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<document id="0C77E164134EC9042A234AD2CD8415C3" ID-DOI="10.1016/j.rbe.2015.11.013" ID-ISSN="1806-9665" ID-Zenodo-Dep="13195155" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722634454983" checkinUser="felipe" docAuthor="Castro, Ivan, Bravo, Freddy &amp; Vieira, Rodrigo" docDate="2016" docId="A52B87D8FFB4FFF6DB0DFA19FF2DD9F8" docLanguage="en" docName="RevBrasEntomol.60.2.137-149.pdf" docOrigin="Revista Brasileira de Entomologia 60 (2)" docSource="http://dx.doi.org/10.1016/j.rbe.2015.11.013" docStyle="DocumentStyle:0303D37C566E4E2A42C6591146119864.3:RevBrasEntomol.2015-.journal_article" docStyleId="0303D37C566E4E2A42C6591146119864" docStyleName="RevBrasEntomol.2015-.journal_article" docStyleVersion="3" docTitle="Triorla beckeri Castro, Bravo and Vieira 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="141" masterDocId="5912FFA0FFB7FFF2DB58FFCBFF94DF04" masterDocTitle="Four new species of Triorla Parks (Diptera, Asilidae, Asilinae) from Brazil" masterLastPageNumber="149" masterPageNumber="137" pageNumber="140" updateTime="1722681803857" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B8537ADBA5AF5E9ACC3831E33821AC8B">Four new species of Triorla Parks (Diptera, Asilidae, Asilinae) from Brazil</mods:title>
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<mods:affiliation id="AA04D6F3C350DECC5F17E8D7A0CE3092">Post-graduate Program in Zoology, Universidade Estadual de Feira de Santana, Feira de Santana, BA, Brazil</mods:affiliation>
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<mods:namePart id="53A21CE4C0795DD84CE1B233709A062D">Vieira, Rodrigo</mods:namePart>
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<paragraph id="2D3D36CEFFB4FFF1DB0DFA19FDB9DAE2" blockId="3.[85,557,1490,1510]" box="[85,557,1490,1510]" pageId="3" pageNumber="140">
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<taxonomicName id="EA824D4DFFB4FFF1DB0DFA19FE4DDAE2" authority="Castro, Bravo and Vieira" authorityName="Castro, Bravo and Vieira" authorityYear="2016" box="[85,473,1490,1510]" class="Insecta" family="Asilidae" genus="Triorla" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="1FF6EADCFFB4FFF1DB0DFA19FF76DAE2" box="[85,226,1490,1510]" italics="true" pageId="3" pageNumber="140">Triorla beckeri</emphasis>
Castro, Bravo and Vieira
</taxonomicName>
<emphasis id="1FF6EADCFFB4FFF1DA87FA19FDB9DAE2" bold="true" box="[479,557,1490,1510]" italics="true" pageId="3" pageNumber="140">
<taxonomicNameLabel id="04C557A7FFB4FFF1DA87FA19FDB9DAE2" box="[479,557,1490,1510]" pageId="3" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="65986545FFB4FFF6DB2DF9C0FF2DD9F8" lastPageId="4" lastPageNumber="141" pageId="3" pageNumber="140" type="description">
<paragraph id="2D3D36CEFFB4FFF1DB2DF9C0FEAFD91A" blockId="3.[85,756,1547,1985]" box="[117,315,1547,1566]" pageId="3" pageNumber="140">
<figureCitation id="B5B92A4BFFB4FFF1DB2DF9C0FF27D91A" box="[117,179,1547,1566]" captionStart="Figures 15" captionStartId="2.[114,177,1946,1960]" captionTargetBox="[391,1225,886,1914]" captionTargetId="figure-243@2.[391,1226,886,1915]" captionTargetPageId="2" captionText="Figures 15. Head in frontal view.1. Triorla beckeri sp. nov.; 2.Triorla milineae sp. nov.; 3.Triorla spatulata sp. nov.; 4.Triorla paraensis sp. nov.; 5.Triorla parastriola Pamplona and Aires, 1999." figureDoi="http://doi.org/10.5281/zenodo.13195157" httpUri="https://zenodo.org/record/13195157/files/figure.png" pageId="3" pageNumber="140">Figs. 1</figureCitation>
,
<figureCitation id="B5B92A4BFFB4FFF1DBE7F9C0FF59D91A" box="[191,205,1547,1566]" captionStart="Figures 610" captionStartId="3.[85,148,1163,1177]" captionTargetBox="[365,1194,154,1132]" captionTargetId="figure-14@3.[364,1194,150,1132]" captionTargetPageId="3" captionText="Figures 610. Head in lateral view. 6. Triorla beckeri sp. nov.; 7. Triorla milineae sp. nov.; 8. Triorla spatulata sp. nov.; 9. Triorla paraensis sp. nov.; 10. Triorla parastriola Pamplona and Aires, 1999." figureDoi="http://doi.org/10.5281/zenodo.13195161" httpUri="https://zenodo.org/record/13195161/files/figure.png" pageId="3" pageNumber="140">6</figureCitation>
,
<figureCitation id="B5B92A4BFFB4FFF1DB8EF9C0FF64D91A" box="[214,240,1547,1566]" captionStart="Figures 1115" captionStartId="4.[114,177,1099,1113]" captionTargetBox="[506,1110,153,1052]" captionTargetId="figure-14@4.[506,1110,150,1053]" captionTargetPageId="4" captionText="Figures 1115. Wing. 11. Triorla beckeri sp. nov.; 12. Triorla milineae sp. nov.; 13. Triorla paraensis sp. nov.; 14. Triorla parastriola Pamplona and Aires, 1999; 15. Triorla spatulata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13195163" httpUri="https://zenodo.org/record/13195163/files/figure.png" pageId="3" pageNumber="140">11</figureCitation>
,
<figureCitation id="B5B92A4BFFB4FFF1DBA3F9C0FEAFD91A" box="[251,315,1547,1566]" captionStart="Figures 2229" captionStartId="6.[114,177,1253,1267]" captionTargetBox="[396,1219,151,1223]" captionTargetId="figure-14@6.[393,1225,150,1222]" captionTargetPageId="6" captionText="Figures 2229. Triorla beckeri sp. nov. 22. Apex of aedeagal prongs; 23. Gonocoxite; 24. Hypandrium; 25. Male abdomen; 26. Macrosetae of epandrium; 27. Gonostylus in lateral view; 28. Aedeagus and ejaculatory apodeme in lateral view; 29. Male terminalia in lateral view. Abbreviations: abd: abdomen; mcs: macrosetae; m ter: male terminalia." figureDoi="http://doi.org/10.5281/zenodo.13195165" httpUri="https://zenodo.org/record/13195165/files/figure.png" pageId="3" pageNumber="140">2229</figureCitation>
</paragraph>
<paragraph id="2D3D36CEFFB4FFF1DB2DF9ECFF4FD831" blockId="3.[85,756,1547,1985]" pageId="3" pageNumber="140">
Diagnosis: four ocellar setae; face and front yellow pruinose; dorsal margin of anepisternum white setose; 8 pronotal macrosetae; tergite I with 34 black and yellow macrosetae; epandrium with apical concavity in lateral view, macrosetae in mediodorsal region (
<figureCitation id="B5B92A4BFFB4FFF1DB88F95DFE8CD9AD" box="[208,280,1686,1705]" captionStart="Figures 2229" captionStartId="6.[114,177,1253,1267]" captionTargetBox="[396,1219,151,1223]" captionTargetId="figure-14@6.[393,1225,150,1222]" captionTargetPageId="6" captionText="Figures 2229. Triorla beckeri sp. nov. 22. Apex of aedeagal prongs; 23. Gonocoxite; 24. Hypandrium; 25. Male abdomen; 26. Macrosetae of epandrium; 27. Gonostylus in lateral view; 28. Aedeagus and ejaculatory apodeme in lateral view; 29. Male terminalia in lateral view. Abbreviations: abd: abdomen; mcs: macrosetae; m ter: male terminalia." figureDoi="http://doi.org/10.5281/zenodo.13195165" httpUri="https://zenodo.org/record/13195165/files/figure.png" pageId="3" pageNumber="140">Fig. 29</figureCitation>
); gonocoxite triangular, ending in an acute, short projection in posteroventral region (
<figureCitation id="B5B92A4BFFB4FFF1DAA4F979FDAAD9C1" box="[508,574,1714,1733]" captionStart="Figures 2229" captionStartId="6.[114,177,1253,1267]" captionTargetBox="[396,1219,151,1223]" captionTargetId="figure-14@6.[393,1225,150,1222]" captionTargetPageId="6" captionText="Figures 2229. Triorla beckeri sp. nov. 22. Apex of aedeagal prongs; 23. Gonocoxite; 24. Hypandrium; 25. Male abdomen; 26. Macrosetae of epandrium; 27. Gonostylus in lateral view; 28. Aedeagus and ejaculatory apodeme in lateral view; 29. Male terminalia in lateral view. Abbreviations: abd: abdomen; mcs: macrosetae; m ter: male terminalia." figureDoi="http://doi.org/10.5281/zenodo.13195165" httpUri="https://zenodo.org/record/13195165/files/figure.png" pageId="3" pageNumber="140">Fig. 23</figureCitation>
); gonostylus 5.5× longer than wide, bilobed at apex (
<figureCitation id="B5B92A4BFFB4FFF1DAEEF905FE6DD9E5" box="[438,505,1742,1761]" captionStart="Figures 2229" captionStartId="6.[114,177,1253,1267]" captionTargetBox="[396,1219,151,1223]" captionTargetId="figure-14@6.[393,1225,150,1222]" captionTargetPageId="6" captionText="Figures 2229. Triorla beckeri sp. nov. 22. Apex of aedeagal prongs; 23. Gonocoxite; 24. Hypandrium; 25. Male abdomen; 26. Macrosetae of epandrium; 27. Gonostylus in lateral view; 28. Aedeagus and ejaculatory apodeme in lateral view; 29. Male terminalia in lateral view. Abbreviations: abd: abdomen; mcs: macrosetae; m ter: male terminalia." figureDoi="http://doi.org/10.5281/zenodo.13195165" httpUri="https://zenodo.org/record/13195165/files/figure.png" pageId="3" pageNumber="140">Fig. 27</figureCitation>
); distal region of hypandrium without median projection (
<figureCitation id="B5B92A4BFFB4FFF1DA99F921FD91D9F9" box="[449,517,1770,1789]" captionStart="Figures 2229" captionStartId="6.[114,177,1253,1267]" captionTargetBox="[396,1219,151,1223]" captionTargetId="figure-14@6.[393,1225,150,1222]" captionTargetPageId="6" captionText="Figures 2229. Triorla beckeri sp. nov. 22. Apex of aedeagal prongs; 23. Gonocoxite; 24. Hypandrium; 25. Male abdomen; 26. Macrosetae of epandrium; 27. Gonostylus in lateral view; 28. Aedeagus and ejaculatory apodeme in lateral view; 29. Male terminalia in lateral view. Abbreviations: abd: abdomen; mcs: macrosetae; m ter: male terminalia." figureDoi="http://doi.org/10.5281/zenodo.13195165" httpUri="https://zenodo.org/record/13195165/files/figure.png" pageId="3" pageNumber="140">Fig. 24</figureCitation>
); ejaculatory apodeme without spines (
<figureCitation id="B5B92A4BFFB4FFF1DBA1F8CDFEA8D81D" box="[249,316,1798,1817]" captionStart="Figures 2229" captionStartId="6.[114,177,1253,1267]" captionTargetBox="[396,1219,151,1223]" captionTargetId="figure-14@6.[393,1225,150,1222]" captionTargetPageId="6" captionText="Figures 2229. Triorla beckeri sp. nov. 22. Apex of aedeagal prongs; 23. Gonocoxite; 24. Hypandrium; 25. Male abdomen; 26. Macrosetae of epandrium; 27. Gonostylus in lateral view; 28. Aedeagus and ejaculatory apodeme in lateral view; 29. Male terminalia in lateral view. Abbreviations: abd: abdomen; mcs: macrosetae; m ter: male terminalia." figureDoi="http://doi.org/10.5281/zenodo.13195165" httpUri="https://zenodo.org/record/13195165/files/figure.png" pageId="3" pageNumber="140">Fig. 28</figureCitation>
); aedeagal prongs with 1 subapical dilation (
<figureCitation id="B5B92A4BFFB4FFF1DB05F8E9FF5AD831" box="[93,206,1826,1845]" captionStart="Figures 2229" captionStartId="6.[114,177,1253,1267]" captionTargetBox="[396,1219,151,1223]" captionTargetId="figure-14@6.[393,1225,150,1222]" captionTargetPageId="6" captionText="Figures 2229. Triorla beckeri sp. nov. 22. Apex of aedeagal prongs; 23. Gonocoxite; 24. Hypandrium; 25. Male abdomen; 26. Macrosetae of epandrium; 27. Gonostylus in lateral view; 28. Aedeagus and ejaculatory apodeme in lateral view; 29. Male terminalia in lateral view. Abbreviations: abd: abdomen; mcs: macrosetae; m ter: male terminalia." figureDoi="http://doi.org/10.5281/zenodo.13195165" httpUri="https://zenodo.org/record/13195165/files/figure.png" pageId="3" pageNumber="140">Figs. 22, 28</figureCitation>
).
</paragraph>
<paragraph id="2D3D36CEFFB4FFF1DB2DF8F6FC4DD918" blockId="3.[85,756,1547,1985]" lastBlockId="3.[804,1476,1265,1982]" pageId="3" pageNumber="140">
Description. Male. Head (
<figureCitation id="B5B92A4BFFB4FFF1DAD5F8F6FE47D855" box="[397,467,1853,1873]" captionStart="Figures 15" captionStartId="2.[114,177,1946,1960]" captionTargetBox="[391,1225,886,1914]" captionTargetId="figure-243@2.[391,1226,886,1915]" captionTargetPageId="2" captionText="Figures 15. Head in frontal view.1. Triorla beckeri sp. nov.; 2.Triorla milineae sp. nov.; 3.Triorla spatulata sp. nov.; 4.Triorla paraensis sp. nov.; 5.Triorla parastriola Pamplona and Aires, 1999." figureDoi="http://doi.org/10.5281/zenodo.13195157" httpUri="https://zenodo.org/record/13195157/files/figure.png" pageId="3" pageNumber="140">Figs. 1</figureCitation>
,
<figureCitation id="B5B92A4BFFB4FFF1DABFF8F6FE60D854" box="[487,500,1853,1872]" captionStart="Figures 610" captionStartId="3.[85,148,1163,1177]" captionTargetBox="[365,1194,154,1132]" captionTargetId="figure-14@3.[364,1194,150,1132]" captionTargetPageId="3" captionText="Figures 610. Head in lateral view. 6. Triorla beckeri sp. nov.; 7. Triorla milineae sp. nov.; 8. Triorla spatulata sp. nov.; 9. Triorla paraensis sp. nov.; 10. Triorla parastriola Pamplona and Aires, 1999." figureDoi="http://doi.org/10.5281/zenodo.13195161" httpUri="https://zenodo.org/record/13195161/files/figure.png" pageId="3" pageNumber="140">6</figureCitation>
). Ocellar tubercle with 4 yellow setae at apex, 46 brown setae laterally, 1719 yellow postocular macrosetae, 47 black and yellow lateral orbital macrosetae. Mystax: black macrosetae dorsally, yellow macrosetae laterally; median and inferior region with 12 macrosetae and setae, the inferior macrosetae transversally distributed; 34 pairs of subvibrissal macrosetae. Inferior occipital setae white and branched, and 1213 occipital macrosetae. Basal proboscial white setae. Mentum with yellow and apically undulated setae. Palpus brown setose on distal half and yellow to pale on proximal half. Face and front yellow pruinose; face wide in frontal view, face at antennal level 1.2× as wide as eyes. Front with brown transversal vitae; gena and epistomal margin brownish-white pruinose. Scape 2.0× the length of pedicel, black setae dorsally and yellow setae ventrally. Facial gibbosity developed. Lateral width of gena 0.25× the lateral length of compound eye.
</paragraph>
<paragraph id="2D3D36CEFFB4FFF1D81CF9EFFB69D836" blockId="3.[804,1476,1265,1982]" pageId="3" pageNumber="140">Thorax. Eight pronotal macrosetae. Postpronotal lobe white setose; 2 pairs of black notopleural macrosetae; 2 pairs of black supraalar macrosetae; 2 pairs black postalar macrosetae; 6 black scutellar macrosetae apically; scutellar disc yellow, setose yellow basally, black apically, undulated at apex; posterior acrostichal setae replaced by black apically undulated setae; 4pairs dorsocentral black macrosetae. Mesonotum mainly brownish. Anepisternum with white setae on dorsal margin; basalar sclerite black setose; katepisternum with few white setae; anepimeron with some white setae; katepimeron glabrous.</paragraph>
<paragraph id="2D3D36CEFFB4FFF1D81CF8F0FAE8D84A" blockId="3.[804,1476,1265,1982]" box="[836,1404,1851,1870]" pageId="3" pageNumber="140">
Wing. (
<figureCitation id="B5B92A4BFFB4FFF1D8D6F8F0FC44D84A" box="[910,976,1851,1870]" captionStart="Figures 1115" captionStartId="4.[114,177,1099,1113]" captionTargetBox="[506,1110,153,1052]" captionTargetId="figure-14@4.[506,1110,150,1053]" captionTargetPageId="4" captionText="Figures 1115. Wing. 11. Triorla beckeri sp. nov.; 12. Triorla milineae sp. nov.; 13. Triorla paraensis sp. nov.; 14. Triorla parastriola Pamplona and Aires, 1999; 15. Triorla spatulata sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13195163" httpUri="https://zenodo.org/record/13195163/files/figure.png" pageId="3" pageNumber="140">Fig. 11</figureCitation>
); membrane yellowish. Halters brownish.
</paragraph>
<paragraph id="2D3D36CEFFB4FFF1D81CF89CFA56D8B9" blockId="3.[804,1476,1265,1982]" pageId="3" pageNumber="140">
Abdomen (
<figureCitation id="B5B92A4BFFB4FFF1D8EAF89CFC67D86E" box="[946,1011,1879,1898]" captionStart="Figures 2229" captionStartId="6.[114,177,1253,1267]" captionTargetBox="[396,1219,151,1223]" captionTargetId="figure-14@6.[393,1225,150,1222]" captionTargetPageId="6" captionText="Figures 2229. Triorla beckeri sp. nov. 22. Apex of aedeagal prongs; 23. Gonocoxite; 24. Hypandrium; 25. Male abdomen; 26. Macrosetae of epandrium; 27. Gonostylus in lateral view; 28. Aedeagus and ejaculatory apodeme in lateral view; 29. Male terminalia in lateral view. Abbreviations: abd: abdomen; mcs: macrosetae; m ter: male terminalia." figureDoi="http://doi.org/10.5281/zenodo.13195165" httpUri="https://zenodo.org/record/13195165/files/figure.png" pageId="3" pageNumber="140">Fig.25</figureCitation>
). Tergite I with 34 black and yellow macrosetae; posterior margin fringed with yellow macrosetae, remaining surfaces yellow setose. Lateral margins of tergite II yellow setose; dorsal surface yellow setose. Lateral margins of tergite III with short
</paragraph>
<paragraph id="2D3D36CEFFB3FFF6DF45FBD6FBBBDB28" blockId="4.[1053,1071,1053,1068]" box="[1053,1071,1053,1068]" pageId="4" pageNumber="141">15</paragraph>
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<paragraph id="2D3D36CEFFB3FFF6DB2AFB80FF6BDB74" blockId="4.[114,1503,1098,1136]" pageId="4" pageNumber="141">
<emphasis id="1FF6EADCFFB3FFF6DB2AFB80FF7ADB5D" bold="true" box="[114,238,1099,1113]" pageId="4" pageNumber="141">Figures 1115.</emphasis>
Wing. 11.
<taxonomicName id="EA824D4DFFB3FFF6DA14FB81FE2ADB5D" authorityName="Castro &amp; Bravo &amp; Vieira" authorityYear="2016" box="[332,446,1098,1113]" class="Insecta" family="Asilidae" genus="Triorla" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="1FF6EADCFFB3FFF6DA14FB81FE2ADB5D" box="[332,446,1098,1113]" italics="true" pageId="4" pageNumber="141">Triorla beckeri</emphasis>
</taxonomicName>
<emphasis id="1FF6EADCFFB3FFF6DA9BFB80FD92DB5D" bold="true" box="[451,518,1099,1113]" pageId="4" pageNumber="141">
<taxonomicNameLabel id="04C557A7FFB3FFF6DA9BFB80FD92DB5D" box="[451,518,1099,1113]" pageId="4" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 12.
<taxonomicName id="EA824D4DFFB3FFF6D968FB81FD39DB5D" authorityName="Castro &amp; Bravo &amp; Vieira" authorityYear="2016" box="[560,685,1098,1113]" class="Insecta" family="Asilidae" genus="Triorla" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="milineae" status="sp. nov.">
<emphasis id="1FF6EADCFFB3FFF6D968FB81FD39DB5D" box="[560,685,1098,1113]" italics="true" pageId="4" pageNumber="141">Triorla milineae</emphasis>
</taxonomicName>
<emphasis id="1FF6EADCFFB3FFF6D9EAFB80FD60DB5D" bold="true" box="[690,756,1099,1113]" pageId="4" pageNumber="141">
<taxonomicNameLabel id="04C557A7FFB3FFF6D9EAFB80FD60DB5D" box="[690,756,1099,1113]" pageId="4" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 13.
<taxonomicName id="EA824D4DFFB3FFF6D847FB81FC36DB5D" authorityName="Castro &amp; Bravo &amp; Vieira" authorityYear="2016" box="[799,930,1098,1113]" class="Insecta" family="Asilidae" genus="Triorla" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="paraensis" status="sp. nov.">
<emphasis id="1FF6EADCFFB3FFF6D847FB81FC36DB5D" box="[799,930,1098,1113]" italics="true" pageId="4" pageNumber="141">Triorla paraensis</emphasis>
</taxonomicName>
<emphasis id="1FF6EADCFFB3FFF6D8F0FB80FC7EDB5D" bold="true" box="[936,1002,1099,1113]" pageId="4" pageNumber="141">
<taxonomicNameLabel id="04C557A7FFB3FFF6D8F0FB80FC7EDB5D" box="[936,1002,1099,1113]" pageId="4" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 14.
<taxonomicName id="EA824D4DFFB3FFF6DF4DFB81FA15DB5D" authority="Pamplona and Aires, 1999" authorityName="Pamplona and Aires" authorityYear="1999" box="[1045,1409,1098,1113]" class="Insecta" family="Asilidae" genus="Triorla" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="parastriola">
<emphasis id="1FF6EADCFFB3FFF6DF4DFB81FB37DB5D" box="[1045,1187,1098,1113]" italics="true" pageId="4" pageNumber="141">Triorla parastriola</emphasis>
<bibRefCitation id="49134B3FFFB3FFF6DFF1FB80FA15DB5D" author="Pamplona, D. M. &amp; Aires, C." box="[1193,1409,1099,1113]" pageId="4" pageNumber="141" pagination="1127 - 1134" refId="ref11297" refString="Pamplona, D. M., Aires, C., 1999. Revision of Triorla Parks and description of a new species emphasizing the systematic value of spermatheca (Diptera, Asilidae). Revista Brasileira de Zoologia 16, 1127 - 1134." type="journal article" year="1999">Pamplona and Aires, 1999</bibRefCitation>
</taxonomicName>
; 15.
<taxonomicName id="EA824D4DFFB3FFF6DEF3FB81FF2FDB74" authorityName="Castro &amp; Bravo &amp; Vieira" authorityYear="2016" class="Insecta" family="Asilidae" genus="Triorla" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="spatulata" status="sp. nov.">
<emphasis id="1FF6EADCFFB3FFF6DEF3FB81FF2FDB74" italics="true" pageId="4" pageNumber="141">Triorla spatulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="04C557A7FFB3FFF6DBE7FBA9FF6BDB74" box="[191,255,1122,1136]" pageId="4" pageNumber="141" rank="species">
<emphasis id="1FF6EADCFFB3FFF6DBE7FBA9FF6FDB74" bold="true" box="[191,251,1122,1136]" pageId="4" pageNumber="141">sp. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="2D3D36CEFFB3FFF6DB2AFB54FC86DBEE" blockId="4.[114,786,1183,1788]" pageId="4" pageNumber="141">yellow setae. Tergites IVV dark brown to black with transversal posterior band light-brown. Tergite VI white, with a median and triangular black spot. Tergite VII black, with lateral white margins.</paragraph>
<paragraph id="2D3D36CEFFB3FFF6DBCAFB38FEC8D919" blockId="4.[114,786,1183,1788]" pageId="4" pageNumber="141">
Terminalia (
<figureCitation id="B5B92A4BFFB3FFF6DA4BFB38FE4FDA02" box="[275,475,1267,1286]" captionStart="Figures 2229" captionStartId="6.[114,177,1253,1267]" captionTargetBox="[396,1219,151,1223]" captionTargetId="figure-14@6.[393,1225,150,1222]" captionTargetPageId="6" captionText="Figures 2229. Triorla beckeri sp. nov. 22. Apex of aedeagal prongs; 23. Gonocoxite; 24. Hypandrium; 25. Male abdomen; 26. Macrosetae of epandrium; 27. Gonostylus in lateral view; 28. Aedeagus and ejaculatory apodeme in lateral view; 29. Male terminalia in lateral view. Abbreviations: abd: abdomen; mcs: macrosetae; m ter: male terminalia." figureDoi="http://doi.org/10.5281/zenodo.13195165" httpUri="https://zenodo.org/record/13195165/files/figure.png" pageId="4" pageNumber="141">Figs. 2224, 2629</figureCitation>
). Epandrium in lateral view, with mediodorsal region bearing differentiated macrosetae (
<figureCitation id="B5B92A4BFFB3FFF6DB22FAE0FF66DA3A" box="[122,242,1323,1342]" captionStart="Figures 2229" captionStartId="6.[114,177,1253,1267]" captionTargetBox="[396,1219,151,1223]" captionTargetId="figure-14@6.[393,1225,150,1222]" captionTargetPageId="6" captionText="Figures 2229. Triorla beckeri sp. nov. 22. Apex of aedeagal prongs; 23. Gonocoxite; 24. Hypandrium; 25. Male abdomen; 26. Macrosetae of epandrium; 27. Gonostylus in lateral view; 28. Aedeagus and ejaculatory apodeme in lateral view; 29. Male terminalia in lateral view. Abbreviations: abd: abdomen; mcs: macrosetae; m ter: male terminalia." figureDoi="http://doi.org/10.5281/zenodo.13195165" httpUri="https://zenodo.org/record/13195165/files/figure.png" pageId="4" pageNumber="141">Figs. 26, 29</figureCitation>
); cercus elliptic, ending at same level as alate projections of subepandrial sclerite. Hypandrium with rounded distal margin, without median projection (
<figureCitation id="B5B92A4BFFB3FFF6DA86FAA9FD8ADA71" box="[478,542,1378,1397]" captionStart="Figures 2229" captionStartId="6.[114,177,1253,1267]" captionTargetBox="[396,1219,151,1223]" captionTargetId="figure-14@6.[393,1225,150,1222]" captionTargetPageId="6" captionText="Figures 2229. Triorla beckeri sp. nov. 22. Apex of aedeagal prongs; 23. Gonocoxite; 24. Hypandrium; 25. Male abdomen; 26. Macrosetae of epandrium; 27. Gonostylus in lateral view; 28. Aedeagus and ejaculatory apodeme in lateral view; 29. Male terminalia in lateral view. Abbreviations: abd: abdomen; mcs: macrosetae; m ter: male terminalia." figureDoi="http://doi.org/10.5281/zenodo.13195165" httpUri="https://zenodo.org/record/13195165/files/figure.png" pageId="4" pageNumber="141">Fig.24</figureCitation>
). Gonocoxite triangular, ending in acute, short projection in posteroventral region (
<figureCitation id="B5B92A4BFFB3FFF6D99AFAB5FC90DA95" box="[706,772,1406,1425]" captionStart="Figures 2229" captionStartId="6.[114,177,1253,1267]" captionTargetBox="[396,1219,151,1223]" captionTargetId="figure-14@6.[393,1225,150,1222]" captionTargetPageId="6" captionText="Figures 2229. Triorla beckeri sp. nov. 22. Apex of aedeagal prongs; 23. Gonocoxite; 24. Hypandrium; 25. Male abdomen; 26. Macrosetae of epandrium; 27. Gonostylus in lateral view; 28. Aedeagus and ejaculatory apodeme in lateral view; 29. Male terminalia in lateral view. Abbreviations: abd: abdomen; mcs: macrosetae; m ter: male terminalia." figureDoi="http://doi.org/10.5281/zenodo.13195165" httpUri="https://zenodo.org/record/13195165/files/figure.png" pageId="4" pageNumber="141">Fig. 23</figureCitation>
). Gonostylus 5.5× longer than wide and 1.5× the length of gonocoxite, bilobed at apex (
<figureCitation id="B5B92A4BFFB3FFF6DA35FA7DFE26DACD" box="[365,434,1462,1481]" captionStart="Figures 2229" captionStartId="6.[114,177,1253,1267]" captionTargetBox="[396,1219,151,1223]" captionTargetId="figure-14@6.[393,1225,150,1222]" captionTargetPageId="6" captionText="Figures 2229. Triorla beckeri sp. nov. 22. Apex of aedeagal prongs; 23. Gonocoxite; 24. Hypandrium; 25. Male abdomen; 26. Macrosetae of epandrium; 27. Gonostylus in lateral view; 28. Aedeagus and ejaculatory apodeme in lateral view; 29. Male terminalia in lateral view. Abbreviations: abd: abdomen; mcs: macrosetae; m ter: male terminalia." figureDoi="http://doi.org/10.5281/zenodo.13195165" httpUri="https://zenodo.org/record/13195165/files/figure.png" pageId="4" pageNumber="141">Fig. 27</figureCitation>
). Aedeagal prongs with subapical dilation (
<figureCitation id="B5B92A4BFFB3FFF6DB97FA19FED6DAE1" box="[207,322,1490,1509]" captionStart="Figures 2229" captionStartId="6.[114,177,1253,1267]" captionTargetBox="[396,1219,151,1223]" captionTargetId="figure-14@6.[393,1225,150,1222]" captionTargetPageId="6" captionText="Figures 2229. Triorla beckeri sp. nov. 22. Apex of aedeagal prongs; 23. Gonocoxite; 24. Hypandrium; 25. Male abdomen; 26. Macrosetae of epandrium; 27. Gonostylus in lateral view; 28. Aedeagus and ejaculatory apodeme in lateral view; 29. Male terminalia in lateral view. Abbreviations: abd: abdomen; mcs: macrosetae; m ter: male terminalia." figureDoi="http://doi.org/10.5281/zenodo.13195165" httpUri="https://zenodo.org/record/13195165/files/figure.png" pageId="4" pageNumber="141">Figs. 22, 28</figureCitation>
); aedeagal sheath as long as prongs (
<figureCitation id="B5B92A4BFFB3FFF6D998FA19FC90DAE1" box="[704,772,1490,1509]" captionStart="Figures 2229" captionStartId="6.[114,177,1253,1267]" captionTargetBox="[396,1219,151,1223]" captionTargetId="figure-14@6.[393,1225,150,1222]" captionTargetPageId="6" captionText="Figures 2229. Triorla beckeri sp. nov. 22. Apex of aedeagal prongs; 23. Gonocoxite; 24. Hypandrium; 25. Male abdomen; 26. Macrosetae of epandrium; 27. Gonostylus in lateral view; 28. Aedeagus and ejaculatory apodeme in lateral view; 29. Male terminalia in lateral view. Abbreviations: abd: abdomen; mcs: macrosetae; m ter: male terminalia." figureDoi="http://doi.org/10.5281/zenodo.13195165" httpUri="https://zenodo.org/record/13195165/files/figure.png" pageId="4" pageNumber="141">Fig. 28</figureCitation>
), dilation of median prong situated more anteriorly than same on lateral prongs (
<figureCitation id="B5B92A4BFFB3FFF6DA54F9C1FEDBD919" box="[268,335,1546,1565]" captionStart="Figures 2229" captionStartId="6.[114,177,1253,1267]" captionTargetBox="[396,1219,151,1223]" captionTargetId="figure-14@6.[393,1225,150,1222]" captionTargetPageId="6" captionText="Figures 2229. Triorla beckeri sp. nov. 22. Apex of aedeagal prongs; 23. Gonocoxite; 24. Hypandrium; 25. Male abdomen; 26. Macrosetae of epandrium; 27. Gonostylus in lateral view; 28. Aedeagus and ejaculatory apodeme in lateral view; 29. Male terminalia in lateral view. Abbreviations: abd: abdomen; mcs: macrosetae; m ter: male terminalia." figureDoi="http://doi.org/10.5281/zenodo.13195165" httpUri="https://zenodo.org/record/13195165/files/figure.png" pageId="4" pageNumber="141">Fig. 22</figureCitation>
).
</paragraph>
<paragraph id="2D3D36CEFFB3FFF6DBCAF9EDFE05D93D" blockId="4.[114,786,1183,1788]" box="[146,401,1573,1593]" pageId="4" pageNumber="141">
Size. Body length
<quantity id="EA7A9B2BFFB3FFF6DA1DF9EEFE19D93D" box="[325,397,1573,1593]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="141" unit="mm" value="15.0">15 mm</quantity>
.
</paragraph>
<paragraph id="2D3D36CEFFB3FFF6DBCAF98AFEC4D950" blockId="4.[114,786,1183,1788]" box="[146,336,1601,1620]" pageId="4" pageNumber="141">Female. Unknown.</paragraph>
<paragraph id="2D3D36CEFFB3FFF6DBCAF996FF6BD988" blockId="4.[114,786,1183,1788]" pageId="4" pageNumber="141">Etymology. This species is named for Johann Becker, a Brazilian entomologist.</paragraph>
<paragraph id="2D3D36CEFFB3FFF6DBCAF95EFDA7D9AC" blockId="4.[114,786,1183,1788]" box="[146,563,1685,1704]" pageId="4" pageNumber="141">
Geographical records.
<collectingCountry id="5595765EFFB3FFF6DA2CF95EFE27D9AC" box="[372,435,1685,1704]" name="Brazil" pageId="4" pageNumber="141">Brazil</collectingCountry>
:
<collectingRegion id="EF46F82CFFB3FFF6DAE3F95EFDBAD9AC" box="[443,558,1685,1704]" country="Brazil" name="Bahia" pageId="4" pageNumber="141">Bahia State</collectingRegion>
.
</paragraph>
<paragraph id="2D3D36CEFFB3FFF6DBCAF979FF2DD9F8" blockId="4.[114,786,1183,1788]" pageId="4" pageNumber="141">
Type material.
<materialsCitation id="9DEA3C93FFB3FFF6DA7DF97AFF2DD9F8" collectingDate="1993-12" collectionCode="MZFS" collectorName="de Jaua &amp; J. Becker" country="Brazil" location="Dunas de Jaua" municipality="Camacari" pageId="4" pageNumber="141" specimenCount="1" specimenCount-male="1" stateProvince="Bahia" typeStatus="holotype">
<typeStatus id="F239886CFFB3FFF6DA7DF97AFE13D9C0" box="[293,391,1713,1732]" pageId="4" pageNumber="141" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="5595765EFFB3FFF6DAD6F97AFE43D9C0" box="[398,471,1713,1732]" name="Brazil" pageId="4" pageNumber="141">BRASIL</collectingCountry>
,
<collectingRegion id="EF46F82CFFB3FFF6DA86F97AFD8CD9C0" box="[478,536,1713,1732]" country="Brazil" name="Bahia" pageId="4" pageNumber="141">Bahia</collectingRegion>
,
<collectingMunicipality id="CD59ACB4FFB3FFF6D947F97AFDEBD9C0" box="[543,639,1713,1732]" pageId="4" pageNumber="141">Camaçari</collectingMunicipality>
,
<location id="285D6015FFB3FFF6D9DEF979FC85D9C0" LSID="urn:lsid:plazi:treatment:A52B87D8FFB4FFF6DB0DFA19FF2DD9F8:285D6015FFB3FFF6D9DEF979FC85D9C0" box="[646,785,1713,1733]" country="Brazil" municipality="Camacari" name="Dunas de Jaua" pageId="4" pageNumber="141" stateProvince="Bahia">
Dunas
<collectorName id="80775318FFB3FFF6D991F97AFC85D9C0" box="[713,785,1713,1732]" pageId="4" pageNumber="141">de Jauá</collectorName>
</location>
[12
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46
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S, 38
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21
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W],
<date id="593C100EFFB3FFF6DAD3F906FE4BD9E4" box="[395,479,1741,1760]" pageId="4" pageNumber="141" value="1993-12">
<collectingDate id="4978E9E6FFB3FFF6DAD3F906FE4BD9E4" box="[395,479,1741,1760]" pageId="4" pageNumber="141" value="1993-12">xii.1993</collectingDate>
</date>
.
<collectorName id="80775318FFB3FFF6DABFF906FDA8D9E4" box="[487,572,1741,1760]" pageId="4" pageNumber="141">J. Becker</collectorName>
col., MZFS#29005 (
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<collectionCode id="4B93AE0BFFB3FFF6DB2AF922FF3BD9F8" box="[114,175,1769,1788]" pageId="4" pageNumber="141">MZFS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>