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179 lines
25 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zse.94.26859" ID-GBIF-Dataset="30d1d279-f033-4868-b0c9-925a406be2c5" ID-GBIF-Taxon="182236651" ID-Pensoft-Pub="1860-0743-2-369" ID-Pensoft-UUID="83D45F0665FE5C729C47210ACD710944" ID-Zenodo-Dep="1344764" ID-ZooBank="C354D7DF198F4D4FA2A8F763FD03018D" LSID="urn:lsid:zoobank.org:act:7CA90D40-0E5F-47CF-B85B-04321A516EED" ModsDocID="1860-0743-94-2-369" checkinTime="1621423395062" checkinUser="pensoft" docAuthor="Song, Zhi-Shun, Malenovsky, Igor, Chen, Jian-Qin, Deckert, Juergen & Liang, Ai-Ping" docDate="2018" docId="C6B12FD2C57E4B285F432B31F80B79E3" docLanguage="en" docName="ZoosystEvol 94(2): 369-391" docOrigin="Zoosystematics and Evolution 94 (2)" docPubDate="2018-08-06" docSource="http://dx.doi.org/10.3897/zse.94.26859" docTitle="Orthopagus bartletti Song, Malenovsky & Deckert 2018, sp. n." docType="treatment" docUuid="7CA90D40-0E5F-47CF-B85B-04321A516EED" docUuidSource="ZooBank" docVersion="3" id="83D45F0665FE5C729C47210ACD710944" lastPageNumber="371" masterDocId="83D45F0665FE5C729C47210ACD710944" masterDocTitle="Taxonomic review of the planthopper genus Orthopagus (Hemiptera, Fulgoromorpha, Dictyopharidae), with descriptions of two new species" masterLastPageNumber="391" masterPageNumber="369" pageId="2" pageNumber="371" updateTime="1632016297512" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic review of the planthopper genus Orthopagus (Hemiptera, Fulgoromorpha, Dictyopharidae), with descriptions of two new species</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Song, Zhi-Shun</mods:namePart>
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<mods:affiliation>Jiangsu Key Laboratory of Biofunctional Molecules, School of Life Sciences, Chemistry & Chemical Engineering, Ji & angsu Second Normal University, Nanjing 210013, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Malenovsky, Igor</mods:namePart>
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<mods:affiliation>Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar</mods:affiliation>
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<mods:nameIdentifier type="email">imalenovsky@mzm.cz</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Chen, Jian-Qin</mods:namePart>
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<mods:affiliation>Jiangsu Key Laboratory of Biofunctional Molecules, School of Life Sciences, Chemistry & Chemical Engineering, Ji</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Deckert, Juergen</mods:namePart>
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<mods:affiliation>Department of Botany and Zoology, Faculty of Science, Masaryk University, Kotlarska 2, 611 37 Brno, Czech Republic</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Liang, Ai-Ping</mods:namePart>
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<mods:affiliation>Museum fuer Naturkunde, Leibniz Institute for Research on Evolution and Biodiversity Science, Invalidenstrasse 43, Berlin 10115, Germany & Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:nameIdentifier type="email">liangap@ioz.ac.cn</mods:nameIdentifier>
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<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2018-08-06</mods:number>
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<mods:detail type="volume">
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<mods:number>94</mods:number>
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<mods:number>2</mods:number>
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<mods:start>369</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zse.94.26859</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.94.26859</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-2-369</mods:identifier>
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<treatment ID-GBIF-Taxon="182236651" LSID="urn:lsid:zoobank.org:act:7CA90D40-0E5F-47CF-B85B-04321A516EED" httpUri="http://treatment.plazi.org/id/C6B12FD2C57E4B285F432B31F80B79E3" lastPageNumber="371" pageId="2" pageNumber="371">
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<subSubSection pageId="2" pageNumber="371" type="nomenclature">
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<paragraph pageId="2" pageNumber="371">
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<taxonomicName LSID="http://zoobank.org/7CA90D40-0E5F-47CF-B85B-04321A516EED" authority="Song, Malenovský & Deckert" authorityName="Song, Malenovsky & Deckert" authorityYear="2018" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus bartletti" order="Hemiptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="bartletti" status="sp. n.">
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Orthopagus bartletti Song,
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<normalizedToken originalValue="Malenovský">Malenovsky</normalizedToken>
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& Deckert
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</taxonomicName>
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<taxonomicNameLabel pageId="2" pageNumber="371">sp. n.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Orthopagus species. (A, B) O. bartletti sp. n., male, holotype, Karnataka, India; (C, D) O. exoletus (Melichar), male and female, Sri Lanka; (E, F) O. hainanensis sp. n., male and female, paratypes, Hainan, China; (G, H) O. lunulifer Uhler, male and female, China; (I, J) O. philippinus Melichar, male and female, Philippines; (K, L) O. splendens (Germar), male and female, China." figureDoi="10.3897/zse.94.26859.figure2" httpUri="https://binary.pensoft.net/fig/221398" pageId="2" pageNumber="371">Figs 2A-B</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Forewing of Orthopagus species. (A) O. bartletti sp. n.; (B) O. exoletus (Melichar); (C) O. hainanensis sp. n.; (D) O. lunulifer Uhler; (E) O. philippinus Melichar; (F) O. splendens (Germar)." figureDoi="10.3897/zse.94.26859.figure3" httpUri="https://binary.pensoft.net/fig/221399" pageId="2" pageNumber="371">, 3A</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">, 4A-I</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="371" type="type material">
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<paragraph pageId="2" pageNumber="371">Type material.</paragraph>
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<paragraph pageId="2" pageNumber="371">
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Holotype male,
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<emphasis bold="true" pageId="2" pageNumber="371">INDIA</emphasis>
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: Karnataka: Shimoga district, Someshwari Wildlife Sanctuary, 10 km W Agumbe,
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<geoCoordinate degrees="13" direction="north" minutes="28" orientation="latitude" precision="15" seconds="24" value="13.473333">13°28′24″N</geoCoordinate>
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;
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<geoCoordinate degrees="75" direction="east" minutes="00" orientation="longitude" precision="15" seconds="40" value="75.01111">75°00′40″E</geoCoordinate>
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, alt. 372 ft, early successional, wet evergreen forest, 22.ix.2005, C. R. Bartlett leg. (UDCC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="371" type="diagnosis">
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<paragraph pageId="2" pageNumber="371">Diagnosis.</paragraph>
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<paragraph pageId="2" pageNumber="371">
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<taxonomicName authorityName="Song, Malenovsky & Deckert" authorityYear="2018" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus bartletti" order="Hemiptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="bartletti">
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<emphasis italics="true" pageId="2" pageNumber="371">Orthopagus bartletti</emphasis>
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</taxonomicName>
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sp. n. can be separated from all other
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<taxonomicName authorityName="Uhler" authorityYear="1897" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus" order="Hemiptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="2" pageNumber="371">Orthopagus</emphasis>
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</taxonomicName>
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species by the very short head, hardly produced in front of eyes; the forewings with a brownish streak along nodal line connecting the pterostigmal area with the distal sublunate streak; the dorso-posterior margin of the male pygofer with a large and broad process forming two distinct angles; the robust male segment X; and the lobes of the phallobase muricate apically.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="371" type="description">
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<paragraph pageId="2" pageNumber="371">Description.</paragraph>
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<paragraph pageId="2" pageNumber="371">Measurements (1 male). Body length (from apex of head to tip of forewings): 11.5 mm; head length (from apex of cephalic process to base of eyes): 1.2 mm; head width (including eyes): 1.5 mm; forewing length: 9.6 mm.</paragraph>
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<paragraph pageId="2" pageNumber="371">
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Coloration (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Orthopagus species. (A, B) O. bartletti sp. n., male, holotype, Karnataka, India; (C, D) O. exoletus (Melichar), male and female, Sri Lanka; (E, F) O. hainanensis sp. n., male and female, paratypes, Hainan, China; (G, H) O. lunulifer Uhler, male and female, China; (I, J) O. philippinus Melichar, male and female, Philippines; (K, L) O. splendens (Germar), male and female, China." figureDoi="10.3897/zse.94.26859.figure2" httpUri="https://binary.pensoft.net/fig/221398" pageId="2" pageNumber="371">2A-B</figureCitation>
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). General colour brownish ochraceous marked with dark brown on dorsum. Head greenish ochraceous, vertex with basal corners, a pair of round patches in basal third, and apical diamond-shaped spot dark brown (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4A</figureCitation>
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); frons with median area between intermediate carinae extensively dark brown anteriorly and with series of small pale fuscous spots along intermediate and lateral carinae and narrow ivory white band basally (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4C</figureCitation>
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). Clypeus ivory white, with two small spots at base and apex dark brown. Compound eyes fuscous with posterior margin ochraceous; ocelli purplish-red. Pronotum brownish ochraceous, median carina, apical marginal areas of ventral lobes, and posterior lateral angles ivory white. Mesonotum brownish ochraceous (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4A</figureCitation>
