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<document id="3923A66D3FBDDEE91BFCA6061E234525" ID-CLB-Dataset="21708" ID-DOI="10.11646/zootaxa.4790.2.1" ID-GBIF-Dataset="312733bb-486b-4f4c-b8b3-260aea3285a9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3889396" ID-ZooBank="F10A4BCE-899A-4EED-9211-343BB3E2BEB8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591859331748" checkinUser="plazi" docAuthor="Wood, Thomas James, Cross, Ian &amp; Baldock, David W." docDate="2020" docId="9F7A87C3FFD9FFF10392FF0AF5E6F9E1" docLanguage="en" docName="zootaxa.4790.2.1.pdf" docOrigin="Zootaxa 4790 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stelis (Heterostelis) hispanica Dusmet y Alonso 1921" docType="treatment" docVersion="8" lastPageNumber="215" masterDocId="6343FFBBFFD7FFFF0305FF92F177FFB3" masterDocTitle="Updates to the bee fauna of Portugal with the description of three new Iberian Andrena species (Hymenoptera: Apoidea: Anthophila)" masterLastPageNumber="228" masterPageNumber="201" pageNumber="215" updateTime="1698842308630" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F317315F893F61AF2906343A8FF1043A">Updates to the bee fauna of Portugal with the description of three new Iberian Andrena species (Hymenoptera: Apoidea: Anthophila)</mods:title>
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<mods:namePart id="D95B944B903D4903259DA947E1EF8917">Wood, Thomas James</mods:namePart>
<mods:affiliation id="E6848CF7B876BFC477D5B6BB71E8EE50">Laboratoire de Zoologie, Université de Mons, 7000, Mons, Belgium.</mods:affiliation>
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<mods:namePart id="79196007EFE2432981666404C97B0957">Cross, Ian</mods:namePart>
<mods:affiliation id="F56F36478ADAC8DE097AC876AE69FE0C">16 Briantspuddle, Dorchester, Dorset, DT 2 7 HS, United Kingdom.</mods:affiliation>
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<mods:namePart id="E4A91A66AAD22F8D2C64AECED8E5F9D4">Baldock, David W.</mods:namePart>
<mods:affiliation id="D9FC0103D35AE5D8C5DCEEC77CF9635F">Nightingales, Haslemere Road, Milford, Surrey, GU 8 5 BN, United Kingdom.</mods:affiliation>
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<treatment id="9F7A87C3FFD9FFF10392FF0AF5E6F9E1" ID-DOI="http://doi.org/10.5281/zenodo.5610240" ID-GBIF-Taxon="164501126" ID-Zenodo-Dep="5610240" LSID="urn:lsid:plazi:treatment:9F7A87C3FFD9FFF10392FF0AF5E6F9E1" httpUri="http://treatment.plazi.org/id/9F7A87C3FFD9FFF10392FF0AF5E6F9E1" lastPageNumber="215" pageId="14" pageNumber="215">
<subSubSection id="5FC9655EFFD9FFF10392FF0AF272FF01" box="[151,773,152,179]" pageId="14" pageNumber="215" type="nomenclature">
<paragraph id="176C36D5FFD9FFF10392FF0AF272FF01" blockId="14.[151,1437,152,1619]" box="[151,773,152,179]" pageId="14" pageNumber="215">
<heading id="4C2481B9FFD9FFF10392FF0AF272FF01" bold="true" box="[151,773,152,179]" fontSize="11" level="1" pageId="14" pageNumber="215" reason="1">
<taxonomicName id="D0D34D56FFD9FFF10392FF0AF272FF01" ID-CoL="8SFZ7" authority="Dusmet y Alonso, 1921" authorityName="Dusmet y Alonso" authorityYear="1921" box="[151,773,152,179]" class="Insecta" family="Megachilidae" genus="Stelis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="hispanica" subGenus="Heterostelis">
<emphasis id="25A7EAC7FFD9FFF10392FF0AF272FF01" bold="true" box="[151,773,152,179]" pageId="14" pageNumber="215">
<emphasis id="25A7EAC7FFD9FFF10392FF0AF1A2FF00" bold="true" box="[151,213,152,179]" italics="true" pageId="14" pageNumber="215">Stelis</emphasis>
(
<emphasis id="25A7EAC7FFD9FFF103E3FF0AF018FF00" bold="true" box="[230,367,152,179]" italics="true" pageId="14" pageNumber="215">Heterostelis</emphasis>
)
<emphasis id="25A7EAC7FFD9FFF1027AFF0AF098FF00" bold="true" box="[383,495,152,179]" italics="true" pageId="14" pageNumber="215">hispanica</emphasis>
Dusmet y Alonso, 1921
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5FC9655EFFD9FFF10392FF2EF330FDA9" pageId="14" pageNumber="215" type="distribution">
<paragraph id="176C36D5FFD9FFF10392FF2EF28EFF48" blockId="14.[151,1437,152,1619]" pageId="14" pageNumber="215">
<emphasis id="25A7EAC7FFD9FFF10392FF2EF044FF64" bold="true" box="[151,307,188,215]" pageId="14" pageNumber="215">Distribution:</emphasis>
Known only from the
<typeStatus id="C8688877FFD9FFF10136FF2EF3E2FF65" box="[563,661,188,214]" pageId="14" pageNumber="215" type="holotype">holotype</typeStatus>
female collected from Valadol 14.vi.[18]92, presumably Valladolid in northwestern
<collectingCountry id="6FC47645FFD9FFF10279FF72F0CBFF49" box="[380,444,224,250]" name="Spain" pageId="14" pageNumber="215">Spain</collectingCountry>
(
<bibRefCitation id="73424B24FFD9FFF102CFFF72F242FF48" author="Schwarz, M. &amp; Gusenleitner, F." box="[458,821,224,251]" pageId="14" pageNumber="215" pagination="1311 - 1321" refId="ref22885" refString="Schwarz, M. &amp; Gusenleitner, F. (2010) Beitrag zur Kenntnis der Stelis - Arten Spaniens (Hymenoptera, Apidae, Megachilinae). Linzer biologischer Beitrage, 42, 1311 - 1321." type="journal article" year="2010">Schwarz and Gusenleitner 2010</bibRefCitation>
;
<bibRefCitation id="73424B24FFD9FFF10045FF72F29AFF48" author="Kasparek, M." box="[832,1005,224,251]" pageId="14" pageNumber="215" pagination="1 - 144" refId="ref21187" refString="Kasparek, M. (2015) The cuckoo bees of the genus Stelis Panzer, 1806 in Europe, North Africa and the Middle East. Entomofauna, Supplement 18, 1 - 144." type="journal article" year="2015">Kasparek 2015</bibRefCitation>
).
</paragraph>
<paragraph id="176C36D5FFD9FFF10392FE97F3A6FEF0" blockId="14.[151,1437,152,1619]" pageId="14" pageNumber="215">
<emphasis id="25A7EAC7FFD9FFF10392FE97F064FEAC" bold="true" box="[151,275,261,287]" pageId="14" pageNumber="215">New data:</emphasis>
<materialsCitation id="A7BB3C88FFD9FFF10219FE97F3BAFEF0" ID-GBIF-Occurrence="2632885314" collectingDate="2019-06-20" collectionCode="R, OOLM" collectorName="A. Soares &amp; Santos &amp; Felix &amp; Ramirez. Deposited" country="Portugal" determinerName="Wood" location="Beira Atla" municipality="Sabugal" pageId="14" pageNumber="215" specimenCount="1" specimenCount-male="1">
<collectingCountry id="6FC47645FFD9FFF10219FE97F0C3FEAC" box="[284,436,261,287]" name="Portugal" pageId="14" pageNumber="215">PORTUGAL</collectingCountry>
:
<location id="120C600EFFD9FFF102C5FE96F340FEAD" LSID="urn:lsid:plazi:treatment:9F7A87C3FFD9FFF10392FF0AF5E6F9E1:120C600EFFD9FFF102C5FE96F340FEAD" box="[448,567,260,286]" country="Portugal" municipality="Sabugal" name="Beira Atla" pageId="14" pageNumber="215">Beira Atla</location>
,
<collectingMunicipality id="F708ACAFFFD9FFF10141FE96F3D6FEAD" box="[580,673,260,286]" pageId="14" pageNumber="215">Sabugal</collectingMunicipality>
,
<date id="636D1015FFD9FFF101ABFE96F25FFEAD" box="[686,808,260,286]" pageId="14" pageNumber="215" value="2019-06-20">
<collectingDate id="7329E9FDFFD9FFF101ABFE96F25FFEAD" box="[686,808,260,286]" pageId="14" pageNumber="215" value="2019-06-20">20.vi.2019</collectingDate>
</date>
,
<specimenCount id="01D5FD5CFFD9FFF10030FE97F22DFEAC" box="[821,858,261,287]" pageId="14" pageNumber="215" type="male">1♂</specimenCount>
, det.
<determinerName id="E2B0DA11FFD9FFF1009CFE96F2A9FEAD" box="[921,990,260,286]" pageId="14" pageNumber="215">Wood</determinerName>
, leg.
<collectorName id="BA265303FFD9FFF10719FE97F5FAFEAC" box="[1052,1165,261,287]" pageId="14" pageNumber="215">A. Soares</collectorName>
,
<collectionCode id="71C2AE10FFD9FFF1079FFE97F5C7FEAC" box="[1178,1200,261,287]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="215">R</collectionCode>
.
<collectorName id="BA265303FFD9FFF107B9FE97F47EFEAC" box="[1212,1289,261,287]" pageId="14" pageNumber="215">Santos</collectorName>
,
<collectionCode id="71C2AE10FFD9FFF10612FE97F45AFEAC" box="[1303,1325,261,287]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="215">R</collectionCode>
.
<collectorName id="BA265303FFD9FFF1063CFE96F402FEAD" box="[1337,1397,260,286]" pageId="14" pageNumber="215">Felix</collectorName>
,
<collectionCode id="71C2AE10FFD9FFF10686FE97F4EEFEAC" box="[1411,1433,261,287]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="215">R</collectionCode>
.
<collectorName id="BA265303FFD9FFF103C3FEBAF0D2FEF1" box="[198,421,296,322]" pageId="14" pageNumber="215">Ramirez. Deposited</collectorName>
in the
<collectionCode id="71C2AE10FFD9FFF102F4FEBAF334FEF0" box="[497,579,296,323]" pageId="14" pageNumber="215">OÖLM</collectionCode>
(
<figureCitation id="8FE82A50FFD9FFF10157FEBAF3B3FEF0" box="[594,708,296,323]" captionStart="FIGURES 2330" captionStartId="19.[151,264,1944,1969]" captionTargetBox="[219,1369,181,1918]" captionTargetId="figure@19.[219,1369,181,1918]" captionTargetPageId="19" captionText="FIGURES 2330. Stelis hispanica Dusmet y Alonso, 1921, 23. male dorsal profile, 24. male tergita 46, 25. male mesosoma, 26. male sterna; Stelis annulata (Lepeletier, 1841), 27. male mesosoma, 28. male sterna; Stelis gigantea Friese, 1921, 29. male mesosoma, 30. male sterna." figureDoi="http://doi.org/10.5281/zenodo.3889404" httpUri="https://zenodo.org/record/3889404/files/figure.png" pageId="14" pageNumber="215">Figure 23</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="176C36D5FFD9FFF10392FEDFF330FDA9" blockId="14.[151,1437,152,1619]" pageId="14" pageNumber="215">
<emphasis id="25A7EAC7FFD9FFF10392FEDFF194FED4" bold="true" box="[151,227,333,359]" pageId="14" pageNumber="215">Notes:</emphasis>
<taxonomicName id="D0D34D56FFD9FFF103E8FEDEF0D7FED5" authorityName="Dusmet y Alonso" authorityYear="1921" box="[237,416,332,358]" class="Insecta" family="Megachilidae" genus="Stelis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="hispanica">
<emphasis id="25A7EAC7FFD9FFF103E8FEDEF0D7FED5" box="[237,416,332,358]" italics="true" pageId="14" pageNumber="215">Stelis hispanica</emphasis>
</taxonomicName>
was resurrected from synonymy with
<taxonomicName id="D0D34D56FFD9FFF10058FEDEF5ACFED4" authority="(Lepeletier, 1841)" baseAuthorityName="Lepeletier" baseAuthorityYear="1841" box="[861,1243,332,359]" class="Insecta" family="Megachilidae" genus="Stelis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="annulata">
<emphasis id="25A7EAC7FFD9FFF10058FEDEF570FED5" box="[861,1031,332,358]" italics="true" pageId="14" pageNumber="215">Stelis annulata</emphasis>
(Lepeletier, 1841)
</taxonomicName>
by
<bibRefCitation id="73424B24FFD9FFF1060FFEDEF0D0FE38" author="Schwarz, M. &amp; Gusenleitner, F." pageId="14" pageNumber="215" pagination="1311 - 1321" refId="ref22885" refString="Schwarz, M. &amp; Gusenleitner, F. (2010) Beitrag zur Kenntnis der Stelis - Arten Spaniens (Hymenoptera, Apidae, Megachilinae). Linzer biologischer Beitrage, 42, 1311 - 1321." type="journal article" year="2010">Schwarz and Gusenleitner (2010)</bibRefCitation>
who citied multiple differences. However, as only the
<typeStatus id="C8688877FFD9FFF1070EFEE2F51AFE39" box="[1035,1133,368,394]" pageId="14" pageNumber="215" type="holotype">holotype</typeStatus>
was known it was difficult to judge potential variation within this species, and whether it might fall within the variation seen in
<taxonomicName id="D0D34D56FFD9FFF1063BFE07F185FE61" baseAuthorityName="Lepeletier" baseAuthorityYear="1841" class="Insecta" family="Megachilidae" genus="Stelis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="annulata">
<emphasis id="25A7EAC7FFD9FFF1063BFE07F185FE61" italics="true" pageId="14" pageNumber="215">S. annulata</emphasis>
</taxonomicName>
. Examination of newly caught male material approximately
<quantity id="D02B9B30FFD9FFF100A1FE2AF28DFE61" box="[932,1018,440,467]" metricMagnitude="5" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="215" unit="km" value="250.0">250 km</quantity>
from the
<emphasis id="25A7EAC7FFD9FFF10769FE2AF589FE61" box="[1132,1278,440,466]" italics="true" pageId="14" pageNumber="215">locus typicus</emphasis>
shows that
<taxonomicName id="D0D34D56FFD9FFF10682FE2BF044FE45" authorityName="Dusmet y Alonso" authorityYear="1921" class="Insecta" family="Megachilidae" genus="Stelis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="hispanica">
<emphasis id="25A7EAC7FFD9FFF10682FE2BF044FE45" italics="true" pageId="14" pageNumber="215">S. hispanica</emphasis>
</taxonomicName>
is clearly distinct from
<taxonomicName id="D0D34D56FFD9FFF10147FE4FF3B4FE45" baseAuthorityName="Lepeletier" baseAuthorityYear="1841" box="[578,707,476,503]" class="Insecta" family="Megachilidae" genus="Stelis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="annulata">
<emphasis id="25A7EAC7FFD9FFF10147FE4FF3B4FE45" box="[578,707,476,503]" italics="true" pageId="14" pageNumber="215">S. annulata</emphasis>
</taxonomicName>
, supporting the position of Schwarz and Gusenleitner. The male is described here for the first time.
</paragraph>
</subSubSection>
<subSubSection id="5FC9655EFFD9FFF10392FDB6F5E6F9E1" pageId="14" pageNumber="215" type="description">
<paragraph id="176C36D5FFD9FFF10392FDB6F033FA80" blockId="14.[151,1437,152,1619]" pageId="14" pageNumber="215">
<emphasis id="25A7EAC7FFD9FFF10392FDB6F05CFD8C" bold="true" box="[151,299,548,575]" pageId="14" pageNumber="215">Description:</emphasis>
Male: Body length
<quantity id="D02B9B30FFD9FFF1010FFDB7F320FD8C" box="[522,599,549,575]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="14" pageNumber="215" unit="mm" value="11.0">11 mm</quantity>
(
<figureCitation id="8FE82A50FFD9FFF10160FDB6F3A0FD8C" box="[613,727,548,575]" captionStart="FIGURES 2330" captionStartId="19.[151,264,1944,1969]" captionTargetBox="[219,1369,181,1918]" captionTargetId="figure@19.[219,1369,181,1918]" captionTargetPageId="19" captionText="FIGURES 2330. Stelis hispanica Dusmet y Alonso, 1921, 23. male dorsal profile, 24. male tergita 46, 25. male mesosoma, 26. male sterna; Stelis annulata (Lepeletier, 1841), 27. male mesosoma, 28. male sterna; Stelis gigantea Friese, 1921, 29. male mesosoma, 30. male sterna." figureDoi="http://doi.org/10.5281/zenodo.3889404" httpUri="https://zenodo.org/record/3889404/files/figure.png" pageId="14" pageNumber="215">Figure 23</figureCitation>
).
<emphasis id="25A7EAC7FFD9FFF101EFFDB6F258FD8D" box="[746,815,548,574]" italics="true" pageId="14" pageNumber="215">Head:</emphasis>
Head as broad as long, round in frontal view. Gena and vertex enlarged, gena near occiput 1.2 times wider than maximal width of compound eye. Vertex greatly enlarged, longer than ocellocular distance. Paraocular area with thin yellow mark that widens slightly near lateral margin of clypeus; greatest width less than half antennocular distance. Clypeus centrally with a transverse yellow marking that does not reach paraocular markings, the three yellow marks on the face therefore separated. Face, gena, and vertex with fine whitish to brownish pubescence, longest hairs equalling the length of scape. Facial integument clearly and consistently punctured, punctures almost touching, interspaces near lateral ocelli separated by 0.5 puncture diameters. Malar space linear. Mandibles black, bidentate.
<taxonomicName id="D0D34D56FFD9FFF1079DFCB3F4EBFC88" authority=": Mesosoma" authorityName="Mesosoma" box="[1176,1436,801,827]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="25A7EAC7FFD9FFF1079DFCB3F46EFC89" box="[1176,1305,801,826]" italics="true" pageId="14" pageNumber="215">Mesosoma:</emphasis>
Mesosoma
</taxonomicName>
entirely black (
<figureCitation id="8FE82A50FFD9FFF10268FCD6F0A9FCEC" box="[365,478,836,863]" captionStart="FIGURES 2330" captionStartId="19.[151,264,1944,1969]" captionTargetBox="[219,1369,181,1918]" captionTargetId="figure@19.[219,1369,181,1918]" captionTargetPageId="19" captionText="FIGURES 2330. Stelis hispanica Dusmet y Alonso, 1921, 23. male dorsal profile, 24. male tergita 46, 25. male mesosoma, 26. male sterna; Stelis annulata (Lepeletier, 1841), 27. male mesosoma, 28. male sterna; Stelis gigantea Friese, 1921, 29. male mesosoma, 30. male sterna." figureDoi="http://doi.org/10.5281/zenodo.3889404" httpUri="https://zenodo.org/record/3889404/files/figure.png" pageId="14" pageNumber="215">Figure 25</figureCitation>
). Axillae rounded, slightly protruding, outer margin not smoothly contiguous with the scutellum.
<taxonomicName id="D0D34D56FFD9FFF10246FCFBF0CBFC30" box="[323,444,873,899]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="215" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
with moderately long brownish pubescence, shorter than that of face, individual hairs less than length of scape. Scutum, scutellum, axillae, episternum, and propodeum clearly and densely punctate, punctures almost touching, interspaces shiny, except propodeal declivity with central impunctate section. Femora black with apex orange, tibiae and tarsi orange with yellow highlights at base and apex of tibiae. Tarsal claws dark brown, apically bifid. Wings generally dark, infumate, with black venation.
<emphasis id="25A7EAC7FFD9FFF10795FC6BF47AFBA1" box="[1168,1293,1017,1042]" italics="true" pageId="14" pageNumber="215">Metasoma:</emphasis>
Terga black, T13 with pairs of yellow markings clearly interrupted medially (
<figureCitation id="8FE82A50FFD9FFF100B7FB8EF553FB84" box="[946,1060,1052,1079]" captionStart="FIGURES 2330" captionStartId="19.[151,264,1944,1969]" captionTargetBox="[219,1369,181,1918]" captionTargetId="figure@19.[219,1369,181,1918]" captionTargetPageId="19" captionText="FIGURES 2330. Stelis hispanica Dusmet y Alonso, 1921, 23. male dorsal profile, 24. male tergita 46, 25. male mesosoma, 26. male sterna; Stelis annulata (Lepeletier, 1841), 27. male mesosoma, 28. male sterna; Stelis gigantea Friese, 1921, 29. male mesosoma, 30. male sterna." figureDoi="http://doi.org/10.5281/zenodo.3889404" httpUri="https://zenodo.org/record/3889404/files/figure.png" pageId="14" pageNumber="215">Figure 23</figureCitation>
), T4 with markings touching, T5 with markings contiguous. T6 with a central yellow marking (
<figureCitation id="8FE82A50FFD9FFF10080FBD2F28FFBE8" box="[901,1016,1088,1115]" captionStart="FIGURES 2330" captionStartId="19.[151,264,1944,1969]" captionTargetBox="[219,1369,181,1918]" captionTargetId="figure@19.[219,1369,181,1918]" captionTargetPageId="19" captionText="FIGURES 2330. Stelis hispanica Dusmet y Alonso, 1921, 23. male dorsal profile, 24. male tergita 46, 25. male mesosoma, 26. male sterna; Stelis annulata (Lepeletier, 1841), 27. male mesosoma, 28. male sterna; Stelis gigantea Friese, 1921, 29. male mesosoma, 30. male sterna." figureDoi="http://doi.org/10.5281/zenodo.3889404" httpUri="https://zenodo.org/record/3889404/files/figure.png" pageId="14" pageNumber="215">Figure 24</figureCitation>
). T7 weakly tripartite with two weak lateral lobes and a central projection with a longitudinal carina. All terga densely punctured, punctures on tergal margins almost touching, those on disc separated by 0.751 puncture diameters. Sterna black, S3 and S4 with long fringes of orange setae (
<figureCitation id="8FE82A50FFD9FFF10100FB3EF303FB74" box="[517,628,1196,1223]" captionStart="FIGURES 2330" captionStartId="19.[151,264,1944,1969]" captionTargetBox="[219,1369,181,1918]" captionTargetId="figure@19.[219,1369,181,1918]" captionTargetPageId="19" captionText="FIGURES 2330. Stelis hispanica Dusmet y Alonso, 1921, 23. male dorsal profile, 24. male tergita 46, 25. male mesosoma, 26. male sterna; Stelis annulata (Lepeletier, 1841), 27. male mesosoma, 28. male sterna; Stelis gigantea Friese, 1921, 29. male mesosoma, 30. male sterna." figureDoi="http://doi.org/10.5281/zenodo.3889404" httpUri="https://zenodo.org/record/3889404/files/figure.png" pageId="14" pageNumber="215">Figure 26</figureCitation>
), S4 medially and apically with black comb made up of short, thick hairs. Comb 3 times wider than long, occupying approximately a quarter of the sternal margin. S5 deeply emarginate, forming a V shape. Terga and sterna with general brownish pubescence, at most equalling length of hairs on the
<taxonomicName id="D0D34D56FFD9FFF103C3FA8BF037FA80" box="[198,320,1305,1331]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="215" phylum="Arthropoda" rank="genus">mesosoma</taxonomicName>
.
</paragraph>
<paragraph id="176C36D5FFD9FFF10392FAAFF5E6F9E1" blockId="14.[151,1437,152,1619]" pageId="14" pageNumber="215">
<emphasis id="25A7EAC7FFD9FFF10392FAAFF194FAE4" bold="true" box="[151,227,1341,1367]" pageId="14" pageNumber="215">Notes:</emphasis>
Using the key of
<bibRefCitation id="73424B24FFD9FFF102B5FAAEF318FAE4" author="Kasparek, M." box="[432,623,1340,1367]" pageId="14" pageNumber="215" pagination="1 - 144" refId="ref21187" refString="Kasparek, M. (2015) The cuckoo bees of the genus Stelis Panzer, 1806 in Europe, North Africa and the Middle East. Entomofauna, Supplement 18, 1 - 144." type="journal article" year="2015">Kasparek (2015)</bibRefCitation>
, the male of
<taxonomicName id="D0D34D56FFD9FFF10006FAAFF2FAFAE5" authorityName="Dusmet y Alonso" authorityYear="1921" box="[771,909,1340,1367]" class="Insecta" family="Megachilidae" genus="Stelis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="hispanica">
<emphasis id="25A7EAC7FFD9FFF10006FAAFF2FAFAE5" box="[771,909,1340,1367]" italics="true" pageId="14" pageNumber="215">S. hispanica</emphasis>
</taxonomicName>
keys to couplet 21 due to its yellow tergal colouration, its lack of a claw-like inner spine on the fore tibia, its large size, and its deeply emarginate S5. Here it can be separated from
<taxonomicName id="D0D34D56FFD9FFF102BAFA17F349FA2D" baseAuthorityName="Lepeletier" baseAuthorityYear="1841" box="[447,574,1412,1439]" class="Insecta" family="Megachilidae" genus="Stelis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="annulata">
<emphasis id="25A7EAC7FFD9FFF102BAFA17F349FA2D" box="[447,574,1412,1439]" italics="true" pageId="14" pageNumber="215">S. annulata</emphasis>
</taxonomicName>
as the latter has the axillae and scutellum marked with yellow (
<figureCitation id="8FE82A50FFD9FFF107FDFA16F411FA2C" box="[1272,1382,1412,1439]" captionStart="FIGURES 2330" captionStartId="19.[151,264,1944,1969]" captionTargetBox="[219,1369,181,1918]" captionTargetId="figure@19.[219,1369,181,1918]" captionTargetPageId="19" captionText="FIGURES 2330. Stelis hispanica Dusmet y Alonso, 1921, 23. male dorsal profile, 24. male tergita 46, 25. male mesosoma, 26. male sterna; Stelis annulata (Lepeletier, 1841), 27. male mesosoma, 28. male sterna; Stelis gigantea Friese, 1921, 29. male mesosoma, 30. male sterna." figureDoi="http://doi.org/10.5281/zenodo.3889404" httpUri="https://zenodo.org/record/3889404/files/figure.png" pageId="14" pageNumber="215">Figure 27</figureCitation>
) and the dark comb of S4 is narrow, longer than wide (
<figureCitation id="8FE82A50FFD9FFF101F4FA3AF214FA70" box="[753,867,1448,1475]" captionStart="FIGURES 2330" captionStartId="19.[151,264,1944,1969]" captionTargetBox="[219,1369,181,1918]" captionTargetId="figure@19.[219,1369,181,1918]" captionTargetPageId="19" captionText="FIGURES 2330. Stelis hispanica Dusmet y Alonso, 1921, 23. male dorsal profile, 24. male tergita 46, 25. male mesosoma, 26. male sterna; Stelis annulata (Lepeletier, 1841), 27. male mesosoma, 28. male sterna; Stelis gigantea Friese, 1921, 29. male mesosoma, 30. male sterna." figureDoi="http://doi.org/10.5281/zenodo.3889404" httpUri="https://zenodo.org/record/3889404/files/figure.png" pageId="14" pageNumber="215">Figure 28</figureCitation>
). It can be separated from
<taxonomicName id="D0D34D56FFD9FFF10789FA3BF4EBFA70" authority="Friese, 1921" authorityName="Friese" authorityYear="1921" box="[1164,1436,1448,1475]" class="Insecta" family="Megachilidae" genus="Stelis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis id="25A7EAC7FFD9FFF10789FA3BF47CFA71" box="[1164,1291,1449,1475]" italics="true" pageId="14" pageNumber="215">S. gigantea</emphasis>
Friese, 1921
</taxonomicName>
because the axillae are not produced into blunt teeth (
<figureCitation id="8FE82A50FFD9FFF1002EFA5EF2ECFA54" box="[811,923,1484,1511]" captionStart="FIGURES 2330" captionStartId="19.[151,264,1944,1969]" captionTargetBox="[219,1369,181,1918]" captionTargetId="figure@19.[219,1369,181,1918]" captionTargetPageId="19" captionText="FIGURES 2330. Stelis hispanica Dusmet y Alonso, 1921, 23. male dorsal profile, 24. male tergita 46, 25. male mesosoma, 26. male sterna; Stelis annulata (Lepeletier, 1841), 27. male mesosoma, 28. male sterna; Stelis gigantea Friese, 1921, 29. male mesosoma, 30. male sterna." figureDoi="http://doi.org/10.5281/zenodo.3889404" httpUri="https://zenodo.org/record/3889404/files/figure.png" pageId="14" pageNumber="215">Figure 29</figureCitation>
) and whilst the dark comb of S4 is similarly wide, the hind margin of S2 is strongly emarginate in
<taxonomicName id="D0D34D56FFD9FFF10024FA63F2E8F9B9" authorityName="Friese" authorityYear="1921" box="[801,927,1521,1547]" class="Insecta" family="Megachilidae" genus="Stelis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis id="25A7EAC7FFD9FFF10024FA63F2E8F9B9" box="[801,927,1521,1547]" italics="true" pageId="14" pageNumber="215">S. gigantea</emphasis>
</taxonomicName>
(
<figureCitation id="8FE82A50FFD9FFF100ABFA62F56AF9B8" box="[942,1053,1520,1547]" captionStart="FIGURES 2330" captionStartId="19.[151,264,1944,1969]" captionTargetBox="[219,1369,181,1918]" captionTargetId="figure@19.[219,1369,181,1918]" captionTargetPageId="19" captionText="FIGURES 2330. Stelis hispanica Dusmet y Alonso, 1921, 23. male dorsal profile, 24. male tergita 46, 25. male mesosoma, 26. male sterna; Stelis annulata (Lepeletier, 1841), 27. male mesosoma, 28. male sterna; Stelis gigantea Friese, 1921, 29. male mesosoma, 30. male sterna." figureDoi="http://doi.org/10.5281/zenodo.3889404" httpUri="https://zenodo.org/record/3889404/files/figure.png" pageId="14" pageNumber="215">Figure 30</figureCitation>
) whereas the hind margin of S2 is only weakly emarginate in
<taxonomicName id="D0D34D56FFD9FFF102F0F987F309F99D" authorityName="Dusmet y Alonso" authorityYear="1921" box="[501,638,1556,1583]" class="Insecta" family="Megachilidae" genus="Stelis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="hispanica">
<emphasis id="25A7EAC7FFD9FFF102F0F987F309F99D" box="[501,638,1556,1583]" italics="true" pageId="14" pageNumber="215">S. hispanica</emphasis>
</taxonomicName>
(
<figureCitation id="8FE82A50FFD9FFF10188F986F389F99C" box="[653,766,1556,1583]" captionStart="FIGURES 2330" captionStartId="19.[151,264,1944,1969]" captionTargetBox="[219,1369,181,1918]" captionTargetId="figure@19.[219,1369,181,1918]" captionTargetPageId="19" captionText="FIGURES 2330. Stelis hispanica Dusmet y Alonso, 1921, 23. male dorsal profile, 24. male tergita 46, 25. male mesosoma, 26. male sterna; Stelis annulata (Lepeletier, 1841), 27. male mesosoma, 28. male sterna; Stelis gigantea Friese, 1921, 29. male mesosoma, 30. male sterna." figureDoi="http://doi.org/10.5281/zenodo.3889404" httpUri="https://zenodo.org/record/3889404/files/figure.png" pageId="14" pageNumber="215">Figure 26</figureCitation>
). Nothing is known about its host, but given its similarity to
<taxonomicName id="D0D34D56FFD9FFF103C3F9ABF031F9E1" baseAuthorityName="Lepeletier" baseAuthorityYear="1841" box="[198,326,1592,1619]" class="Insecta" family="Megachilidae" genus="Stelis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="annulata">
<emphasis id="25A7EAC7FFD9FFF103C3F9ABF031F9E1" box="[198,326,1592,1619]" italics="true" pageId="14" pageNumber="215">S. annulata</emphasis>
</taxonomicName>
and
<taxonomicName id="D0D34D56FFD9FFF10279F9ABF08DF9E1" authorityName="Friese" authorityYear="1921" box="[380,506,1593,1619]" class="Insecta" family="Megachilidae" genus="Stelis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis id="25A7EAC7FFD9FFF10279F9ABF08DF9E1" box="[380,506,1593,1619]" italics="true" pageId="14" pageNumber="215">S. gigantea</emphasis>
</taxonomicName>
, presumably a species of
<taxonomicName id="D0D34D56FFD9FFF1001EF9AAF541F9E0" box="[795,1078,1592,1619]" class="Insecta" family="Megachilidae" genus="Trachusa" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="genus">
<emphasis id="25A7EAC7FFD9FFF1001EF9AAF2F4F9E1" box="[795,899,1592,1618]" italics="true" pageId="14" pageNumber="215">Trachusa</emphasis>
(Megachilidae)
</taxonomicName>
is used.
</paragraph>
</subSubSection>
</treatment>
</document>