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<mods:title id="6B374B9B14E06B93C00EC1C521BDDAE4">The bats of the Congo and of Rwanda and Burundi revisited (Mammalia: Chiroptera)</mods:title>
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<mods:affiliation id="5875925D29E0D6235FDA058F0569F0F1">University of Antwerp, Department of Biology, Functional Morphology, Campus Drie Eiken, Universiteitsplein, 1, B- 2610 Antwerpen (Wilrijk), Belgium. &amp; AfricanBats NPC, 357 Botha Ave, Kloofsig, 0157, Republic of South Africa. &amp; urn: lsid: zoobank. org: author: 9 A 0 E 1 AF 5 - C 248 - 4648 - 9 D 64 - 443112890346 &amp; Corresponding author: Victor. VanCakenberghe @ uantwerpen. be</mods:affiliation>
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<mods:affiliation id="70D5A17678A8CF69BB2B1471580DC13F">Faculté des Sciences, Université de Kisangani, Democratic Republic of the Congo. &amp; Email: gembuguycrispin @ gmail. com &amp; urn: lsid: zoobank. org: author: 2 AE 2 F 96 F-A 138 - 419 B-B 650 - 6 DB 44 D 535 D 14 &amp; Faculté des Sciences, Université de Kisangani, Democratic Republic of the Congo. &amp; Email: prescottmusaba @ yahoo. fr &amp; urn: lsid: zoobank. org: author: 4 C 40421 C- 1 D 84 - 4 BD 1 - A 444 - 360300 ECBEEC</mods:affiliation>
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<mods:affiliation id="2F142FF4F21FD79394B068854A916C52">AfricanBats NPC, 357 Botha Ave, Kloofsig, 0157, Republic of South Africa. &amp; Centre for Wildlife Management, University of Pretoria, Private Bag X 20 Hatfield, Pretoria 0028, Republic of South Africa. &amp; Email: ernest. seamark @ africanbats. org &amp; urn: lsid: zoobank. org: author: B 60 CA 9 A 1 - D 288 - 468 E-AEEB-D 0 F 136 E 96 F 5 B</mods:affiliation>
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<paragraph id="8B9F369198315A33DB86FB3AD659FB2B" blockId="45.[487,1100,1201,1265]" box="[487,1100,1201,1228]" pageId="45" pageNumber="46">
<heading id="D0D781FD98315A33DB86FB3AD659FB2B" box="[487,1100,1201,1228]" centered="true" fontSize="11" level="2" pageId="45" pageNumber="46" reason="2">
<taxonomicName id="4C204D1298315A33DB86FB3AD1B0FB2B" ID-CoL="7WZCS" authority="E. Geoffroy St." authorityName="E. Geoffroy Saint-Hilaire" authorityYear="1818" box="[487,933,1201,1228]" class="Mammalia" family="Emballonuridae" genus="Taphozous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="perforatus">
<emphasis id="B954EA8398315A33DB86FB3AD0F8FB2B" bold="true" box="[487,749,1201,1228]" italics="true" pageId="45" pageNumber="46">Taphozous perforatus</emphasis>
E. Geoffroy St.
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-Hilaire, 1818
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<paragraph id="8B9F369198315A33D8B5FB5CD14AFB11" blockId="45.[487,1100,1201,1265]" box="[724,863,1239,1265]" pageId="45" pageNumber="46">
<figureCitation id="131B2A1498315A33D8B5FB5CD132FB11" box="[724,807,1239,1265]" captionStart="Fig" captionStartId="46.[189,232,1930,1956]" captionTargetBox="[233,1353,307,1878]" captionTargetId="figure@46.[225,1363,299,1888]" captionTargetPageId="46" captionText="Fig. 20. Distribution maps. AB. Taphozous nudiventris Cretzschmar, 1830. CD. Taphozous perforatus E. Geoffroy St.-Hilaire, 1818. EF. Nycteris arge Thomas, 1903. A, C, E. Distribution in the CRB area. B, D, F. Pan-African distribution." figureDoi="http://doi.org/10.5281/zenodo.3860127" httpUri="https://zenodo.org/record/3860127/files/figure.png" pageId="45" pageNumber="46">Fig. 20</figureCitation>
CD
</paragraph>
<paragraph id="8B9F369198315A33DADCFA98D16CFACD" blockId="45.[189,889,1299,1364]" box="[189,889,1299,1326]" pageId="45" pageNumber="46">
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<taxonomicName id="4C204D1298315A33DADCFA98D160FACD" ID-CoL="7WZCS" authority="E. Geoffroy Saint-Hilaire, 1818: 126" authorityName="E. Geoffroy Saint-Hilaire" authorityPageNumber="126" authorityYear="1818" box="[189,885,1299,1326]" class="Mammalia" family="Emballonuridae" genus="Taphozous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="perforatus">
<emphasis id="B954EA8398315A33DADCFA98D3AEFACE" box="[189,443,1299,1326]" italics="true" pageId="45" pageNumber="46">Taphozous perforatus</emphasis>
E. Geoffroy Saint-Hilaire, 1818: 126
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.
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<paragraph id="8B9F369198315A33DADCFAB1D082FAB4" blockId="45.[189,889,1299,1364]" box="[189,663,1337,1364]" pageId="45" pageNumber="46">
<treatmentCitationGroup id="AB3011BF98315A33DADCFAB1D082FAB4" box="[189,663,1337,1364]" pageId="45" pageNumber="46">
<emphasis id="B954EA8398315A33DADCFAB1D2D9FAB4" box="[189,204,1338,1364]" italics="true" pageId="45" pageNumber="46">*</emphasis>
<treatmentCitation id="0A81108098315A33DAB2FAB2D086FAB4" author="Thomas O." box="[211,659,1337,1364]" page="561" pageId="45" pageNumber="46" year="1915">
<taxonomicName id="4C204D1298315A33DAB2FAB2D086FAB4" ID-CoL="54S69" authority="Thomas, 1915: 561" authorityName="Thomas" authorityPageNumber="561" authorityYear="1915" box="[211,659,1337,1364]" class="Mammalia" family="Emballonuridae" genus="Taphozous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="sudani">
<emphasis id="B954EA8398315A33DAB2FAB2D3B0FAB3" box="[211,421,1337,1363]" italics="true" pageId="45" pageNumber="46">Taphozous sudani</emphasis>
<bibRefCitation id="EFB14B6098315A33DBCDFAB2D086FAB4" author="Thomas" box="[428,659,1337,1364]" firstAuthor="Thomas" pageId="45" pageNumber="46" pagination="465 - 481" refId="ref273943" refString="Thomas O. 1915. List of mammals (exclusive of Ungulata) collected on the Upper Congo by Dr. Christy for the Congo Museum, Tervueren. Annals and Magazine of Natural History, Ser. 8, 16 (96): 465 - 481. Available from https: // www. biodiversitylibrary. org / page / 22131562 # page / 477 / mode / 1 up [accessed 29 Nov. 2017]." type="journal article" year="1915">Thomas, 1915: 561</bibRefCitation>
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.
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<subSubSection id="C33A651A98315A33DADCFAF7D04EF902" pageId="45" pageNumber="46" type="discussion">
<paragraph id="8B9F369198315A33DADCFAF7D134F9CD" blockId="45.[189,1399,1403,1581]" pageId="45" pageNumber="46">
<bibRefCitation id="EFB14B6098315A33DADCFAF7D3ECFA76" author="Rosevear" box="[189,505,1403,1430]" firstAuthor="Rosevear" pageId="45" pageNumber="46" refId="ref272471" refString="Rosevear D. R. 1965. The Bats of West Africa. Trustees of the British Museum (Natural History), London." type="book" year="1965">Rosevear (1965: 150151)</bibRefCitation>
compared material of both
<taxonomicName id="4C204D1298315A33D929FAF7D1F4FA76" authorityName="E. Geoffroy Saint-Hilaire" authorityYear="1818" box="[840,993,1404,1430]" class="Mammalia" family="Emballonuridae" genus="Taphozous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="perforatus">
<emphasis id="B954EA8398315A33D929FAF7D1F4FA76" box="[840,993,1404,1430]" italics="true" pageId="45" pageNumber="46">T. perforatus</emphasis>
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and
<taxonomicName id="4C204D1298315A33DE7FFAF7D69EFA75" authorityName="Thomas" authorityYear="1915" box="[1054,1163,1403,1429]" class="Mammalia" family="Emballonuridae" genus="Taphozous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="sudani">
<emphasis id="B954EA8398315A33DE7FFAF7D69EFA75" box="[1054,1163,1403,1429]" italics="true" pageId="45" pageNumber="46">T. sudani</emphasis>
</taxonomicName>
, and found that the only character distinguishing the two taxa is the darker pelage in
<taxonomicName id="4C204D1298315A33D9D0FA2AD1EAFA5B" authorityName="Thomas" authorityYear="1915" box="[945,1023,1441,1467]" class="Mammalia" family="Emballonuridae" genus="Taphozous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="sudani">
<emphasis id="B954EA8398315A33D9D0FA2AD1EAFA5B" box="[945,1023,1441,1467]" italics="true" pageId="45" pageNumber="46">sudani</emphasis>
</taxonomicName>
, generally combined with white wings. He also found a specimen with intermediate characters, which led him to retain
<taxonomicName id="4C204D1298315A33DED8FA4CD712FA01" authorityName="Thomas" authorityYear="1915" box="[1209,1287,1479,1505]" class="Mammalia" family="Emballonuridae" genus="Taphozous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="sudani">
<emphasis id="B954EA8398315A33DED8FA4CD712FA01" box="[1209,1287,1479,1505]" italics="true" pageId="45" pageNumber="46">sudani</emphasis>
</taxonomicName>
only as a mere subspecies of
<taxonomicName id="4C204D1298315A33DBC6FA66D034F9E7" authorityName="E. Geoffroy Saint-Hilaire" authorityYear="1818" box="[423,545,1517,1543]" class="Mammalia" family="Emballonuridae" genus="Taphozous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="perforatus">
<emphasis id="B954EA8398315A33DBC6FA66D034F9E7" box="[423,545,1517,1543]" italics="true" pageId="45" pageNumber="46">perforatus</emphasis>
</taxonomicName>
. This view was followed by most subsequent authors (e.g.,
<bibRefCitation id="EFB14B6098315A33DE8DFA66D2CBF9CD" author="Kock" firstAuthor="Kock" pageId="45" pageNumber="46" pagination="1 - 238" refId="ref268151" refString="Kock D. 1969. Die Fledermaus-Fauna des Sudan (Mammalia, Chiroptera). Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 521: 1 - 238." type="journal article" year="1969">Kock 1969: 74</bibRefCitation>
;
<bibRefCitation id="EFB14B6098315A33DA8BF998D007F9CC" author="Hayman &amp; Hill" box="[234,530,1554,1581]" firstAuthor="Hayman" pageId="45" pageNumber="46" refId="ref265812" refString="Hayman R. W. &amp; Hill J. E. 1971. Order Chiroptera. In: Meester J. &amp; Setzer H. W. (eds) The Mammals of Africa, an Identification Manual. Part 2. Smithsonian Institution Press, Washington." type="book" year="1971">Hayman &amp; Hill 1971: 16</bibRefCitation>
;
<bibRefCitation id="EFB14B6098315A33D87EF998D100F9CD" author="Koopman" box="[543,789,1554,1581]" firstAuthor="Koopman" pageId="45" pageNumber="46" pagination="353 - 444" refId="ref268702" refString="Koopman K. F. 1975. Bats of the Sudan. Bulletin of the American Museum of Natural History 154 (4): 353 - 444. Available from http: // hdl. handle. net / 2246 / 609 [accessed 29 Nov. 2017]." type="journal article" year="1975">Koopman 1975: 370</bibRefCitation>
).
</paragraph>
<paragraph id="8B9F369198315A33DADCF9DCD04EF902" blockId="45.[189,1398,1623,1763]" pageId="45" pageNumber="46">
With the exception of a few extra specimens from the same localities, all the material reported in this paper was already mentioned by
<bibRefCitation id="EFB14B6098315A33D82EF9F6D193F977" author="Hayman" box="[591,902,1660,1687]" etAl="et al." firstAuthor="Hayman" pageId="45" pageNumber="46" pagination="1 - 105" refId="ref265860" refString="Hayman R. W., Misonne X. &amp; Verheyen W. N. 1966. The bats of the Congo and of Rwanda and Burundi. Annalen van het Koninklijk Museum voor Midden Afrika, Zoologische Wetenschappen, Ser. 8, 154: 1 - 105." type="journal article" year="1966">
Hayman
<emphasis id="B954EA8398315A33D8DCF9F6D0EFF976" box="[701,762,1660,1686]" italics="true" pageId="45" pageNumber="46">et al.</emphasis>
(1966: 32)
</bibRefCitation>
. The SDM map shows that
<taxonomicName id="4C204D1298315A33DEBDF9F6D763F977" authorityName="E. Geoffroy Saint-Hilaire" authorityYear="1818" box="[1244,1398,1661,1687]" class="Mammalia" family="Emballonuridae" genus="Taphozous" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="perforatus">
<emphasis id="B954EA8398315A33DEBDF9F6D763F977" box="[1244,1398,1661,1687]" italics="true" pageId="45" pageNumber="46">T. perforatus</emphasis>
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primarily occurs in sub-Saharan Africa, above the equator, with a few records reported from northern
<collectingCountry id="F337760198315A33DADCF942D324F903" box="[189,305,1737,1763]" name="Botswana" pageId="45" pageNumber="46">Botswana</collectingCountry>
and southern
<collectingCountry id="F337760198315A33DBB7F943D042F902" box="[470,599,1736,1762]" name="Zimbabwe" pageId="45" pageNumber="46">Zimbabwe</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>