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). Forewings hyaline, veins ochraceous, pterostigmal area, a streak along nodal line, and a wide sublunate streak on distal third dull ochraceous; posterior (claval) margin broadly faintly brown (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Forewing of Orthopagus species. (A) O. bartletti sp. n.; (B) O. exoletus (Melichar); (C) O. hainanensis sp. n.; (D) O. lunulifer Uhler; (E) O. philippinus Melichar; (F) O. splendens (Germar)." figureDoi="10.3897/zse.94.26859.figure3" httpUri="https://binary.pensoft.net/fig/221399" pageId="2" pageNumber="371">3A</figureCitation>
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). Hind wings hyaline, veins and an apical spot dull ochraceous. Legs pale brown; fore femora subapically and hind tibiae at base and apex (including lateral and apical spines) blackish. Abdomen dorsally and ventrally brownish ochraceous.
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</paragraph>
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<paragraph pageId="2" pageNumber="371">
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Structure. Head (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4A-C</figureCitation>
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) very short, cephalic process practically absent. Vertex (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4A</figureCitation>
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) with ratio of length at midline to width between eyes 1.4. Frons with base slightly inflated anteriad in dorsal view (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4A</figureCitation>
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), with transition to vertex broadly rounded in lateral view (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4B</figureCitation>
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); in ventral view, frons with ratio of length at midline to maximum width 2.2; median carina more or less obscure at base (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4C</figureCitation>
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).
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</paragraph>
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<paragraph pageId="2" pageNumber="371">
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Male genitalia. Pygofer, in lateral view, with dorso-posterior margin forming a large, broad, biangular lobe (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4E</figureCitation>
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); in ventral view (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4F</figureCitation>
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) much longer than in dorsal view (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4D</figureCitation>
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) with ratio of ventral to dorsal width about 3.4. Gonostyles (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4E, F</figureCitation>
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) elongate, relatively narrow in basal half, with strongly sinuate dorsal margin medially. Aedeagus (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4G-I</figureCitation>
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) with endosomal processes relatively short and robust, without distinct minute superficial spines, and directed laterad; phallobase with a pair of large, elongate, thumb-like ventral lobes, curved dorso-posteriad and muricate apically (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4G-H</figureCitation>
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); and a pair of shorter lateral lobes, directed posteriad (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4H</figureCitation>
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). Segment X, in lateral view, relatively short and robust, with ventral margin gradually widening from base to broadly truncate apex (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4E</figureCitation>
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); in dorsal view broad, broadest medially, with ratio of length to maximum width 1.1 (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orthopagus bartletti sp. n. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure4" httpUri="https://binary.pensoft.net/fig/221400" pageId="2" pageNumber="371">4D</figureCitation>
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).
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</paragraph>
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<paragraph pageId="2" pageNumber="371">Female genitalia unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="371" type="etymology">
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<paragraph pageId="2" pageNumber="371">Etymology.</paragraph>
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<paragraph pageId="2" pageNumber="371">The new species is named after Dr. Charles R. Bartlett, collector of the type specimen and curator of the insect collection at the Department of Entomology and Wildlife Ecology, University of Delaware, USA, in recognition of his kindest help and support to the first author when he visited UDCC in 2017. The species name is to be treated as a noun in genitive case.</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="371" type="distribution">
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<paragraph pageId="2" pageNumber="371">Distribution.</paragraph>
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<paragraph pageId="2" pageNumber="371">
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So far only known from southwestern India (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Geographical distribution of the Orthopagus species." figureDoi="10.3897/zse.94.26859.figure11" httpUri="https://binary.pensoft.net/fig/221408" pageId="2" pageNumber="371">11</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |