474 lines
56 KiB
XML
474 lines
56 KiB
XML
<document id="113DF37A021FFFE2A57C92B218D701C8" ID-CLB-Dataset="29992" ID-DOI="http://dx.doi.org/10.3897/zookeys.763.21252" ID-GBIF-Dataset="6162ab6f-6b50-40ee-b28e-c72eee01e99e" ID-PMC="PMC5996013" ID-Pensoft-Pub="1313-2970-763-1" ID-PubMed="29896045" ID-ZooBank="90B772554C5E436CA793D924892B5B14" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-763-1" ModsDocOrigin="ZooKeys 763" ModsDocTitle="Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae)" checkinTime="1528130281812" checkinUser="pensoft" docAuthor="Goulding, Tricia C., Khalil, Munawar, Tan, Shau Hwai & Dayrat, Benoit" docDate="2018" docId="A505118479BC5E7CF886F5D33ADD1D3A" docLanguage="en" docName="ZooKeys 763: 1-111" docOrigin="ZooKeys 763" docSource="http://dx.doi.org/10.3897/zookeys.763.21252" docTitle="Wallaconchis buetschlii Stantschinsky 1907, comb. n." docType="treatment" docVersion="6" lastPageNumber="1" masterDocId="FFF78434FFEAFFF5FFBBF264B1300C6B" masterDocTitle="Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae)" masterLastPageNumber="111" masterPageNumber="1" pageNumber="1" updateTime="1701377276478" updateUser="ExternalLinkService">
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<mods:title id="571FFFAB31AB908A5C47455468A38C3C">Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda, Pulmonata, Onchidiidae)</mods:title>
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<mods:roleTerm id="7D9BCD8E08B334BCF663085B64D73A1D">Author</mods:roleTerm>
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<mods:namePart id="A179F514CA48B88C789C90ED20D9AA02">Goulding, Tricia C.</mods:namePart>
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<mods:namePart id="ABC6B302646599A85D3885A4820BB131">Khalil, Munawar</mods:namePart>
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<mods:namePart id="3A4379B23DBA5FEC874EDBE831A32E1A">Tan, Shau Hwai</mods:namePart>
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<mods:roleTerm id="884B57CCB1BF3DC289A9BBEE03AE9080">Author</mods:roleTerm>
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<mods:namePart id="B6A0D8E75E12956A7AA5A32ED71435FF">Dayrat, Benoit</mods:namePart>
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<mods:typeOfResource id="525A082562B1625EE59205A0F79633EF">text</mods:typeOfResource>
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<mods:title id="29C586EEEF1F66104E034717D74ED609">ZooKeys</mods:title>
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<mods:date id="AB65B439794B7773EDF6DD6E87F69120">2018</mods:date>
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<mods:number id="E2A1E09B89593474B498CD90553CE337">763</mods:number>
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<mods:url id="A71E2342D5F7D0B6345326CE592B935C">http://dx.doi.org/10.3897/zookeys.763.21252</mods:url>
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<mods:classification id="E16CE577A4FDD69BE3DEDD0D310E51EA">journal article</mods:classification>
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<mods:identifier id="9613D774CE58B3B0764FEDECA5727622" type="DOI">http://dx.doi.org/10.3897/zookeys.763.21252</mods:identifier>
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<mods:identifier id="6318EA03E52B1A1AB9A9A130F19D06DB" type="Pensoft-Pub">1313-2970-763-1</mods:identifier>
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<treatment id="A505118479BC5E7CF886F5D33ADD1D3A" ID-GBIF-Taxon="144234918" LSID="urn:lsid:plazi:treatment:A505118479BC5E7CF886F5D33ADD1D3A" httpUri="http://treatment.plazi.org/id/A505118479BC5E7CF886F5D33ADD1D3A" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection id="99FCB6E66BFC84C32876E09A88BB9A88" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="42F2C05FEEB61CA01372AA799715BAA5" pageId="0" pageNumber="1">
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<taxonomicName id="88A1BE00EB9EA577DFDDCA7CCC9768CF" ID-CoL="7G6Q8" authority="Stantschinsky, 1907" authorityName="Stantschinsky" authorityYear="1907" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis buetschlii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="buetschlii">Wallaconchis buetschlii (Stantschinsky, 1907)</taxonomicName>
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<taxonomicNameLabel id="4A69D847F25BB1082B4DA2EE037D1B51" pageId="0" pageNumber="1">comb. n.</taxonomicNameLabel>
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Figs 19, 20, 21, 22, 23
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</paragraph>
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</subSubSection>
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<subSubSection id="5E6B593573F7917F66E1EC5E832AF31E" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="CE69628B08D97D1A1C1240D9EE19EC62" pageId="0" pageNumber="1">
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<taxonomicName id="3B1223AF2FBAAE3D20F61B3DAB84C375" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium buetschlii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="buetschlii">Onchidium buetschlii</taxonomicName>
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Stantschinsky, 1907: 383-386, pl. 12, figs 10-12, pl. 13, figs 20a, 20b, 22, 35;
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<bibRefCitation id="B818689FBBC263E5370F0AB9DD43A131" author="Bretnall, W" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="303 - 328" title="Onchidiidae from Australia and the South-Western Pacific Islands." url="https://doi.org/10.3853/j.0067-1975.12.1919.888" volume="12" year="1919">Bretnall 1919</bibRefCitation>
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: 316 (as
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<taxonomicName id="B3C7EC7862271409EC2B53E301639B2C" class="Gastropoda" family="Onchidiidae" genus="Oncidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oncidium buetschlii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="buetschlii">Oncidium buetschlii</taxonomicName>
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).
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</paragraph>
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<paragraph id="D85ED7A2FD7C6DD78245788207356D8E" pageId="0" pageNumber="1">
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<taxonomicName id="0ADA0D44DBB943A6239D11E2FBDFC33F" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium gracile" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gracile">Onchidium gracile</taxonomicName>
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:
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<bibRefCitation id="1B09F7B9330936662ACA6BCF59791B34" author="Stantschinsky, W" journalOrPublisher="Zoologische Jahrbuecher, Systematik" pageId="0" pageNumber="1" pagination="353 - 402" title="Zur Anatomie und Systematik der Gattung Oncidium." url="https://doi.org/10.5962/bhl.part.11959" volume="25" year="1907">Stantschinsky 1907</bibRefCitation>
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: pl. 13, fig. 34 [non
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<taxonomicName id="2468D74F834CA4D06F55285BD51D8F40" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium gracile" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gracile">Onchidium gracile</taxonomicName>
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Stantschinsky, 1907].
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</paragraph>
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</subSubSection>
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<subSubSection id="FE43C833736D72720599A22B599E3693" pageId="0" pageNumber="1" type="type locality">
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<paragraph id="CAFE6D8A76D0EC3CE55022EE765A2B05" pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph id="17A0D9B0FB61E89B31A34114FEC1615A" pageId="0" pageNumber="1">Australien: Queensland [Queensland, Australia]. Stantschinsky indicated that the syntypes were collected from the state of Queensland by Professor Simroth, but no detailed information about the locality is available in either the original description or the jar of the type material. Fresh material was collected from the type locality (see below, additional material).</paragraph>
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</subSubSection>
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<subSubSection id="4702BFAA41B9FF1A1DD7C7FD37B86C69" pageId="0" pageNumber="1" type="type material">
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<paragraph id="BAD401C6F3EED68A7FD2BF51D9807D0E" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="67D6D6DE078411DD89C35FFEBC2E18E9" pageId="0" pageNumber="1">Lectotype, 19/17 mm, designated here (SMF 333595/2). The lectotype was dissected prior to the present study and its posterior end is missing. A lectotype is designated here because its remaining anterior, male parts can be used for species identification. The two other syntypes (30/22 mm and 30/29 mm) become paralectotypes (SMF 333595/2). Both paralectotypes were dissected prior to the present study and their male parts are missing. According to the original description, there were four syntypes, so one syntype has been lost or destroyed.</paragraph>
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</subSubSection>
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<subSubSection id="313B056134BB6B098B7A9382E2143B9A" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="1C282D1E244284E9EC5C167A21078B8C" pageId="0" pageNumber="1">Additional material examined.</paragraph>
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<paragraph id="AC6358CFB4573D26CEFC976EEB858B3D" pageId="0" pageNumber="1">
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Indonesia, North Sulawesi, Wori,
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<geoCoordinate id="A62CDE4DE989178E9D7D344750E9D11C" direction="north" orientation="latitude" precision="9" value="1.6010001">01°36.06'N</geoCoordinate>
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,
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<geoCoordinate id="43BC3406DA9F18957B6D6FF52368E93A" direction="east" orientation="longitude" precision="9" value="124.86217">124°51.73'E</geoCoordinate>
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, 2 specimens 31/14 mm [2120] and 27/14 mm [2122], st 84, old
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<taxonomicName id="58CE654BC52418E48CC1ADDBDAC68781" class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
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and
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<taxonomicName id="F9B5FF946B320CED6145B95D16DCCB2C" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
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mangrove (UMIZ 00021); North Sulawesi, Bahoi,
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<geoCoordinate id="43428A6FAA2B5FE5069C94DFE3DCB405" direction="north" orientation="latitude" precision="9" value="1.7226667">01°43.36'N</geoCoordinate>
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,
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<geoCoordinate id="144A668C54A7A3753249B4980F5BE1F9" direction="east" orientation="longitude" precision="9" value="125.0205">125°01.23'E</geoCoordinate>
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, 1 specimen 22/18 mm [2189], st 85, sand and small rocks outside a mangrove (UMIZ 00022); North Sulawesi, Wori,
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<geoCoordinate id="9B7695037624C1359290BED69A043271" direction="north" orientation="latitude" precision="9" value="1.6010001">01°36.06'N</geoCoordinate>
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,
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<geoCoordinate id="1BF049AC20A9777A5DCBE7462A765474" direction="east" orientation="longitude" precision="9" value="124.86217">124°51.73'E</geoCoordinate>
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, 1 specimen 23/14 mm [2249], st 90, old mangrove forest with
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<taxonomicName id="7CEF6885B20CDEF18DC038EACBC1A29B" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
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,
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<taxonomicName id="CF10D359365C30C341068C8BAC1F1793" class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
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, and
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<taxonomicName id="EBD0487871B564F184A8B91117B7E13E" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhizophora" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Rhizophora</taxonomicName>
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, with rocks (UMIZ 00024); North Sulawesi, Mantehage Island,
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<geoCoordinate id="9CFC91EC4CFA5872CA9472AC581FDF13" direction="north" orientation="latitude" precision="9" value="1.698">01°41.88'N</geoCoordinate>
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,
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<geoCoordinate id="1CB452D946C0C19D3B7544426D814A3A" direction="east" orientation="longitude" precision="9" value="124.779">124°46.74'E</geoCoordinate>
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, 1 specimen 32/22 mm [2310], st 91, rocks behind a mangrove of
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<taxonomicName id="D8B98E917B260165211DDA2FB2F6B59D" class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
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and
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<taxonomicName id="DC01075DDCDBCF3DCE8B69FC8922AC32" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhizophora" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Rhizophora</taxonomicName>
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(UMIZ 00025); Ambon, Haruku Island,
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<geoCoordinate id="20DF4C15B516E04BE931C45B6F4A1112" direction="south" orientation="latitude" precision="9" value="-3.6086667">03°36.52'S</geoCoordinate>
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,
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<geoCoordinate id="76F6D87D207B0C14FCB9F950695A66E5" direction="east" orientation="longitude" precision="9" value="128.41783">128°25.07'E</geoCoordinate>
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, 1 specimen 46/21 mm [2722], st 127, rocky
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<taxonomicName id="5F2B5A4AD47FF4BB31E41093B6DC3A3B" class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
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mangrove with coral rubble (UMIZ 00026); Ambon, Wai,
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<geoCoordinate id="3444658A1724006449C1EBA542A8277D" direction="south" orientation="latitude" precision="9" value="-3.5775">03°34.65'S</geoCoordinate>
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,
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<geoCoordinate id="4730DA8850BBBDFC1AFCAE84990FEC12" direction="east" orientation="longitude" precision="9" value="128.3255">128°19.53'E</geoCoordinate>
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, 1 specimen 31/16 mm [2853], st 132, narrow band of old
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<taxonomicName id="2324B48D767F11D644A660FD2ED24452" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
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trees on sandy mud, old logs on ground (UMIZ 00027); Seram,
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<geoCoordinate id="2AAA688472761D83C0F4A56965FF6DB7" direction="south" orientation="latitude" precision="9" value="-2.9706666">02°58.24'S</geoCoordinate>
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,
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<geoCoordinate id="8CB87C03549A5C12D495E01C6D0A9E4C" direction="east" orientation="longitude" precision="9" value="128.11783">128°07.07'E</geoCoordinate>
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, 1 specimen 21/17 mm [2872], st 135, mud next to a mangrove (UMIZ 00028); Maluku, Kei Islands, Tual City, Fiditan,
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<geoCoordinate id="F92DC97EEA622AC65E4CA5F2B55F20F2" direction="south" orientation="latitude" precision="9" value="-5.5993333">05°35.96'S</geoCoordinate>
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,
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<geoCoordinate id="D980ADA02D470A5E52D589498F280B05" direction="east" orientation="longitude" precision="9" value="132.75183">132°45.11'E</geoCoordinate>
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, 2 specimens 30/16 mm [2928] and 30/15 mm [2933], st 144, rocks behind muddy mangrove of
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<taxonomicName id="252ABDFC3709E0CA77A78B3BBBBB617A" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhizophora" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Rhizophora</taxonomicName>
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(UMIZ 00029); Lombok, Don Don,
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<geoCoordinate id="5AC85A47ACABFBD116A59C4EE5C21699" direction="south" orientation="latitude" precision="9" value="-8.909">08°54.54'S</geoCoordinate>
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,
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<geoCoordinate id="0ADB894EA2EDCEEEF902A006DFE5AF7F" direction="east" orientation="longitude" precision="9" value="116.35833">116°21.50'E</geoCoordinate>
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, 2 specimens 43/23 mm [2982] and 43/23 mm [2989] st 149, old,
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<taxonomicName id="144005FB37908B190881A75316ED0301" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
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forest with coral rubble (UMIZ 00030); Bali, Gilimanuk,
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<geoCoordinate id="3D96AE14B0DC9801D7B3469B0EAA83C6" direction="south" orientation="latitude" precision="9" value="-8.169333">08°10.16'S</geoCoordinate>
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,
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<geoCoordinate id="8B40DECDA092157D5C5EE1EBAC7780C8" direction="east" orientation="longitude" precision="9" value="114.44417">114°26.65'E</geoCoordinate>
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, 1 specimen 24/15 mm [3123], st 156, sandy mudflat outside
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<taxonomicName id="E42B67D7406400674F5D43B8B66BA727" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhizophora" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Rhizophora</taxonomicName>
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and
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<taxonomicName id="309F737D047E776067E293B08330E115" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
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mangrove (UMIZ 00032); Bali, Pemuteran, Labuhan Lalang Harbor,
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||
<geoCoordinate id="F918EE91D0B00EA18486175CACCC65B5" direction="south" orientation="latitude" precision="9" value="-8.1435">08°08.61'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="9D06713928FCF42F652971F99DF554A6" direction="east" orientation="longitude" precision="9" value="114.53883">114°32.33'E</geoCoordinate>
|
||
, 1 specimen 27/17 mm [3128], st 157, coral rubble, rocks and mud with a few
|
||
<taxonomicName id="0ECDD2FC839C864566B3D3515472439B" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
(UMIZ 00033); North Maluku, Ternate, Bastiong,
|
||
<geoCoordinate id="CD6FA1F3B96E4C195B0BEBC99DA70B32" direction="north" orientation="latitude" precision="9" value="0.7735">00°46.41'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2E2C65A46490B13D189DC774E175AB88" direction="east" orientation="longitude" precision="9" value="127.37933">127°22.76'E</geoCoordinate>
|
||
, 1 specimen 32/19 mm [5016], st 203, muddy rocks near a mangrove (UMIZ 00034); Halmahera, Sofifi,
|
||
<geoCoordinate id="2BA7E20B3651701F428E532494303F0E" direction="north" orientation="latitude" precision="9" value="0.75783336">00°45.47'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B51A62A4483509A98BF96DAF092F5B7E" direction="east" orientation="longitude" precision="9" value="127.598335">127°35.90'E</geoCoordinate>
|
||
, 1 specimen 37/20 mm [5067], st 205,
|
||
<taxonomicName id="F13D8D259EA6C5A91EA1965DD833B1D5" class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
|
||
mangrove (UMIZ 00035); Timor, Oesapa,
|
||
<geoCoordinate id="46F4845753E624ED833EE706866E9390" direction="south" orientation="latitude" precision="9" value="-10.1455">10°08.73'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="9660E0E45535B8EF807C14E126A19C24" direction="east" orientation="longitude" precision="9" value="123.635">123°38.10'E</geoCoordinate>
|
||
, 1 specimen 44/26 mm [5898], st 250, sandy area with
|
||
<taxonomicName id="43D8DBDFE75B3AABCD96504DE47E10E3" class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
|
||
and
|
||
<taxonomicName id="39DA5C70AEF42F6007242973ABCB7C10" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
(UMIZ 00072). Australia, Queensland, Cairns, Yule Point,
|
||
<geoCoordinate id="6EF3B316DC6F776F66AD34B6941E4EE9" direction="south" orientation="latitude" precision="9" value="-16.5705">16°34.23'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="937B2CBAF222C1BA286973F49DB0F398" direction="east" orientation="longitude" precision="9" value="145.50967">145°30.58'E</geoCoordinate>
|
||
, 2 specimens 43/23 mm [2554] and 16/10 mm [2555], st 100, sand flat outside mangrove with dense
|
||
<taxonomicName id="6CB643C1C087B55306D98E78465832AB" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhizophora" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Rhizophora</taxonomicName>
|
||
(st100, MTQ). Philippines, Mindanao, [no information on the collecting date], 1 specimen 28/20 mm, paralectotype of
|
||
<taxonomicName id="7BB653C31FCCD54CAB1E40FF4CC05CCC" lsidName="W. gracile" pageId="0" pageNumber="1" rank="species" species="gracile">W. gracile</taxonomicName>
|
||
, [no information on the collector] (ZMB 103082b); Luzon, Batangas, Lian,
|
||
<geoCoordinate id="B2F07757AD44B07A414FB77AE65378A3" direction="north" orientation="latitude" precision="9" value="13.996">13°59.76'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C29498B3DCB7F791950B7B5C30C8846E" direction="east" orientation="longitude" precision="9" value="120.62383">120°37.43'E</geoCoordinate>
|
||
, 1 specimen 52/34 mm [3159], st 181, sandy, open
|
||
<taxonomicName id="63CCACECA8288AE986999905D9CDCE5F" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
forest (PNM 041209); Bohol, Guindulman,
|
||
<geoCoordinate id="4E25B40D6539AEF6650563E5081C8813" direction="north" orientation="latitude" precision="9" value="9.734333">09°44.06'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="8F46612B34B6498F4DB558E3C6023C9C" direction="east" orientation="longitude" precision="9" value="124.4605">124°27.63'E</geoCoordinate>
|
||
, 1 specimen 31/25 mm [3631], st 197, rocks, coral rubble, and sand near a few
|
||
<taxonomicName id="95FAA6622879F5634CC5E888E8EEEE34" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
trees (PNM 041210); Bohol, Loay,
|
||
<geoCoordinate id="5CFD1220EBFB43187C95442F382233CD" direction="north" orientation="latitude" precision="9" value="9.603833">09°36.23'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="445EC0DD9FCF4E0D8FC2CC816540656F" direction="east" orientation="longitude" precision="9" value="123.99533">123°59.72'E</geoCoordinate>
|
||
, 1 specimen 35/20 mm [3637], st 198, mostly sand, and a few
|
||
<taxonomicName id="63838E487FB338D6626AA0CA5879E0E7" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
(PNM 041211); Bohol, Maribojoc,
|
||
<geoCoordinate id="5B565D8152740A91D033A582334CE218" direction="north" orientation="latitude" precision="9" value="9.738">09°44.28'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="9E9BB67306818F5FB79E4E28EF561481" direction="east" orientation="longitude" precision="9" value="123.823166">123°49.39'E</geoCoordinate>
|
||
, 2 specimens 36/17 mm [3409] and 18/13 mm [3410], st 202, coral rubble with sand and algae, near
|
||
<taxonomicName id="6BD495BA9CD89CD61DB9DB09AF346C72" class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
|
||
(PNM 041212).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="78C55A5FF593811C1036305171EBF425" pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph id="1D0E9CE79A10EB41381FC557D34CF683" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph id="5361CCD8E01DF7C3777F824FAB548E2E" pageId="0" pageNumber="1">Australia: Queensland (type locality). Indonesia: Ambon, Bali, Halmahera, Kei Islands, Lombok, Seram, Sulawesi, and Timor. Philippines: Bohol and Luzon. All records, except the type locality, are new.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="32C4027B3D17E716FB0FDB3135AE77AC" pageId="0" pageNumber="1" type="habitat">
|
||
<paragraph id="7AE77C7B044A27A4FFE3CCA8F213C0E9" pageId="0" pageNumber="1">Habitat</paragraph>
|
||
<paragraph id="06ED849E042D0CD631229208A995E3A8" pageId="0" pageNumber="1">
|
||
(Fig. 19, Table 3).
|
||
<taxonomicName id="C2541D2C23F6971D2865F87D9CCF8D6A" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis buetschlii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="buetschlii">Wallaconchis buetschlii</taxonomicName>
|
||
is found on firm mud or coarse sand in mangroves (i.e., mud which is not saturated in water) along with
|
||
<taxonomicName id="30A0A4269B4405D120834CBB0AA9E90D" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis uncinus" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="uncinus">Wallaconchis uncinus</taxonomicName>
|
||
. It also lives on fine, sandy mud along with
|
||
<taxonomicName id="E93C0365F8C0DB925FED123047279284" lsidName="W. gracile" pageId="0" pageNumber="1" rank="species" species="gracile">W. gracile</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="59EB402041935DB01A65ACF72A28C8B5" pageId="0" pageNumber="1">
|
||
<paragraph id="420AB0D2870FE5D0A12A51C01EA264D2" pageId="0" pageNumber="1">
|
||
Figure 19. Habitats,
|
||
<taxonomicName id="19B38DA376014CDF2812A99E5DC41BBC" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis buetschlii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="buetschlii">Wallaconchis buetschlii</taxonomicName>
|
||
. A Indonesia, Sulawesi, Wori, tall mangrove forest of
|
||
<taxonomicName id="2CAE2ECADBDFFF11A207C18ACFE7FE2D" class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
|
||
and some
|
||
<taxonomicName id="9F71408C0AD06EAA3B92FFC681B4B90F" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
with a rocky area and dead logs (st 84) B Australia, Queensland, Yule Point, sandy area with a few
|
||
<taxonomicName id="305A733A62F73088F8A9195089D1264B" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhizophora" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Rhizophora</taxonomicName>
|
||
and
|
||
<taxonomicName id="6FE2FF31B7706CEDA33F12C01FECD2A8" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
trees (st 100) C Indonesia, Ambon, Wai, narrow band of old
|
||
<taxonomicName id="ED0EAF26B7906FC4C790FEAFCEB1A887" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
trees on sandy mud, with old logs on ground (st 132) D Indonesia, Lombok, Don Don, old
|
||
<taxonomicName id="E3BAED32FF4B34D228A4DB0C6E67105E" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
forest, with coral rubble (st 149) E Indonesia, Sulawesi, Mantehage Island, rocks behind a mangrove of
|
||
<taxonomicName id="33D6C0EA0093F4EE9DCF7F6918963220" class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sonneratia" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sonneratia</taxonomicName>
|
||
and
|
||
<taxonomicName id="21A73E6E45BC342BD9371C5294A6D7B9" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhizophora" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Rhizophora</taxonomicName>
|
||
(st 91) F Indonesia, Bali, Gilimanuk, sandy mudflat outside
|
||
<taxonomicName id="5F0EE67B8CDED1109751CB9DBD867079" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhizophora" order="Malpighiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Rhizophora</taxonomicName>
|
||
and
|
||
<taxonomicName id="DDC5777E3C3394C5B765B1BB6047B589" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Avicennia" order="Lamiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Avicennia</taxonomicName>
|
||
mangrove (st 156).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="EAA1AFA5AF7FC44BBD14300C13C46F5A" pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph id="85C8B4F4E301F12EC3F50D269A678477" pageId="0" pageNumber="1">Diagnosis</paragraph>
|
||
<paragraph id="11DB7FF3DE6F32A93AD3A82F8941DC1D" pageId="0" pageNumber="1">
|
||
(Table 5). The slugs that are part of
|
||
<taxonomicName id="B553C73E1FA1C31676A2F0D9FC513151" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis buetschlii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="buetschlii">Wallaconchis buetschlii</taxonomicName>
|
||
are the largest of the genus (the largest live animal measured 60 mm). However, species cannot reliably be distinguished based on individual length. Red individuals cannot be distinguished from several species (usually
|
||
<taxonomicName id="F5A1F496AEC2D25D109998963624E18C" lsidName="W. nangkauriense" pageId="0" pageNumber="1" rank="species" species="nangkauriense">W. nangkauriense</taxonomicName>
|
||
and
|
||
<taxonomicName id="C62B35655C2D94389E57FAFED08953DF" lsidName="W. ater" pageId="0" pageNumber="1" rank="species" species="ater">W. ater</taxonomicName>
|
||
), and grey or brown individuals cannot be distinguished from other species. Dorsal papillae with yellow tips can be prominent in
|
||
<taxonomicName id="1B3429AA2C357FE45B8022DE998FA815" lsidName="W. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">W. buetschlii</taxonomicName>
|
||
, but also can be visible in other
|
||
<taxonomicName id="A54E687C03E276D42DDB10049646F48A" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
species (particularly in
|
||
<taxonomicName id="4A288BA71BBC5A95EAEAC6E60FA5B9D2" lsidName="W. melanesiensis" pageId="0" pageNumber="1" rank="species" species="melanesiensis">W. melanesiensis</taxonomicName>
|
||
). In addition, the dorsal papillae with yellow tips found on the notum of
|
||
<taxonomicName id="D735F91B940AF90D006E8F4058FB00E1" lsidName="W. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">W. buetschlii</taxonomicName>
|
||
are nearly identical to those on the notum of several species in other genera. The color of the hyponotum is helpful to distinguish
|
||
<taxonomicName id="0A9BABDD985C01A6094CD09518F70D1D" lsidName="W. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">W. buetschlii</taxonomicName>
|
||
(grey or yellow-grey hyponotum) from species in other genera (orange hyponotum) that live in the same habitat but is not fully reliable and should only be used with caution (i.e., the orange hyponotum in other onchidiids is occasionally a dull orange-grey similar to the yellow-grey hyponotum of
|
||
<taxonomicName id="F0928AE03889AFDB79060B416CD3FBB1" lsidName="W. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">W. buetschlii</taxonomicName>
|
||
). As a result,
|
||
<taxonomicName id="98686A25E1B3BCCCEE867F750B9DBA63" lsidName="W. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">W. buetschlii</taxonomicName>
|
||
cannot be reliably distinguished externally from other onchidiid species.
|
||
</paragraph>
|
||
<paragraph id="310E5955A6983ED8C166D27894A04231" pageId="0" pageNumber="1">
|
||
Internally, the penis is diagnostic of
|
||
<taxonomicName id="50D19C62162267842A8BB55A847E8353" lsidName="W. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">W. buetschlii</taxonomicName>
|
||
: it is not protected by a penial sheath or a vestibule and bears internal longitudinal ridges. The oviduct attached to the posterior wall of the visceral cavity is peculiar but it also occurs in
|
||
<taxonomicName id="AC6514070011AB2EB5A7A3CE4D7BEC44" lsidName="W. gracile" pageId="0" pageNumber="1" rank="species" species="gracile">W. gracile</taxonomicName>
|
||
. However, the penis of
|
||
<taxonomicName id="B893EFA0C34550636BF25A8E52C41315" lsidName="W. gracile" pageId="0" pageNumber="1" rank="species" species="gracile">W. gracile</taxonomicName>
|
||
is protected by a penial sheath and extends into the vestibule.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4C49C7A5F5876F8C4128B63ECFE0173A" pageId="0" pageNumber="1" type="color and morphology of live animals">
|
||
<paragraph id="A989BD1BFCABC6E722EF81FE22BD730F" pageId="0" pageNumber="1">Color and morphology of live animals</paragraph>
|
||
<paragraph id="5CC7E431C4E6CA60EEC9FA20FA9FDD96" pageId="0" pageNumber="1">(Fig. 20). The dorsal coloration is variable. It is usually brown or grey, but occasionally brown with red or yellow patches, or even completely red. In some individuals, many small yellow dots are present on the dorsal notum. The color of the ocular tentacles also varies, and may be yellow-orange, brown, or reddish brown. The hyponotum color can be grey, light grey, or yellow-grey. The foot is also grey or yellow-grey. Crawling slugs tend to be elongated, but they contract when disturbed.</paragraph>
|
||
<caption id="927ABE7558EC9024727F5250E84D3E4D" pageId="0" pageNumber="1">
|
||
<paragraph id="18DC1A67FEED29A9E34D251FD86AA94F" pageId="0" pageNumber="1">
|
||
Figure 20. Live specimens,
|
||
<taxonomicName id="CB3C37F4CED3CEFABD2A9AC02ACA2F07" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis buetschlii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="buetschlii">Wallaconchis buetschlii</taxonomicName>
|
||
. A Dorsal view, 46 mm long [2722], Indonesia, Ambon (UMIZ 00026) B Dorsal view, 31 mm long [2853], Indonesia, Ambon (UMIZ 00027) C Dorsal view, 23 mm long [2249], Indonesia, Sulawesi (UMIZ 00024) D Dorsal view, 30 mm long [2928], Indonesia, Kei Islands (UMIZ 00029) E Dorsal view, 52 mm long [3159], Philippines, Luzon (PNM 041209) F Dorsal view, 36 mm long, [3409], Philippines, Bohol (PNM 041212) G Ventral view, 44 mm long [5898], Indonesia, Timor (UMIZ 00072) H Ventral view, same as F I Ventral view, 43 mm long [2982], Indonesia, Lombok (UMIZ 00030).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="88EF71FC112B22DF17E14B8490601147" pageId="0" pageNumber="1" type="external morphology">
|
||
<paragraph id="719684B907143C4359E8CD0B581DA25C" pageId="0" pageNumber="1">External morphology.</paragraph>
|
||
<paragraph id="38A323CF329B9369787BD69BA3CE0440" pageId="0" pageNumber="1">There are many papillae on the dorsal surface, which are usually bright yellow in color, or occasionally brown. Between six and 14 papillae bear eyes, with three or four eyes per papilla. There is a large, retractable papilla with eyes in the center of the dorsal notum, and it is often raised several millimeters above the dorsal surface.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6346264A753DF448522E3C2E981955AA" pageId="0" pageNumber="1" type="digestive system">
|
||
<paragraph id="5EE2F95258ED8D07A10BD1D8990E89EC" pageId="0" pageNumber="1">Digestive system</paragraph>
|
||
<paragraph id="697E56BDD56CAD71DAF748A4549330C1" pageId="0" pageNumber="1">
|
||
(Fig. 21, Table 4). Examples of radular formulae are presented in Table 4. The length of the rachidian teeth is approximately 20
|
||
<normalizedToken id="0C98A3F199533CE3C2E12FEE24666FA7" originalValue="µm">µm</normalizedToken>
|
||
, significantly smaller than that of the lateral teeth. The length of the hook of the lateral teeth gradually increases, from 40 to 75
|
||
<normalizedToken id="C3D17EEADCE29D9ABED610580FF7F812" originalValue="µm">µm</normalizedToken>
|
||
, from the inner to the outer teeth, excluding the innermost and outermost lateral teeth which are significantly smaller. The intestinal loops are of type I.
|
||
</paragraph>
|
||
<caption id="B6580E6C701781672EB432AD8FBE15A9" pageId="0" pageNumber="1">
|
||
<paragraph id="1F9E1B69170B8F560E8D878238139FBB" pageId="0" pageNumber="1">
|
||
Figure 21. Radula,
|
||
<taxonomicName id="F6487DCC58B303B5F9A541C7799C1B0A" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis buetschlii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="buetschlii">Wallaconchis buetschlii</taxonomicName>
|
||
(
|
||
<normalizedToken id="C4AD3A0DB92AFAB2F3A53D2393A2B36D" originalValue="A–D">A-D</normalizedToken>
|
||
) Indonesia, Sulawesi [2120] (UMIZ 00021) (E) Australia, Queensland [2554] (MTQ st100) (F) Australia, Queensland [2555] (MTQ st100). A Rachidian and innermost lateral teeth, scale bar 10
|
||
<normalizedToken id="AF208FB4531AD9DF67AF177FDBE1E134" originalValue="μm">μm</normalizedToken>
|
||
B Rachidian and innermost lateral teeth, scale bar 40
|
||
<normalizedToken id="9F3FC1CA1B83A40C03145D18F600060D" originalValue="μm">μm</normalizedToken>
|
||
C Inner lateral teeth, scale bar 30
|
||
<normalizedToken id="2BA4D97495B2EB1DD0861C824E463069" originalValue="μm">μm</normalizedToken>
|
||
D Outermost lateral teeth, scale bar 20
|
||
<normalizedToken id="04F7667F2D65E7B75FC7DDEC45FAF1AF" originalValue="μm">μm</normalizedToken>
|
||
E Outermost lateral teeth, scale bar 10
|
||
<normalizedToken id="C7983794BD28380BFAC59912A25B34EA" originalValue="μm">μm</normalizedToken>
|
||
F Transition between inner lateral teeth and outer lateral teeth, with the basal lateral spine revealed on the underside of the inner lateral teeth at the bottom, scale bar 30
|
||
<normalizedToken id="D06EC42BA61A28A69E5D5176D56D5BFA" originalValue="μm">μm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="27499AEA4026603837B63FE8F5852AB5" pageId="0" pageNumber="1" type="reproductive system">
|
||
<paragraph id="BC1F4187A510427B9581E07FA348D7B3" pageId="0" pageNumber="1">Reproductive system</paragraph>
|
||
<paragraph id="D0132F3C6AFB73077F563E284B2FECB4" pageId="0" pageNumber="1">(Fig. 22A). The distal region of the oviduct forms a loop, approximately 1/2 to 3/4 of a circle, attached by fibers of tissue to the inner wall of the visceral cavity (Fig. 22A). The oviduct is narrow and approximately of the same width as the deferent duct. The spermatheca is apple-shaped and joins the middle part of the oviduct through a short duct.</paragraph>
|
||
<caption id="7FD0FADEE4997A8D16AB296E70EE5B28" pageId="0" pageNumber="1">
|
||
<paragraph id="6DC3EEECF8B7789878DE3CE5CE907977" pageId="0" pageNumber="1">
|
||
Figure 22. Reproductive system,
|
||
<taxonomicName id="E7140D98E0E71B2761A5C8EBB5B05C7D" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis buetschlii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="buetschlii">Wallaconchis buetschlii</taxonomicName>
|
||
. A Hermaphroditic (female), posterior parts, Indonesia, Sulawesi, scale bar 2 mm [2120] (UMIZ 00021) B Anterior, male copulatory parts, Australia, Queensland, scale bar 1 mm [2555] (MTQ st100) C Anterior, male copulatory parts, Indonesia, Kei, scale bar 1 mm [2933] (UMIZ 00029). Abbreviations: dd deferent duct fgm female gland mass hd hermaphroditic duct hg hermaphroditic gland of oviduct fibers attaching to the inner wall of the visceral cavity ov oviduct ps penial sheath rm retractor muscle rs receptaculum seminis sp spermatheca.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="746640392E6A2309109EC9717CDB6113" pageId="0" pageNumber="1" type="copulatory apparatus">
|
||
<paragraph id="6FC2844CF9B0AC02848F45F703078DC2" pageId="0" pageNumber="1">Copulatory apparatus</paragraph>
|
||
<paragraph id="D396D93AD58C1FC194A6163FDC518494" pageId="0" pageNumber="1">(Figs 22B, C, 23). The anterior, male copulatory apparatus exclusively consists of the deferent duct, the penis, and the retractor muscle of the penis. The penis, distal to the deferent duct, is a long and smooth tube, which can evaginate like a glove. It is not protected by a distinct penial sheath and, strictly speaking, there is no vestibule either. The penis is narrow proximally, widens gradually, with three longitudinal, internal ridges distally (one ridge shown in Fig. 23). In small individuals, the penis is narrow for its entire length. The relative length of the penis compared to that of the retractor muscle varies greatly, from both being of similar length to the penis being five times longer than the retractor muscle. The retractor muscle inserts near the rectum at the posterior end of the visceral cavity. In mature specimens, the deferent duct is thicker than the penis and convoluted distally (Fig. 22C). In immature individuals, the deferent duct is thinner, and less convoluted (Fig. 22B).</paragraph>
|
||
<caption id="ABFF3C6EC44BD9AC978CB96AB1D02B50" pageId="0" pageNumber="1">
|
||
<paragraph id="F2AB1883A0AE494D41382E31A9D9F70F" pageId="0" pageNumber="1">
|
||
Figure 23. Penis,
|
||
<taxonomicName id="48F4B368A3B2D6125E746CD2545E95DF" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis buetschlii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="buetschlii">Wallaconchis buetschlii</taxonomicName>
|
||
, Indonesia. A Longitudinal ridge on the internal surface of the penis, Kei Islands, scale bar 60
|
||
<normalizedToken id="25DF414B9A0E1046E7BE71DE61B14C4B" originalValue="μm">μm</normalizedToken>
|
||
[2933] (UMIZ 00029) B Close-up of a groove between two longitudinal ridges showing smooth internal surface (no hooks), Sulawesi, scale bar 20
|
||
<normalizedToken id="C5E69A05791687AD67784013D9C1540B" originalValue="μm">μm</normalizedToken>
|
||
[2120] (UMIZ 00021).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="BE6624A20CF31A9100D17BBE6309BFA9" pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph id="D7E2E66B164AF35AC97EF727EDDE2377" pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph id="E88AE9019551F4663DAA74E2B0502F59" pageId="0" pageNumber="1">
|
||
<normalizedToken id="EA4EC4B7E37186830B2801E53CCE10EC" originalValue="Stantschinsky’s">Stantschinsky's</normalizedToken>
|
||
<taxonomicName id="39F2B808E634C9B373B71AC7A79F93D0" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium buetschlii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="buetschlii">Onchidium buetschlii</taxonomicName>
|
||
refers to a
|
||
<taxonomicName id="7CA6EF2C27949CDDD7D04F14EF3FCF08" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
species because of the unique combination of characters of the lectotype (intestine of type I, no penial accessory gland, no rectal gland).
|
||
<normalizedToken id="14142E1460D8AE2E8EC7AC76B106C163" originalValue="Stantschinsky’s">Stantschinsky's</normalizedToken>
|
||
original description of the penis (confirmed by our examination of the lectotype) matches exactly the penial anatomy of the species described here. The unique attachment of the oviduct to the posterior body wall was not described by Stantschinsky but is clearly present in the lectotype as well. Because the male parts of the two paralectotypes of
|
||
<taxonomicName id="5EF5891488CB7B6B6A9B78CB1DA2F8A3" lsidName="W. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">W. buetschlii</taxonomicName>
|
||
are missing, it cannot be confirmed that they are part of
|
||
<taxonomicName id="919B401CB47D37568EF01FB3FB3D589F" lsidName="W. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">W. buetschlii</taxonomicName>
|
||
. The anatomy of the posterior (female) reproductive parts of the paralectotypes matches that of the lectotype (with an oviduct attached to posterior body wall), indicating that they could be part of either
|
||
<taxonomicName id="35A77F3EB36128B8629BD5DF7426F5EC" lsidName="W. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">W. buetschlii</taxonomicName>
|
||
or
|
||
<taxonomicName id="085A2D7823105953E70DC05E3B53E8A8" lsidName="W. gracile" pageId="0" pageNumber="1" rank="species" species="gracile">W. gracile</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9CA7C50E72D1F82E37869F53223C3EB6" pageId="0" pageNumber="1">
|
||
The two syntypes used by Stantschinsky to describe
|
||
<taxonomicName id="18CA4E08A3E35CD1AE73C84ABA29E7FF" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium gracile" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gracile">Onchidium gracile</taxonomicName>
|
||
are part of two distinct species; therefore, one specimen is designated as the lectotype (see
|
||
<taxonomicName id="B7E4825B9D52AD569C27061E59920AC5" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis gracile" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gracile">Wallaconchis gracile</taxonomicName>
|
||
). The paralectotype of
|
||
<taxonomicName id="7A4B13000E89B71BE88D0EF63198464E" lsidName="O. gracile" pageId="0" pageNumber="1" rank="species" species="gracile">O. gracile</taxonomicName>
|
||
is part of
|
||
<taxonomicName id="C8F313CD47507C0FA0CA8F2F74DBE9AC" lsidName="W. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">W. buetschlii</taxonomicName>
|
||
. Both species share the unique attachment of the oviduct to the posterior body wall and are anatomically extremely similar. However, the penis (of the lectotype) of
|
||
<taxonomicName id="D7FC734F2E1A0DB1CC3393A8A90E4796" lsidName="W. gracile" pageId="0" pageNumber="1" rank="species" species="gracile">W. gracile</taxonomicName>
|
||
is within a penial sheath, while the penis (of the lectotype) of
|
||
<taxonomicName id="6B2E6138430EB52F3301F453D9863F0A" lsidName="W. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">W. buetschlii</taxonomicName>
|
||
lacks a penial sheath. The penis of the paralectotype of
|
||
<taxonomicName id="C2F47AE24FC19FEA5A04A560CB11F0DF" lsidName="W. gracile" pageId="0" pageNumber="1" rank="species" species="gracile">W. gracile</taxonomicName>
|
||
matches perfectly that of (the lectotype of)
|
||
<taxonomicName id="ECE1F105AEC373614E5B82187C1A652F" lsidName="W. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">W. buetschlii</taxonomicName>
|
||
. The written description of
|
||
<taxonomicName id="F2BA198255E26B7E1C34644F7C0023B0" lsidName="O. gracile" pageId="0" pageNumber="1" rank="species" species="gracile">O. gracile</taxonomicName>
|
||
(
|
||
<bibRefCitation id="A2FE20630E3DE64B0B8D9B87E4C201E9" author="Stantschinsky, W" journalOrPublisher="Zoologische Jahrbuecher, Systematik" pageId="0" pageNumber="1" pagination="353 - 402" title="Zur Anatomie und Systematik der Gattung Oncidium." url="https://doi.org/10.5962/bhl.part.11959" volume="25" year="1907">Stantschinsky 1907</bibRefCitation>
|
||
: 380-383) is confusing because it likely is a mix of both
|
||
<taxonomicName id="BF7F4CD62C5D2603CD39C8CCEEB924CE" lsidName="W. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">W. buetschlii</taxonomicName>
|
||
(paralectotype) and
|
||
<taxonomicName id="8185BB4690817E0B531BDC30CD22EEE5" lsidName="W. gracile" pageId="0" pageNumber="1" rank="species" species="gracile">W. gracile</taxonomicName>
|
||
(lectotype). However,
|
||
<bibRefCitation id="F53FACAF97F118C72E30D96955018014" author="Stantschinsky, W" journalOrPublisher="Zoologische Jahrbuecher, Systematik" pageId="0" pageNumber="1" pagination="353 - 402" title="Zur Anatomie und Systematik der Gattung Oncidium." url="https://doi.org/10.5962/bhl.part.11959" volume="25" year="1907">
|
||
<normalizedToken id="5DDFE1BB82C8D9CFD300E882601B73DA" originalValue="Stantschinsky’s">Stantschinsky's</normalizedToken>
|
||
(1907
|
||
</bibRefCitation>
|
||
: pl. 13, fig. 34) illustration of
|
||
<taxonomicName id="DFE5556963D87AABF5375B7452C1DDAA" lsidName="O. gracile" pageId="0" pageNumber="1" rank="species" species="gracile">O. gracile</taxonomicName>
|
||
matches the copulatory apparatus of the paralectotype and is thus an illustration of
|
||
<taxonomicName id="689745DB32ED1D0480704F619E92FAE3" lsidName="W. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">W. buetschlii</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="BC422F5D929584FA50D986CD4AA9308F" pageId="0" pageNumber="1">
|
||
<bibRefCitation id="87DC004FC17AFF38C54108BCF24D7FB5" author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher (Jena)" pageId="0" pageNumber="1" pagination="29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann (1928)</bibRefCitation>
|
||
commented that
|
||
<taxonomicName id="ABFFF7693609060B7B8FD8677E1CCE6E" lsidName="O. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">O. buetschlii</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="9D7928C02C388202C3446344FD2392EA" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium palaense" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="palaense">Onchidium palaense</taxonomicName>
|
||
, but he did not list any new material for either species, and he does not seem to have examined the type material (he does not mention that he examined it).
|
||
<normalizedToken id="422C7D019D870B750E423E182E2BB91B" originalValue="Semper’s">Semper's</normalizedToken>
|
||
description of the position of the male opening in
|
||
<taxonomicName id="080992F1E984F36023CD6CDE0DA99925" lsidName="O. palaense" pageId="0" pageNumber="1" rank="species" species="palaense">O. palaense</taxonomicName>
|
||
as "between the two very small tentacles, almost in the middle between these" (1880: 276) differs from all
|
||
<taxonomicName id="D0E3C7286E9017CD156059966673675C" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
species and indicates that
|
||
<taxonomicName id="5BAED2299440C9D1D55514A383930876" lsidName="O. palaense" pageId="0" pageNumber="1" rank="species" species="palaense">O. palaense</taxonomicName>
|
||
does not belong to
|
||
<taxonomicName id="C81764CD2CBF8CCF0AB0E64FBCFA30F7" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
. The application of the name
|
||
<taxonomicName id="C2E3111131038B7ABA64F8CC80579373" lsidName="O. palaense" pageId="0" pageNumber="1" rank="species" species="palaense">O. palaense</taxonomicName>
|
||
will be discussed in the revision of another onchidiid genus.
|
||
</paragraph>
|
||
<paragraph id="232C162AE975D90B2EEA47CEB384AFE0" pageId="0" pageNumber="1">
|
||
<bibRefCitation id="A466F21081D865F31C37F709C0E5E9D8" author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique de Monaco" pageId="0" pageNumber="1" pagination="173 - 246" title="Les Silicodermes (Labbe) du Museum d'Histoire Naturelle de Paris. Premiere partie: Classification, formes nouvelles ou peu connues." volume="14" year="1934">
|
||
<normalizedToken id="AE6988A9A7C42983634BD6D61EF877E8" originalValue="Labbé">Labbe</normalizedToken>
|
||
(1934)
|
||
</bibRefCitation>
|
||
identified two specimens from New Guinea as
|
||
<taxonomicName id="9E334931A349D4720D738F6A7162376D" class="Gastropoda" family="Onchidiidae" genus="Paraoncidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraoncidium buetschlii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="buetschlii">Paraoncidium buetschlii</taxonomicName>
|
||
, but he also indicated they might be part of
|
||
<taxonomicName id="F0F2EA61768F7F4CF8E96D7F2E876216" lsidName="O. palaense" pageId="0" pageNumber="1" rank="species" species="palaense">O. palaense</taxonomicName>
|
||
, both of which were originally described with no rectal gland. The lack of rectal gland combined with traits described by
|
||
<normalizedToken id="89472ECC6ED1BCCCE835C10EF53E4459" originalValue="Labbé">Labbe</normalizedToken>
|
||
(intestinal loops of type I, no accessory penial gland) indicate that the specimens he examined are part of a
|
||
<taxonomicName id="2CCD865B8D67260BB9B04B17B24063C3" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
species. However,
|
||
<normalizedToken id="15BFAEB32D32A571C211A21907AED065" originalValue="Labbé’s">Labbe's</normalizedToken>
|
||
brief description does not provide enough information to determine whether the specimens he examined are part of
|
||
<taxonomicName id="98CCED7E41DBA742AF95E9DF29448B7E" lsidName="W. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">W. buetschlii</taxonomicName>
|
||
or another
|
||
<taxonomicName id="84BC83FA823BEDA1F02DD1B958B5CA81" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
species without penial hooks. The fact that
|
||
<normalizedToken id="8EA1EC7D8FB8358F854FDD82D1A5B7FA" originalValue="Labbé">Labbe</normalizedToken>
|
||
did not see any dorsal eyes was most likely due to preservation because dorsal eyes are present in all known
|
||
<taxonomicName id="E681F0025043077AC5C698A8A8B6FF22" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Wallaconchis</taxonomicName>
|
||
species. Finally,
|
||
<bibRefCitation id="8BF2B7EBFABE0CF1F25E2BE866721DE6" author="Bretnall, W" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="303 - 328" title="Onchidiidae from Australia and the South-Western Pacific Islands." url="https://doi.org/10.3853/j.0067-1975.12.1919.888" volume="12" year="1919">Bretnall (1919)</bibRefCitation>
|
||
summarized
|
||
<normalizedToken id="0C66D9FEE858B49A822D4F9B055342B4" originalValue="Stantschinsky’s">Stantschinsky's</normalizedToken>
|
||
original description of
|
||
<taxonomicName id="5CC8AE2AF7BB937940A2292169D832CE" lsidName="W. buetschlii" pageId="0" pageNumber="1" rank="species" species="buetschlii">W. buetschlii</taxonomicName>
|
||
but did not examine any new material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |