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105 lines
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<document id="2606C3F01CD290314F66D7A93132B67C" ID-DOI="10.1206/0003-0082(2000)287<0001:ROTNTT>2.0.CO;2" ID-ISSN="0003-0082" ID-Zenodo-Dep="10110744" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630265040447" checkinUser="carolina" docAuthor="SMETANA, ALEŠ & DAVIES, ANTHONY" docDate="2000" docId="0384878FD83FFB32FC4F6E4B0B1CFBD7" docLanguage="en" docName="N3287.pdf" docOrigin="American Museum Novitates 3287" docSource="http://www.bioone.org/doi/abs/10.1206/0003-0082%282000%29287%3C0001%3AROTNTT%3E2.0.CO%3B2" docStyle="DocumentStyle:059C9B7006157ECE636E30DEF6793B3A.3:AmMusNovit.2000-2010.journal_article.type2" docStyleId="059C9B7006157ECE636E30DEF6793B3A" docStyleName="AmMusNovit.2000-2010.journal_article.type2" docStyleVersion="3" docTitle="Abemus Mulsant and Rey 1876" docType="treatment" docVersion="3" masterDocId="FFBDFFF7D829FB25FFD86B530951FFD4" masterDocTitle="Reclassification of the North Temperate Taxa Associated with Staphylinus Sensu Lato, Including Comments on Relevant Subtribes of Staphylinini (Coleoptera: Staphylinidae)" masterLastPageNumber="1" masterPageNumber="1" pageNumber="1" updateTime="1699682540511" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="A892FFC620178CF9E36FB17DDDB490EF">
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<mods:title id="39B2642A4B9A6E69AB97A0405D811A92">Reclassification of the North Temperate Taxa Associated with Staphylinus Sensu Lato, Including Comments on Relevant Subtribes of Staphylinini (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="8A73D70A16624EA90FFD377FA43A98BD">SMETANA, ALEŠ</mods:namePart>
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<mods:roleTerm id="08C2FD1306D5272033CA80D1B7C34FAF">Author</mods:roleTerm>
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<mods:namePart id="68A29DB104EC6CE4326AC1BF2B37990A">DAVIES, ANTHONY</mods:namePart>
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<mods:typeOfResource id="2EFA6B51CCEB2925907D6D5541891663">text</mods:typeOfResource>
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<mods:title id="BD81AABDFDD3C09927D136DEEC5B22AE">American Museum Novitates</mods:title>
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</mods:titleInfo>
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<mods:part id="99CDECDADE043D5B52A70399531C07F2">
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<mods:date id="211D7E1DACA6D385B3F5722FED758079">2000</mods:date>
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<mods:number id="B9AE6B6B52EA1EDD9A75DEFDBB9D2FB8">2000-02-02</mods:number>
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<mods:detail id="6EBD9266AE585D6229610214FBB76ED6" type="volume">
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<mods:number id="9F71907ED86EB823552FB927757B2701">3287</mods:number>
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<mods:url id="120F7BA8F8CB943966608479C15B1A03">http://www.bioone.org/doi/abs/10.1206/0003-0082%282000%29287%3C0001%3AROTNTT%3E2.0.CO%3B2</mods:url>
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<mods:classification id="8F48AEBDADE69BD9BFD93CD8B655FEDC">journal article</mods:classification>
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<mods:identifier id="6C7BF1DF868611FE5A8B8D03843089BB" type="DOI">10.1206/0003-0082(2000)287<0001:ROTNTT>2.0.CO;2</mods:identifier>
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<mods:identifier id="2A40054CDD5561D9E20EC5CCF57B1899" type="ISSN">0003-0082</mods:identifier>
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<mods:identifier id="276822B40EBDE947F191EB756504B43D" type="Zenodo-Dep">10110744</mods:identifier>
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<subSubSection id="C3376512D83FFB33FC4F6E4B0C49FAFB" box="[919,1304,1304,1328]" pageId="22" type="nomenclature">
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<paragraph id="8B923699D83FFB33FC4F6E4B0C49FAFB" blockId="22.[919,1304,1304,1328]" box="[919,1304,1304,1328]" pageId="22">
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<heading id="D0DA81F5D83FFB33FC4F6E4B0C49FAFB" box="[919,1304,1304,1328]" centered="true" fontSize="36" level="2" pageId="22" reason="8">
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<taxonomicName id="4C2D4D1AD83FFB33FC4F6E4B0C49FAFB" ID-CoL="8XJ7P" authority="Mulsant and Rey, 1876" authorityName="Mulsant and Rey" authorityYear="1876" box="[919,1304,1304,1328]" class="Insecta" family="Staphylinidae" genus="Abemus" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="genus">
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<emphasis id="B959EA8BD83FFB33FC4F6E4B0AA9FAE4" bold="true" box="[919,1016,1304,1328]" italics="true" pageId="22">Abemus</emphasis>
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Mulsant and Rey, 1876
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3376512D83FFB33FCB56E1F0ABEFA54" pageId="22" type="materials_examined">
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<paragraph id="8B923699D83FFB33FCB56E1F0ABEFA54" blockId="22.[848,1377,1356,1731]" pageId="22">
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<materialsCitation id="3B453CC4D83FFB33FCB56E1F0ABDFA54" collectionCode="SPECIES" pageId="22" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5496883BD83FFB33FCB56E1F0AF6FAB6" box="[877,935,1356,1379]" pageId="22">TYPE</typeStatus>
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<collectionCode id="ED3CAE5CD83FFB33FC646E1F0D42FAB6" box="[956,1043,1356,1379]" pageId="22">SPECIES</collectionCode>
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:
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<taxonomicName id="4C2D4D1AD83FFB33FBE86E1F0ABDFA54" authority="Panzer, 1796" authorityName="Panzer" authorityYear="1796" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="chloropterus">
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<emphasis id="B959EA8BD83FFB33FBE86E1F0C30FAB7" box="[1072,1377,1356,1379]" italics="true" pageId="22">Staphylinus chloropterus</emphasis>
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Panzer, 1796
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</taxonomicName>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3376512D83FFB32FCB56ED50B1CFBD7" lastPageId="23" pageId="22" type="discussion">
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<paragraph id="8B923699D83FFB32FCB56ED50BA9FE4F" blockId="22.[848,1377,1356,1731]" lastBlockId="23.[238,767,182,1027]" lastPageId="23" pageId="22">
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COMMENTS: Members of this genus share the following character states: (
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<bibRefCitation id="EFBC4B68D83FFB33FB116EF70D84FA6F" author="Natural History Museum, London" box="[1225,1237,1444,1467]" pageId="22" pagination="204" refId="ref25242" refString="1. Ocypus almorensis (Cameron, 1932). Cameron (1932: 204) described the species (in Staphylinus) from specimens from '' West Almora: Kali Valley, alt. 9000 feet' ' and '' Punjab: Triun, alt. 9300 feet. '' We were able to study one of the male syntypes in the Natural History Museum, London, England, labelled as follows: '' Triun, 9,300 ft Dharm- sala. Punjab. H. G. C. / M. Cameron. Bequest. B. M. 1955 - 147. / Staphylinus (Ocypus) almorensis. '' The specimen is in very good shape. It was dissected, and the aedeagus and the genital segment were glued to the plate with the beetle. It is hereby designated as the lectotype of Staphylinus almorensis; the label '' LECTOTYPE Staphylinus almorensis Cameron A. Smetana des. 1999 '' was attached to it. The specimen cannot be distinguished from those of Ocypus helleni J. Muller that occur in northern Pakistan and in Kashmir. The name O. almorensis becomes a junior synonym of O. helleni (syn. nov.). The corresponding determination label was attached to the specimen." type="journal article" year="1955">1</bibRefCitation>
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) mandibles with similarly developed teeth, each with large tridentate middle tooth (individual dents not at same plane) and a small tooth basad of it in a deep, subrectangular indentation (
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<figureCitation id="13162A1CD83FFB33FC726D650D40F999" box="[938,1041,1590,1613]" captionStart="Figs" captionStartId="81.[264,312,1587,1608]" captionTargetBox="[246,1334,176,1565]" captionTargetId="figure-34@81.[246,1334,176,1565]" captionTargetPageId="81" captionText="Figs. 123–128. 123. Abemus chloropterus, labium and mentum. 124. Abemus chloropterus, left mandible, dorsal view. 125. Miobdelus sp., posterolateral portion of head, showing pitlike depressions. 126. Miobdelus sp., first three visible abdominal tergites, showing bisinuate posterior basal line. 127. Ocypus picipennis picipennis, left mandible, ventral view. 128. Ocypus picipennis picipennis, right mandible, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10110792" httpUri="https://zenodo.org/record/10110792/files/figure.png" pageId="22">fig. 124</figureCitation>
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); (
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<bibRefCitation id="EFBC4B68D83FFB33FBEE6D650D17F999" author="Ocypus" box="[1078,1094,1590,1613]" pageId="22" pagination="170" refId="ref25465" refString="2. Ocypus auricomus (Lindberg, 1953: 4), described in Staphylinus, is preoccupied by Staphylinus auricomus Brulle', 1842: pl. 5, fig. 6, now a synonym of Glenus chrysis (Gravenhorst, 1806), and by Staphylinus auricomus Cameron, 1929: 65. The name Ocypus mateui (Coiffait, 1954: 170), listed at present as a junior synonym of O. auricomus, becomes the valid name of this species (stat. nov.)." type="journal article" year="1842">2</bibRefCitation>
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) mandibular prostheca appearing bilobed or even multilobed, with extensive basal group, and long distal group of ciliae originating on separate supporting arm (
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<figureCitation id="13162A1CD83FFB33FC486DFF0AB3F917" box="[912,994,1708,1731]" captionStart="Figs" captionStartId="62.[298,346,1373,1394]" captionTargetBox="[280,1368,176,1352]" captionTargetId="figure-36@62.[280,1368,176,1352]" captionTargetPageId="62" captionText="Figs. 11–16. 11. Platydracus stercorarius, right mandibular prostheca, ventral view. 12. Abemus chloropterus, left mandibular prostheca, ventral view. 13. Ocypus picipennis, left mandibular prostheca, ventral view. 14. Tasgius ater, left mandibular prostheca, ventral view. 15. Staphylinus dimidiaticornis, left mandibular prostheca, ventral view. 16. Tasgius melanarius, left mandibular prostheca, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10110754" httpUri="https://zenodo.org/record/10110754/files/figure.png" pageId="22">fig. 12</figureCitation>
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); (
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<bibRefCitation id="EFBC4B68D83FFB33FBDD6DFF0D45F917" author="Lordithon" box="[1029,1044,1708,1731]" pageId="22" refId="ref25558" refString="3. Ocypus bimaculatus (Cameron, 1932: 207), described in Staphylinus, is preoccupied by several names in Staphylinus, the oldest one being Staphylinus bimaculatus Schrank, 1798: 644, now a synonym of Lordithon trinotatus (Erichson, 1839 b: 409). The replacement name Ocypus cameroni, nom. nov., is hereby proposed for Ocypus bimaculatus (Cameron, 1932)." type="journal volume" year="1839">3</bibRefCitation>
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) mentum with anterolateral portions (bearing two anterolateral setae) each obliquely deflected at an angle and therefore not at same level as disc of mentum, separated from it by obtuse carina (
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<figureCitation id="13162A1CD83EFB32FD0F6A5D084CFE97" captionStart="Figs" captionStartId="81.[264,312,1587,1608]" captionTargetBox="[246,1334,176,1565]" captionTargetId="figure-34@81.[246,1334,176,1565]" captionTargetPageId="81" captionText="Figs. 123–128. 123. Abemus chloropterus, labium and mentum. 124. Abemus chloropterus, left mandible, dorsal view. 125. Miobdelus sp., posterolateral portion of head, showing pitlike depressions. 126. Miobdelus sp., first three visible abdominal tergites, showing bisinuate posterior basal line. 127. Ocypus picipennis picipennis, left mandible, ventral view. 128. Ocypus picipennis picipennis, right mandible, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10110792" httpUri="https://zenodo.org/record/10110792/files/figure.png" pageId="23">fig. 123</figureCitation>
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); and (
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<bibRefCitation id="EFBC4B68D83EFB32FEA36A7F08DAFE97" author="Naturhistorisches Museum in Wien" box="[379,395,300,323]" pageId="23" pagination="172" refId="ref25632" refString="4. Ocypus fulvotomentosus Eppelsheim, 1889: 172. The collection at the Naturhistorisches Museum in Wien, Austria, contains one male specimen under this name. It is labelled as follows: '' Kan-ssa 1885 G. Patanin / c. Eppelsh. Steind. d. / fulvotomentosus det. Bernhauer / fulvotomentosus det. J. Muller / TYPUS. '' The specimen is in good condition; however, the entire left middle leg, out- er three segments on left hind tarsus, and the entire right middle tarsus are missing. It was dissected and the aedeagus was glued to a plate attached to the pin, with the beetle. The specimen is hereby designated as the lectotype of O. fulvotomentosus; the label '' LEC- TOTYPE Ocypus fulvotomentosus Eppelsheim A. Smetana des. 1997 '' was attached to it." type="journal article" year="1997">4</bibRefCitation>
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) postmandibular ridge almost reaching infraorbital puncture, running close to eye basally, infraorbital puncture situated three to four diameters from margin of eye.
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</paragraph>
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<paragraph id="8B923699D83EFB32FED36AF20B1CFBD7" blockId="23.[238,767,182,1027]" pageId="23">
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Further character states include: maxillary and labial palpi both with last segment asetose, fusiform; mandibles each with dorsolateral ridge as in
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<taxonomicName id="4C2D4D1AD83EFB32FE0E6AAA0B0EFDC4" authorityName="Ganglbauer" authorityYear="1895" box="[470,607,505,528]" class="Insecta" family="Staphylinidae" genus="Ontholestes" kingdom="Animalia" order="Coleoptera" pageId="23" phylum="Arthropoda" rank="genus">
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<emphasis id="B959EA8BD83EFB32FE0E6AAA0B0EFDC4" box="[470,607,505,528]" italics="true" pageId="23">Ontholestes</emphasis>
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</taxonomicName>
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or
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<taxonomicName id="4C2D4D1AD83EFB32FD4A6AAA0836FDFA" authorityName="Bernhauer" authorityYear="1915" class="Insecta" family="Staphylinidae" genus="Thoracostrongylus" kingdom="Animalia" order="Coleoptera" pageId="23" phylum="Arthropoda" rank="genus">
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<emphasis id="B959EA8BD83EFB32FD4A6AAA0836FDFA" italics="true" pageId="23">Thoracostrongylus</emphasis>
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</taxonomicName>
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; submentum with anterior margin not beaded; lateral end of nuchal ridge distinctly, but moderately, removed from nuchal constriction, directed toward level of transverse postgenal ridge; epimeron of prothorax well developed, projecting triangularly (
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<figureCitation id="13162A1CD83EFB32FD0E69FA084CFD09" captionStart="Figs" captionStartId="79.[264,312,1523,1544]" captionTargetBox="[272,1307,179,1502]" captionTargetId="figure-34@79.[272,1307,179,1502]" captionTargetPageId="79" captionText="Figs. 111–116. 111. Dinothenarus pubescens, pronotum showing both marginal lines of pronotal hypomeron, and two lateral macrosetae, lateral view. 112. Staphylinus erythropterus, pronotum showing both marginal lines of pronotal hypomeron, and two lateral macrosetae, lateral view. 113. Ocypus ophthalmicus ophthalmicus, pronotum showing both marginal lines of pronotal hypomeron, and two lateral macrosetae, ventrolateral view. 114. Ocypus nitens semialatus, pronotum showing both marginal lines of pronotal hypomeron, and two lateral macrosetae, lateral view. 115. Physetops sp., dorsal face of neck, with dorsal portion of nuchal ridge missing. 116. Ontholestes cingulatus, dorsal face of neck, with nuchal ridge." figureDoi="http://doi.org/10.5281/zenodo.10110788" httpUri="https://zenodo.org/record/10110788/files/figure.png" pageId="23">fig. 111</figureCitation>
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); prosternum moderately prominent with broadly rounded slope anteriad, with prominent, arcuate carina continuing evenly onto furcasternum; mesosternum with long serial setae originating behind small elevations in moreorless Vshaped pattern (
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<figureCitation id="13162A1CD83EFB32FD85680A0BEDFCA4" box="[605,700,857,880]" captionStart="Figs" captionStartId="80.[298,346,1603,1624]" captionTargetBox="[280,1368,184,1581]" captionTargetId="figure-36@80.[280,1368,184,1581]" captionTargetPageId="80" captionText="Figs. 117–122. 117. Ontholestes murinus, mesosternum with fully developed medial carina. 118. Thoracostrongylus sp., mesosternum with short medial carina at base. 119. Dinothenarus fossor, labium with emarginate ligula, ventral view. 120. Dinothenarus pubescens, labium and mentum. 121. Dinothenarus pubescens, right mandible and right maxilla, ventral view. 122. Abemus chloropterus, mesosternum, showing sockets of serial setae originating behind small elevations." figureDoi="http://doi.org/10.5281/zenodo.10110790" httpUri="https://zenodo.org/record/10110790/files/figure.png" pageId="23">fig. 122</figureCitation>
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), mesosternal intercoxal process rather acute apically, mesocoxal acetabulum moderately impressed behind; metasternum more or less horizontal, with little difference in convexity between projections and disc.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |