476 lines
74 KiB
XML
476 lines
74 KiB
XML
<document id="AFCF8DA4625C374806C44D3FDDC8787F" ID-CLB-Dataset="199799" ID-DOI="10.11646/zootaxa.5214.3.4" ID-GBIF-Dataset="9c8b2ed9-a132-4df0-afa9-b153547b9027" ID-ISSN="1175-5326" ID-Zenodo-Dep="7389267" ID-ZooBank="395A3FB5-DA5E-4145-B9EA-7801C67D6CE3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669970496187" checkinUser="plazi" docAuthor="Huynh, Cuong" docDate="2022" docId="038987F7BC1608669495F9300E319C40" docLanguage="en" docName="zootaxa.5214.3.4.pdf" docOrigin="Zootaxa 5214 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Monographis cattienensis Huynh 2022, new species" docType="treatment" docVersion="7" lastPageNumber="403" masterDocId="FFB0FF8FBC12086C9402FF890C769D01" masterDocTitle="Four new species of Monographis Attems, 1907 (Diplopoda, Polyxenida, Polyxenidae) from Vietnam" masterLastPageNumber="420" masterPageNumber="393" pageNumber="397" updateTime="1699412757114" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A9BC13F5C8B6701F78273A8E21856D55">Four new species of Monographis Attems, 1907 (Diplopoda, Polyxenida, Polyxenidae) from Vietnam</mods:title>
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<mods:name id="626C04E208C2D02D69B0B8A2C398F77B" type="personal">
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<mods:roleTerm id="DA409537B7D64C01CCAF2328E81F3ECF">Author</mods:roleTerm>
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<mods:namePart id="2900A2D25C6CE3966AAD2460E999641F">Huynh, Cuong</mods:namePart>
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<mods:affiliation id="9DE5B1A4D25726B8AE9B525D1287853E">Centre for Cellular and Molecular Biology (CCMB), Deakin University, 221 Burwood Hwy, Burwood, Melbourne, Australia 3125</mods:affiliation>
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<mods:typeOfResource id="58A3EC7C3190BB4CB8D44308B6170CCD">text</mods:typeOfResource>
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<mods:title id="955EE27DEFAA012CDFDC3899777B6531">Zootaxa</mods:title>
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<mods:part id="B5D4FB2E5171EDBF38D135CF9DF8CB75">
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<mods:date id="94EE0EE96B14AD951B94A4BFA1C8EB18">2022</mods:date>
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<mods:number id="5B9A0D7EFE5DCFB307F4C15DCBA7DF65">2022-12-02</mods:number>
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<mods:number id="88F7EC562010E2BB281BB93D062FD42F">5214</mods:number>
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<mods:number id="303B01514D38D40AD0F0634347B2F8D8">3</mods:number>
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<mods:identifier id="B0848E0A2E168E7AF57474882ECC438A" type="CLB-Dataset">199799</mods:identifier>
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<mods:identifier id="A97CDFA6524BB82F0E33D186A565983F" type="DOI">10.11646/zootaxa.5214.3.4</mods:identifier>
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<treatment id="038987F7BC1608669495F9300E319C40" ID-DOI="http://doi.org/10.5281/zenodo.7389466" ID-GBIF-Taxon="205697561" ID-Zenodo-Dep="7389466" LSID="urn:lsid:plazi:treatment:038987F7BC1608669495F9300E319C40" httpUri="http://treatment.plazi.org/id/038987F7BC1608669495F9300E319C40" lastPageId="10" lastPageNumber="403" pageId="4" pageNumber="397">
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<subSubSection id="C33A656ABC1608689495F9300E119BD2" box="[151,615,1721,1748]" pageId="4" pageNumber="397" type="nomenclature">
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<paragraph id="8B9F36E1BC1608689495F9300E119BD2" blockId="4.[151,615,1721,1784]" box="[151,615,1721,1748]" pageId="4" pageNumber="397">
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<heading id="D0D7818DBC1608689495F9300E119BD2" bold="true" box="[151,615,1721,1748]" fontSize="11" level="1" pageId="4" pageNumber="397" reason="1">
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<emphasis id="B954EAF3BC1608689495F9300E119BD2" bold="true" box="[151,615,1721,1748]" pageId="4" pageNumber="397">
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<taxonomicName id="4C204D62BC1608689495F9300DA69BD5" authority="Huynh, 2022" authorityName="Huynh" authorityYear="2022" box="[151,464,1721,1748]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="397" phylum="Arthropoda" rank="species" species="cattienensis" status="sp. nov.">
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<emphasis id="B954EAF3BC1608689495F9300DA69BD5" bold="true" box="[151,464,1721,1748]" italics="true" pageId="4" pageNumber="397">Monographis cattienensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2675788BC16086895D5F9330E119BD2" box="[471,615,1721,1748]" pageId="4" pageNumber="397" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A656ABC1608689495F9570D6B9BF9" box="[151,285,1758,1784]" pageId="4" pageNumber="397" type="description">
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<paragraph id="8B9F36E1BC1608689495F9570D6B9BF9" blockId="4.[151,615,1721,1784]" box="[151,285,1758,1784]" pageId="4" pageNumber="397">
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<figureCitation id="131B2A64BC1608689495F9570D6B9BF9" box="[151,285,1758,1784]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStartId-0="5.[152,255,1823,1847]" captionStartId-1="6.[152,255,1527,1551]" captionStartId-2="7.[152,255,1467,1491]" captionStartId-3="8.[152,255,1818,1842]" captionStartId-4="9.[152,255,1865,1889]" captionTargetBox-0="[184,1403,191,1791]" captionTargetBox-1="[154,1433,424,1492]" captionTargetBox-2="[157,1432,674,1440]" captionTargetBox-3="[221,1360,193,1780]" captionTargetBox-4="[214,1372,525,1842]" captionTargetId-0="figure-19@5.[184,1404,181,1800]" captionTargetId-1="figure-121@6.[151,1436,417,1504]" captionTargetId-2="figure-243@7.[151,1436,669,1444]" captionTargetId-3="figure-29@8.[216,1371,181,1795]" captionTargetId-4="figure-185@9.[214,1374,525,1842]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="FIGURE 3. Holotype male Monographis cattienensis sp. n.. A. The head capsule showing eye patterns (each side with eight ommatidia); trichobothria: trichobothrium a, b and c; anterior vertex (av) and posterior vertex (pv). B. The enlarge trichome sockets in posterior vertex, pv (L): left side, pv (R): right side. C. Trichobothria (left side): trichobothrium a (interior position), b (lateral) and c (exterior); D. Right gnathochilarium showing lateral palp (Lp) with 11 conical sensilla and medial palp (Mp) with 22; E. Clypeo-labrum showing setose surface, with two lateral lamellae (Ll), two lamellae (l) and a median cleft (mc), anterior setae (sa); posterior setae (Se); a row of backward-pointing setae (sb)." captionText-1="FIGURE 4. Holotype male Monographis cattienensis sp. n.. A. Antennal articles VI, VII, VIII and sensory cones (sc); B. and C. showing the arrangements of sensilla on articles VI and VII. (T: thick bacilliform sensillum (Ta: anterior, Tp: posterior position), t: thin bacilliform sensillum, ts: short thin bacilliform sensillum, S: setiform and C: conical sensillum)" captionText-2="FIGURE 5. Holotype male Monographis cattienensis sp. n. with typical trichome socket patterns on the collum and tergites of Monographis species. A. Collum (Col)—the first tergite showing trichome patterns and the first pair of pleural projections—lateral protuberances (Lp); B. Trichome socket patterns present on tergites 2 (T2) to 9 (T9), and pleural projections (pp); C. Trichome socket patterns present on tergite 10 (T10)—the last tergite, and its pleural projections, the trichome sockets showing dense and smaller than other tergite." captionText-3="FIGURE 6. Holotype male Monographis cattienensis sp. n.. A. The second leg showing the segments and the chaetotaxy— hairs on its segments: Coxa (co), prefemur (pf), femur (f), postfemur (pof), tibia (tib), tarsus 1 (T1), tarsus 2 (T2) and claw (c); B. Typical seta present on leg articles; C. Spine on Tarsus 1 & 2; D. Telotarsus: Claw (c), lamella (la), setiform (s) and anterior (a), posterior (p) lateral processes. E. Two pairs of coxal glands present on coxae VIII and IX; F. Dorsal ornamental trichome sockets showing both sides, trichome sockets a: a1 to a18 (Left), with 17 sockets (Right); trichome socket b; trichome sockets c: c1, c2 and c3. Circular indentation d; G. Female’s nest trichome; H. A caudal bundle trichome." captionText-4="FIGURE 7. SEM images of Monographis cattienensis sp. n.. A. antennal articles VI, VII and VIII; B. and C. showing the arrangement of sensilla on the article VI; D. Clypeo-labrum showing setose surface with anterior setae (sa), lateral lamella (Ll), median cleft (mc), row of backward-pointing setae (sb) and posterior setae (Se); E. Leg articles and two spines (pointed arrows) which are typical characters of Monographis; F. Telotarsus showing processes: anterior lateral process (a), posterior lateral process (p), claw (c), anterior setiform process (s) and lamella (la)." figureDoi-0="http://doi.org/10.5281/zenodo.7389283" figureDoi-1="http://doi.org/10.5281/zenodo.7389285" figureDoi-2="http://doi.org/10.5281/zenodo.7389289" figureDoi-3="http://doi.org/10.5281/zenodo.7389293" figureDoi-4="http://doi.org/10.5281/zenodo.7389299" httpUri-0="https://zenodo.org/record/7389283/files/figure.png" httpUri-1="https://zenodo.org/record/7389285/files/figure.png" httpUri-2="https://zenodo.org/record/7389289/files/figure.png" httpUri-3="https://zenodo.org/record/7389293/files/figure.png" httpUri-4="https://zenodo.org/record/7389299/files/figure.png" pageId="4" pageNumber="397">Figures 3–7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A656ABC1608689495F8AC0FBB9AF5" pageId="4" pageNumber="397" type="materials_examined">
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<paragraph id="8B9F36E1BC1608689495F8AC0FBB9AF5" blockId="4.[151,1437,1829,2036]" pageId="4" pageNumber="397">
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<emphasis id="B954EAF3BC1608689495F8AC0D0C9A3E" bold="true" box="[151,378,1829,1855]" pageId="4" pageNumber="397">Material examined</emphasis>
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: Type specimens.
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<typeStatus id="549B8843BC160868964EF8AF0EC29A41" box="[588,692,1830,1856]" pageId="4" pageNumber="397" type="holotype">Holotype</typeStatus>
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adult male, to be returned to the Zoological Museum of the
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<collectingRegion id="49E4F803BC1608689163F8AF0CB09A65" country="Russia" name="Moscow" pageId="4" pageNumber="397">Moscow</collectingRegion>
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<materialsCitation id="3B483CBCBC16086894CEF8C30E0E9A65" ID-GBIF-Occurrence="3981599309" box="[204,632,1866,1892]" collectionCode="ZMUM" country="Russia" location="Moscow" pageId="4" pageNumber="397" specimenCount="1" stateProvince="Moscow">
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University (
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<collectionCode id="ED31AE24BC1608689552F8C30DDB9A65" box="[336,429,1866,1892]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="4" pageNumber="397">ZMUM</collectionCode>
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),
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<collectingRegion id="49E4F803BC16086895BDF8C30E579A65" box="[447,545,1866,1892]" country="Russia" name="Moscow" pageId="4" pageNumber="397">Moscow</collectingRegion>
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,
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<collectingCountry id="F3377671BC1608689628F8C30E0E9A65" box="[554,632,1866,1892]" name="Russia" pageId="4" pageNumber="397">Russia</collectingCountry>
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</materialsCitation>
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.
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<materialsCitation id="3B483CBCBC1608689680F8C30F4F9AAD" ID-GBIF-Occurrence="3981599306" collectionCode="ZMUM" country="Vietnam" location="Nam Cat Tien National Park" pageId="4" pageNumber="397" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Dong Nai" typeStatus="paratype">
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<typeStatus id="549B8843BC1608689680F8C30E849A65" box="[642,754,1866,1892]" pageId="4" pageNumber="397" type="paratype">Paratypes</typeStatus>
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,
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<specimenCount id="9D26FD68BC16086896FEF8C30F279A65" box="[764,849,1866,1892]" pageId="4" pageNumber="397" type="male">2 males</specimenCount>
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and
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<specimenCount id="9D26FD68BC1608689784F8C30F879A65" box="[902,1009,1866,1892]" pageId="4" pageNumber="397" type="female">2 females</specimenCount>
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(
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<collectionCode id="ED31AE24BC16086897FDF8C3082A9A65" box="[1023,1116,1866,1892]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="4" pageNumber="397">ZMUM</collectionCode>
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), all collected from the study of microarthropods and macrofauna in the
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<location id="8EFF603ABC160868967AF8E70FCE9A89" LSID="urn:lsid:plazi:treatment:038987F7BC1608669495F9300E319C40:8EFF603ABC160868967AF8E70FCE9A89" box="[632,952,1902,1928]" country="Vietnam" name="Nam Cat Tien National Park" pageId="4" pageNumber="397" stateProvince="Dong Nai">Nam Cat Tien National Park</location>
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during 2004–2005 (Anchkin
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<emphasis id="B954EAF3BC1608689104F8E709499A89" box="[1286,1343,1902,1928]" italics="true" pageId="4" pageNumber="397">et. al</emphasis>
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., 2007), a south-eastern lowland forest,
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<collectingRegion id="49E4F803BC16086895F1F81B0EBA9AAD" box="[499,716,1938,1964]" country="Vietnam" name="Dong Nai" pageId="4" pageNumber="397">Dong Nai Province</collectingRegion>
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,
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<collectingCountry id="F3377671BC16086896D5F81B0F4F9AAD" box="[727,825,1938,1964]" name="Vietnam" pageId="4" pageNumber="397">Vietnam</collectingCountry>
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</materialsCitation>
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;
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<geoCoordinate id="EE145026BC1608689746F8180FC39AAD" box="[836,949,1937,1964]" degrees="11.4475" direction="north" orientation="latitude" pageId="4" pageNumber="397" precision="5" value="11.4475">11.4475N</geoCoordinate>
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,
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<geoCoordinate id="EE145026BC16086897BDF81B082E9AAD" box="[959,1112,1938,1964]" degrees="107.442778" direction="east" orientation="longitude" pageId="4" pageNumber="397" precision="1" value="107.44278">107.442778E</geoCoordinate>
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, elevation
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<quantity id="4CD89B04BC16086890D2F81B096F9AAD" box="[1232,1305,1938,1964]" metricMagnitude="2" metricUnit="m" metricValue="1.31" pageId="4" pageNumber="397" unit="m" value="131.0">131 m</quantity>
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. Specimens were sent to authors by Professor Sergei Golovatch, the collections were preserved in 80% ethanol, the specimens appeared brownish in colour, and most trichomes and antennae were lost.
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</paragraph>
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</subSubSection>
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<caption id="DF5F6669BC170869949AF8960F2A9AED" ID-DOI="http://doi.org/10.5281/zenodo.7389283" ID-Zenodo-Dep="7389283" httpUri="https://zenodo.org/record/7389283/files/figure.png" pageId="5" pageNumber="398" startId="5.[152,255,1823,1847]" targetBox="[184,1403,191,1791]" targetPageId="5">
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<paragraph id="8B9F36E1BC170869949AF8960F2A9AED" blockId="5.[151,1437,1823,2028]" pageId="5" pageNumber="398">
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<emphasis id="B954EAF3BC170869949AF8960D6F9A36" bold="true" box="[152,281,1823,1847]" pageId="5" pageNumber="398">FIGURE 3.</emphasis>
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Holotype male
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<taxonomicName id="4C204D62BC17086995BFF8A90EB39A36" authority="Huynh, 2022" authorityName="Huynh" authorityYear="2022" box="[445,709,1824,1848]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="5" pageNumber="398" phylum="Arthropoda" rank="species" species="cattienensis" status="sp. nov.">
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<emphasis id="B954EAF3BC17086995BFF8A90EB39A36" box="[445,709,1824,1848]" italics="true" pageId="5" pageNumber="398">Monographis cattienensis</emphasis>
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</taxonomicName>
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||
<emphasis id="B954EAF3BC17086996C9F8A90F739A39" bold="true" box="[715,773,1824,1848]" pageId="5" pageNumber="398">
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||
<taxonomicNameLabel id="A2675788BC17086996C9F8A90F739A39" box="[715,773,1824,1848]" pageId="5" pageNumber="398" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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.
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<emphasis id="B954EAF3BC1708699710F8960F5D9A36" bold="true" box="[786,811,1823,1847]" pageId="5" pageNumber="398">A.</emphasis>
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The head capsule showing eye patterns (each side with eight ommatidia); trichobothria: trichobothrium
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<emphasis id="B954EAF3BC170869964DF8CD0E2A9A5A" bold="true" box="[591,604,1860,1883]" italics="true" pageId="5" pageNumber="398">a</emphasis>
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,
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<emphasis id="B954EAF3BC1708699669F8CA0E0E9A5A" bold="true" box="[619,632,1859,1883]" italics="true" pageId="5" pageNumber="398">b</emphasis>
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and
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<emphasis id="B954EAF3BC17086996ACF8CD0ECF9A5A" bold="true" box="[686,697,1860,1883]" italics="true" pageId="5" pageNumber="398">c</emphasis>
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||
; anterior vertex (av) and posterior vertex (pv).
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<emphasis id="B954EAF3BC17086990A5F8CD08C89A5D" bold="true" box="[1191,1214,1860,1884]" pageId="5" pageNumber="398">B.</emphasis>
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The enlarge trichome sockets in posterior vertex, pv (L): left side, pv (R): right side.
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<emphasis id="B954EAF3BC1708699704F8EE0F699A7E" bold="true" box="[774,799,1895,1919]" pageId="5" pageNumber="398">C.</emphasis>
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Trichobothria (left side): trichobothrium
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<emphasis id="B954EAF3BC17086990BDF8E108BA9A7E" bold="true" box="[1215,1228,1896,1919]" italics="true" pageId="5" pageNumber="398">a</emphasis>
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(interior position),
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||
<emphasis id="B954EAF3BC1708699192F8EE09EB9A7E" bold="true" box="[1424,1437,1895,1919]" italics="true" pageId="5" pageNumber="398">b</emphasis>
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(lateral) and
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<emphasis id="B954EAF3BC1708699515F8050D549AA2" bold="true" box="[279,290,1932,1955]" italics="true" pageId="5" pageNumber="398">c</emphasis>
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(exterior);
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<emphasis id="B954EAF3BC1708699596F8050DDB9AA5" bold="true" box="[404,429,1932,1956]" pageId="5" pageNumber="398">D.</emphasis>
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Right gnathochilarium showing lateral palp (Lp) with 11 conical sensilla and medial palp (Mp) with 22;
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<emphasis id="B954EAF3BC17086994BCF8390CA39AC9" bold="true" box="[190,213,1968,1992]" pageId="5" pageNumber="398">E.</emphasis>
|
||
Clypeo-labrum showing setose surface, with two lateral lamellae (Ll), two lamellae (l) and a median cleft (mc), anterior setae (sa); posterior setae (Se); a row of backward-pointing setae (sb).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33A656ABC14086A94C5FF1F09509C1C" pageId="6" pageNumber="399" type="diagnosis">
|
||
<paragraph id="8B9F36E1BC14086A94C5FF1F09509C1C" blockId="6.[151,1437,150,393]" pageId="6" pageNumber="399">
|
||
<emphasis id="B954EAF3BC14086A94C5FF1F0D4C9DB1" bold="true" box="[199,314,150,176]" pageId="6" pageNumber="399">Diagnosis</emphasis>
|
||
: Body length
|
||
<quantity id="4CD89B04BC14086A95E7FF1E0E049DB0" box="[485,626,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" metricValueMax="3.6" metricValueMin="2.6" pageId="6" pageNumber="399" unit="mm" value="3.1" valueMax="3.6" valueMin="2.6">2.6–3.6 mm</quantity>
|
||
. Antennal article VI with 14–16 bacilliform sensilla forming crescent shape, setiform sensillum located anterior position, conical sensillum present near centre of crescent. Labrum setose, with long slender anterior setae on anterior margin. Telotarsus with posterior lateral process less than half length of claw, anterior lateral and lamella processes present, anterior setiform process longer than claw.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A656ABC14086594C5FEAF09539CF4" lastPageId="9" lastPageNumber="402" pageId="6" pageNumber="399" type="description">
|
||
<paragraph id="8B9F36E1BC14086A94C5FEAF0D2C9C40" blockId="6.[151,1437,150,393]" box="[199,346,294,321]" pageId="6" pageNumber="399">
|
||
<emphasis id="B954EAF3BC14086A94C5FEAF0D279C41" bold="true" box="[199,337,294,320]" pageId="6" pageNumber="399">Description</emphasis>
|
||
:
|
||
</paragraph>
|
||
<paragraph id="8B9F36E1BC14086A94C5FEC20E899C88" blockId="6.[151,1437,150,393]" pageId="6" pageNumber="399">
|
||
<typeStatus id="549B8843BC14086A94C5FEC20D599C64" box="[199,303,331,357]" pageId="6" pageNumber="399" type="holotype">Holotype</typeStatus>
|
||
male body
|
||
<quantity id="4CD89B04BC14086A95B6FEC20E7B9C64" box="[436,525,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="399" unit="mm" value="2.6">2.6 mm</quantity>
|
||
;
|
||
<typeStatus id="549B8843BC14086A961BFEC20EFF9C64" box="[537,649,331,357]" pageId="6" pageNumber="399" type="paratype">paratypes</typeStatus>
|
||
: males
|
||
<quantity id="4CD89B04BC14086A96DEFEC20F139C64" box="[732,869,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="6" pageNumber="399" unit="mm" value="2.7" valueMax="2.8" valueMin="2.6">2.6–2.8 mm</quantity>
|
||
, females
|
||
<quantity id="4CD89B04BC14086A97CDFEC2082C9C64" box="[975,1114,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="3.6" metricValueMin="3.4" pageId="6" pageNumber="399" unit="mm" value="3.5" valueMax="3.6" valueMin="3.4">3.4–3.6 mm</quantity>
|
||
; caudal bundle in both sexes same length
|
||
<quantity id="4CD89B04BC14086A9527FEE70D0B9C88" box="[293,381,366,393]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="399" unit="mm" value="0.5">0.5 mm</quantity>
|
||
, nest trichomes in female
|
||
<quantity id="4CD89B04BC14086A96A0FEE60E8D9C88" box="[674,763,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="399" unit="mm" value="0.2">0.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF5F6669BC14086A949AFA7E08AD9B56" ID-DOI="http://doi.org/10.5281/zenodo.7389285" ID-Zenodo-Dep="7389285" httpUri="https://zenodo.org/record/7389285/files/figure.png" pageId="6" pageNumber="399" startId="6.[152,255,1527,1551]" targetBox="[154,1433,424,1492]" targetPageId="6">
|
||
<paragraph id="8B9F36E1BC14086A949AFA7E08AD9B56" blockId="6.[151,1437,1527,1623]" pageId="6" pageNumber="399">
|
||
<emphasis id="B954EAF3BC14086A949AFA7E0D6E9B0E" bold="true" box="[152,280,1527,1551]" pageId="6" pageNumber="399">FIGURE 4.</emphasis>
|
||
Holotype male
|
||
<taxonomicName id="4C204D62BC14086A95BEFA7E0EB59B0E" authority="Huynh, 2022" authorityName="Huynh" authorityYear="2022" box="[444,707,1527,1551]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="6" pageNumber="399" phylum="Arthropoda" rank="species" species="cattienensis" status="sp. nov.">
|
||
<emphasis id="B954EAF3BC14086A95BEFA7E0EB59B0E" box="[444,707,1527,1551]" italics="true" pageId="6" pageNumber="399">Monographis cattienensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EAF3BC14086A96CBFA7E0F749B0E" bold="true" box="[713,770,1527,1551]" pageId="6" pageNumber="399">
|
||
<taxonomicNameLabel id="A2675788BC14086A96CBFA7E0F749B0E" box="[713,770,1527,1551]" pageId="6" pageNumber="399" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B954EAF3BC14086A970DFA7E0F5E9B0E" bold="true" box="[783,808,1527,1551]" pageId="6" pageNumber="399">A.</emphasis>
|
||
Antennal articles VI, VII, VIII and sensory cones (sc);
|
||
<emphasis id="B954EAF3BC14086A9158FA7E09079B0E" bold="true" box="[1370,1393,1527,1551]" pageId="6" pageNumber="399">B.</emphasis>
|
||
and
|
||
<emphasis id="B954EAF3BC14086A9495F9920CC69B32" bold="true" box="[151,176,1563,1587]" pageId="6" pageNumber="399">C.</emphasis>
|
||
showing the arrangements of sensilla on articles VI and VII. (T: thick bacilliform sensillum (Ta: anterior, Tp: posterior position), t: thin bacilliform sensillum, ts: short thin bacilliform sensillum, S: setiform and C: conical sensillum)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F36E1BC14086B94C5F90F0E009C40" blockId="6.[151,1437,1669,2020]" lastBlockId="7.[151,1437,151,645]" lastPageId="7" lastPageNumber="400" pageId="6" pageNumber="399">
|
||
<emphasis id="B954EAF3BC14086A94C5F90F0D719BA1" bold="true" box="[199,263,1670,1696]" pageId="6" pageNumber="399">Head</emphasis>
|
||
: Each side 8 ommatidia: 4 dorsal, 4 lateral (1 anterior, 2 medial and 1 posterior). Vertex with 2 posterior trichome groups (pv) and a large medial gap. Each group with 2 rows: Anterior row curved slightly, on an oblique angle with large trichome sockets in central position, gradually reducing the size at both ends (
|
||
<typeStatus id="549B8843BC14086A90D5F944094D9BE6" box="[1239,1339,1741,1767]" pageId="6" pageNumber="399" type="holotype">holotype</typeStatus>
|
||
12 each side); posterior row with few small sockets (
|
||
<typeStatus id="549B8843BC14086A9687F9780E9F9A0A" box="[645,745,1777,1803]" pageId="6" pageNumber="399" type="holotype">holotype</typeStatus>
|
||
2 each side); a narrow space between these rows. (
|
||
<figureCitation id="131B2A64BC14086A911DF9780CA19A31" captionStart="FIGURE 3" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[184,1403,191,1791]" captionTargetId="figure-19@5.[184,1404,181,1800]" captionTargetPageId="5" captionText="FIGURE 3. Holotype male Monographis cattienensis sp. n.. A. The head capsule showing eye patterns (each side with eight ommatidia); trichobothria: trichobothrium a, b and c; anterior vertex (av) and posterior vertex (pv). B. The enlarge trichome sockets in posterior vertex, pv (L): left side, pv (R): right side. C. Trichobothria (left side): trichobothrium a (interior position), b (lateral) and c (exterior); D. Right gnathochilarium showing lateral palp (Lp) with 11 conical sensilla and medial palp (Mp) with 22; E. Clypeo-labrum showing setose surface, with two lateral lamellae (Ll), two lamellae (l) and a median cleft (mc), anterior setae (sa); posterior setae (Se); a row of backward-pointing setae (sb)." figureDoi="http://doi.org/10.5281/zenodo.7389283" httpUri="https://zenodo.org/record/7389283/files/figure.png" pageId="6" pageNumber="399">Figures 3A & 3B</figureCitation>
|
||
).
|
||
<typeStatus id="549B8843BC14086A94E9F89F0D2E9A31" box="[235,344,1814,1840]" pageId="6" pageNumber="399" type="paratype">Paratypes</typeStatus>
|
||
showed that variation is common in this species, regardless of sex, ranging from 12–16 sockets in anterior rows,
|
||
<quantity id="4CD89B04BC14086A9538F8B30DF49A52" box="[314,386,1849,1876]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="6" pageNumber="399" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">2–3 in</quantity>
|
||
posterior rows.
|
||
<emphasis id="B954EAF3BC14086A963AF8B00EA99A52" bold="true" box="[568,735,1849,1875]" pageId="6" pageNumber="399">Trichobothria</emphasis>
|
||
: typically thin, sensory hairs with narrow cylindrical funicles; trichobothria equal in socket size forming an isosceles triangle with equal distance
|
||
<emphasis id="B954EAF3BC14086A9058F8D408009A79" bold="true" box="[1114,1142,1885,1912]" italics="true" pageId="6" pageNumber="399">ab</emphasis>
|
||
and
|
||
<emphasis id="B954EAF3BC14086A90B3F8D408BD9A79" bold="true" box="[1201,1227,1885,1912]" italics="true" pageId="6" pageNumber="399">bc</emphasis>
|
||
, trichobothrium
|
||
<emphasis id="B954EAF3BC14086A918CF8D709EA9A79" bold="true" box="[1422,1436,1886,1912]" italics="true" pageId="6" pageNumber="399">a</emphasis>
|
||
located in interior position, trichobothrium
|
||
<emphasis id="B954EAF3BC14086A9684F8080EE29A9D" bold="true" box="[646,660,1921,1948]" italics="true" pageId="6" pageNumber="399">b</emphasis>
|
||
in lateral position, trichobothrium
|
||
<emphasis id="B954EAF3BC14086A9027F80B08479A9D" bold="true" box="[1061,1073,1922,1948]" italics="true" pageId="6" pageNumber="399">c</emphasis>
|
||
in exterior position of the head capsule (
|
||
<figureCitation id="131B2A64BC14086A94FFF82C0DA49AC1" box="[253,466,1957,1984]" captionStart="FIGURE 3" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[184,1403,191,1791]" captionTargetId="figure-19@5.[184,1404,181,1800]" captionTargetPageId="5" captionText="FIGURE 3. Holotype male Monographis cattienensis sp. n.. A. The head capsule showing eye patterns (each side with eight ommatidia); trichobothria: trichobothrium a, b and c; anterior vertex (av) and posterior vertex (pv). B. The enlarge trichome sockets in posterior vertex, pv (L): left side, pv (R): right side. C. Trichobothria (left side): trichobothrium a (interior position), b (lateral) and c (exterior); D. Right gnathochilarium showing lateral palp (Lp) with 11 conical sensilla and medial palp (Mp) with 22; E. Clypeo-labrum showing setose surface, with two lateral lamellae (Ll), two lamellae (l) and a median cleft (mc), anterior setae (sa); posterior setae (Se); a row of backward-pointing setae (sb)." figureDoi="http://doi.org/10.5281/zenodo.7389283" httpUri="https://zenodo.org/record/7389283/files/figure.png" pageId="6" pageNumber="399">Figures. 3A & 3C</figureCitation>
|
||
).
|
||
<emphasis id="B954EAF3BC14086A95EBF82C0ECE9ABE" bold="true" box="[489,696,1957,1983]" pageId="6" pageNumber="399">Gnathochilarium</emphasis>
|
||
: Lateral palp 1.25 times medial palp; 11 conical sensilla on the lateral palp and 22 on the medial palp, same in
|
||
<typeStatus id="549B8843BC14086A96BCF8400F569AE2" box="[702,800,1993,2019]" pageId="6" pageNumber="399" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="549B8843BC14086A9758F8430FB39AE5" box="[858,965,1994,2020]" pageId="6" pageNumber="399" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<figureCitation id="131B2A64BC14086A97D4F84008269AE5" box="[982,1104,1993,2020]" captionStart="FIGURE 3" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[184,1403,191,1791]" captionTargetId="figure-19@5.[184,1404,181,1800]" captionTargetPageId="5" captionText="FIGURE 3. Holotype male Monographis cattienensis sp. n.. A. The head capsule showing eye patterns (each side with eight ommatidia); trichobothria: trichobothrium a, b and c; anterior vertex (av) and posterior vertex (pv). B. The enlarge trichome sockets in posterior vertex, pv (L): left side, pv (R): right side. C. Trichobothria (left side): trichobothrium a (interior position), b (lateral) and c (exterior); D. Right gnathochilarium showing lateral palp (Lp) with 11 conical sensilla and medial palp (Mp) with 22; E. Clypeo-labrum showing setose surface, with two lateral lamellae (Ll), two lamellae (l) and a median cleft (mc), anterior setae (sa); posterior setae (Se); a row of backward-pointing setae (sb)." figureDoi="http://doi.org/10.5281/zenodo.7389283" httpUri="https://zenodo.org/record/7389283/files/figure.png" pageId="6" pageNumber="399">Figure 3D</figureCitation>
|
||
).
|
||
<emphasis id="B954EAF3BC14086A9064F840096A9AE2" bold="true" box="[1126,1308,1993,2019]" pageId="6" pageNumber="399">Clypeo-labrum</emphasis>
|
||
:
|
||
<typeStatus id="549B8843BC14086A912FF84009EE9AE2" box="[1325,1432,1993,2019]" pageId="6" pageNumber="399" type="holotype">Holotype</typeStatus>
|
||
, labrum posterior margin with 12 setae (Se), setae equal in length to labrum width;
|
||
<typeStatus id="549B8843BC15086B9040FF1E08C49DB0" box="[1090,1202,151,177]" pageId="7" pageNumber="400" type="paratype">Paratypes</typeStatus>
|
||
, 10–12 setae in both sexes. Labrum surface setose (l), labrum posterior margin has a row of minute backward-pointing setae (sb). Lateral lamella (Ll) and single broad lamella each side of median cleft (mc); 7 (Left: L) and 12 (Right: R) slender setae (sa) present on anterior margin of labrum. The number of these anterior slender setae varies among
|
||
<typeStatus id="549B8843BC15086B90CFFE8A094D9C1C" box="[1229,1339,259,285]" pageId="7" pageNumber="400" type="paratype">paratypes</typeStatus>
|
||
, ranging from 18–20 across both sides. (
|
||
<figureCitation id="131B2A64BC15086B95F7FEAE0E1C9C40" box="[501,618,295,321]" captionStart="FIGURE 3" captionStartId="5.[152,255,1823,1847]" captionTargetBox="[184,1403,191,1791]" captionTargetId="figure-19@5.[184,1404,181,1800]" captionTargetPageId="5" captionText="FIGURE 3. Holotype male Monographis cattienensis sp. n.. A. The head capsule showing eye patterns (each side with eight ommatidia); trichobothria: trichobothrium a, b and c; anterior vertex (av) and posterior vertex (pv). B. The enlarge trichome sockets in posterior vertex, pv (L): left side, pv (R): right side. C. Trichobothria (left side): trichobothrium a (interior position), b (lateral) and c (exterior); D. Right gnathochilarium showing lateral palp (Lp) with 11 conical sensilla and medial palp (Mp) with 22; E. Clypeo-labrum showing setose surface, with two lateral lamellae (Ll), two lamellae (l) and a median cleft (mc), anterior setae (sa); posterior setae (Se); a row of backward-pointing setae (sb)." figureDoi="http://doi.org/10.5281/zenodo.7389283" httpUri="https://zenodo.org/record/7389283/files/figure.png" pageId="7" pageNumber="400">Figure 3E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F36E1BC15086B94C5FEC30D399F18" blockId="7.[151,1437,151,645]" pageId="7" pageNumber="400">
|
||
<emphasis id="B954EAF3BC15086B94C5FEC30D4C9C65" bold="true" box="[199,314,330,356]" pageId="7" pageNumber="400">Antennae</emphasis>
|
||
:
|
||
<materialsCitation id="3B483CBCBC15086B954AFEC20D399F18" ID-GBIF-Occurrence="3981599303" collectionCode="VI, T" pageId="7" pageNumber="400" specimenCount="1" typeStatus="holotype">
|
||
Antennal article
|
||
<collectionCode id="ED31AE24BC15086B9601FEC20E579C64" box="[515,545,331,357]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="7" pageNumber="400">VI</collectionCode>
|
||
with 15–16 bacilliform sensilla.
|
||
<typeStatus id="549B8843BC15086B9794FEC20F889C64" box="[918,1022,331,357]" pageId="7" pageNumber="400" type="holotype">Holotype</typeStatus>
|
||
with 16 bacilliform sensilla forming a crescent shape with 3 thick (
|
||
<collectionCode id="ED31AE24BC15086B95F0FEE60E739C88" box="[498,517,367,393]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="400">T</collectionCode>
|
||
) and 8 thin bacilliform sensilla (t), 5 short thin bacilliform sensilla (ts); setiform sensillum (S) located in anterior position, conical sensillum (C) present near the centre of the crescent (
|
||
<figureCitation id="131B2A64BC15086B911AFE1A09EE9CAC" box="[1304,1432,403,429]" captionStart="FIGURE 4" captionStartId="6.[152,255,1527,1551]" captionTargetBox="[154,1433,424,1492]" captionTargetId="figure-121@6.[151,1436,417,1504]" captionTargetPageId="6" captionText="FIGURE 4. Holotype male Monographis cattienensis sp. n.. A. Antennal articles VI, VII, VIII and sensory cones (sc); B. and C. showing the arrangements of sensilla on articles VI and VII. (T: thick bacilliform sensillum (Ta: anterior, Tp: posterior position), t: thin bacilliform sensillum, ts: short thin bacilliform sensillum, S: setiform and C: conical sensillum)" figureDoi="http://doi.org/10.5281/zenodo.7389285" httpUri="https://zenodo.org/record/7389285/files/figure.png" pageId="7" pageNumber="400">Figures 4C</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A64BC15086B9495FE3E0D759CD0" box="[151,259,439,465]" captionStart="FIGURE 7" captionStartId="9.[152,255,1865,1889]" captionTargetBox="[214,1372,525,1842]" captionTargetId="figure-185@9.[214,1374,525,1842]" captionTargetPageId="9" captionText="FIGURE 7. SEM images of Monographis cattienensis sp. n.. A. antennal articles VI, VII and VIII; B. and C. showing the arrangement of sensilla on the article VI; D. Clypeo-labrum showing setose surface with anterior setae (sa), lateral lamella (Ll), median cleft (mc), row of backward-pointing setae (sb) and posterior setae (Se); E. Leg articles and two spines (pointed arrows) which are typical characters of Monographis; F. Telotarsus showing processes: anterior lateral process (a), posterior lateral process (p), claw (c), anterior setiform process (s) and lamella (la)." figureDoi="http://doi.org/10.5281/zenodo.7389299" httpUri="https://zenodo.org/record/7389299/files/figure.png" pageId="7" pageNumber="400">7B & 7C</figureCitation>
|
||
). Antennal article VII with 2 thick bacilliform sensilla (
|
||
<collectionCode id="ED31AE24BC15086B977CFE3E0FE79CD0" box="[894,913,439,465]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="400">T</collectionCode>
|
||
, Ta and Tp), setiform sensillum (S) located in between these two sensilla (
|
||
<collectionCode id="ED31AE24BC15086B95D2FE520D929CF4" box="[464,484,475,501]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="400">T</collectionCode>
|
||
), conical sensillum (C) located next to Tp (
|
||
<figureCitation id="131B2A64BC15086B97C4FE52084D9CF4" box="[966,1083,475,501]" captionStart="FIGURE 4" captionStartId="6.[152,255,1527,1551]" captionTargetBox="[154,1433,424,1492]" captionTargetId="figure-121@6.[151,1436,417,1504]" captionTargetPageId="6" captionText="FIGURE 4. Holotype male Monographis cattienensis sp. n.. A. Antennal articles VI, VII, VIII and sensory cones (sc); B. and C. showing the arrangements of sensilla on articles VI and VII. (T: thick bacilliform sensillum (Ta: anterior, Tp: posterior position), t: thin bacilliform sensillum, ts: short thin bacilliform sensillum, S: setiform and C: conical sensillum)" figureDoi="http://doi.org/10.5281/zenodo.7389285" httpUri="https://zenodo.org/record/7389285/files/figure.png" pageId="7" pageNumber="400">Figure 4B</figureCitation>
|
||
), which is an arrangement typical of the genus.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9F36E1BC15086B94C5FDAA0F7C9F84" blockId="7.[151,1437,151,645]" pageId="7" pageNumber="400">
|
||
<emphasis id="B954EAF3BC15086B94C5FDAA0D7D9F3C" bold="true" box="[199,267,547,573]" pageId="7" pageNumber="400">Body.</emphasis>
|
||
Collum—the first tergite with a pair of pleural projections called lateral protuberances (Lp) (
|
||
<figureCitation id="131B2A64BC15086B9115FDAA09F99F3C" box="[1303,1423,546,573]" captionStart="FIGURE 5" captionStartId="7.[152,255,1467,1491]" captionTargetBox="[157,1432,674,1440]" captionTargetId="figure-243@7.[151,1436,669,1444]" captionTargetPageId="7" captionText="FIGURE 5. Holotype male Monographis cattienensis sp. n. with typical trichome socket patterns on the collum and tergites of Monographis species. A. Collum (Col)—the first tergite showing trichome patterns and the first pair of pleural projections—lateral protuberances (Lp); B. Trichome socket patterns present on tergites 2 (T2) to 9 (T9), and pleural projections (pp); C. Trichome socket patterns present on tergite 10 (T10)—the last tergite, and its pleural projections, the trichome sockets showing dense and smaller than other tergite." figureDoi="http://doi.org/10.5281/zenodo.7389289" httpUri="https://zenodo.org/record/7389289/files/figure.png" pageId="7" pageNumber="400">Figure 5A</figureCitation>
|
||
); tergites 2 (T2) to 10 (T10), each connected with a pair of pleural projections (pp) (
|
||
<figureCitation id="131B2A64BC15086B902EFDCE08D79F60" box="[1068,1185,582,609]" captionStart="FIGURE 5" captionStartId="7.[152,255,1467,1491]" captionTargetBox="[157,1432,674,1440]" captionTargetId="figure-243@7.[151,1436,669,1444]" captionTargetPageId="7" captionText="FIGURE 5. Holotype male Monographis cattienensis sp. n. with typical trichome socket patterns on the collum and tergites of Monographis species. A. Collum (Col)—the first tergite showing trichome patterns and the first pair of pleural projections—lateral protuberances (Lp); B. Trichome socket patterns present on tergites 2 (T2) to 9 (T9), and pleural projections (pp); C. Trichome socket patterns present on tergite 10 (T10)—the last tergite, and its pleural projections, the trichome sockets showing dense and smaller than other tergite." figureDoi="http://doi.org/10.5281/zenodo.7389289" httpUri="https://zenodo.org/record/7389289/files/figure.png" pageId="7" pageNumber="400">Figure 5B</figureCitation>
|
||
); tergite T10—the last tergite, trichome sockets dense and smaller (
|
||
<figureCitation id="131B2A64BC15086B9685FDE20E889F84" box="[647,766,618,645]" captionStart="FIGURE 5" captionStartId="7.[152,255,1467,1491]" captionTargetBox="[157,1432,674,1440]" captionTargetId="figure-243@7.[151,1436,669,1444]" captionTargetPageId="7" captionText="FIGURE 5. Holotype male Monographis cattienensis sp. n. with typical trichome socket patterns on the collum and tergites of Monographis species. A. Collum (Col)—the first tergite showing trichome patterns and the first pair of pleural projections—lateral protuberances (Lp); B. Trichome socket patterns present on tergites 2 (T2) to 9 (T9), and pleural projections (pp); C. Trichome socket patterns present on tergite 10 (T10)—the last tergite, and its pleural projections, the trichome sockets showing dense and smaller than other tergite." figureDoi="http://doi.org/10.5281/zenodo.7389289" httpUri="https://zenodo.org/record/7389289/files/figure.png" pageId="7" pageNumber="400">Figure 5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF5F6669BC15086B949AFA320E739B62" ID-DOI="http://doi.org/10.5281/zenodo.7389289" ID-Zenodo-Dep="7389289" httpUri="https://zenodo.org/record/7389289/files/figure.png" pageId="7" pageNumber="400" startId="7.[152,255,1467,1491]" targetBox="[157,1432,674,1440]" targetPageId="7">
|
||
<paragraph id="8B9F36E1BC15086B949AFA320E739B62" blockId="7.[151,1437,1467,1635]" pageId="7" pageNumber="400">
|
||
<emphasis id="B954EAF3BC15086B949AFA320D6198D5" bold="true" box="[152,279,1467,1492]" pageId="7" pageNumber="400">FIGURE 5.</emphasis>
|
||
Holotype male
|
||
<taxonomicName id="4C204D62BC15086B95BAFA320EC998D2" authority="Huynh, 2022" authorityName="Huynh" authorityYear="2022" box="[440,703,1467,1491]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="7" pageNumber="400" phylum="Arthropoda" rank="species" species="cattienensis" status="sp. nov.">
|
||
<emphasis id="B954EAF3BC15086B95BAFA320EC998D2" box="[440,703,1467,1491]" italics="true" pageId="7" pageNumber="400">Monographis cattienensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EAF3BC15086B96C7FA350E8B98D5" bold="true" box="[709,765,1468,1492]" pageId="7" pageNumber="400">
|
||
<taxonomicNameLabel id="A2675788BC15086B96C7FA350E8B98D5" box="[709,765,1468,1492]" pageId="7" pageNumber="400" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
with typical trichome socket patterns on the collum and tergites of
|
||
<taxonomicName id="4C204D62BC15086B9495FA560DFB98F6" box="[151,397,1503,1527]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="7" pageNumber="400" phylum="Arthropoda" rank="species" species="undefined-a">
|
||
<emphasis id="B954EAF3BC15086B9495FA560D5698F6" box="[151,288,1503,1527]" italics="true" pageId="7" pageNumber="400">Monographis</emphasis>
|
||
species.
|
||
<emphasis id="B954EAF3BC15086B957BFA560DFB98F6" bold="true" box="[377,397,1503,1527]" pageId="7" pageNumber="400">A</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EAF3BC15086B958FFA560DE498F6" bold="true" box="[397,402,1503,1527]" pageId="7" pageNumber="400">.</emphasis>
|
||
Collum (Col)—the first tergite showing trichome patterns and the first pair of pleural projections—lateral protuberances (Lp);
|
||
<emphasis id="B954EAF3BC15086B9596F98D0DDD9B1D" bold="true" box="[404,427,1540,1564]" pageId="7" pageNumber="400">B.</emphasis>
|
||
Trichome socket patterns present on tergites 2 (T2) to 9 (T9), and pleural projections (pp);
|
||
<emphasis id="B954EAF3BC15086B9154F98A09199B1A" bold="true" box="[1366,1391,1539,1563]" pageId="7" pageNumber="400">C.</emphasis>
|
||
Trichome socket patterns present on tergite 10 (T10)—the last tergite, and its pleural projections, the trichome sockets showing dense and smaller than other tergite.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F36E1BC15086B94C5F91B0CA09A85" blockId="7.[151,1437,1681,2032]" pageId="7" pageNumber="400">
|
||
<emphasis id="B954EAF3BC15086B94C5F91B0D709BAD" bold="true" box="[199,262,1682,1708]" pageId="7" pageNumber="400">Legs.</emphasis>
|
||
Leg segmentation following
|
||
<bibRefCitation id="EFB14B10BC15086B964DF91B0E819BAD" author="Manton, S. M." box="[591,759,1681,1708]" pageId="7" pageNumber="400" pagination="153 - 187" refId="ref11807" refString="Manton, S. M. (1956) The Evolution of Arthropodan Locomotory Mechanisms - Part 5: The Structure, Habits and Evolution of the Pselaphognatha (Diplopoda). Journal of the Linnean Society London, 43, 153 - 187. https: // doi. org / 10.1111 / j. 1096 - 3642.1957. tb 02516. x" type="journal article" year="1956">Manton (1956)</bibRefCitation>
|
||
and with articles typical of
|
||
<taxonomicName id="4C204D62BC15086B902CF91B08C79BAD" authorityName="Verhoeff" authorityYear="1934" box="[1070,1201,1682,1708]" class="Diplopoda" kingdom="Animalia" order="Polyxenida" pageId="7" pageNumber="400" phylum="Arthropoda" rank="order">Polyxenida</taxonomicName>
|
||
: legs 1 and 2 without trochanter, leg 1 without tarsus 1. Chaetotaxy (setae on leg articles): coxa 1 and 2: 2 setae, coxae 3–13: 2–4 setae; pre-femur, post-femur, tibia with 1 seta, except femur with 1–4 setae, tarsus 1 and 2 with a spine. Coxa, pre-femur and femur with biarticulated seta and ridged funicle. 1–3 smaller biarticulated setae in posterior position of femur, some also on penis. Post-femur and tibia distally with setiform seta. The spine on tarsus 2 is sharply pointed and twice the length of a spine in the antero-sternal position of tarsus 1 (
|
||
<figureCitation id="131B2A64BC15086B97AFF8CF08249A61" box="[941,1106,1862,1888]" captionStart="FIGURE 6" captionStartId="8.[152,255,1818,1842]" captionTargetBox="[221,1360,193,1780]" captionTargetId="figure-29@8.[216,1371,181,1795]" captionTargetPageId="8" captionText="FIGURE 6. Holotype male Monographis cattienensis sp. n.. A. The second leg showing the segments and the chaetotaxy— hairs on its segments: Coxa (co), prefemur (pf), femur (f), postfemur (pof), tibia (tib), tarsus 1 (T1), tarsus 2 (T2) and claw (c); B. Typical seta present on leg articles; C. Spine on Tarsus 1 & 2; D. Telotarsus: Claw (c), lamella (la), setiform (s) and anterior (a), posterior (p) lateral processes. E. Two pairs of coxal glands present on coxae VIII and IX; F. Dorsal ornamental trichome sockets showing both sides, trichome sockets a: a1 to a18 (Left), with 17 sockets (Right); trichome socket b; trichome sockets c: c1, c2 and c3. Circular indentation d; G. Female’s nest trichome; H. A caudal bundle trichome." figureDoi="http://doi.org/10.5281/zenodo.7389293" httpUri="https://zenodo.org/record/7389293/files/figure.png" pageId="7" pageNumber="400">Figures 6A–C</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A64BC15086B905DF8CF08F79A61" box="[1119,1153,1862,1888]" captionStart="FIGURE 7" captionStartId="9.[152,255,1865,1889]" captionTargetBox="[214,1372,525,1842]" captionTargetId="figure-185@9.[214,1374,525,1842]" captionTargetPageId="9" captionText="FIGURE 7. SEM images of Monographis cattienensis sp. n.. A. antennal articles VI, VII and VIII; B. and C. showing the arrangement of sensilla on the article VI; D. Clypeo-labrum showing setose surface with anterior setae (sa), lateral lamella (Ll), median cleft (mc), row of backward-pointing setae (sb) and posterior setae (Se); E. Leg articles and two spines (pointed arrows) which are typical characters of Monographis; F. Telotarsus showing processes: anterior lateral process (a), posterior lateral process (p), claw (c), anterior setiform process (s) and lamella (la)." figureDoi="http://doi.org/10.5281/zenodo.7389299" httpUri="https://zenodo.org/record/7389299/files/figure.png" pageId="7" pageNumber="400">7E</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A64BC15086B908DF8CF08B79A61" box="[1167,1217,1862,1888]" captionStart="FIGURE 10" captionStartId="12.[152,255,1912,1936]" captionTargetBox="[155,1430,752,1879]" captionTargetId="figure-282@12.[151,1436,741,1889]" captionTargetPageId="12" captionText="FIGURE 10. Holotype female Monographis konkakinhensis sp. n.. A. The second leg showing its segments and its chaetotaxy, vulva (v); B. Telotarsus and its processes, dotted line indicating setiform (s) shorter than the claw (c); C. Dorsal ornamental trichome sockets showing both sides with 25 sockets a, one b, three c (c1, c2 & c3) and the indentation d on each side." figureDoi="http://doi.org/10.5281/zenodo.7389313" httpUri="https://zenodo.org/record/7389313/files/figure.png" pageId="7" pageNumber="400">10A</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A64BC15086B90CDF8CF09769A61" box="[1231,1280,1862,1888]" captionStart="FIGURE 12" captionStartId="14.[152,255,1228,1252]" captionTargetBox="[151,1436,181,1204]" captionTargetId="figure-29@14.[151,1436,181,1204]" captionTargetPageId="14" captionText="FIGURE 12. SEM images of a female Monographis konkakinhensis sp. n.. A. The vulvae (v) in coxae II; B. Telson (Te) showing the ornamental trichome sockets (ots) dorsally, trichome bundle structures with the main caudal barbate trichome sockets (cbts), few barbate trichome sockets (bts) in the centre of the bundle, two bundles of nest trichome sockets (nts) ventrally; C. Leg segments and two spines (pointed arrows) typical Monographis; D. Telotarsus and its processes (a: anterior lateral process, p: posterior lateral process, c: claw, s: anterior setiform process, la; lamella process)." figureDoi="http://doi.org/10.5281/zenodo.7389321" httpUri="https://zenodo.org/record/7389321/files/figure.png" pageId="7" pageNumber="400">12C</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A64BC15086B910CF8CC09379A61" box="[1294,1345,1861,1888]" captionStart="FIGURE 15" captionStartId="17.[152,255,1104,1128]" captionTargetBox="[290,1295,188,1070]" captionTargetId="figure-19@17.[289,1298,181,1080]" captionTargetPageId="17" captionText="FIGURE 15. Holotype male Monographis tamdaoensis sp. n.. A. The second leg showing its segments and chaetotaxy, penis (p). B. Telotarsus showing the claw (c) and processes: anterior (a), posterior (p), lamella (la) and anterior setiform process (s) shorter than half length of the claw (dot line). C. The ornamental trichome sockets with trichomes a, b, three c and an indented d." figureDoi="http://doi.org/10.5281/zenodo.7389333" httpUri="https://zenodo.org/record/7389333/files/figure.png" pageId="7" pageNumber="400">15A</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A64BC15086B914DF8CF09089A61" box="[1359,1406,1862,1888]" captionStart="FIGURE 20" captionStartId="21.[152,255,1896,1920]" captionTargetBox="[163,1413,191,1870]" captionTargetId="figure-19@21.[158,1430,181,1872]" captionTargetPageId="21" captionText="FIGURE 20. Holotype male Monographis thatsonensis sp. n.. A. Second leg showing seven articles: coxa (co), prefemur (pf), femur (f), postfemur (pof), tibia (tib), tarsus 1 (T1), tarsus 2 (T2); claw (c) and penis (p). B. Telotarsus showing claw (c) and all processes: anterior (a), posterior (p), setiform (s) and the lamella (la). C. Ornamental trichome sockets showing the arrangements on both sides: trichome sockets a, b, c, and indentation d." figureDoi="http://doi.org/10.5281/zenodo.7389352" httpUri="https://zenodo.org/record/7389352/files/figure.png" pageId="7" pageNumber="400">20A</figureCitation>
|
||
&
|
||
<figureCitation id="131B2A64BC15086B9495F8E30CBC9A85" box="[151,202,1898,1924]" captionStart="FIGURE 23" captionStartId="23.[152,255,1170,1194]" captionTargetBox="[152,1436,181,1146]" captionTargetId="figure-19@23.[151,1436,181,1146]" captionTargetPageId="23" captionText="FIGURE 23. SEM images of Monographis thatsonensis sp. n.. A. Male’s caudal bundle with ornamental trichome sockets (ots) a, b, c and d; and telson (Te), and the bundle with caudal bundle trichome sockets (cbts), barbate trichome sockets (bts); the last pleural projection (pp). B. Female’s caudal bundle structure showing the same dorsally, except the presence of two bundles of nest trichome sockets ventrally (nts). C. Presence of two spines—Tarsus 1 and 2, typical of Monographis species. D. Telotarsus showing a claw (c) and all processes: anterior (a), posterior (p), anterior setiform process (s) and lamella (la)." figureDoi="http://doi.org/10.5281/zenodo.7389361" httpUri="https://zenodo.org/record/7389361/files/figure.png" pageId="7" pageNumber="400">23C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F36E1BC15086B94C5F8070CB39AF1" blockId="7.[151,1437,1681,2032]" pageId="7" pageNumber="400">
|
||
<emphasis id="B954EAF3BC15086B94C5F8070D3A9AA9" bold="true" box="[199,332,1934,1960]" pageId="7" pageNumber="400">Telotarsus:</emphasis>
|
||
Claw slender with anterior lateral (a) and posterior lateral (p) processes less than half length of the claw (c); the anterior setiform process (s) is longer than the claw; lamella process (la) is present (
|
||
<figureCitation id="131B2A64BC15086B9102F83B09099ACD" box="[1280,1407,1970,1996]" captionStart="FIGURE 6" captionStartId="8.[152,255,1818,1842]" captionTargetBox="[221,1360,193,1780]" captionTargetId="figure-29@8.[216,1371,181,1795]" captionTargetPageId="8" captionText="FIGURE 6. Holotype male Monographis cattienensis sp. n.. A. The second leg showing the segments and the chaetotaxy— hairs on its segments: Coxa (co), prefemur (pf), femur (f), postfemur (pof), tibia (tib), tarsus 1 (T1), tarsus 2 (T2) and claw (c); B. Typical seta present on leg articles; C. Spine on Tarsus 1 & 2; D. Telotarsus: Claw (c), lamella (la), setiform (s) and anterior (a), posterior (p) lateral processes. E. Two pairs of coxal glands present on coxae VIII and IX; F. Dorsal ornamental trichome sockets showing both sides, trichome sockets a: a1 to a18 (Left), with 17 sockets (Right); trichome socket b; trichome sockets c: c1, c2 and c3. Circular indentation d; G. Female’s nest trichome; H. A caudal bundle trichome." figureDoi="http://doi.org/10.5281/zenodo.7389293" httpUri="https://zenodo.org/record/7389293/files/figure.png" pageId="7" pageNumber="400">Figures 6D</figureCitation>
|
||
&
|
||
<figureCitation id="131B2A64BC15086B9495F85F0CCE9AF1" box="[151,184,2006,2032]" captionStart="FIGURE 7" captionStartId="9.[152,255,1865,1889]" captionTargetBox="[214,1372,525,1842]" captionTargetId="figure-185@9.[214,1374,525,1842]" captionTargetPageId="9" captionText="FIGURE 7. SEM images of Monographis cattienensis sp. n.. A. antennal articles VI, VII and VIII; B. and C. showing the arrangement of sensilla on the article VI; D. Clypeo-labrum showing setose surface with anterior setae (sa), lateral lamella (Ll), median cleft (mc), row of backward-pointing setae (sb) and posterior setae (Se); E. Leg articles and two spines (pointed arrows) which are typical characters of Monographis; F. Telotarsus showing processes: anterior lateral process (a), posterior lateral process (p), claw (c), anterior setiform process (s) and lamella (la)." figureDoi="http://doi.org/10.5281/zenodo.7389299" httpUri="https://zenodo.org/record/7389299/files/figure.png" pageId="7" pageNumber="400">7F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF5F6669BC1A0864949AF89308059AE6" ID-DOI="http://doi.org/10.5281/zenodo.7389293" ID-Zenodo-Dep="7389293" httpUri="https://zenodo.org/record/7389293/files/figure.png" pageId="8" pageNumber="401" startId="8.[152,255,1818,1842]" targetBox="[221,1360,193,1780]" targetPageId="8">
|
||
<paragraph id="8B9F36E1BC1A0864949AF89308059AE6" blockId="8.[151,1437,1818,2023]" pageId="8" pageNumber="401">
|
||
<emphasis id="B954EAF3BC1A0864949AF8930D6F9A33" bold="true" box="[152,281,1818,1842]" pageId="8" pageNumber="401">FIGURE 6.</emphasis>
|
||
Holotype male
|
||
<taxonomicName id="4C204D62BC1A086495C0F8920EBC9A33" authority="Huynh, 2022" authorityName="Huynh" authorityYear="2022" box="[450,714,1819,1843]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="8" pageNumber="401" phylum="Arthropoda" rank="species" species="cattienensis" status="sp. nov.">
|
||
<emphasis id="B954EAF3BC1A086495C0F8920EBC9A33" box="[450,714,1819,1843]" italics="true" pageId="8" pageNumber="401">Monographis cattienensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EAF3BC1A086496D3F8920F7D9A32" bold="true" box="[721,779,1819,1843]" pageId="8" pageNumber="401">
|
||
<taxonomicNameLabel id="A2675788BC1A086496D3F8920F7D9A32" box="[721,779,1819,1843]" pageId="8" pageNumber="401" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B954EAF3BC1A0864971AF8930F479A33" bold="true" box="[792,817,1818,1842]" pageId="8" pageNumber="401">A.</emphasis>
|
||
The second leg showing the segments and the chaetotaxy— hairs on its segments: Coxa (co), prefemur (pf), femur (f), postfemur (pof), tibia (tib), tarsus 1 (T1), tarsus 2 (T2) and claw (c);
|
||
<emphasis id="B954EAF3BC1A08649495F8EA0CD89A7A" bold="true" box="[151,174,1891,1915]" pageId="8" pageNumber="401">B.</emphasis>
|
||
Typical seta present on leg articles;
|
||
<emphasis id="B954EAF3BC1A0864961DF8EB0E4E9A7B" bold="true" box="[543,568,1890,1914]" pageId="8" pageNumber="401">C.</emphasis>
|
||
Spine on Tarsus 1 & 2;
|
||
<emphasis id="B954EAF3BC1A08649724F8EA0F499A7A" bold="true" box="[806,831,1891,1915]" pageId="8" pageNumber="401">D.</emphasis>
|
||
Telotarsus: Claw (c), lamella (la), setiform (s) and anterior (a), posterior (p) lateral processes.
|
||
<emphasis id="B954EAF3BC1A086495FCF80E0E639A9E" bold="true" box="[510,533,1927,1951]" pageId="8" pageNumber="401">E.</emphasis>
|
||
Two pairs of coxal glands present on coxae VIII and IX;
|
||
<emphasis id="B954EAF3BC1A08649067F80E080E9A9E" bold="true" box="[1125,1144,1927,1951]" pageId="8" pageNumber="401">F.</emphasis>
|
||
Dorsal ornamental trichome sockets showing both sides, trichome sockets
|
||
<emphasis id="B954EAF3BC1A0864966AF8220E039AC3" bold="true" box="[616,629,1963,1986]" italics="true" pageId="8" pageNumber="401">a</emphasis>
|
||
:
|
||
<emphasis id="B954EAF3BC1A08649681F8220EE69AC3" bold="true" box="[643,656,1963,1986]" italics="true" pageId="8" pageNumber="401">a</emphasis>
|
||
1 to
|
||
<emphasis id="B954EAF3BC1A086496BFF8220EBC9AC3" bold="true" box="[701,714,1963,1986]" italics="true" pageId="8" pageNumber="401">a</emphasis>
|
||
18 (Left), with 17 sockets (Right); trichome socket
|
||
<emphasis id="B954EAF3BC1A086490D4F82308959AC3" bold="true" box="[1238,1251,1962,1986]" italics="true" pageId="8" pageNumber="401">b</emphasis>
|
||
; trichome sockets
|
||
<emphasis id="B954EAF3BC1A08649495F8460CD49AE7" bold="true" box="[151,162,1999,2022]" italics="true" pageId="8" pageNumber="401">c</emphasis>
|
||
:
|
||
<emphasis id="B954EAF3BC1A086494B2F8460CCD9AE7" bold="true" box="[176,187,1999,2022]" italics="true" pageId="8" pageNumber="401">c</emphasis>
|
||
1,
|
||
<emphasis id="B954EAF3BC1A086494D6F8460CA99AE7" bold="true" box="[212,223,1999,2022]" italics="true" pageId="8" pageNumber="401">c</emphasis>
|
||
2 and
|
||
<emphasis id="B954EAF3BC1A0864951CF8460D5F9AE7" bold="true" box="[286,297,1999,2022]" italics="true" pageId="8" pageNumber="401">c</emphasis>
|
||
3. Circular indentation
|
||
<emphasis id="B954EAF3BC1A08649616F8470E579AE7" bold="true" box="[532,545,1998,2022]" italics="true" pageId="8" pageNumber="401">d</emphasis>
|
||
;
|
||
<emphasis id="B954EAF3BC1A0864962CF8470E3E9AE7" bold="true" box="[558,584,1998,2022]" pageId="8" pageNumber="401">G.</emphasis>
|
||
Female’s nest trichome;
|
||
<emphasis id="B954EAF3BC1A08649745F8460F179AE6" bold="true" box="[839,865,1999,2023]" pageId="8" pageNumber="401">H.</emphasis>
|
||
A caudal bundle trichome.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F36E1BC1B086594C5FF1F0DB09DD4" blockId="9.[151,1437,150,501]" pageId="9" pageNumber="402">
|
||
<emphasis id="B954EAF3BC1B086594C5FF1F0DAC9DB0" bold="true" box="[199,474,150,177]" pageId="9" pageNumber="402">Sex organs in the male:</emphasis>
|
||
A pair of penes on coxae 2 and 2 pairs of coxal glands are located on the 8th–9th coxal plates (
|
||
<figureCitation id="131B2A64BC1B086594E5FF320D159DD4" box="[231,355,187,213]" captionStart="FIGURE 6" captionStartId="8.[152,255,1818,1842]" captionTargetBox="[221,1360,193,1780]" captionTargetId="figure-29@8.[216,1371,181,1795]" captionTargetPageId="8" captionText="FIGURE 6. Holotype male Monographis cattienensis sp. n.. A. The second leg showing the segments and the chaetotaxy— hairs on its segments: Coxa (co), prefemur (pf), femur (f), postfemur (pof), tibia (tib), tarsus 1 (T1), tarsus 2 (T2) and claw (c); B. Typical seta present on leg articles; C. Spine on Tarsus 1 & 2; D. Telotarsus: Claw (c), lamella (la), setiform (s) and anterior (a), posterior (p) lateral processes. E. Two pairs of coxal glands present on coxae VIII and IX; F. Dorsal ornamental trichome sockets showing both sides, trichome sockets a: a1 to a18 (Left), with 17 sockets (Right); trichome socket b; trichome sockets c: c1, c2 and c3. Circular indentation d; G. Female’s nest trichome; H. A caudal bundle trichome." figureDoi="http://doi.org/10.5281/zenodo.7389293" httpUri="https://zenodo.org/record/7389293/files/figure.png" pageId="9" pageNumber="402">Figures 6E</figureCitation>
|
||
&
|
||
<figureCitation id="131B2A64BC1B08659585FF320DCF9DD4" box="[391,441,187,213]" captionStart="FIGURE 22" captionStartId="22.[152,255,1902,1926]" captionTargetBox="[297,1291,1236,1879]" captionTargetId="figure-94@22.[297,1291,1236,1879]" captionTargetPageId="22" captionText="FIGURE 22. SEM images of Monographis thatsonensis sp. n.. A. The head—right side showing the posterior vertex (pv) trichome sockets and trichobothria (a, b, and c). B. Coxal glands on legs VIII and IX (pointed arrows). C. Clypeo-labrum showing structure of labrum (l) with asetose surface, lateral labrum (Ll), medial cleft (mc), anterior setae (sa), posterior setae (Se), backward-pointing setae (sb). D. Gnathochilaria (g): lateral palps (Lp) and median palps (Mp), the frons (f), labrum (l)." figureDoi="http://doi.org/10.5281/zenodo.7389358" httpUri="https://zenodo.org/record/7389358/files/figure.png" pageId="9" pageNumber="402">22B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F36E1BC1B086594C5FF570E0C9C40" blockId="9.[151,1437,150,501]" pageId="9" pageNumber="402">
|
||
<emphasis id="B954EAF3BC1B086594C5FF570DFD9DF8" bold="true" box="[199,395,222,249]" pageId="9" pageNumber="402">Caudal bundles:</emphasis>
|
||
Similar to congeners (
|
||
<bibRefCitation id="EFB14B10BC1B08659681FF560FE49DF8" author="Huynh, C. & Veenstra, A." box="[643,914,223,249]" pageId="9" pageNumber="402" pagination="1 - 30" refId="ref11435" refString="Huynh, C. & Veenstra, A. (2020) Three new species of penicillate millipedes (Diplopoda, Lophoproctidae and Polyxenidae) from the Con Dao Islands of southeast Vietnam. Zootaxa, 4759 (1), 1 - 30. https: // doi. org / 10.11646 / zootaxa. 4759.1.1" type="journal article" year="2020">Huynh & Veenstra 2020</bibRefCitation>
|
||
) and classified as caudal bundle
|
||
<typeStatus id="549B8843BC1B086590F8FF56095C9DF8" box="[1274,1322,223,249]" pageId="9" pageNumber="402">type</typeStatus>
|
||
I arrangement (
|
||
<bibRefCitation id="EFB14B10BC1B086594DCFE8A0EDF9C1C" author="Conde, B. & Nguyen Duy-Jacquemin, M." box="[222,681,259,285]" pageId="9" pageNumber="402" pagination="291 - 302" refId="ref11057" refString="Conde, B. & Nguyen Duy-Jacquemin, M. (2008) Classification actuelle des Diplopodes Penicillates (Myriapodes) avec nouvelles d e finition des taxa. Bulletin de la Societe Zoologique de France, 133 (4), 291 - 302." type="journal article" year="2008">Condé & Nguyen Duy-Jacquemin, 2008</bibRefCitation>
|
||
). Caudal trichomes with a series of 2–6 backward hooks (
|
||
<bibRefCitation id="EFB14B10BC1B08659133FE8A0D4F9C40" author="Huynh, C. & Veenstra, A." pageId="9" pageNumber="402" pagination="573 - 588" refId="ref11164" refString="Huynh, C. & Veenstra, A. (2013) Taxonomy and biology of a new species of Pincushion Millipede of the genus Monographis (Diplopoda: Polyxenida) from Australia. Zootaxa, 3721 (6), 573 - 588. https: // doi. org / 10.11646 / zootaxa. 3721.6.4" type="journal article" year="2013">Huynh & Veenstra 2013</bibRefCitation>
|
||
) (
|
||
<figureCitation id="131B2A64BC1B0865954CFEAE0DAB9C40" box="[334,477,295,321]" captionStart="FIGURE 12" captionStartId="14.[152,255,1228,1252]" captionTargetBox="[151,1436,181,1204]" captionTargetId="figure-29@14.[151,1436,181,1204]" captionTargetPageId="14" captionText="FIGURE 12. SEM images of a female Monographis konkakinhensis sp. n.. A. The vulvae (v) in coxae II; B. Telson (Te) showing the ornamental trichome sockets (ots) dorsally, trichome bundle structures with the main caudal barbate trichome sockets (cbts), few barbate trichome sockets (bts) in the centre of the bundle, two bundles of nest trichome sockets (nts) ventrally; C. Leg segments and two spines (pointed arrows) typical Monographis; D. Telotarsus and its processes (a: anterior lateral process, p: posterior lateral process, c: claw, s: anterior setiform process, la; lamella process)." figureDoi="http://doi.org/10.5281/zenodo.7389321" httpUri="https://zenodo.org/record/7389321/files/figure.png" pageId="9" pageNumber="402">Figures 12B</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A64BC1B086595EAFEAE0E1B9C40" box="[488,621,295,321]" captionStart="FIGURE 23" captionStartId="23.[152,255,1170,1194]" captionTargetBox="[152,1436,181,1146]" captionTargetId="figure-19@23.[151,1436,181,1146]" captionTargetPageId="23" captionText="FIGURE 23. SEM images of Monographis thatsonensis sp. n.. A. Male’s caudal bundle with ornamental trichome sockets (ots) a, b, c and d; and telson (Te), and the bundle with caudal bundle trichome sockets (cbts), barbate trichome sockets (bts); the last pleural projection (pp). B. Female’s caudal bundle structure showing the same dorsally, except the presence of two bundles of nest trichome sockets ventrally (nts). C. Presence of two spines—Tarsus 1 and 2, typical of Monographis species. D. Telotarsus showing a claw (c) and all processes: anterior (a), posterior (p), anterior setiform process (s) and lamella (la)." figureDoi="http://doi.org/10.5281/zenodo.7389361" httpUri="https://zenodo.org/record/7389361/files/figure.png" pageId="9" pageNumber="402">23A & 23B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F36E1BC1B086594C5FEC209539CF4" blockId="9.[151,1437,150,501]" pageId="9" pageNumber="402">
|
||
<emphasis id="B954EAF3BC1B086594C5FEC20D629C64" bold="true" box="[199,276,331,357]" pageId="9" pageNumber="402">Telson</emphasis>
|
||
: Dorsal ornamental trichome sockets symmetrically arranged either side of telson; 17 (L) and 18 (R) sockets of trichomes
|
||
<emphasis id="B954EAF3BC1B08659584FEE60DE29C88" bold="true" box="[390,404,367,393]" italics="true" pageId="9" pageNumber="402">a</emphasis>
|
||
on each side of the telson in the
|
||
<typeStatus id="549B8843BC1B08659705FEE60F1F9C88" box="[775,873,367,393]" pageId="9" pageNumber="402" type="holotype">holotype</typeStatus>
|
||
(
|
||
<typeStatus id="549B8843BC1B0865977AFEE60F939C88" box="[888,997,367,393]" pageId="9" pageNumber="402" type="paratype">paratypes</typeStatus>
|
||
with 16–18). Trichome
|
||
<emphasis id="B954EAF3BC1B086590F5FEE609739C88" bold="true" box="[1271,1285,367,393]" italics="true" pageId="9" pageNumber="402">a</emphasis>
|
||
sockets form 2 rows, top row with small sockets, bottom row with larger sockets. Socket of trichome
|
||
<emphasis id="B954EAF3BC1B08659081FE1B08E79CAC" bold="true" box="[1155,1169,402,429]" italics="true" pageId="9" pageNumber="402">b</emphasis>
|
||
and 3 large sockets trichomes
|
||
<emphasis id="B954EAF3BC1B086594F6FE3E0D769CD0" bold="true" box="[244,256,439,465]" italics="true" pageId="9" pageNumber="402">c</emphasis>
|
||
with protruding base:
|
||
<emphasis id="B954EAF3BC1B086595FFFE3E0E619CD0" bold="true" box="[509,535,439,465]" italics="true" pageId="9" pageNumber="402">c1</emphasis>
|
||
,
|
||
<emphasis id="B954EAF3BC1B08659627FE3E0E499CD0" bold="true" box="[549,575,439,465]" italics="true" pageId="9" pageNumber="402">c2</emphasis>
|
||
and
|
||
<emphasis id="B954EAF3BC1B08659674FE3E0EE69CD0" bold="true" box="[630,656,439,465]" italics="true" pageId="9" pageNumber="402">c3</emphasis>
|
||
, forming triangular shape located in ventroposterior positions in each arrangement. Circular indentation
|
||
<emphasis id="B954EAF3BC1B0865961BFE530E519CF4" bold="true" box="[537,551,474,501]" italics="true" pageId="9" pageNumber="402">d</emphasis>
|
||
apparent each side near the exterior side of trichomes
|
||
<emphasis id="B954EAF3BC1B08659088FE5208E09CF4" bold="true" box="[1162,1174,475,501]" italics="true" pageId="9" pageNumber="402">c</emphasis>
|
||
(
|
||
<figureCitation id="131B2A64BC1B086590A7FE52096E9CF4" box="[1189,1304,475,501]" captionStart="FIGURE 6" captionStartId="8.[152,255,1818,1842]" captionTargetBox="[221,1360,193,1780]" captionTargetId="figure-29@8.[216,1371,181,1795]" captionTargetPageId="8" captionText="FIGURE 6. Holotype male Monographis cattienensis sp. n.. A. The second leg showing the segments and the chaetotaxy— hairs on its segments: Coxa (co), prefemur (pf), femur (f), postfemur (pof), tibia (tib), tarsus 1 (T1), tarsus 2 (T2) and claw (c); B. Typical seta present on leg articles; C. Spine on Tarsus 1 & 2; D. Telotarsus: Claw (c), lamella (la), setiform (s) and anterior (a), posterior (p) lateral processes. E. Two pairs of coxal glands present on coxae VIII and IX; F. Dorsal ornamental trichome sockets showing both sides, trichome sockets a: a1 to a18 (Left), with 17 sockets (Right); trichome socket b; trichome sockets c: c1, c2 and c3. Circular indentation d; G. Female’s nest trichome; H. A caudal bundle trichome." figureDoi="http://doi.org/10.5281/zenodo.7389293" httpUri="https://zenodo.org/record/7389293/files/figure.png" pageId="9" pageNumber="402">Figure 6F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5F6669BC1B0865949AF8C00F799AF0" ID-DOI="http://doi.org/10.5281/zenodo.7389299" ID-Zenodo-Dep="7389299" httpUri="https://zenodo.org/record/7389299/files/figure.png" pageId="9" pageNumber="402" startId="9.[152,255,1865,1889]" targetBox="[214,1372,525,1842]" targetPageId="9">
|
||
<paragraph id="8B9F36E1BC1B0865949AF8C00F799AF0" blockId="9.[151,1437,1865,2034]" pageId="9" pageNumber="402">
|
||
<emphasis id="B954EAF3BC1B0865949AF8C00D6E9A63" bold="true" box="[152,280,1865,1890]" pageId="9" pageNumber="402">FIGURE 7.</emphasis>
|
||
SEM images of
|
||
<taxonomicName id="4C204D62BC1B086595C6F8C00EBD9A60" authority="Huynh, 2022" authorityName="Huynh" authorityYear="2022" box="[452,715,1865,1889]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="402" phylum="Arthropoda" rank="species" species="cattienensis" status="sp. nov.">
|
||
<emphasis id="B954EAF3BC1B086595C6F8C00EBD9A60" box="[452,715,1865,1889]" italics="true" pageId="9" pageNumber="402">Monographis cattienensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EAF3BC1B086596D3F8C30F7C9A63" bold="true" box="[721,778,1866,1890]" pageId="9" pageNumber="402">
|
||
<taxonomicNameLabel id="A2675788BC1B086596D3F8C30F7C9A63" box="[721,778,1866,1890]" pageId="9" pageNumber="402" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B954EAF3BC1B08659715F8C00F469A60" bold="true" box="[791,816,1865,1889]" pageId="9" pageNumber="402">A.</emphasis>
|
||
antennal articles VI, VII and VIII;
|
||
<emphasis id="B954EAF3BC1B08659095F8C308D89A63" bold="true" box="[1175,1198,1866,1890]" pageId="9" pageNumber="402">B.</emphasis>
|
||
and
|
||
<emphasis id="B954EAF3BC1B086590E2F8C0088E9A60" bold="true" box="[1248,1272,1865,1889]" pageId="9" pageNumber="402">C.</emphasis>
|
||
showing the arrangement of sensilla on the article VI;
|
||
<emphasis id="B954EAF3BC1B0865962FF8E70E309A87" bold="true" box="[557,582,1902,1926]" pageId="9" pageNumber="402">D.</emphasis>
|
||
Clypeo-labrum showing setose surface with anterior setae (sa), lateral lamella (Ll), median cleft (mc), row of backward-pointing setae (sb) and posterior setae (Se);
|
||
<emphasis id="B954EAF3BC1B086597C3F81B0FAE9AAB" bold="true" box="[961,984,1938,1962]" pageId="9" pageNumber="402">E.</emphasis>
|
||
Leg articles and two spines (pointed arrows) which are typical characters of
|
||
<taxonomicName id="4C204D62BC1B086595CFF83C0E209ACC" authorityName="Attems" authorityYear="1907" box="[461,598,1973,1997]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="9" pageNumber="402" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EAF3BC1B086595CFF83C0E209ACC" box="[461,598,1973,1997]" italics="true" pageId="9" pageNumber="402">Monographis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B954EAF3BC1B08659660F83F0E039ACF" bold="true" box="[610,629,1974,1998]" pageId="9" pageNumber="402">F.</emphasis>
|
||
Telotarsus showing processes: anterior lateral process (a), posterior lateral process (p), claw (c), anterior setiform process (s) and lamella (la).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33A656ABC18086694C5FF1E09EB9DF8" pageId="10" pageNumber="403" type="discussion">
|
||
<paragraph id="8B9F36E1BC18086694C5FF1E09EB9DF8" blockId="10.[151,1437,151,321]" pageId="10" pageNumber="403">
|
||
<emphasis id="B954EAF3BC18086694C5FF1E0D429DB0" bold="true" box="[199,308,151,177]" pageId="10" pageNumber="403">Remarks</emphasis>
|
||
:
|
||
<taxonomicName id="4C204D62BC1808669541FF1E0E139DB1" authority="Huynh, 2022" authorityName="Huynh" authorityYear="2022" box="[323,613,151,177]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="403" phylum="Arthropoda" rank="species" species="cattienensis" status="sp. nov.">
|
||
<emphasis id="B954EAF3BC1808669541FF1E0E139DB1" box="[323,613,151,177]" italics="true" pageId="10" pageNumber="403">Monographis cattienensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EAF3BC180866966EFF1E0EDD9DB0" bold="true" box="[620,683,151,177]" pageId="10" pageNumber="403">
|
||
<taxonomicNameLabel id="A2675788BC180866966EFF1E0EDD9DB0" box="[620,683,151,177]" pageId="10" pageNumber="403" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, differs from other species in the genus in several characters. It has a distinctive number of bacilliform sensilla arranged in a crescent shape, setose labrum surface and slender setae present on lamellae along the anterior margin of the labrum, plus an anterior setiform process longer than the claw.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A656ABC18086694C5FE8A0E319C40" pageId="10" pageNumber="403" type="etymology">
|
||
<paragraph id="8B9F36E1BC18086694C5FE8A0E319C40" blockId="10.[151,1437,151,321]" pageId="10" pageNumber="403">
|
||
<emphasis id="B954EAF3BC18086694C5FE8A0D3E9C1C" bold="true" box="[199,328,259,285]" pageId="10" pageNumber="403">Etymology</emphasis>
|
||
:
|
||
<taxonomicName id="4C204D62BC180866955AFE8A0E0C9C1D" authorityName="Huynh" authorityYear="2022" box="[344,634,259,285]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="403" phylum="Arthropoda" rank="species" species="cattienensis">
|
||
<emphasis id="B954EAF3BC180866955AFE8A0E0C9C1D" box="[344,634,259,285]" italics="true" pageId="10" pageNumber="403">Monographis cattienensis</emphasis>
|
||
</taxonomicName>
|
||
is named after the Cat Tien National Park in
|
||
<collectingRegion id="49E4F803BC180866907FFE8A092F9C1C" box="[1149,1369,259,285]" country="Vietnam" name="Dong Nai" pageId="10" pageNumber="403">Dong Nai Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3377671BC1808669166FE8A0CBC9C40" name="Vietnam" pageId="10" pageNumber="403">Vietnam</collectingCountry>
|
||
, where this species was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |