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<document ID-DOI="http://dx.doi.org/10.3897/jhr.33.5078" ID-GBIF-Taxon="182249306" ID-Pensoft-Pub="1314-2607-33-1" ID-Pensoft-UUID="5F20FFE2E95AAC21FFD1FFA0D8160112" ID-Zenodo-Dep="574825" ID-ZooBank="88E03F8FD9B947F1B6E90456D47B1E37" ModsDocID="1314-2607-33-1" checkinTime="1621553083643" checkinUser="pensoft" docAuthor="Kimsey, Lynn S. &amp; Wasbauer, Marius S." docDate="2013" docId="7AFA489E9D0E7126D0A77B560C7E30B0" docLanguage="en" docOrigin="Journal of Hymenoptera Research 33" docPubDate="2013-08-01" docSource="http://dx.doi.org/10.3897/jhr.33.5078" docTitle="Colocistis Krombein 1942" docType="treatment" docVersion="2" id="5F20FFE2E95AAC21FFD1FFA0D8160112" lastPageId="4" lastPageNumber="3" masterDocId="5F20FFE2E95AAC21FFD1FFA0D8160112" masterDocTitle="Revision of the brachycistidine genus Colocistis Krombein, 1942 (Hymenoptera, Tiphiidae)" masterLastPageNumber="24" masterPageNumber="1" pageId="1" pageNumber="2" updateTime="1643536911076" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the brachycistidine genus Colocistis Krombein, 1942 (Hymenoptera, Tiphiidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kimsey, Lynn S.</mods:namePart>
<mods:affiliation>Bohart Museum of Entomology, Department of Entomology, University of California, Davis, 95616 USA</mods:affiliation>
<mods:nameIdentifier type="email">lskimsey@ucdavis.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wasbauer, Marius S.</mods:namePart>
<mods:affiliation>Bohart Museum of Entomology, Department of Entomology, University of California, Davis, 95616 USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-08-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>33</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>24</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/jhr.33.5078</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.33.5078</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2607-33-1</mods:identifier>
<mods:identifier type="ZooBank">88E03F8FD9B947F1B6E90456D47B1E37</mods:identifier>
<mods:identifier type="Pensoft-UUID">5F20FFE2E95AAC21FFD1FFA0D8160112</mods:identifier>
<mods:identifier type="Zenodo-Dep">574825</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182249306" LSID="urn:lsid:plazi:treatment:7AFA489E9D0E7126D0A77B560C7E30B0" httpUri="http://treatment.plazi.org/id/7AFA489E9D0E7126D0A77B560C7E30B0" lastPageId="4" lastPageNumber="3" pageId="1" pageNumber="2">
<subSubSection pageId="1" pageNumber="2" type="nomenclature">
<paragraph pageId="1" pageNumber="2">
<taxonomicName LSID="http://species-id.net/wiki/Colocistis" authority="Krombein" authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Colocistis" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Colocistis Krombein</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="2" type="reference_group">
<paragraph pageId="1" pageNumber="2">
<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Colocistis</taxonomicName>
Krombein, 1942 (April):65. Type species:
<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis italics="true" pageId="1" pageNumber="2">Colocistis pilosa</emphasis>
</taxonomicName>
<bibRefCitation author="Krombein, KV" journalOrPublisher="Proceedings of theEntomological Society of Washington" pageId="18" pageNumber="19" pagination="67 - 68" publicationUrl="10.2307/2420897" refId="B7" refString="Krombein, KV, 1942. A new genus and species of Brachycistidinae. Proceedings of theEntomological Society of Washington 44: 67 - 68, 10.2307/2420897" title="A new genus and species of Brachycistidinae." url="10.2307/2420897" volume="44" year="1942">Krombein 1942</bibRefCitation>
:66. Original designation.
</paragraph>
<paragraph pageId="1" pageNumber="2">
<taxonomicName authorityName="Mickel &amp; Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Aglyptacros" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Aglyptacros</taxonomicName>
Mickel &amp; Krombein, 1942 (November):669. Type species:
<taxonomicName genus="Glyptometopa" lsidName="" pageId="1" pageNumber="2" rank="species" species="eureka">
<emphasis italics="true" pageId="1" pageNumber="2">Glyptometopa eureka</emphasis>
</taxonomicName>
<bibRefCitation author="Banks, N" journalOrPublisher="The Canadian Entomologist" pageId="18" pageNumber="19" pagination="197 - 203" publicationUrl="10.4039/Ent44197-7" refId="B1" refString="Banks, N, 1912. New Scolioidea. The Canadian Entomologist 44: 197 - 203, 10.4039/Ent44197-7" title="New Scolioidea." url="10.4039/Ent44197-7" volume="44" year="1912">Banks 1912</bibRefCitation>
. Original designation. Synonymized by
<bibRefCitation author="Kimsey, LS" journalOrPublisher="Zootaxa" pageId="18" pageNumber="19" pagination="21 - 34" refId="B5" refString="Kimsey, LS, 2006. Competing taxonomies: reexamination of the female-based genera of Brachycistidinae (Hymenoptera: Tiphiidae). Zootaxa 1121: 21 - 34" title="Competing taxonomies: reexamination of the female-based genera of Brachycistidinae (Hymenoptera: Tiphiidae)." volume="1121" year="2006">Kimsey 2006</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="diagnosis">
<paragraph pageId="1" pageNumber="2">Diagnosis.</paragraph>
<paragraph lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
The genus
<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="3">
<pageBreakToken pageId="2" pageNumber="3" start="start">Colocistis</pageBreakToken>
</emphasis>
</taxonomicName>
exhibits a number of unique traits in both sexes. Features in the female, include the ventrally angulate occipital carina, the ventrally angulate mid- and hindtrochanters, hindcoxal carina on both the inner, ventral margin and dorsobasal surface. Because a number of these features, including the angulate occipital carina and hindcoxal carina are shared with males of
<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="3">Colocistis</emphasis>
</taxonomicName>
and the distributions of the two genera coincide
<taxonomicName authorityName="Mickel &amp; Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Aglyptacros" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="3">Aglyptacros</emphasis>
</taxonomicName>
was made the junior synonym of
<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="3">Colocistis</emphasis>
</taxonomicName>
(
<bibRefCitation author="Kimsey, LS" journalOrPublisher="Zootaxa" pageId="18" pageNumber="19" pagination="21 - 34" refId="B5" refString="Kimsey, LS, 2006. Competing taxonomies: reexamination of the female-based genera of Brachycistidinae (Hymenoptera: Tiphiidae). Zootaxa 1121: 21 - 34" title="Competing taxonomies: reexamination of the female-based genera of Brachycistidinae (Hymenoptera: Tiphiidae)." volume="1121" year="2006">Kimsey 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="description">
<paragraph pageId="2" pageNumber="3">Description.</paragraph>
<paragraph pageId="2" pageNumber="3">
<emphasis bold="true" pageId="2" pageNumber="3">Male</emphasis>
Body length 5-16 mm.
</paragraph>
<paragraph pageId="2" pageNumber="3">
<emphasis italics="true" pageId="2" pageNumber="3">Head</emphasis>
. Compound eyes and ocelli enlarged (except quite small in
<taxonomicName authorityName="Kimsey &amp; Wasbauer" authorityYear="2013" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="chemsaki">
<emphasis italics="true" pageId="2" pageNumber="3">Colocistis chemsaki</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis italics="true" pageId="2" pageNumber="3">Colocistis pilosa</emphasis>
</taxonomicName>
); gular carina either not elevated or elevated gradually at level of posterior mandibular condyle, not suddenly produced apically, clypeal length versus width ratio not exceeding 0.35; maxillary and labial palpi not reduced, maxillary palpi six-segmented, exceeding posterior border of gular orifice by half length of palpus; labial palpi four-segmented.
</paragraph>
<paragraph pageId="2" pageNumber="3">
<emphasis italics="true" pageId="2" pageNumber="3">Mesosoma</emphasis>
. Fore wing with costa extending beyond stigma for a distance not over half length of stigma; forewing with two discoidal cells and two or three submarginal cells, second submarginal cell completely or almost completely underlying first submarginal cell, first transverse cubital vein arising at or before basal third of second submarginal cell (
<figureCitation captionStart="Figures 13–24" captionStartId="F2" captionText="Figures 13 - 24. 13, 14 Male forewing. 15 Ventral view of male hindcoxa 16 Dorsal view of male propodeum 17 dorsal view of male mandible 18 lateral view of mandible 19 Ventral view of male metasomal sterna I-II 20, 22 Lateral view of male metasomal segments I and II 21, 23, 24 Ventral view of female metasomal sternum I. Abbreviations: SM- 1, 2, 3 = submarginal cells 1 - 3." httpUri="https://binary.pensoft.net/fig/11788" pageId="2" pageNumber="3">Fig. 14</figureCitation>
); hind wing with cubitus arcuately curved, longer that first transverse cubital vein and meeting it at an angle of less than 135 degrees; hind wing with jugal lobe shorter than submedial cell; forecoxa with admesal stridulatory area; hindcoxa with longitudinal carina on inner margin.
</paragraph>
<paragraph pageId="2" pageNumber="3">
<emphasis italics="true" pageId="2" pageNumber="3">Metasoma</emphasis>
. Sternum II without basal longitudinal carina (one species with transverse carina); digitus of genitalia without process extending distad of articulum; volsellar plate without spines or teeth, but with flexible hairs on mesal surface.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/11788" pageId="2" pageNumber="3" start="Figures 13–24" startId="F2">
<paragraph pageId="2" pageNumber="3">
<emphasis bold="true" pageId="2" pageNumber="3">Figures 13-24.</emphasis>
<emphasis bold="true" pageId="2" pageNumber="3">13, 14</emphasis>
Male forewing.
<emphasis bold="true" pageId="2" pageNumber="3">15</emphasis>
Ventral view of male hindcoxa
<emphasis bold="true" pageId="2" pageNumber="3">16</emphasis>
Dorsal view of male propodeum
<emphasis bold="true" pageId="2" pageNumber="3">17</emphasis>
dorsal view of male mandible
<emphasis bold="true" pageId="2" pageNumber="3">18</emphasis>
lateral view of mandible
<emphasis bold="true" pageId="2" pageNumber="3">19</emphasis>
Ventral view of male metasomal sterna I-II
<emphasis bold="true" pageId="2" pageNumber="3">20, 22</emphasis>
Lateral view of male metasomal segments I and II
<emphasis bold="true" pageId="2" pageNumber="3">21, 23, 24</emphasis>
Ventral view of female metasomal sternum I. Abbreviations: SM-1, 2, 3 = submarginal cells 1-3.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="description">
<paragraph pageId="2" pageNumber="3">Description.</paragraph>
<paragraph pageId="2" pageNumber="3">
<emphasis bold="true" pageId="2" pageNumber="3">Female</emphasis>
Body length 5-12 mm.
</paragraph>
<paragraph pageId="2" pageNumber="3">
<emphasis italics="true" pageId="2" pageNumber="3">Head</emphasis>
. Mandible broadest subapically, with two distinct apical teeth, inner margin with large, obtuse tooth at broadest point, obtuse tooth obscured by scutate bristles, inner margin below tooth with three small teeth, outer carina obsolescent, ventral carina distinct, sharp; maxillary palp 5-segmented; occipital carina well-developed, angulate ventrolaterally, forming straight line ventrally, often subtended by secondary carina; vertex with submedial impressions, lateral setose groove represented by row of large punctures or foveae.
</paragraph>
<paragraph pageId="2" pageNumber="3">
<emphasis italics="true" pageId="2" pageNumber="3">Mesosoma</emphasis>
. Propleural ventral margin strongly elevated, with sharp, transverse carina; hindcoxa with longitudinal carina on inner margin; hindtibial calcar slightly curved and acute apically.
</paragraph>
<paragraph pageId="2" pageNumber="3">
<emphasis italics="true" pageId="2" pageNumber="3">Metasoma</emphasis>
. Sternum I with transverse subbasal carina well-developed and protruding, subtended by medial pit, and with two foveae basad; tergum I with well-developed lateral ridge.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="distribution">
<paragraph pageId="2" pageNumber="3">Distribution.</paragraph>
<paragraph pageId="2" pageNumber="3">
<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="3">Colocistis</emphasis>
</taxonomicName>
species are found in southeastern California, southern Nevada, southern Arizona, southern Colorado, western Texas and as far south as Oaxaca, Mexico.
</paragraph>
<paragraph pageId="2" pageNumber="3">
<emphasis bold="true" pageId="2" pageNumber="3">
Key to species of
<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="3">Colocistis</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
<table inLine="true" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
<tr pageId="2" pageNumber="3">
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">1</td>
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">Wingless, metasoma 6-segmented; females</td>
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">2</td>
</tr>
<tr pageId="2" pageNumber="3">
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">-</td>
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">Winged, metasoma 7-segmented; males</td>
<td colspan="1" pageId="2" pageNumber="3" rowspan="1">5</td>
</tr>
<tr pageId="3" pageNumber="4">
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
<pageBreakToken pageId="3" pageNumber="4" start="start">2</pageBreakToken>
</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
Gular carina strongly elevated or angulate on either side of midline, meeting occipital carina (as in
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="3" pageNumber="4">Fig. 11</figureCitation>
); meso-metapleural suture obsolescent; metasomal sternum I with medial, longitudinal sulcus present on basal third
</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">3</td>
</tr>
<tr pageId="3" pageNumber="4">
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">-</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
Gular carina linear or crenulate, not elevated or angulate on either side of midline, interrupted medially occipital carina incomplete laterally, not meeting gular carina (as in
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="3" pageNumber="4">Fig. 12</figureCitation>
); meso-metapleural suture present, distinct; metasomal sternum I with medial, longitudinal sulcus present on more than half of length
</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">4</td>
</tr>
<tr pageId="3" pageNumber="4">
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">3</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
Gular carina interrupted medially, with strongly developed submedial tooth or angle, meeting occipital carina at distinct angle (
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="3" pageNumber="4">Fig. 11</figureCitation>
); transverse carina posterior to gular carina medially interrupted, without curved angles or lobes; meso-metapleural suture absent or only faintly visible in certain views
</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
<taxonomicName class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="segredentata">
<emphasis italics="true" pageId="3" pageNumber="4">Colocistis segredentata</emphasis>
</taxonomicName>
Mickel &amp; Krombein
</td>
</tr>
<tr pageId="3" pageNumber="4">
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">-</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
Gular carina not interrupted medially, strongly developed submedial rounded lobe (
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="3" pageNumber="4">Fig. 10</figureCitation>
); gular carina meeting occipital carina in even curve; transverse carina posterior to gular carina with two low, rounded prominences behind lobes of gular carina; meso-metapleural suture faint but distinct
</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
<taxonomicName class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="eureka">
<emphasis italics="true" pageId="3" pageNumber="4">Colocistis eureka</emphasis>
</taxonomicName>
(Banks)
</td>
</tr>
<tr pageId="3" pageNumber="4">
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">4</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
Gular carina extending laterally considerably beyond where occipital carina would terminate if present (
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="3" pageNumber="4">Fig. 12</figureCitation>
); vertex foveae linear, consisting of row of contiguous pits; metasomal sternum I medial, longitudinal sulcus extending complete length of sternum, sulcus narrow and deep anteriorly, broad and shallow posteriorly
</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
<taxonomicName class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sulcata">
<emphasis italics="true" pageId="3" pageNumber="4">Colocistis sulcata</emphasis>
</taxonomicName>
Mickel &amp; Krombein
</td>
</tr>
<tr pageId="3" pageNumber="4">
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">-</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">Gular carina not extending laterally beyond where occipital carina would terminate if present; vertex foveae short, elliptical; metasomal sternum I medial, longitudinal sulcus same width throughout, deeper anteriorly than posteriorly</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
<taxonomicName class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="paxillata">
<emphasis italics="true" pageId="3" pageNumber="4">Colocistis paxillata</emphasis>
</taxonomicName>
Mickel &amp; Krombein
</td>
</tr>
<tr pageId="3" pageNumber="4">
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">5</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
Forewing with two submarginal cells (as in
<figureCitation captionStart="Figures 13–24" captionStartId="F2" captionText="Figures 13 - 24. 13, 14 Male forewing. 15 Ventral view of male hindcoxa 16 Dorsal view of male propodeum 17 dorsal view of male mandible 18 lateral view of mandible 19 Ventral view of male metasomal sterna I-II 20, 22 Lateral view of male metasomal segments I and II 21, 23, 24 Ventral view of female metasomal sternum I. Abbreviations: SM- 1, 2, 3 = submarginal cells 1 - 3." httpUri="https://binary.pensoft.net/fig/11788" pageId="3" pageNumber="4">Fig. 13</figureCitation>
); hindcoxal carina low and indistinct
</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">6</td>
</tr>
<tr pageId="3" pageNumber="4">
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">-</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
Forewing with three submarginal cells (as in
<figureCitation captionStart="Figures 13–24" captionStartId="F2" captionText="Figures 13 - 24. 13, 14 Male forewing. 15 Ventral view of male hindcoxa 16 Dorsal view of male propodeum 17 dorsal view of male mandible 18 lateral view of mandible 19 Ventral view of male metasomal sterna I-II 20, 22 Lateral view of male metasomal segments I and II 21, 23, 24 Ventral view of female metasomal sternum I. Abbreviations: SM- 1, 2, 3 = submarginal cells 1 - 3." httpUri="https://binary.pensoft.net/fig/11788" pageId="3" pageNumber="4">Fig. 14</figureCitation>
); hindcoxal carina prominent
</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">8</td>
</tr>
<tr pageId="3" pageNumber="4">
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">6</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
Gular carina elevated anteriorly (as in
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="3" pageNumber="4">Fig. 9</figureCitation>
)
</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
<taxonomicName class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="eremi">
<emphasis italics="true" pageId="3" pageNumber="4">Colocistis eremi</emphasis>
</taxonomicName>
(Bradley)
</td>
</tr>
<tr pageId="3" pageNumber="4">
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">-</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">Gular carina not elevated anteriorly, nearly linear</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">7</td>
</tr>
<tr pageId="3" pageNumber="4">
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">7</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
OOL 1 ocellar diameter or less; vertex not visible behind ocelli in full frontal view (
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="3" pageNumber="4">Fig. 1</figureCitation>
); gonostylar apical margin evenly curved to acute apex (
<figureCitation captionStart="Figures 49–56" captionStartId="F7" captionText="Figures 49 - 56. Lateral view of male genital capsule." httpUri="https://binary.pensoft.net/fig/11793" pageId="3" pageNumber="4">Fig. 49</figureCitation>
)
</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
<taxonomicName class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="brevis">
<emphasis italics="true" pageId="3" pageNumber="4">Colocistis brevis</emphasis>
</taxonomicName>
(Cresson)
</td>
</tr>
<tr pageId="3" pageNumber="4">
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">-</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
OOL 2 ocellar diameters or more; vertex clearly visible behind ocelli in front view (
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="3" pageNumber="4">Fig. 3</figureCitation>
); gonostylar apical margin truncate or slightly emarginate (
<figureCitation captionStart="Figures 49–56" captionStartId="F7" captionText="Figures 49 - 56. Lateral view of male genital capsule." httpUri="https://binary.pensoft.net/fig/11793" pageId="3" pageNumber="4">Fig. 51</figureCitation>
)
</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
<taxonomicName authorityName="Kimsey &amp; Wasbauer" authorityYear="2013" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="chemsaki">
<emphasis italics="true" pageId="3" pageNumber="4">Colocistis chemsaki</emphasis>
</taxonomicName>
sp. n.
</td>
</tr>
<tr pageId="3" pageNumber="4">
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">8</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
Metasomal sternum II basally slightly raised and separated from posterior part by transverse declivity (as in
<figureCitation captionStart="Figures 13–24" captionStartId="F2" captionText="Figures 13 - 24. 13, 14 Male forewing. 15 Ventral view of male hindcoxa 16 Dorsal view of male propodeum 17 dorsal view of male mandible 18 lateral view of mandible 19 Ventral view of male metasomal sterna I-II 20, 22 Lateral view of male metasomal segments I and II 21, 23, 24 Ventral view of female metasomal sternum I. Abbreviations: SM- 1, 2, 3 = submarginal cells 1 - 3." httpUri="https://binary.pensoft.net/fig/11788" pageId="3" pageNumber="4">Figs 19, 22</figureCitation>
)
</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">9</td>
</tr>
<tr pageId="3" pageNumber="4">
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">-</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
Metasomal sternum II without raised area basally (as in
<figureCitation captionStart="Figures 13–24" captionStartId="F2" captionText="Figures 13 - 24. 13, 14 Male forewing. 15 Ventral view of male hindcoxa 16 Dorsal view of male propodeum 17 dorsal view of male mandible 18 lateral view of mandible 19 Ventral view of male metasomal sterna I-II 20, 22 Lateral view of male metasomal segments I and II 21, 23, 24 Ventral view of female metasomal sternum I. Abbreviations: SM- 1, 2, 3 = submarginal cells 1 - 3." httpUri="https://binary.pensoft.net/fig/11788" pageId="3" pageNumber="4">Fig. 20</figureCitation>
)
</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">10</td>
</tr>
<tr pageId="3" pageNumber="4">
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">9</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
Head and body black; ocelli small, diameter of lateral ocellus 0.5
<normalizedToken originalValue="×">x</normalizedToken>
OOL (
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="3" pageNumber="4">Fig. 7</figureCitation>
); clypeus with median tubercle (
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="3" pageNumber="4">Fig. 7</figureCitation>
); aedeagus in lateral view evenly curved, simple at apex (
<figureCitation captionStart="Figures 49–56" captionStartId="F7" captionText="Figures 49 - 56. Lateral view of male genital capsule." httpUri="https://binary.pensoft.net/fig/11793" pageId="3" pageNumber="4">Fig. 55</figureCitation>
)
</td>
<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis italics="true" pageId="3" pageNumber="4">Colocistis pilosa</emphasis>
</taxonomicName>
Krombein
</td>
</tr>
<tr pageId="4" pageNumber="5">
<td colspan="1" pageId="4" pageNumber="5" rowspan="1">
<pageBreakToken pageId="4" pageNumber="5" start="start">-</pageBreakToken>
</td>
<td colspan="1" pageId="4" pageNumber="5" rowspan="1">
Head and body castaneus; ocelli enlarged, diameter of lateral ocellus at least 0.7
<normalizedToken originalValue="×">x</normalizedToken>
OOL (
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="4" pageNumber="5">Fig. 4</figureCitation>
); clypeus without median tubercle (
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="4" pageNumber="5">Fig. 4</figureCitation>
); aedeagus in lateral view suddenly decurved, apex with a dorsally projecting process
<figureCitation captionStart="Figures 49–56" captionStartId="F7" captionText="Figures 49 - 56. Lateral view of male genital capsule." httpUri="https://binary.pensoft.net/fig/11793" pageId="4" pageNumber="5">Fig. 52</figureCitation>
)
</td>
<td colspan="1" pageId="4" pageNumber="5" rowspan="1">
<taxonomicName class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="crassa">
<emphasis italics="true" pageId="4" pageNumber="5">Colocistis crassa</emphasis>
</taxonomicName>
(Bradley)
</td>
</tr>
<tr pageId="4" pageNumber="5">
<td colspan="1" pageId="4" pageNumber="5" rowspan="1">10</td>
<td colspan="1" pageId="4" pageNumber="5" rowspan="1">
Clypeus with a projecting tubercle, plainly visible above mandibles in full lateral view, area between tubercle and apex appearing beveled (
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="4" pageNumber="5">Fig. 8</figureCitation>
); genitalia in lateral view with gonostyli straight or slightly concave at apex; aedeagus decurved before apex (
<figureCitation captionStart="Figures 49–56" captionStartId="F7" captionText="Figures 49 - 56. Lateral view of male genital capsule." httpUri="https://binary.pensoft.net/fig/11793" pageId="4" pageNumber="5">Fig. 56</figureCitation>
)
</td>
<td colspan="1" pageId="4" pageNumber="5" rowspan="1">
<taxonomicName class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="thermarum">
<emphasis italics="true" pageId="4" pageNumber="5">Colocistis thermarum</emphasis>
</taxonomicName>
(Bradley)
</td>
</tr>
<tr pageId="4" pageNumber="5">
<td colspan="1" pageId="4" pageNumber="5" rowspan="1">-</td>
<td colspan="1" pageId="4" pageNumber="5" rowspan="1">
Clypeus somewhat flattened and depressed, not visible above mandibles in full lateral view, apex not beveled; genitalia in lateral view with gonostylus strongly excavate at apex; aedeagus not decurved before apex (as in
<figureCitation captionStart="Figures 49–56" captionStartId="F7" captionText="Figures 49 - 56. Lateral view of male genital capsule." httpUri="https://binary.pensoft.net/fig/11793" pageId="4" pageNumber="5">Figs 50, 54</figureCitation>
)
</td>
<td colspan="1" pageId="4" pageNumber="5" rowspan="1">11</td>
</tr>
<tr pageId="4" pageNumber="5">
<td colspan="1" pageId="4" pageNumber="5" rowspan="1">11</td>
<td colspan="1" pageId="4" pageNumber="5" rowspan="1">
Integument of head, mesosoma and first metasomal segment lighter in color than remainder of metasoma; head in full frontal view (
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="4" pageNumber="5">Fig. 6</figureCitation>
) with vertex arched far above ocelli; genitalia in lateral view (
<figureCitation captionStart="Figures 49–56" captionStartId="F7" captionText="Figures 49 - 56. Lateral view of male genital capsule." httpUri="https://binary.pensoft.net/fig/11793" pageId="4" pageNumber="5">Fig. 54</figureCitation>
) with apex of gonostylus nearly straight
</td>
<td colspan="1" pageId="4" pageNumber="5" rowspan="1">
<taxonomicName authorityName="Kimsey &amp; Wasbauer" authorityYear="2013" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="oaxacana">
<emphasis italics="true" pageId="4" pageNumber="5">Colocistis oaxacana</emphasis>
</taxonomicName>
sp. n.
</td>
</tr>
<tr pageId="4" pageNumber="5">
<td colspan="1" pageId="4" pageNumber="5" rowspan="1">-</td>
<td colspan="1" pageId="4" pageNumber="5" rowspan="1">
Integument of entire body concolorous; head in full frontal view with vertex arched only slightly, if at all, above ocelli (
<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="4" pageNumber="5">Fig. 2</figureCitation>
); genitalia in lateral view (
<figureCitation captionStart="Figures 49–56" captionStartId="F7" captionText="Figures 49 - 56. Lateral view of male genital capsule." httpUri="https://binary.pensoft.net/fig/11793" pageId="4" pageNumber="5">Fig. 50</figureCitation>
) with apex of gonostylus strongly excavate
</td>
<td colspan="1" pageId="4" pageNumber="5" rowspan="1">
<taxonomicName class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="castanea">
<emphasis italics="true" pageId="4" pageNumber="5">Colocistis castanea</emphasis>
</taxonomicName>
(Cresson)
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>
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