeth (distal half only pointed) (see
<bibRefCitation id="EFB1448DE943FFB510A01163FBBC2185" author="Mahnert, V." box="[942,1087,631,653]" pageId="8" pageNumber="325" pagination="719 - 810" refId="ref14603" refString="Mahnert, V. (1979) Pseudoskorpione (Arachnida) aus dem Amazonas-Gebiet (Brasilien). Revue Suisse de Zoologie, 86, 719 - 810. https: // doi. org / 10.5962 / bhl. part. 82338" type="journal article" year="1979">Mahnert, 1979</bibRefCitation>
; fig. 88b)......................................................................................................
<taxonomicName id="4C2042FFE943FFB517B81180FA1F21A3" authority="Mahnert" authorityName="Mahnert" authorityYear="1979" box="[1206,1436,660,682]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="8" pageNumber="325" phylum="Arthropoda" rank="species" species="amazonicus">
<emphasis id="B954E56EE943FFB517B81180FB4421A0" box="[1206,1223,660,681]" italics="true" pageId="8" pageNumber="325">G</emphasis>
.
<emphasis id="B954E56EE943FFB517DA1181FAC621A3" box="[1236,1349,661,682]" italics="true" pageId="8" pageNumber="325">amazonicus</emphasis>
Mahnert
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F397CE943FFB5139911A6FA1F200B" blockId="8.[151,1437,220,770]" pageId="8" pageNumber="325">
<keyLead id="30D19A49E943FFB5139911A6FA1F200B" pageId="8" pageNumber="325">
- Femur of pedipalp 3.2×, patella 2.3× as long as wide (see
<bibRefCitation id="EFB1448DE943FFB5100911A5FC0021CE" author="Beier, M." box="[775,899,689,711]" pageId="8" pageNumber="325" pagination="1 - 258" refId="ref13542" refString="Beier, M. (1932 a) Pseudoscorpionidea I. Subord. Chthoniinea et Neobisiinea. Tierreich, 57, i - xx + 1 - 258. https: // doi. org / 10.1515 / 9783111435107.1" type="journal article" year="1932">Beier, 1932a</bibRefCitation>
; fig. 258); chela finger (
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) as long as hand (0.49) with pedicel; fixed finger with about 27 teeth, movable finger with 22 teeth (more or less clearly pointed on the base).......................................................................................
<taxonomicName id="4C2042FFE943FFB517B211F8FA1F200B" authority="Beier" authorityName="Beier" authorityYear="1931" box="[1212,1436,748,770]" class="Arachnida" family="Geogarypidae" genus="Geogarypus" kingdom="Animalia" order="Pseudoscorpiones" pageId="8" pageNumber="325" phylum="Arthropoda" rank="species" species="paraguayanus">
<emphasis id="B954E56EE943FFB517B211F8FB4E2008" box="[1212,1229,748,769]" italics="true" pageId="8" pageNumber="325">G</emphasis>
.
<emphasis id="B954E56EE943FFB517D711F9FAE1200B" box="[1241,1378,749,770]" italics="true" pageId="8" pageNumber="325">paraguayanus</emphasis>
Beier
</taxonomicName>
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