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<mods:title id="C36D0EC996728EA484053E3F5F92DDFF">New euagrid spider species from Thailand and Malaysia, and new localities of Leptothele bencha (Arachnida: Araneae)</mods:title>
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<mods:namePart id="74BCDF0D352DDD0462525037F50CE434">Schwendinger, Peter J.</mods:namePart>
<mods:affiliation id="3415DCA735C24678A1214E78B0EA692A">Muséum d’histoire naturelle de Genève, C. P. 6434, CH- 1211 Genève 6, Switzerland</mods:affiliation>
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<mods:namePart id="57BA5DB5617BEDDA7F518CF378CB99DB">Lehmann-Graber, Christina</mods:namePart>
<mods:affiliation id="B6733DF2B7CF02D6A88FBBF879E3D70B">Muséum d’histoire naturelle de Genève, C. P. 6434, CH- 1211 Genève 6, Switzerland</mods:affiliation>
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<mods:namePart id="134BFE42830FE90ABF16BE6323DA83D2">Hongpadharakiree, Komson</mods:namePart>
<mods:affiliation id="26F49F7635D74DFBCCE592C26F842DA4">Sirinath Rajini Mangrove Ecosystem Learning Center, Pranburi, Prachuab Khiri Khan, Thailand</mods:affiliation>
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<mods:namePart id="D6D65C7EF73FB40EE3635BC38EB9A896">Syuhadah, Nurul</mods:namePart>
<mods:affiliation id="7E5B3A74BF7E926A2C8359284DBEE928">The Liphistius Project, Petaling Jaya, Malaysia</mods:affiliation>
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<paragraph id="E56F36A2C62AFF9413FAFA17532EFA0D" blockId="3.[906,1362,1482,1506]" box="[906,1362,1482,1506]" pageId="3" pageNumber="425">
<heading id="BE2781CEC62AFF9413FAFA17532EFA0D" bold="true" box="[906,1362,1482,1506]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="425" reason="2">
<taxonomicName id="22D04D21C62AFF9413FAFA17532EFA0D" authority="Raven &amp; Schwendinger, 1995" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[906,1362,1482,1506]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="425" phylum="Arthropoda" rank="genus">
<emphasis id="D7A4EAB0C62AFF9413FAFA17532EFA0D" bold="true" box="[906,1362,1482,1506]" pageId="3" pageNumber="425">
<emphasis id="D7A4EAB0C62AFF9413FAFA175582FA0D" bold="true" box="[906,1022,1482,1506]" italics="true" pageId="3" pageNumber="425">Leptothele</emphasis>
<bibRefCitation id="81414B53C62AFF941474FA16532EFA0D" author="Raven R. J. &amp; Schwendinger P. J." box="[1028,1362,1483,1506]" pageId="3" pageNumber="425" pagination="623 - 641" refId="ref23253" refString="Raven R. J., Schwendinger P. J. 1995. Three new mygalomorph spider genera from Thailand and China (Araneae). Memoirs of the Queensland Museum 38 (2): 623 - 641." type="journal article" year="1995">Raven &amp; Schwendinger, 1995</bibRefCitation>
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<paragraph id="E56F36A2C62AFF94134FFA215353F9FE" blockId="3.[831,1327,1532,1553]" box="[831,1327,1532,1553]" pageId="3" pageNumber="425">
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<treatmentCitation id="647110B3C62AFF94134FFA215357F9FE" author="Raven R. J. &amp; Schwendinger P. J." box="[831,1323,1532,1553]" page="637" pageId="3" pageNumber="425" year="1995">
<taxonomicName id="22D04D21C62AFF94134FFA215357F9FE" authority="Raven &amp; Schwendinger, 1995: 636 - 637" authorityName="Raven &amp; Schwendinger" authorityPageNumber="636 - 637" authorityYear="1995" box="[831,1323,1532,1553]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="425" phylum="Arthropoda" rank="genus">
<emphasis id="D7A4EAB0C62AFF94134FFA2155D8F9FE" box="[831,932,1532,1553]" italics="true" pageId="3" pageNumber="425">Leptothele</emphasis>
<bibRefCitation id="81414B53C62AFF9413DBFA215357F9FE" author="Raven R. J. &amp; Schwendinger P. J." box="[939,1323,1532,1553]" pageId="3" pageNumber="425" pagination="623 - 641" refId="ref23253" refString="Raven R. J., Schwendinger P. J. 1995. Three new mygalomorph spider genera from Thailand and China (Araneae). Memoirs of the Queensland Museum 38 (2): 623 - 641." type="journal article" year="1995">Raven &amp; Schwendinger, 1995: 636-637</bibRefCitation>
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.
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<typeStatus id="3A6B8800C62AFF94134FF9F4550BF9AF" box="[831,887,1577,1600]" pageId="3" pageNumber="425">Type</typeStatus>
species:
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<taxonomicName id="22D04D21C62AFF94139CF9F455B9F98F" authority="Raven &amp; Schwendinger, 1995" authorityName="Raven &amp; Schwendinger" authorityYear="1995" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="425" phylum="Arthropoda" rank="species" species="bencha">
<emphasis id="D7A4EAB0C62AFF94139CF9F452C6F9AF" box="[1004,1210,1577,1600]" italics="true" pageId="3" pageNumber="425">Leptothele bencha</emphasis>
<bibRefCitation id="81414B53C62AFF9414B8F9F455B9F98F" author="Raven R. J. &amp; Schwendinger P. J." pageId="3" pageNumber="425" pagination="623 - 641" refId="ref23253" refString="Raven R. J., Schwendinger P. J. 1995. Three new mygalomorph spider genera from Thailand and China (Araneae). Memoirs of the Queensland Museum 38 (2): 623 - 641." type="journal article" year="1995">Raven &amp; Schwendinger, 1995</bibRefCitation>
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by monotypy and by designation.
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<emphasis id="D7A4EAB0C62AFF94134FF9A55267F960" bold="true" box="[831,1051,1656,1679]" pageId="3" pageNumber="425">Species included:</emphasis>
<taxonomicName id="22D04D21C62AFF941444F9A55265F940" authority="Raven &amp; Schwendinger, 1995" authorityName="Raven &amp; Schwendinger" authorityYear="1995" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="425" phylum="Arthropoda" rank="species" species="bencha">
<emphasis id="D7A4EAB0C62AFF941444F9A5536CF960" box="[1076,1296,1656,1679]" italics="true" pageId="3" pageNumber="425">Leptothele bencha</emphasis>
<bibRefCitation id="81414B53C62AFF941559F9A55265F940" author="Raven R. J. &amp; Schwendinger P. J." pageId="3" pageNumber="425" pagination="623 - 641" refId="ref23253" refString="Raven R. J., Schwendinger P. J. 1995. Three new mygalomorph spider genera from Thailand and China (Araneae). Memoirs of the Queensland Museum 38 (2): 623 - 641." type="journal article" year="1995">Raven &amp; Schwendinger, 1995</bibRefCitation>
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and 
<taxonomicName id="22D04D21C62AFF94143DF94552D7F940" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[1101,1195,1688,1711]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="425" phylum="Arthropoda" rank="species" species="chang" status="sp. nov.">
<emphasis id="D7A4EAB0C62AFF94143DF94552D7F940" box="[1101,1195,1688,1711]" italics="true" pageId="3" pageNumber="425">L. chang</emphasis>
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<taxonomicNameLabel id="CC9757CBC62AFF9414C1F9455378F940" box="[1201,1284,1688,1711]" pageId="3" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="E56F36A2C62AFF94134FF91A53E0F7F2" blockId="3.[831,1437,1735,2077]" pageId="3" pageNumber="425">
<emphasis id="D7A4EAB0C62AFF94134FF91A526BF931" bold="true" box="[831,1047,1735,1758]" pageId="3" pageNumber="425">Revised diagnosis:</emphasis>
Distinguished from 
<taxonomicName id="22D04D21C62AFF941571F91A53E0F931" authorityName="Simon" authorityYear="1889" box="[1281,1436,1735,1758]" class="Arachnida" family="Dipluridae" genus="Phyxioschema" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="425" phylum="Arthropoda" rank="genus">
<emphasis id="D7A4EAB0C62AFF941571F91A53E0F931" box="[1281,1436,1735,1758]" italics="true" pageId="3" pageNumber="425">Phyxioschema</emphasis>
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by males lacking a prolateral band of elongated spinules and by possessing a much shorter ventral spur (
<figureCitation id="7DEB2A27C62AFF941530F8DA552EF8D2" captionStart="Fig" captionStartId="7.[152,187,1970,1991]" captionTargetBox="[157,1426,704,1913]" captionTargetId="figure-219@7.[151,1435,699,1917]" captionTargetPageId="7" captionText="Fig. 3. Leptothele bencha; males from Ko Sirey (A-B), Ko Tarutao (C-F, G, showing two specimens), Ko Libong (H) and Ko Sukon (I- J). (A, D, H) Distal part of palp, ventral view. (B, E) Right tibia and metatarsus II, prolateral view. (C) Left tibia and metatarsus II, ventral view. (F) Right tibia and metatarsus II, ventral view. (G) Palpal organ, ventral view. (I) Left tibia and metatarsus II of aberrant specimen, prolateral view. (J) Same, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5743960" httpUri="https://zenodo.org/record/5743960/files/figure.png" pageId="3" pageNumber="425">Figs 3B, E</figureCitation>
, 
<figureCitation id="7DEB2A27C62AFF94132FF8FB5503F8D2" box="[863,895,1830,1853]" captionStart="Fig" captionStartId="10.[151,186,1968,1989]" captionTargetBox="[159,1427,610,1932]" captionTargetId="figure-192@10.[153,1434,605,1947]" captionTargetPageId="10" captionText="Fig. 5. Leptothele chang sp. nov.; male holotype (A-B, E-H), female allotype (C-D) and male paratype (I). (A, C) Habitus, dorsal view. (B, D) Habitus, ventral view. (E) Femur to metatarsus of right leg II, prolateral view. (F) Right femur II, proventral view. (G) Right femur I, retrodorsal view. (H) Tarsus I, lateral view. (I) Distal part of median article and entire distal article of PLS, lateral view; arrow indicating pseudosegmentation. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743964" httpUri="https://zenodo.org/record/5743964/files/figure.png" pageId="3" pageNumber="425">5E</figureCitation>
, 
<figureCitation id="7DEB2A27C62AFF9413FCF8FB55D0F8D2" box="[908,940,1830,1853]" captionStart="Fig" captionStartId="11.[151,186,1879,1900]" captionTargetBox="[163,1428,257,1828]" captionTargetId="figure-6@11.[151,1436,254,1838]" captionTargetPageId="11" captionText="Fig. 6. Leptothele chang sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A) Right palpal organ, ventral view. (B) Left palpal organ and palpal tarsus, prolateral view. (C) Right palpal organ and palpal tarsus, ventral view. (D) Left palpal organ and palpal tarsus, ventral view. (E) Right palpal organ and palpal tarsus, prolateral view. (F) Patella to metatarsus of left leg I, ventral view. (G) Patella to metatarsus of left leg II, prolateral view. (H) Same, ventral view. (I) Same of right leg II, ventral view. (J) Right tibia II, ventral view. (K) Left tibia II, ventral view. Scale lines 1.0 mm (A-E; F-K)." figureDoi="http://doi.org/10.5281/zenodo.5743966" httpUri="https://zenodo.org/record/5743966/files/figure.png" pageId="3" pageNumber="425">6G</figureCitation>
cf. Raven, 1981: fig. 7; 
<bibRefCitation id="81414B53C62AFF9414C9F8FB53E4F8D2" author="Schwendinger P. J." box="[1209,1432,1830,1853]" pageId="3" pageNumber="425" pagination="1 - 40" refId="ref23293" refString="Schwendinger P. J. 2009. A taxonomic revision of the genus Phyxioschema (Araneae, Dipluridae), I: species from Thailand. Zootaxa 2126: 1 - 40." type="journal article" year="2009">Schwendinger, 2009</bibRefCitation>
: figs 2D, 4C, 5C, 9C, 12C, 15C) and a low transversal subdistal ventral ridge on tibia II (
<figureCitation id="7DEB2A27C62AFF941490F8BB5357F892" box="[1248,1323,1894,1917]" captionStart="Fig" captionStartId="7.[152,187,1970,1991]" captionTargetBox="[157,1426,704,1913]" captionTargetId="figure-219@7.[151,1435,699,1917]" captionTargetPageId="7" captionText="Fig. 3. Leptothele bencha; males from Ko Sirey (A-B), Ko Tarutao (C-F, G, showing two specimens), Ko Libong (H) and Ko Sukon (I- J). (A, D, H) Distal part of palp, ventral view. (B, E) Right tibia and metatarsus II, prolateral view. (C) Left tibia and metatarsus II, ventral view. (F) Right tibia and metatarsus II, ventral view. (G) Palpal organ, ventral view. (I) Left tibia and metatarsus II of aberrant specimen, prolateral view. (J) Same, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5743960" httpUri="https://zenodo.org/record/5743960/files/figure.png" pageId="3" pageNumber="425">Figs 3</figureCitation>
B-C, E-F, I-J, 5E, 6G-K), and by possessing a single, widely conical or mound-like ventral process instead of 2-3 longitudinal ridges on metatarsus II (
<figureCitation id="7DEB2A27C62AFF941497F81B5352F832" box="[1255,1326,1990,2013]" captionStart="Fig" captionStartId="7.[152,187,1970,1991]" captionTargetBox="[157,1426,704,1913]" captionTargetId="figure-219@7.[151,1435,699,1917]" captionTargetPageId="7" captionText="Fig. 3. Leptothele bencha; males from Ko Sirey (A-B), Ko Tarutao (C-F, G, showing two specimens), Ko Libong (H) and Ko Sukon (I- J). (A, D, H) Distal part of palp, ventral view. (B, E) Right tibia and metatarsus II, prolateral view. (C) Left tibia and metatarsus II, ventral view. (F) Right tibia and metatarsus II, ventral view. (G) Palpal organ, ventral view. (I) Left tibia and metatarsus II of aberrant specimen, prolateral view. (J) Same, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5743960" httpUri="https://zenodo.org/record/5743960/files/figure.png" pageId="3" pageNumber="425">Figs 3</figureCitation>
B-C, E-F, I-J, 5E, 6G-I cf. 
<bibRefCitation id="81414B53C62AFF941473F83B52E3F812" author="Raven R. J." box="[1027,1183,2022,2045]" pageId="3" pageNumber="425" pagination="225 - 231" refId="ref23102" refString="Raven R. J. 1981 a. The mygalomorph spider genera Phyxioschema Simon and Stenygrocerus Simon (Dipluridae: Ischnothelinae). Bulletin of the British Arachnological Society 5 (5): 225 - 231." type="journal article" year="1981">Raven, 1981a</bibRefCitation>
: fig. 7; 
<bibRefCitation id="81414B53C62AFF94148DF83B5505F7F2" author="Schwendinger P. J." pageId="3" pageNumber="425" pagination="1 - 40" refId="ref23293" refString="Schwendinger P. J. 2009. A taxonomic revision of the genus Phyxioschema (Araneae, Dipluridae), I: species from Thailand. Zootaxa 2126: 1 - 40." type="journal article" year="2009">Schwendinger, 2009</bibRefCitation>
: figs 2D, 6J, 10J, 13J, 16K, 19L). Females with
</paragraph>
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<paragraph id="E56F36A2C62DFF9310EBF8E052E3F7F4" blockId="4.[155,1439,1853,2075]" pageId="4" pageNumber="426">
Fig. 2. Localities of 
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<emphasis id="D7A4EAB0C62DFF931114F8E057B5F8BD" box="[356,457,1853,1874]" italics="true" pageId="4" pageNumber="426">Leptothele</emphasis>
spp.
</taxonomicName>
and 
<taxonomicName id="22D04D21C62DFF931254F8E054B6F8BD" box="[548,714,1853,1874]" class="Arachnida" family="Euagridae" genus="Malayathele" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="D7A4EAB0C62DFF931254F8E054E0F8BD" box="[548,668,1853,1874]" italics="true" pageId="4" pageNumber="426">Malayathele</emphasis>
spp.
</taxonomicName>
on the Thai-Malay Peninsula. The coast of Sumatra is omitted. 1 - Ko Sirey (
<taxonomicName id="22D04D21C62DFF93109FF8875732F880" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[239,334,1882,1903]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="bencha">
<emphasis id="D7A4EAB0C62DFF93109FF8875732F880" box="[239,334,1882,1903]" italics="true" pageId="4" pageNumber="426">L. bencha</emphasis>
</taxonomicName>
); 2 - Ko Yao Noi (
<taxonomicName id="22D04D21C62DFF93127CF8875410F880" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[524,620,1882,1903]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="bencha">
<emphasis id="D7A4EAB0C62DFF93127CF8875410F880" box="[524,620,1882,1903]" italics="true" pageId="4" pageNumber="426">L. bencha</emphasis>
</taxonomicName>
); 3 - Khao Phanom Bencha (
<taxonomicName id="22D04D21C62DFF9313FEF8875592F880" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[910,1006,1882,1903]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="bencha">
<emphasis id="D7A4EAB0C62DFF9313FEF8875592F880" box="[910,1006,1882,1903]" italics="true" pageId="4" pageNumber="426">L. bencha</emphasis>
</taxonomicName>
, type locality); 4 - Thab Khaek - Hang Nak Hill and Khlong Jilat (
<taxonomicName id="22D04D21C62DFF9311CEF8AB5460F864" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[446,540,1910,1931]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="bencha">
<emphasis id="D7A4EAB0C62DFF9311CEF8AB5460F864" box="[446,540,1910,1931]" italics="true" pageId="4" pageNumber="426">L. bencha</emphasis>
</taxonomicName>
); 5 - Ban Chong Phlie (
<taxonomicName id="22D04D21C62DFF931372F8AB551DF864" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[770,865,1910,1931]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="bencha">
<emphasis id="D7A4EAB0C62DFF931372F8AB551DF864" box="[770,865,1910,1931]" italics="true" pageId="4" pageNumber="426">L. bencha</emphasis>
</taxonomicName>
); 6 - Ko Lanta (
<taxonomicName id="22D04D21C62DFF93138DF8AB5227F864" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[1021,1115,1910,1931]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="bencha">
<emphasis id="D7A4EAB0C62DFF93138DF8AB5227F864" box="[1021,1115,1910,1931]" italics="true" pageId="4" pageNumber="426">L. bencha</emphasis>
</taxonomicName>
); 7 - Ko Muk (
<taxonomicName id="22D04D21C62DFF93149EF8AB5330F864" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[1262,1356,1910,1931]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="bencha">
<emphasis id="D7A4EAB0C62DFF93149EF8AB5330F864" box="[1262,1356,1910,1931]" italics="true" pageId="4" pageNumber="426">L. bencha</emphasis>
</taxonomicName>
); 8 - Ko Libong (
<taxonomicName id="22D04D21C62DFF93114BF84E57E6F847" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[315,410,1939,1960]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="bencha">
<emphasis id="D7A4EAB0C62DFF93114BF84E57E6F847" box="[315,410,1939,1960]" italics="true" pageId="4" pageNumber="426">L. bencha</emphasis>
</taxonomicName>
); 9 - Ko Sukon (
<taxonomicName id="22D04D21C62DFF931232F84E54DDF847" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[578,673,1939,1960]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="bencha">
<emphasis id="D7A4EAB0C62DFF931232F84E54DDF847" box="[578,673,1939,1960]" italics="true" pageId="4" pageNumber="426">L. bencha</emphasis>
</taxonomicName>
); 10 - Ko Tarutao (
<taxonomicName id="22D04D21C62DFF931310F84E55C3F847" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[864,959,1939,1960]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="bencha">
<emphasis id="D7A4EAB0C62DFF931310F84E55C3F847" box="[864,959,1939,1960]" italics="true" pageId="4" pageNumber="426">L. bencha</emphasis>
</taxonomicName>
); 11 - Tham Khao Chang Hai (
<taxonomicName id="22D04D21C62DFF931482F84E533BF847" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[1266,1351,1939,1960]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="chang" status="sp. nov.">
<emphasis id="D7A4EAB0C62DFF931482F84E533BF847" box="[1266,1351,1939,1960]" italics="true" pageId="4" pageNumber="426">L. chang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62DFF93153DF84E53E7F847" box="[1357,1435,1939,1960]" pageId="4" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
, type locality); 12 - Ulu Groh (
<taxonomicName id="22D04D21C62DFF931262F86D542EF82A" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[530,594,1968,1989]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="ulu" status="sp. nov.">
<emphasis id="D7A4EAB0C62DFF931262F86D542EF82A" box="[530,594,1968,1989]" italics="true" pageId="4" pageNumber="426">M. ulu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62DFF93122AF86D54D4F82A" box="[602,680,1968,1989]" pageId="4" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
, type locality); 13 - Chenderiang (
<taxonomicName id="22D04D21C62DFF93138BF86D52E5F82A" box="[1019,1177,1968,1989]" class="Arachnida" family="Euagridae" genus="Malayathele" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="D7A4EAB0C62DFF93138BF86D520FF82A" box="[1019,1139,1968,1989]" italics="true" pageId="4" pageNumber="426">Malayathele</emphasis>
sp.
</taxonomicName>
); 14 - Cameron Highlands (
<taxonomicName id="22D04D21C62DFF93109FF81057EDF80D" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[239,401,1997,2018]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="cameronensis" status="sp. nov.">
<emphasis id="D7A4EAB0C62DFF93109FF81057EDF80D" box="[239,401,1997,2018]" italics="true" pageId="4" pageNumber="426">M. cameronensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62DFF9311E7F8105798F80D" box="[407,484,1997,2018]" pageId="4" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
, type locality; 
<taxonomicName id="22D04D21C62DFF931200F81054D3F80D" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[624,687,1997,2018]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="ulu" status="sp. nov.">
<emphasis id="D7A4EAB0C62DFF931200F81054D3F80D" box="[624,687,1997,2018]" italics="true" pageId="4" pageNumber="426">M. ulu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62DFF9312C4F810557EF80D" box="[692,770,1997,2018]" pageId="4" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
); 15 - Fraser’s Hill (
<taxonomicName id="22D04D21C62DFF9313B3F8105243F80D" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[963,1087,1997,2018]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="maculosa" status="sp. nov.">
<emphasis id="D7A4EAB0C62DFF9313B3F8105243F80D" box="[963,1087,1997,2018]" italics="true" pageId="4" pageNumber="426">M. maculosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62DFF931435F81052EDF80D" box="[1093,1169,1997,2018]" pageId="4" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
, type locality); 16 - Genting Highlands (
<taxonomicName id="22D04D21C62DFF931127F83757AFF810" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[343,467,2026,2047]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="maculosa" status="sp. nov.">
<emphasis id="D7A4EAB0C62DFF931127F83757AFF810" box="[343,467,2026,2047]" italics="true" pageId="4" pageNumber="426">M. maculosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62DFF9311A8F837545AF810" box="[472,550,2026,2047]" pageId="4" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
); 17 - Templer Park (
<taxonomicName id="22D04D21C62DFF931281F8375514F810" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[753,872,2026,2047]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="kanching" status="sp. nov.">
<emphasis id="D7A4EAB0C62DFF931281F8375514F810" box="[753,872,2026,2047]" italics="true" pageId="4" pageNumber="426">M. kanching</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62DFF93131EF83755C7F810" box="[878,955,2026,2047]" pageId="4" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
; type locality); 18 - Ulu Gombak (
<taxonomicName id="22D04D21C62DFF931575F8375301F810" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[1285,1405,2026,2047]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="kanching" status="sp. nov.">
<emphasis id="D7A4EAB0C62DFF931575F8375301F810" box="[1285,1405,2026,2047]" italics="true" pageId="4" pageNumber="426">M. kanching</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62DFF9315F3F837576FF7F4" pageId="4" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
); 19 - Jeram Pasu Waterfall (
<taxonomicName id="22D04D21C62DFF931259F7DB54B8F7F4" box="[553,708,2054,2075]" class="Arachnida" family="Euagridae" genus="Malayathele" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="D7A4EAB0C62DFF931259F7DB54DDF7F4" box="[553,673,2054,2075]" italics="true" pageId="4" pageNumber="426">Malayathele</emphasis>
sp.
</taxonomicName>
); 20 - Singapore (
<taxonomicName id="22D04D21C62DFF931300F7DB55AEF7F4" authorityName="Raven" authorityYear="1979" box="[880,978,2054,2075]" class="Arachnida" family="Euagridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="family">Euagridae</taxonomicName>
/ 
<taxonomicName id="22D04D21C62DFF9313A9F7DB523DF7F4" box="[985,1089,2054,2075]" class="Arachnida" family="Dipluridae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="426" phylum="Arthropoda" rank="family">Dipluridae</taxonomicName>
gen. sp.).
</paragraph>
</caption>
<subSubSection id="ADCA6529C62CFF9110E7FEDC5763FB39" lastPageId="6" lastPageNumber="428" pageId="5" pageNumber="427" type="key">
<paragraph id="E56F36A2C62CFF9210E7FEDC542BFEF7" blockId="5.[151,599,257,280]" box="[151,599,257,280]" pageId="5" pageNumber="427">
<emphasis id="D7A4EAB0C62CFF9210E7FEDC542BFEF7" bold="true" box="[151,599,257,280]" pageId="5" pageNumber="427">
Key to the genera of 
<taxonomicName id="22D04D21C62CFF9211F3FEDC5786FEF7" ID-CoL="98S2G" authorityName="Raven" authorityYear="1979" box="[387,506,257,280]" class="Arachnida" family="Euagridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="427" phylum="Arthropoda" rank="family">Euagridae</taxonomicName>
in Asia:
</emphasis>
</paragraph>
<key id="91416193C62CFF9210E7FE9C53E1FAB7" pageId="5" pageNumber="427">
<keyStep id="5E242E07C62CFF9210E7FE9C53E0FC57" pageId="5" pageNumber="427">
<paragraph id="E56F36A2C62CFF9210E7FE9C53E0FD77" blockId="5.[151,1437,321,952]" pageId="5" pageNumber="427">
<keyLead id="5E219597C62CFF9210E7FE9C53E0FD77" pageId="5" pageNumber="427">
1A Males with a band of elongated spinules on prolateral side of tibia II (
<bibRefCitation id="81414B53C62CFF9213A0FE9C5215FEB7" author="Raven R. J." box="[976,1129,321,344]" pageId="5" pageNumber="427" pagination="225 - 231" refId="ref23102" refString="Raven R. J. 1981 a. The mygalomorph spider genera Phyxioschema Simon and Stenygrocerus Simon (Dipluridae: Ischnothelinae). Bulletin of the British Arachnological Society 5 (5): 225 - 231." type="journal article" year="1981">Raven, 1981a</bibRefCitation>
: fig. 7), with a distinct, quite long spur (carrying megaspines) on ventral side of tibia II, and with 2-3 longitudinal keels on ventral side of metatarsus II (proventral one reduced to a small cone in 
<taxonomicName id="22D04D21C62CFF921339FE5C52C3FE77" authority="Schwendinger, 2009" authorityName="Schwendinger" authorityYear="2009" box="[841,1215,385,408]" class="Arachnida" family="Dipluridae" genus="Phyxioschema" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="427" phylum="Arthropoda" rank="species" species="sayamense">
<emphasis id="D7A4EAB0C62CFF921339FE5C55A4FE77" box="[841,984,385,408]" italics="true" pageId="5" pageNumber="427">P. sayamense</emphasis>
<bibRefCitation id="81414B53C62CFF921390FE5C52C3FE77" author="Schwendinger P. J." box="[992,1215,385,408]" pageId="5" pageNumber="427" pagination="1 - 40" refId="ref23293" refString="Schwendinger P. J. 2009. A taxonomic revision of the genus Phyxioschema (Araneae, Dipluridae), I: species from Thailand. Zootaxa 2126: 1 - 40." type="journal article" year="2009">Schwendinger, 2009</bibRefCitation>
</taxonomicName>
: fig. 13I-J); patella I with a row of short, sigmoid or (rarely) curved retroventral-distal spines (
<bibRefCitation id="81414B53C62CFF92138EFE7C52A3FE57" author="Schwendinger P. J." box="[1022,1247,417,440]" pageId="5" pageNumber="427" pagination="1 - 40" refId="ref23293" refString="Schwendinger P. J. 2009. A taxonomic revision of the genus Phyxioschema (Araneae, Dipluridae), I: species from Thailand. Zootaxa 2126: 1 - 40." type="journal article" year="2009">Schwendinger, 2009</bibRefCitation>
: figs 3L, 6F, 10F, 13F, 16F); femur II with a long proventral band of hooked spinules (almost or actually reaching distal margin of leg article; 
<bibRefCitation id="81414B53C62CFF92112BFE3C5445FE17" author="Schwendinger P. J." box="[347,569,481,504]" pageId="5" pageNumber="427" pagination="1 - 40" refId="ref23293" refString="Schwendinger P. J. 2009. A taxonomic revision of the genus Phyxioschema (Araneae, Dipluridae), I: species from Thailand. Zootaxa 2126: 1 - 40." type="journal article" year="2009">Schwendinger, 2009</bibRefCitation>
: figs 2B, 4E, 5E, 9E, 12E, 15E, 18E); palpal tarsus with strong distal bristles (not spines; 
<bibRefCitation id="81414B53C62CFF921147FDDC5464FDF7" author="Schwendinger P. J." box="[311,536,513,536]" pageId="5" pageNumber="427" pagination="1 - 40" refId="ref23293" refString="Schwendinger P. J. 2009. A taxonomic revision of the genus Phyxioschema (Araneae, Dipluridae), I: species from Thailand. Zootaxa 2126: 1 - 40." type="journal article" year="2009">Schwendinger, 2009</bibRefCitation>
: fig. 2C). Females with or without secondary receptacles, if present then situated posterior of median receptacles (
<bibRefCitation id="81414B53C62CFF92123BFDFC5551FDD7" author="Schwendinger P. J." box="[587,813,545,568]" pageId="5" pageNumber="427" pagination="1 - 40" refId="ref23293" refString="Schwendinger P. J. 2009. A taxonomic revision of the genus Phyxioschema (Araneae, Dipluridae), I: species from Thailand. Zootaxa 2126: 1 - 40." type="journal article" year="2009">Schwendinger, 2009</bibRefCitation>
: figs 3A-I, 7, 11, 14, 17, 20; 
<bibRefCitation id="81414B53C62CFF921408FDFC5767FDB7" author="Schwendinger P. J. &amp; Zonstein S. L." pageId="5" pageNumber="427" pagination="28 - 48" refId="ref23412" refString="Schwendinger P. J., Zonstein S. L. 2011. A taxonomic revision of the genus Phyxioschema (Araneae, Dipluridae), II: species from Central Asia. Zootaxa 2815: 28 - 48." type="journal article" year="2011">Schwendinger &amp; Zonstein, 2011</bibRefCitation>
: figs 4-5, 8). Both sexes usually with spines on tarsi of posterior legs (most males of 
<taxonomicName id="22D04D21C62CFF9214DDFD9C57C1FD97" authority="Raven &amp; Schwendinger, 1989" authorityName="Raven &amp; Schwendinger" authorityYear="1989" class="Arachnida" family="Dipluridae" genus="Phyxioschema" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="427" phylum="Arthropoda" rank="species" species="suthepium">
<emphasis id="D7A4EAB0C62CFF9214DDFD9C5349FDB7" box="[1197,1333,577,600]" italics="true" pageId="5" pageNumber="427">P. suthepium</emphasis>
<bibRefCitation id="81414B53C62CFF92154CFD9C57C1FD97" author="Raven R. J. &amp; Schwendinger P. J." pageId="5" pageNumber="427" pagination="55 - 60" refId="ref23210" refString="Raven R. J., Schwendinger P. J. 1989. On a new Phyxioschema (Araneae, Mygalomorphae) from Thailand and its biology. Bulletin of the British Arachnological Society 8 (2): 55 - 60." type="journal article" year="1989">Raven &amp; Schwendinger, 1989</bibRefCitation>
</taxonomicName>
and some 
<taxonomicName id="22D04D21C62CFF921240FDBC55E4FD97" authority="Schwendinger, 2009" authorityName="Schwendinger" authorityYear="2009" box="[560,920,609,632]" class="Arachnida" family="Dipluridae" genus="Phyxioschema" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="427" phylum="Arthropoda" rank="species" species="eripnastes">
<emphasis id="D7A4EAB0C62CFF921240FDBC54C4FD97" box="[560,696,609,632]" italics="true" pageId="5" pageNumber="427">P. eripnastes</emphasis>
<bibRefCitation id="81414B53C62CFF9212CEFDBC55E4FD97" author="Schwendinger P. J." box="[702,920,609,632]" pageId="5" pageNumber="427" pagination="1 - 40" refId="ref23293" refString="Schwendinger P. J. 2009. A taxonomic revision of the genus Phyxioschema (Araneae, Dipluridae), I: species from Thailand. Zootaxa 2126: 1 - 40." type="journal article" year="2009">Schwendinger, 2009</bibRefCitation>
</taxonomicName>
without). Central Asia and all over 
<collectingCountry id="9DC77632C62CFF921566FDBC5309FD97" box="[1302,1397,609,632]" name="Thailand" pageId="5" pageNumber="427">Thailand</collectingCountry>
..... ............................................................................................................................................................. 
<taxonomicName id="22D04D21C62CFF921572FD5C53E0FD77" authorityName="Simon" authorityYear="1889" box="[1282,1436,641,664]" class="Arachnida" family="Dipluridae" genus="Phyxioschema" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="D7A4EAB0C62CFF921572FD5C53E0FD77" box="[1282,1436,641,664]" italics="true" pageId="5" pageNumber="427">Phyxioschema</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="E56F36A2C62CFF9210E7FD7C53E0FC57" blockId="5.[151,1437,321,952]" pageId="5" pageNumber="427">
<keyLead id="5E219597C62CFF9210E7FD7C53E0FC57" pageId="5" pageNumber="427">
1B Males without a band of elongated spinules on prolateral side of tibia II (
<figureCitation id="7DEB2A27C62CFF92138EFD7C522AFD57" box="[1022,1110,673,696]" captionStart="Fig" captionStartId="10.[151,186,1968,1989]" captionTargetBox="[159,1427,610,1932]" captionTargetId="figure-192@10.[153,1434,605,1947]" captionTargetPageId="10" captionText="Fig. 5. Leptothele chang sp. nov.; male holotype (A-B, E-H), female allotype (C-D) and male paratype (I). (A, C) Habitus, dorsal view. (B, D) Habitus, ventral view. (E) Femur to metatarsus of right leg II, prolateral view. (F) Right femur II, proventral view. (G) Right femur I, retrodorsal view. (H) Tarsus I, lateral view. (I) Distal part of median article and entire distal article of PLS, lateral view; arrow indicating pseudosegmentation. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743964" httpUri="https://zenodo.org/record/5743964/files/figure.png" pageId="5" pageNumber="427">Figs 5E</figureCitation>
, 
<figureCitation id="7DEB2A27C62CFF921412FD7C52EFFD57" box="[1122,1171,673,696]" captionStart="Fig" captionStartId="23.[151,186,1913,1934]" captionTargetBox="[157,1433,261,1852]" captionTargetId="figure-85@23.[153,1438,257,1862]" captionTargetPageId="23" captionText="Fig. 12. Malayathele maculosa sp. nov., female paratype from Genting Highlands (A-D), M. kanching sp. nov., male holotype (E-G), female allotype (H-I) and male paratype (J-K). (A, E, H) Habitus, dorsal view. (B, F, I) Habitus, ventral view. (C) Habitus, dorsolateral view. (D) Opisthosoma, posteroventral view. (G) Tibia and metatarsus of right leg II, prolateral view. (J) Distal part of proximal article and entire median and distal articles of PLS, lateral view. (K) Same, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743981" httpUri="https://zenodo.org/record/5743981/files/figure.png" pageId="5" pageNumber="427">12G</figureCitation>
), with a rather indistinct short spur (carrying megaspines) on ventral side of tibia II, and with 1-2 mound-like, conical or tooth-shaped processes on ventral side of metatarsus II (exception: a short retroventral keel in 
<taxonomicName id="22D04D21C62CFF921421FD3C5374FD17" ID-CoL="99B9P" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[1105,1288,737,760]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="427" phylum="Arthropoda" rank="species" species="cameronensis" status="sp. nov.">
<emphasis id="D7A4EAB0C62CFF921421FD3C5374FD17" box="[1105,1288,737,760]" italics="true" pageId="5" pageNumber="427">M. cameronensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62CFF921560FD3C531AFD17" box="[1296,1382,737,760]" pageId="5" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
, 
<figureCitation id="7DEB2A27C62CFF921502FD3C5680FCF7" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="5" pageNumber="427">Fig. 11</figureCitation>
G-J); patella I with a row of long, not sigmoid bristles (
<figureCitation id="7DEB2A27C62CFF921339FCDC55E7FCF7" box="[841,923,769,792]" captionStart="Fig" captionStartId="11.[151,186,1879,1900]" captionTargetBox="[163,1428,257,1828]" captionTargetId="figure-6@11.[151,1436,254,1838]" captionTargetPageId="11" captionText="Fig. 6. Leptothele chang sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A) Right palpal organ, ventral view. (B) Left palpal organ and palpal tarsus, prolateral view. (C) Right palpal organ and palpal tarsus, ventral view. (D) Left palpal organ and palpal tarsus, ventral view. (E) Right palpal organ and palpal tarsus, prolateral view. (F) Patella to metatarsus of left leg I, ventral view. (G) Patella to metatarsus of left leg II, prolateral view. (H) Same, ventral view. (I) Same of right leg II, ventral view. (J) Right tibia II, ventral view. (K) Left tibia II, ventral view. Scale lines 1.0 mm (A-E; F-K)." figureDoi="http://doi.org/10.5281/zenodo.5743966" httpUri="https://zenodo.org/record/5743966/files/figure.png" pageId="5" pageNumber="427">Figs 6F</figureCitation>
, 
<figureCitation id="7DEB2A27C62CFF9213D6FCDC55BEFCF7" box="[934,962,769,792]" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="5" pageNumber="427">8F</figureCitation>
, 
<figureCitation id="7DEB2A27C62CFF9213BDFCDC5589FCF7" box="[973,1013,769,792]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="5" pageNumber="427">11F</figureCitation>
, 
<figureCitation id="7DEB2A27C62CFF921471FCDC5259FCF7" box="[1025,1061,769,792]" captionStart="Fig" captionStartId="24.[149,184,1934,1955]" captionTargetBox="[152,1427,583,1872]" captionTargetId="figure-267@24.[149,1433,576,1879]" captionTargetPageId="24" captionText="Fig. 13. Malayathele maculosa sp. nov., male holotype (A-C, H-L) and two male paratypes (D-E; F-G, M). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C, E, G) Palpal organ of right palp, prolateral view. (D, F) Palpal organ of left palp, prolateral view. (H) Patella to metatarsus of left leg I, prolateral view. (I) Patella to tibia of left leg I, ventral view. (J) Patella to metatarsus of right leg II, ventral view. (K) Same of left leg II, prolateral view. (L) Same of right leg II, retrolateral view. (M) Same of left leg II, retrodorsal view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743983" httpUri="https://zenodo.org/record/5743983/files/figure.png" pageId="5" pageNumber="427">13I</figureCitation>
) or with only a single curved spine retroventrally-distally (
<figureCitation id="7DEB2A27C62CFF9211ACFCFC5442FCD7" box="[476,574,801,824]" captionStart="Fig" captionStartId="27.[151,186,1849,1870]" captionTargetBox="[157,1426,263,1804]" captionTargetId="figure-83@27.[151,1432,257,1810]" captionTargetPageId="27" captionText="Fig. 15. Malayathele kanching sp. nov.; male holotype (A-B, D-G) and male paratype (C, H). (A) Distal part of right palp, ventral view. (B-C) Same of left palp, prolateral view. (D) Patella to metatarsus of left leg I, prolateral view. (E) Patella and tibia of left leg I, ventral view. (F) Patella to metatarsus of left leg II, ventral view. (G-H) Same, prolateral view. Arrow indicates single spine retroventrally-distally on patella I. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743987" httpUri="https://zenodo.org/record/5743987/files/figure.png" pageId="5" pageNumber="427">Fig. 15E</figureCitation>
); femur II with a short proventral band of hooked spinules (distant from distal margin of leg article; 
<figureCitation id="7DEB2A27C62CFF9211BBFC9C5461FCB7" box="[459,541,833,856]" captionStart="Fig" captionStartId="10.[151,186,1968,1989]" captionTargetBox="[159,1427,610,1932]" captionTargetId="figure-192@10.[153,1434,605,1947]" captionTargetPageId="10" captionText="Fig. 5. Leptothele chang sp. nov.; male holotype (A-B, E-H), female allotype (C-D) and male paratype (I). (A, C) Habitus, dorsal view. (B, D) Habitus, ventral view. (E) Femur to metatarsus of right leg II, prolateral view. (F) Right femur II, proventral view. (G) Right femur I, retrodorsal view. (H) Tarsus I, lateral view. (I) Distal part of median article and entire distal article of PLS, lateral view; arrow indicating pseudosegmentation. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743964" httpUri="https://zenodo.org/record/5743964/files/figure.png" pageId="5" pageNumber="427">Figs 5F</figureCitation>
, 
<figureCitation id="7DEB2A27C62CFF921258FC9C5424FCB7" box="[552,600,833,856]" captionStart="Fig" captionStartId="20.[151,186,2012,2033]" captionTargetBox="[157,1434,1019,1973]" captionTargetId="figure-450@20.[153,1438,1015,1976]" captionTargetPageId="20" captionText="Fig. 10. Malayathele cameronensis sp. nov., male holotype. (A) Left leg I, retrolateral view; one ventral spine on tibia fallen off. (B) Left leg II, prolateral view. (C) Unpaired claw and scopuliform hairs of left leg II, prolateral view. (D) Ventrally bulged tarsus of left leg II, prolateral view. (E) Scopuliform hair on left tarsus II. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743975" httpUri="https://zenodo.org/record/5743975/files/figure.png" pageId="5" pageNumber="427">10B</figureCitation>
); palpal tarsus with distinct distal spines (
<figureCitation id="7DEB2A27C62CFF921468FC9C52C8FCB7" box="[1048,1204,833,856]" captionStart="Fig" captionStartId="7.[152,187,1970,1991]" captionTargetBox="[157,1426,704,1913]" captionTargetId="figure-219@7.[151,1435,699,1917]" captionTargetPageId="7" captionText="Fig. 3. Leptothele bencha; males from Ko Sirey (A-B), Ko Tarutao (C-F, G, showing two specimens), Ko Libong (H) and Ko Sukon (I- J). (A, D, H) Distal part of palp, ventral view. (B, E) Right tibia and metatarsus II, prolateral view. (C) Left tibia and metatarsus II, ventral view. (F) Right tibia and metatarsus II, ventral view. (G) Palpal organ, ventral view. (I) Left tibia and metatarsus II of aberrant specimen, prolateral view. (J) Same, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5743960" httpUri="https://zenodo.org/record/5743960/files/figure.png" pageId="5" pageNumber="427">Figs 3A, D, H</figureCitation>
, 
<figureCitation id="7DEB2A27C62CFF9214CFFC9C52B0FCB7" box="[1215,1228,833,856]" captionStart="Fig" captionStartId="11.[151,186,1879,1900]" captionTargetBox="[163,1428,257,1828]" captionTargetId="figure-6@11.[151,1436,254,1838]" captionTargetPageId="11" captionText="Fig. 6. Leptothele chang sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A) Right palpal organ, ventral view. (B) Left palpal organ and palpal tarsus, prolateral view. (C) Right palpal organ and palpal tarsus, ventral view. (D) Left palpal organ and palpal tarsus, ventral view. (E) Right palpal organ and palpal tarsus, prolateral view. (F) Patella to metatarsus of left leg I, ventral view. (G) Patella to metatarsus of left leg II, prolateral view. (H) Same, ventral view. (I) Same of right leg II, ventral view. (J) Right tibia II, ventral view. (K) Left tibia II, ventral view. Scale lines 1.0 mm (A-E; F-K)." figureDoi="http://doi.org/10.5281/zenodo.5743966" httpUri="https://zenodo.org/record/5743966/files/figure.png" pageId="5" pageNumber="427">6</figureCitation>
B-E, 8A-E, 11A-D, 15A-C). Females without secondary receptacles (
<figureCitation id="7DEB2A27C62CFF92128FFCBC553BFC97" box="[767,839,865,888]" captionStart="Fig" captionStartId="8.[151,186,2027,2048]" captionTargetBox="[159,1431,770,1960]" captionTargetId="figure-262@8.[151,1435,763,1970]" captionTargetPageId="8" captionText="Fig. 4. Leptothele bencha; vulvae of six females, dorsal view. (A) Female from Ko Libong. (B-C) Two females from Ko Tarutao. (D) Female from Ko Lanta. (E-F) Two females from Ko Siray. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743962" httpUri="https://zenodo.org/record/5743962/files/figure.png" pageId="5" pageNumber="427">Figs 4</figureCitation>
, 
<figureCitation id="7DEB2A27C62CFF921326FCBC5519FC97" box="[854,869,865,888]" captionStart="Fig" captionStartId="12.[151,186,1998,2019]" captionTargetBox="[155,1430,1261,1937]" captionTargetId="figure-579@12.[151,1435,1255,1940]" captionTargetPageId="12" captionText="Fig. 7. Leptothele chang sp. nov.; vulvae of female allotype (A) and of three female paratypes (B-D), dorsal view. (A) Drawn from exuvia of allotype; cuticular gland duct bases visible. (B-D) Drawn from dissected paratypes; gland duct bases not visible but gland pores instead. (C) Right lateral receptacle carrying a tiny secondary receptacle. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743968" httpUri="https://zenodo.org/record/5743968/files/figure.png" pageId="5" pageNumber="427">7</figureCitation>
, 
<figureCitation id="7DEB2A27C62CFF921303FCBC55FEFC97" box="[883,898,865,888]" captionStart="Fig" captionStartId="17.[151,186,1715,1736]" captionTargetBox="[154,1431,263,1659]" captionTargetId="figure-71@17.[151,1435,259,1666]" captionTargetPageId="17" captionText="Fig. 9. Malayathele ulu sp. nov.; vulvae of seven females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B-C) Paratypes from the type locality (Ulu Groh). (D-G) Paratypes from the Cameron Highlands: Parit Falls, 26.IV.2009 (D); Tanah Rata, 7.VIII.1972 (E); Gunung Perdah, 2.-14.V.2009 (F); trails 4 &amp; 13, 23.III.1993, heads of median receptacles collapsed and right lateral receptacle torn off (G). Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743972" httpUri="https://zenodo.org/record/5743972/files/figure.png" pageId="5" pageNumber="427">9</figureCitation>
, 
<figureCitation id="7DEB2A27C62CFF9213E1FCBC55BCFC97" box="[913,960,865,888]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="5" pageNumber="427">11K</figureCitation>
, 
<figureCitation id="7DEB2A27C62CFF9213BEFCBC5597FC97" box="[974,1003,865,888]" captionStart="Fig" captionStartId="28.[152,187,2025,2046]" captionTargetBox="[157,1428,1213,1987]" captionTargetId="figure-544@28.[151,1431,1208,1994]" captionTargetPageId="28" captionText="Fig. 16. Malayathele kanching sp. nov.; vulvae of four female paratypes, dorsal (A-B) to increasingly more postero-dorsal view (C- D). (A, C-D) Specimens from the Templer Park. (B) Specimen from Ulu Gombak. Scale lines 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5743991" httpUri="https://zenodo.org/record/5743991/files/figure.png" pageId="5" pageNumber="427">16</figureCitation>
), if exceptionally present, then situated anterior of median receptacles (
<figureCitation id="7DEB2A27C62CFF921247FC5C54F0FC77" box="[567,652,897,920]" captionStart="Fig" captionStartId="12.[151,186,1998,2019]" captionTargetBox="[155,1430,1261,1937]" captionTargetId="figure-579@12.[151,1435,1255,1940]" captionTargetPageId="12" captionText="Fig. 7. Leptothele chang sp. nov.; vulvae of female allotype (A) and of three female paratypes (B-D), dorsal view. (A) Drawn from exuvia of allotype; cuticular gland duct bases visible. (B-D) Drawn from dissected paratypes; gland duct bases not visible but gland pores instead. (C) Right lateral receptacle carrying a tiny secondary receptacle. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743968" httpUri="https://zenodo.org/record/5743968/files/figure.png" pageId="5" pageNumber="427">Fig. 7C</figureCitation>
). Both sexes without spines on tarsi of posterior legs. Southern 
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and Peninsular 
<collectingCountry id="9DC77632C62CFF9211F7FC7C5795FC57" box="[391,489,929,952]" name="Malaysia" pageId="5" pageNumber="427">Malaysia</collectingCountry>
........................................................................................................................................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="5E242E07C62CFF9210E7FC3C53E1FAB7" pageId="5" pageNumber="427">
<paragraph id="E56F36A2C62CFF9210E7FC3C53E1FB97" blockId="5.[151,1437,993,1368]" pageId="5" pageNumber="427">
<keyLead id="5E219597C62CFF9210E7FC3C53E1FB97" pageId="5" pageNumber="427">
2A Small body size (carapace length in males 1.52-2.73), distal article of PLS pseudosegmented (
<figureCitation id="7DEB2A27C62CFF9214C5FC3C5281FC17" box="[1205,1277,993,1016]" captionStart="Fig" captionStartId="10.[151,186,1968,1989]" captionTargetBox="[159,1427,610,1932]" captionTargetId="figure-192@10.[153,1434,605,1947]" captionTargetPageId="10" captionText="Fig. 5. Leptothele chang sp. nov.; male holotype (A-B, E-H), female allotype (C-D) and male paratype (I). (A, C) Habitus, dorsal view. (B, D) Habitus, ventral view. (E) Femur to metatarsus of right leg II, prolateral view. (F) Right femur II, proventral view. (G) Right femur I, retrodorsal view. (H) Tarsus I, lateral view. (I) Distal part of median article and entire distal article of PLS, lateral view; arrow indicating pseudosegmentation. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743964" httpUri="https://zenodo.org/record/5743964/files/figure.png" pageId="5" pageNumber="427">Fig. 5I</figureCitation>
), no metatarsal preening combs in both sexes. Males with at least two megaspines and with a transversal subdistal ridge on ventral side of tibia II (
<figureCitation id="7DEB2A27C62CFF9211F3FBFC57BFFBD7" box="[387,451,1057,1080]" captionStart="Fig" captionStartId="7.[152,187,1970,1991]" captionTargetBox="[157,1426,704,1913]" captionTargetId="figure-219@7.[151,1435,699,1917]" captionTargetPageId="7" captionText="Fig. 3. Leptothele bencha; males from Ko Sirey (A-B), Ko Tarutao (C-F, G, showing two specimens), Ko Libong (H) and Ko Sukon (I- J). (A, D, H) Distal part of palp, ventral view. (B, E) Right tibia and metatarsus II, prolateral view. (C) Left tibia and metatarsus II, ventral view. (F) Right tibia and metatarsus II, ventral view. (G) Palpal organ, ventral view. (I) Left tibia and metatarsus II of aberrant specimen, prolateral view. (J) Same, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5743960" httpUri="https://zenodo.org/record/5743960/files/figure.png" pageId="5" pageNumber="427">Figs 3</figureCitation>
B-C, E-F, I-J, 5E, 6G-K); metatarsus II with a single conical or mound-like process on ventral side (
<figureCitation id="7DEB2A27C62CFF92116BFB9C5720FBB7" box="[283,348,1089,1112]" captionStart="Fig" captionStartId="7.[152,187,1970,1991]" captionTargetBox="[157,1426,704,1913]" captionTargetId="figure-219@7.[151,1435,699,1917]" captionTargetPageId="7" captionText="Fig. 3. Leptothele bencha; males from Ko Sirey (A-B), Ko Tarutao (C-F, G, showing two specimens), Ko Libong (H) and Ko Sukon (I- J). (A, D, H) Distal part of palp, ventral view. (B, E) Right tibia and metatarsus II, prolateral view. (C) Left tibia and metatarsus II, ventral view. (F) Right tibia and metatarsus II, ventral view. (G) Palpal organ, ventral view. (I) Left tibia and metatarsus II of aberrant specimen, prolateral view. (J) Same, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5743960" httpUri="https://zenodo.org/record/5743960/files/figure.png" pageId="5" pageNumber="427">Figs 3</figureCitation>
B-C, E-F, I-J, 5E, 6G-I); tarsi I-II not ventrally bulged (
<figureCitation id="7DEB2A27C62CFF9213D4FB9C5586FBB7" box="[932,1018,1089,1112]" captionStart="Fig" captionStartId="10.[151,186,1968,1989]" captionTargetBox="[159,1427,610,1932]" captionTargetId="figure-192@10.[153,1434,605,1947]" captionTargetPageId="10" captionText="Fig. 5. Leptothele chang sp. nov.; male holotype (A-B, E-H), female allotype (C-D) and male paratype (I). (A, C) Habitus, dorsal view. (B, D) Habitus, ventral view. (E) Femur to metatarsus of right leg II, prolateral view. (F) Right femur II, proventral view. (G) Right femur I, retrodorsal view. (H) Tarsus I, lateral view. (I) Distal part of median article and entire distal article of PLS, lateral view; arrow indicating pseudosegmentation. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743964" httpUri="https://zenodo.org/record/5743964/files/figure.png" pageId="5" pageNumber="427">Fig. 5H</figureCitation>
). Females with quite long spermathecal trunks; stalks of median receptacles not sclerotised (
<figureCitation id="7DEB2A27C62CFF92137FFBBC5528FB97" box="[783,852,1121,1144]" captionStart="Fig" captionStartId="8.[151,186,2027,2048]" captionTargetBox="[159,1431,770,1960]" captionTargetId="figure-262@8.[151,1435,763,1970]" captionTargetPageId="8" captionText="Fig. 4. Leptothele bencha; vulvae of six females, dorsal view. (A) Female from Ko Libong. (B-C) Two females from Ko Tarutao. (D) Female from Ko Lanta. (E-F) Two females from Ko Siray. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743962" httpUri="https://zenodo.org/record/5743962/files/figure.png" pageId="5" pageNumber="427">Figs 4</figureCitation>
, 
<figureCitation id="7DEB2A27C62CFF921310FBBC550CFB97" box="[864,880,1121,1144]" captionStart="Fig" captionStartId="12.[151,186,1998,2019]" captionTargetBox="[155,1430,1261,1937]" captionTargetId="figure-579@12.[151,1435,1255,1940]" captionTargetPageId="12" captionText="Fig. 7. Leptothele chang sp. nov.; vulvae of female allotype (A) and of three female paratypes (B-D), dorsal view. (A) Drawn from exuvia of allotype; cuticular gland duct bases visible. (B-D) Drawn from dissected paratypes; gland duct bases not visible but gland pores instead. (C) Right lateral receptacle carrying a tiny secondary receptacle. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743968" httpUri="https://zenodo.org/record/5743968/files/figure.png" pageId="5" pageNumber="427">7</figureCitation>
). Southern 
<collectingCountry id="9DC77632C62CFF92139AFBBC5235FB97" box="[1002,1097,1121,1144]" name="Thailand" pageId="5" pageNumber="427">Thailand</collectingCountry>
................................ 
<taxonomicName id="22D04D21C62CFF92155CFBBC53E1FB97" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[1324,1437,1121,1144]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="D7A4EAB0C62CFF92155CFBBC53E1FB97" box="[1324,1437,1121,1144]" italics="true" pageId="5" pageNumber="427">Leptothele</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="E56F36A2C62CFF9210E7FB5C53E1FAB7" blockId="5.[151,1437,993,1368]" pageId="5" pageNumber="427">
<keyLead id="5E219597C62CFF9210E7FB5C53E1FAB7" pageId="5" pageNumber="427">
2B Very small body size (carapace length in males 1.11-1.44), distal article of PLS not pseudosegmented (
<figureCitation id="7DEB2A27C62CFF92156BFB5C531AFB77" box="[1307,1382,1153,1176]" captionStart="Fig" captionStartId="23.[151,186,1913,1934]" captionTargetBox="[157,1433,261,1852]" captionTargetId="figure-85@23.[153,1438,257,1862]" captionTargetPageId="23" captionText="Fig. 12. Malayathele maculosa sp. nov., female paratype from Genting Highlands (A-D), M. kanching sp. nov., male holotype (E-G), female allotype (H-I) and male paratype (J-K). (A, E, H) Habitus, dorsal view. (B, F, I) Habitus, ventral view. (C) Habitus, dorsolateral view. (D) Opisthosoma, posteroventral view. (G) Tibia and metatarsus of right leg II, prolateral view. (J) Distal part of proximal article and entire median and distal articles of PLS, lateral view. (K) Same, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743981" httpUri="https://zenodo.org/record/5743981/files/figure.png" pageId="5" pageNumber="427">Fig. 12</figureCitation>
J-K), metatarsal preening combs present on legs II-IV in both sexes. Males with only one ventral megaspine and without a subdistal ridge on ventral side of tibia II (
<figureCitation id="7DEB2A27C62CFF9212C4FB1C548BFB37" box="[692,759,1217,1240]" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="5" pageNumber="427">Figs 8</figureCitation>
H-K, 10B, 11G-J, 12G, 13J-M, 15F-H); metatarsus II with two small conical processes, a proventral and a retroventral one (
<figureCitation id="7DEB2A27C62CFF92131BFB3C55C7FB17" box="[875,955,1249,1272]" captionStart="Fig" captionStartId="24.[149,184,1934,1955]" captionTargetBox="[152,1427,583,1872]" captionTargetId="figure-267@24.[149,1433,576,1879]" captionTargetPageId="24" captionText="Fig. 13. Malayathele maculosa sp. nov., male holotype (A-C, H-L) and two male paratypes (D-E; F-G, M). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C, E, G) Palpal organ of right palp, prolateral view. (D, F) Palpal organ of left palp, prolateral view. (H) Patella to metatarsus of left leg I, prolateral view. (I) Patella to tibia of left leg I, ventral view. (J) Patella to metatarsus of right leg II, ventral view. (K) Same of left leg II, prolateral view. (L) Same of right leg II, retrolateral view. (M) Same of left leg II, retrodorsal view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743983" httpUri="https://zenodo.org/record/5743983/files/figure.png" pageId="5" pageNumber="427">Figs 13</figureCitation>
J-K, M, 15F-H), or with only a small conical retroventral process (
<figureCitation id="7DEB2A27C62CFF9211B7FADC5479FAF7" box="[455,517,1281,1304]" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="5" pageNumber="427">Fig. 8</figureCitation>
I-J), or with a small conical proventral and a keel-shaped retroventral processes (
<figureCitation id="7DEB2A27C62CFF921501FADC5680FAD7" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="5" pageNumber="427">Fig. 11</figureCitation>
G-J); tarsi I-II ventrally bulged (
<figureCitation id="7DEB2A27C62CFF921213FAFC54B5FAD7" box="[611,713,1313,1336]" captionStart="Fig" captionStartId="20.[151,186,2012,2033]" captionTargetBox="[157,1434,1019,1973]" captionTargetId="figure-450@20.[153,1438,1015,1976]" captionTargetPageId="20" captionText="Fig. 10. Malayathele cameronensis sp. nov., male holotype. (A) Left leg I, retrolateral view; one ventral spine on tibia fallen off. (B) Left leg II, prolateral view. (C) Unpaired claw and scopuliform hairs of left leg II, prolateral view. (D) Ventrally bulged tarsus of left leg II, prolateral view. (E) Scopuliform hair on left tarsus II. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743975" httpUri="https://zenodo.org/record/5743975/files/figure.png" pageId="5" pageNumber="427">Fig. 10D</figureCitation>
). Females with rather short spermathecal trunks; stalks of median receptacles sclerotised (
<figureCitation id="7DEB2A27C62CFF921190FA9C5459FAB7" box="[480,549,1345,1368]" captionStart="Fig" captionStartId="17.[151,186,1715,1736]" captionTargetBox="[154,1431,263,1659]" captionTargetId="figure-71@17.[151,1435,259,1666]" captionTargetPageId="17" captionText="Fig. 9. Malayathele ulu sp. nov.; vulvae of seven females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B-C) Paratypes from the type locality (Ulu Groh). (D-G) Paratypes from the Cameron Highlands: Parit Falls, 26.IV.2009 (D); Tanah Rata, 7.VIII.1972 (E); Gunung Perdah, 2.-14.V.2009 (F); trails 4 &amp; 13, 23.III.1993, heads of median receptacles collapsed and right lateral receptacle torn off (G). Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743972" httpUri="https://zenodo.org/record/5743972/files/figure.png" pageId="5" pageNumber="427">Figs 9</figureCitation>
, 
<figureCitation id="7DEB2A27C62CFF921241FA9C541CFAB7" box="[561,608,1345,1368]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="5" pageNumber="427">11K</figureCitation>
, 
<figureCitation id="7DEB2A27C62CFF92121BFA9C54F4FAB7" box="[619,648,1345,1368]" captionStart="Fig" captionStartId="25.[151,186,2027,2048]" captionTargetBox="[155,1418,1378,1967]" captionTargetId="figure-608@25.[151,1422,1374,1976]" captionTargetPageId="25" captionText="Fig. 14. Malayathele maculosa sp. nov., vulvae of three females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B) Paratype from Genting Highlands. (C) Juvenile female paratype from Fraser’s Hill. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743985" httpUri="https://zenodo.org/record/5743985/files/figure.png" pageId="5" pageNumber="427">14</figureCitation>
, 
<figureCitation id="7DEB2A27C62CFF9212E3FA9C54CDFAB7" box="[659,689,1345,1368]" captionStart="Fig" captionStartId="28.[152,187,2025,2046]" captionTargetBox="[157,1428,1213,1987]" captionTargetId="figure-544@28.[151,1431,1208,1994]" captionTargetPageId="28" captionText="Fig. 16. Malayathele kanching sp. nov.; vulvae of four female paratypes, dorsal (A-B) to increasingly more postero-dorsal view (C- D). (A, C-D) Specimens from the Templer Park. (B) Specimen from Ulu Gombak. Scale lines 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5743991" httpUri="https://zenodo.org/record/5743991/files/figure.png" pageId="5" pageNumber="427">16</figureCitation>
). Peninsular 
<collectingCountry id="9DC77632C62CFF92134BFA9C55E1FAB7" box="[827,925,1345,1368]" name="Malaysia" pageId="5" pageNumber="427">Malaysia</collectingCountry>
........................................ 
<taxonomicName id="22D04D21C62CFF9214C0FA9C5349FAB7" ID-CoL="98X62" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[1200,1333,1345,1368]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="427" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D7A4EAB0C62CFF9214C0FA9C5349FAB7" box="[1200,1333,1345,1368]" italics="true" pageId="5" pageNumber="427">Malayathele</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62CFF92154BFA9C53E1FAB7" box="[1339,1437,1345,1368]" pageId="5" pageNumber="427" rank="genus">gen. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
<paragraph id="E56F36A2C62CFF9210E7FA1C547DFA37" blockId="5.[151,513,1472,1496]" box="[151,513,1472,1496]" pageId="5" pageNumber="427">
<emphasis id="D7A4EAB0C62CFF9210E7FA1C547DFA37" bold="true" box="[151,513,1472,1496]" pageId="5" pageNumber="427">
Key to the species of 
<taxonomicName id="22D04D21C62CFF9211F5FA1D5785FA37" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[389,505,1472,1496]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="D7A4EAB0C62CFF9211F5FA1D5785FA37" bold="true" box="[389,505,1472,1496]" italics="true" pageId="5" pageNumber="427">Leptothele</emphasis>
</taxonomicName>
:
</emphasis>
</paragraph>
<paragraph id="E56F36A2C62CFF9210E7F9DC53E0F957" blockId="5.[151,1437,1537,1880]" pageId="5" pageNumber="427">
<key id="91416193C62CFF9210E7F9DC53E0F957" pageId="5" pageNumber="427">
<keyLead id="5E219597C62CFF9210E7F9DC53E0F957" pageId="5" pageNumber="427">
<keyStep id="5E242E07C62CFF9210E7F9DC53E0F957" pageId="5" pageNumber="427">
1A Small body size (carapace lengths in males 1.52-2.04). Males normally with two ventral megaspines (
<figureCitation id="7DEB2A27C62CFF921557F9DC53E4F9F7" box="[1319,1432,1537,1560]" captionStart="Fig" captionStartId="7.[152,187,1970,1991]" captionTargetBox="[157,1426,704,1913]" captionTargetId="figure-219@7.[151,1435,699,1917]" captionTargetPageId="7" captionText="Fig. 3. Leptothele bencha; males from Ko Sirey (A-B), Ko Tarutao (C-F, G, showing two specimens), Ko Libong (H) and Ko Sukon (I- J). (A, D, H) Distal part of palp, ventral view. (B, E) Right tibia and metatarsus II, prolateral view. (C) Left tibia and metatarsus II, ventral view. (F) Right tibia and metatarsus II, ventral view. (G) Palpal organ, ventral view. (I) Left tibia and metatarsus II of aberrant specimen, prolateral view. (J) Same, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5743960" httpUri="https://zenodo.org/record/5743960/files/figure.png" pageId="5" pageNumber="427">Fig. 3C, F</figureCitation>
; single asymmetrical exception, 
<figureCitation id="7DEB2A27C62CFF921243F9FC5400F9D7" box="[563,636,1569,1592]" captionStart="Fig" captionStartId="7.[152,187,1970,1991]" captionTargetBox="[157,1426,704,1913]" captionTargetId="figure-219@7.[151,1435,699,1917]" captionTargetPageId="7" captionText="Fig. 3. Leptothele bencha; males from Ko Sirey (A-B), Ko Tarutao (C-F, G, showing two specimens), Ko Libong (H) and Ko Sukon (I- J). (A, D, H) Distal part of palp, ventral view. (B, E) Right tibia and metatarsus II, prolateral view. (C) Left tibia and metatarsus II, ventral view. (F) Right tibia and metatarsus II, ventral view. (G) Palpal organ, ventral view. (I) Left tibia and metatarsus II of aberrant specimen, prolateral view. (J) Same, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5743960" httpUri="https://zenodo.org/record/5743960/files/figure.png" pageId="5" pageNumber="427">Fig. 3J</figureCitation>
) and always without wrinkles behind and in front of subdistal-ventral ridge on tibia II (
<figureCitation id="7DEB2A27C62CFF92112FF99C5795F9B7" box="[351,489,1601,1624]" captionStart="Fig" captionStartId="7.[152,187,1970,1991]" captionTargetBox="[157,1426,704,1913]" captionTargetId="figure-219@7.[151,1435,699,1917]" captionTargetPageId="7" captionText="Fig. 3. Leptothele bencha; males from Ko Sirey (A-B), Ko Tarutao (C-F, G, showing two specimens), Ko Libong (H) and Ko Sukon (I- J). (A, D, H) Distal part of palp, ventral view. (B, E) Right tibia and metatarsus II, prolateral view. (C) Left tibia and metatarsus II, ventral view. (F) Right tibia and metatarsus II, ventral view. (G) Palpal organ, ventral view. (I) Left tibia and metatarsus II of aberrant specimen, prolateral view. (J) Same, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5743960" httpUri="https://zenodo.org/record/5743960/files/figure.png" pageId="5" pageNumber="427">Fig. 3C, F, J</figureCitation>
); ventral process on metatarsus II widely conical (
<figureCitation id="7DEB2A27C62CFF92147DF99C52E4F9B7" box="[1037,1176,1601,1624]" captionStart="Fig" captionStartId="7.[152,187,1970,1991]" captionTargetBox="[157,1426,704,1913]" captionTargetId="figure-219@7.[151,1435,699,1917]" captionTargetPageId="7" captionText="Fig. 3. Leptothele bencha; males from Ko Sirey (A-B), Ko Tarutao (C-F, G, showing two specimens), Ko Libong (H) and Ko Sukon (I- J). (A, D, H) Distal part of palp, ventral view. (B, E) Right tibia and metatarsus II, prolateral view. (C) Left tibia and metatarsus II, ventral view. (F) Right tibia and metatarsus II, ventral view. (G) Palpal organ, ventral view. (I) Left tibia and metatarsus II of aberrant specimen, prolateral view. (J) Same, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5743960" httpUri="https://zenodo.org/record/5743960/files/figure.png" pageId="5" pageNumber="427">Fig. 3B, E, I</figureCitation>
); bulbous part of palpal organ as long as wide or wider than long (
<figureCitation id="7DEB2A27C62CFF9212DFF9BC5511F997" box="[687,877,1633,1656]" captionStart="Fig" captionStartId="7.[152,187,1970,1991]" captionTargetBox="[157,1426,704,1913]" captionTargetId="figure-219@7.[151,1435,699,1917]" captionTargetPageId="7" captionText="Fig. 3. Leptothele bencha; males from Ko Sirey (A-B), Ko Tarutao (C-F, G, showing two specimens), Ko Libong (H) and Ko Sukon (I- J). (A, D, H) Distal part of palp, ventral view. (B, E) Right tibia and metatarsus II, prolateral view. (C) Left tibia and metatarsus II, ventral view. (F) Right tibia and metatarsus II, ventral view. (G) Palpal organ, ventral view. (I) Left tibia and metatarsus II of aberrant specimen, prolateral view. (J) Same, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5743960" httpUri="https://zenodo.org/record/5743960/files/figure.png" pageId="5" pageNumber="427">Fig. 3A, D, H, G</figureCitation>
). Females with quite long and narrow spermathecal trunks; stalks of median receptacles mostly shorter than corresponding heads (
<figureCitation id="7DEB2A27C62CFF92146CF95C5227F977" box="[1052,1115,1665,1688]" captionStart="Fig" captionStartId="8.[151,186,2027,2048]" captionTargetBox="[159,1431,770,1960]" captionTargetId="figure-262@8.[151,1435,763,1970]" captionTargetPageId="8" captionText="Fig. 4. Leptothele bencha; vulvae of six females, dorsal view. (A) Female from Ko Libong. (B-C) Two females from Ko Tarutao. (D) Female from Ko Lanta. (E-F) Two females from Ko Siray. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743962" httpUri="https://zenodo.org/record/5743962/files/figure.png" pageId="5" pageNumber="427">Fig. 4</figureCitation>
A-D), except for females from Ko Siray (
<figureCitation id="7DEB2A27C62CFF921121F97C57ECF957" box="[337,400,1697,1720]" captionStart="Fig" captionStartId="8.[151,186,2027,2048]" captionTargetBox="[159,1431,770,1960]" captionTargetId="figure-262@8.[151,1435,763,1970]" captionTargetPageId="8" captionText="Fig. 4. Leptothele bencha; vulvae of six females, dorsal view. (A) Female from Ko Libong. (B-C) Two females from Ko Tarutao. (D) Female from Ko Lanta. (E-F) Two females from Ko Siray. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743962" httpUri="https://zenodo.org/record/5743962/files/figure.png" pageId="5" pageNumber="427">Fig. 4</figureCitation>
E-F) .................................................................................................................................. 
<taxonomicName id="22D04D21C62CFF921543F97C53E0F957" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[1331,1436,1697,1720]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="427" phylum="Arthropoda" rank="species" species="bencha">
<emphasis id="D7A4EAB0C62CFF921543F97C53E0F957" box="[1331,1436,1697,1720]" italics="true" pageId="5" pageNumber="427">L. bencha</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</key>
</paragraph>
<paragraph id="E56F36A2C62CFF9110E7F91C5763FB39" blockId="5.[151,1437,1537,1880]" lastBlockId="6.[151,757,258,1238]" lastPageId="6" lastPageNumber="428" pageId="5" pageNumber="427">
<keyLead id="5E219597C62CFF9210E7F91C53E1F8B7" pageId="5" pageNumber="427">
<keyStep id="5E242E07C62CFF9210E7F91C53E1F8B7" pageId="5" pageNumber="427">
1B Larger body size (carapace lengths in males 2.30-2.73). Males with 2-3 ventral megaspines and with wrinkles behind and in front of subdistal ridge on tibia II (
<figureCitation id="7DEB2A27C62CFF921367F93C5526F917" box="[791,858,1761,1784]" captionStart="Fig" captionStartId="11.[151,186,1879,1900]" captionTargetBox="[163,1428,257,1828]" captionTargetId="figure-6@11.[151,1436,254,1838]" captionTargetPageId="11" captionText="Fig. 6. Leptothele chang sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A) Right palpal organ, ventral view. (B) Left palpal organ and palpal tarsus, prolateral view. (C) Right palpal organ and palpal tarsus, ventral view. (D) Left palpal organ and palpal tarsus, ventral view. (E) Right palpal organ and palpal tarsus, prolateral view. (F) Patella to metatarsus of left leg I, ventral view. (G) Patella to metatarsus of left leg II, prolateral view. (H) Same, ventral view. (I) Same of right leg II, ventral view. (J) Right tibia II, ventral view. (K) Left tibia II, ventral view. Scale lines 1.0 mm (A-E; F-K)." figureDoi="http://doi.org/10.5281/zenodo.5743966" httpUri="https://zenodo.org/record/5743966/files/figure.png" pageId="5" pageNumber="427">Fig. 6</figureCitation>
H-K); ventral process on metatarsus II less distinct, developed as a low, wide mound (
<figureCitation id="7DEB2A27C62CFF921225F8DC54D0F8F7" box="[597,684,1793,1816]" captionStart="Fig" captionStartId="11.[151,186,1879,1900]" captionTargetBox="[163,1428,257,1828]" captionTargetId="figure-6@11.[151,1436,254,1838]" captionTargetPageId="11" captionText="Fig. 6. Leptothele chang sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A) Right palpal organ, ventral view. (B) Left palpal organ and palpal tarsus, prolateral view. (C) Right palpal organ and palpal tarsus, ventral view. (D) Left palpal organ and palpal tarsus, ventral view. (E) Right palpal organ and palpal tarsus, prolateral view. (F) Patella to metatarsus of left leg I, ventral view. (G) Patella to metatarsus of left leg II, prolateral view. (H) Same, ventral view. (I) Same of right leg II, ventral view. (J) Right tibia II, ventral view. (K) Left tibia II, ventral view. Scale lines 1.0 mm (A-E; F-K)." figureDoi="http://doi.org/10.5281/zenodo.5743966" httpUri="https://zenodo.org/record/5743966/files/figure.png" pageId="5" pageNumber="427">Fig. 6G</figureCitation>
); bulbous part of palpal organ longer than wide (
<figureCitation id="7DEB2A27C62CFF9214CFF8DC5283F8F7" box="[1215,1279,1793,1816]" captionStart="Fig" captionStartId="11.[151,186,1879,1900]" captionTargetBox="[163,1428,257,1828]" captionTargetId="figure-6@11.[151,1436,254,1838]" captionTargetPageId="11" captionText="Fig. 6. Leptothele chang sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A) Right palpal organ, ventral view. (B) Left palpal organ and palpal tarsus, prolateral view. (C) Right palpal organ and palpal tarsus, ventral view. (D) Left palpal organ and palpal tarsus, ventral view. (E) Right palpal organ and palpal tarsus, prolateral view. (F) Patella to metatarsus of left leg I, ventral view. (G) Patella to metatarsus of left leg II, prolateral view. (H) Same, ventral view. (I) Same of right leg II, ventral view. (J) Right tibia II, ventral view. (K) Left tibia II, ventral view. Scale lines 1.0 mm (A-E; F-K)." figureDoi="http://doi.org/10.5281/zenodo.5743966" httpUri="https://zenodo.org/record/5743966/files/figure.png" pageId="5" pageNumber="427">Fig. 6</figureCitation>
A-E). Females with relatively shorter and wider spermathecal trunks; stalks of median receptacles very long and thin, almost twice of corresponding head lengths (
<figureCitation id="7DEB2A27C62CFF921204F89C54C8F8B7" box="[628,692,1857,1880]" captionStart="Fig" captionStartId="12.[151,186,1998,2019]" captionTargetBox="[155,1430,1261,1937]" captionTargetId="figure-579@12.[151,1435,1255,1940]" captionTargetPageId="12" captionText="Fig. 7. Leptothele chang sp. nov.; vulvae of female allotype (A) and of three female paratypes (B-D), dorsal view. (A) Drawn from exuvia of allotype; cuticular gland duct bases visible. (B-D) Drawn from dissected paratypes; gland duct bases not visible but gland pores instead. (C) Right lateral receptacle carrying a tiny secondary receptacle. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743968" httpUri="https://zenodo.org/record/5743968/files/figure.png" pageId="5" pageNumber="427">Fig. 7</figureCitation>
) ................................................................................. 
<taxonomicName id="22D04D21C62CFF921496F89C533FF8B7" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[1254,1347,1857,1880]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="427" phylum="Arthropoda" rank="species" species="chang" status="sp. nov.">
<emphasis id="D7A4EAB0C62CFF921496F89C533FF8B7" box="[1254,1347,1857,1880]" italics="true" pageId="5" pageNumber="427">L. chang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62CFF92153AF89C53E1F8B7" box="[1354,1437,1857,1880]" pageId="5" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
</keyStep>
</keyLead>
vulvae quite similar to those of 
<taxonomicName id="22D04D21C62FFF911187FEDF54EDFEF6" authorityName="Simon" authorityYear="1889" box="[503,657,258,281]" class="Arachnida" family="Dipluridae" genus="Phyxioschema" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="428" phylum="Arthropoda" rank="genus">
<emphasis id="D7A4EAB0C62FFF911187FEDF54EDFEF6" box="[503,657,258,281]" italics="true" pageId="6" pageNumber="428">Phyxioschema</emphasis>
</taxonomicName>
females, but stalks of receptacles not sclerotised and usually no secondary receptacles present [one exception – unsymmetrical and situated anterior of median receptacle – in 
<taxonomicName id="22D04D21C62FFF911135FE5C57D9FE77" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[325,421,385,408]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="428" phylum="Arthropoda" rank="species" species="chang" status="sp. nov.">
<emphasis id="D7A4EAB0C62FFF911135FE5C57D9FE77" box="[325,421,385,408]" italics="true" pageId="6" pageNumber="428">L. chang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62FFF9111C0FE5C547AFE77" box="[432,518,385,408]" pageId="6" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="7DEB2A27C62FFF911268FE5C540CFE77" box="[536,624,385,408]" captionStart="Fig" captionStartId="12.[151,186,1998,2019]" captionTargetBox="[155,1430,1261,1937]" captionTargetId="figure-579@12.[151,1435,1255,1940]" captionTargetPageId="12" captionText="Fig. 7. Leptothele chang sp. nov.; vulvae of female allotype (A) and of three female paratypes (B-D), dorsal view. (A) Drawn from exuvia of allotype; cuticular gland duct bases visible. (B-D) Drawn from dissected paratypes; gland duct bases not visible but gland pores instead. (C) Right lateral receptacle carrying a tiny secondary receptacle. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743968" httpUri="https://zenodo.org/record/5743968/files/figure.png" pageId="6" pageNumber="428">Fig. 7C</figureCitation>
); secondary receptacles in 
<taxonomicName id="22D04D21C62FFF911145FE7D57B3FE58" authorityName="Simon" authorityYear="1889" box="[309,463,416,439]" class="Arachnida" family="Dipluridae" genus="Phyxioschema" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="428" phylum="Arthropoda" rank="genus">
<emphasis id="D7A4EAB0C62FFF911145FE7D57B3FE58" box="[309,463,416,439]" italics="true" pageId="6" pageNumber="428">Phyxioschema</emphasis>
</taxonomicName>
, if present, posterior of or at same level as median receptacle, see 
<bibRefCitation id="81414B53C62FFF911226FE1D56ACFE18" author="Schwendinger P. J." pageId="6" pageNumber="428" pagination="1 - 40" refId="ref23293" refString="Schwendinger P. J. 2009. A taxonomic revision of the genus Phyxioschema (Araneae, Dipluridae), I: species from Thailand. Zootaxa 2126: 1 - 40." type="journal article" year="2009">Schwendinger, 2009</bibRefCitation>
: figs 3A-I, 11, 17 and 
<bibRefCitation id="81414B53C62FFF9111BFFE3D56ACFDF9" author="Schwendinger P. J. &amp; Zonstein S. L." pageId="6" pageNumber="428" pagination="28 - 48" refId="ref23412" refString="Schwendinger P. J., Zonstein S. L. 2011. A taxonomic revision of the genus Phyxioschema (Araneae, Dipluridae), II: species from Central Asia. Zootaxa 2815: 28 - 48." type="journal article" year="2011">Schwendinger &amp; Zonstein, 2011</bibRefCitation>
: figs 4-5, 8]. Metatarsal preening combs absent (present in 
<taxonomicName id="22D04D21C62FFF911166FDC257DEFDD9" authorityName="Raven &amp; Schwendinger" authorityYear="1989" box="[278,418,543,566]" class="Arachnida" family="Dipluridae" genus="Phyxioschema" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="428" phylum="Arthropoda" rank="species" species="suthepium">
<emphasis id="D7A4EAB0C62FFF911166FDC257DEFDD9" box="[278,418,543,566]" italics="true" pageId="6" pageNumber="428">P. suthepium</emphasis>
</taxonomicName>
and 
<taxonomicName id="22D04D21C62FFF9111ADFDC256B3FDB9" authority="Schwendinger, 2009" authorityName="Schwendinger" authorityYear="2009" class="Arachnida" family="Dipluridae" genus="Phyxioschema" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="428" phylum="Arthropoda" rank="species" species="erawan">
<emphasis id="D7A4EAB0C62FFF9111ADFDC25437FDD9" box="[477,587,543,566]" italics="true" pageId="6" pageNumber="428">P. erawan</emphasis>
<bibRefCitation id="81414B53C62FFF911225FDC256B3FDB9" author="Schwendinger P. J." pageId="6" pageNumber="428" pagination="1 - 40" refId="ref23293" refString="Schwendinger P. J. 2009. A taxonomic revision of the genus Phyxioschema (Araneae, Dipluridae), I: species from Thailand. Zootaxa 2126: 1 - 40." type="journal article" year="2009">Schwendinger, 2009</bibRefCitation>
</taxonomicName>
, but not in other 
<taxonomicName id="22D04D21C62FFF9111D6FDE2543DFDB9" authorityName="Simon" authorityYear="1889" box="[422,577,575,598]" class="Arachnida" family="Dipluridae" genus="Phyxioschema" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="428" phylum="Arthropoda" rank="genus">
<emphasis id="D7A4EAB0C62FFF9111D6FDE2543DFDB9" box="[422,577,575,598]" italics="true" pageId="6" pageNumber="428">Phyxioschema</emphasis>
</taxonomicName>
). Distinguished from 
<taxonomicName id="22D04D21C62FFF9110A9FD83571CFD9A" ID-CoL="98X62" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[217,352,606,629]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="428" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D7A4EAB0C62FFF9110A9FD83571CFD9A" box="[217,352,606,629]" italics="true" pageId="6" pageNumber="428">Malayathele</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62FFF91111CFD8357A9FD9A" box="[364,469,606,629]" pageId="6" pageNumber="428" rank="genus">gen. nov.</taxonomicNameLabel>
by distal article of PLS with a pseudosegmentation (
<figureCitation id="7DEB2A27C62FFF911194FDA35449FD7A" box="[484,565,638,661]" captionStart="Fig" captionStartId="10.[151,186,1968,1989]" captionTargetBox="[159,1427,610,1932]" captionTargetId="figure-192@10.[153,1434,605,1947]" captionTargetPageId="10" captionText="Fig. 5. Leptothele chang sp. nov.; male holotype (A-B, E-H), female allotype (C-D) and male paratype (I). (A, C) Habitus, dorsal view. (B, D) Habitus, ventral view. (E) Femur to metatarsus of right leg II, prolateral view. (F) Right femur II, proventral view. (G) Right femur I, retrodorsal view. (H) Tarsus I, lateral view. (I) Distal part of median article and entire distal article of PLS, lateral view; arrow indicating pseudosegmentation. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743964" httpUri="https://zenodo.org/record/5743964/files/figure.png" pageId="6" pageNumber="428">Fig. 5I</figureCitation>
cf. 
<figureCitation id="7DEB2A27C62FFF91121FFDA354BEFD7A" box="[623,706,638,661]" captionStart="Fig" captionStartId="23.[151,186,1913,1934]" captionTargetBox="[157,1433,261,1852]" captionTargetId="figure-85@23.[153,1438,257,1862]" captionTargetPageId="23" captionText="Fig. 12. Malayathele maculosa sp. nov., female paratype from Genting Highlands (A-D), M. kanching sp. nov., male holotype (E-G), female allotype (H-I) and male paratype (J-K). (A, E, H) Habitus, dorsal view. (B, F, I) Habitus, ventral view. (C) Habitus, dorsolateral view. (D) Opisthosoma, posteroventral view. (G) Tibia and metatarsus of right leg II, prolateral view. (J) Distal part of proximal article and entire median and distal articles of PLS, lateral view. (K) Same, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743981" httpUri="https://zenodo.org/record/5743981/files/figure.png" pageId="6" pageNumber="428">Fig. 12</figureCitation>
J-K) and by lacking metatarsal preening combs in both sexes; males with 2-3 (
<figureCitation id="7DEB2A27C62FFF9111E4FD605476FD3B" box="[404,522,701,724]" captionStart="Fig" captionStartId="7.[152,187,1970,1991]" captionTargetBox="[157,1426,704,1913]" captionTargetId="figure-219@7.[151,1435,699,1917]" captionTargetPageId="7" captionText="Fig. 3. Leptothele bencha; males from Ko Sirey (A-B), Ko Tarutao (C-F, G, showing two specimens), Ko Libong (H) and Ko Sukon (I- J). (A, D, H) Distal part of palp, ventral view. (B, E) Right tibia and metatarsus II, prolateral view. (C) Left tibia and metatarsus II, ventral view. (F) Right tibia and metatarsus II, ventral view. (G) Palpal organ, ventral view. (I) Left tibia and metatarsus II of aberrant specimen, prolateral view. (J) Same, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5743960" httpUri="https://zenodo.org/record/5743960/files/figure.png" pageId="6" pageNumber="428">Figs 3C, F</figureCitation>
, 
<figureCitation id="7DEB2A27C62FFF911267FD605458FD3B" box="[535,548,701,724]" captionStart="Fig" captionStartId="11.[151,186,1879,1900]" captionTargetBox="[163,1428,257,1828]" captionTargetId="figure-6@11.[151,1436,254,1838]" captionTargetPageId="11" captionText="Fig. 6. Leptothele chang sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A) Right palpal organ, ventral view. (B) Left palpal organ and palpal tarsus, prolateral view. (C) Right palpal organ and palpal tarsus, ventral view. (D) Left palpal organ and palpal tarsus, ventral view. (E) Right palpal organ and palpal tarsus, prolateral view. (F) Patella to metatarsus of left leg I, ventral view. (G) Patella to metatarsus of left leg II, prolateral view. (H) Same, ventral view. (I) Same of right leg II, ventral view. (J) Right tibia II, ventral view. (K) Left tibia II, ventral view. Scale lines 1.0 mm (A-E; F-K)." figureDoi="http://doi.org/10.5281/zenodo.5743966" httpUri="https://zenodo.org/record/5743966/files/figure.png" pageId="6" pageNumber="428">6</figureCitation>
H-K; exceptionally more, 
<figureCitation id="7DEB2A27C62FFF911092FD00574EFD1B" box="[226,306,733,756]" captionStart="Fig" captionStartId="7.[152,187,1970,1991]" captionTargetBox="[157,1426,704,1913]" captionTargetId="figure-219@7.[151,1435,699,1917]" captionTargetPageId="7" captionText="Fig. 3. Leptothele bencha; males from Ko Sirey (A-B), Ko Tarutao (C-F, G, showing two specimens), Ko Libong (H) and Ko Sukon (I- J). (A, D, H) Distal part of palp, ventral view. (B, E) Right tibia and metatarsus II, prolateral view. (C) Left tibia and metatarsus II, ventral view. (F) Right tibia and metatarsus II, ventral view. (G) Palpal organ, ventral view. (I) Left tibia and metatarsus II of aberrant specimen, prolateral view. (J) Same, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5743960" httpUri="https://zenodo.org/record/5743960/files/figure.png" pageId="6" pageNumber="428">Fig. 3J</figureCitation>
) megaspines (only one in 
<taxonomicName id="22D04D21C62FFF91121DFD005488FD1B" ID-CoL="98X62" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[621,756,733,756]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="428" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D7A4EAB0C62FFF91121DFD005488FD1B" box="[621,756,733,756]" italics="true" pageId="6" pageNumber="428">Malayathele</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62FFF9110E7FD205683FCFB" box="[151,255,765,788]" pageId="6" pageNumber="428" rank="genus">gen. nov.</taxonomicNameLabel>
, 
<figureCitation id="7DEB2A27C62FFF91117DFD20572DFCFB" box="[269,337,765,788]" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="6" pageNumber="428">Figs 8</figureCitation>
I-J, 11I-J, 13J, 15F) and a transversal subdistal ridge (
<figureCitation id="7DEB2A27C62FFF91113DFCC157E9FCDC" box="[333,405,796,819]" captionStart="Fig" captionStartId="7.[152,187,1970,1991]" captionTargetBox="[157,1426,704,1913]" captionTargetId="figure-219@7.[151,1435,699,1917]" captionTargetPageId="7" captionText="Fig. 3. Leptothele bencha; males from Ko Sirey (A-B), Ko Tarutao (C-F, G, showing two specimens), Ko Libong (H) and Ko Sukon (I- J). (A, D, H) Distal part of palp, ventral view. (B, E) Right tibia and metatarsus II, prolateral view. (C) Left tibia and metatarsus II, ventral view. (F) Right tibia and metatarsus II, ventral view. (G) Palpal organ, ventral view. (I) Left tibia and metatarsus II of aberrant specimen, prolateral view. (J) Same, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5743960" httpUri="https://zenodo.org/record/5743960/files/figure.png" pageId="6" pageNumber="428">Figs 3</figureCitation>
B-C, E-F, I-J, 6G-K; absent in 
<taxonomicName id="22D04D21C62FFF9110E7FCE15761FCBC" ID-CoL="98X62" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[151,285,828,851]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="428" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D7A4EAB0C62FFF9110E7FCE15761FCBC" box="[151,285,828,851]" italics="true" pageId="6" pageNumber="428">Malayathele</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62FFF911158FCE157F2FCBC" box="[296,398,828,851]" pageId="6" pageNumber="428" rank="genus">gen. nov.</taxonomicNameLabel>
) on ventral side of tibia II, and with a single, widely conical or mound-like median process on metatarsus II (
<figureCitation id="7DEB2A27C62FFF9111DEFCA0578EFC7B" box="[430,498,893,916]" captionStart="Fig" captionStartId="7.[152,187,1970,1991]" captionTargetBox="[157,1426,704,1913]" captionTargetId="figure-219@7.[151,1435,699,1917]" captionTargetPageId="7" captionText="Fig. 3. Leptothele bencha; males from Ko Sirey (A-B), Ko Tarutao (C-F, G, showing two specimens), Ko Libong (H) and Ko Sukon (I- J). (A, D, H) Distal part of palp, ventral view. (B, E) Right tibia and metatarsus II, prolateral view. (C) Left tibia and metatarsus II, ventral view. (F) Right tibia and metatarsus II, ventral view. (G) Palpal organ, ventral view. (I) Left tibia and metatarsus II of aberrant specimen, prolateral view. (J) Same, ventral view. Scale line 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5743960" httpUri="https://zenodo.org/record/5743960/files/figure.png" pageId="6" pageNumber="428">Figs 3</figureCitation>
B-C, E-F, I-J, 6G-I; 1-2 small conical processes or one conical process plus a short keel-shaped process in 
<taxonomicName id="22D04D21C62FFF9111BCFC60542DFC3B" ID-CoL="98X62" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[460,593,957,980]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="428" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D7A4EAB0C62FFF9111BCFC60542DFC3B" box="[460,593,957,980]" italics="true" pageId="6" pageNumber="428">Malayathele</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62FFF911228FC6054C0FC3B" box="[600,700,957,980]" pageId="6" pageNumber="428" rank="genus">gen. nov.</taxonomicNameLabel>
, 
<figureCitation id="7DEB2A27C62FFF9112B6FC6056DFFC1B" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="6" pageNumber="428">Figs 8</figureCitation>
H-K, 11G-J, 13J-M, 15F-H); tarsi I-II not ventrally bulged (
<figureCitation id="7DEB2A27C62FFF911084FC235730FBFA" box="[244,332,1022,1045]" captionStart="Fig" captionStartId="10.[151,186,1968,1989]" captionTargetBox="[159,1427,610,1932]" captionTargetId="figure-192@10.[153,1434,605,1947]" captionTargetPageId="10" captionText="Fig. 5. Leptothele chang sp. nov.; male holotype (A-B, E-H), female allotype (C-D) and male paratype (I). (A, C) Habitus, dorsal view. (B, D) Habitus, ventral view. (E) Femur to metatarsus of right leg II, prolateral view. (F) Right femur II, proventral view. (G) Right femur I, retrodorsal view. (H) Tarsus I, lateral view. (I) Distal part of median article and entire distal article of PLS, lateral view; arrow indicating pseudosegmentation. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743964" httpUri="https://zenodo.org/record/5743964/files/figure.png" pageId="6" pageNumber="428">Fig. 5H</figureCitation>
cf. 
<figureCitation id="7DEB2A27C62FFF91110FFC235794FBFA" box="[383,488,1022,1045]" captionStart="Fig" captionStartId="20.[151,186,2012,2033]" captionTargetBox="[157,1434,1019,1973]" captionTargetId="figure-450@20.[153,1438,1015,1976]" captionTargetPageId="20" captionText="Fig. 10. Malayathele cameronensis sp. nov., male holotype. (A) Left leg I, retrolateral view; one ventral spine on tibia fallen off. (B) Left leg II, prolateral view. (C) Unpaired claw and scopuliform hairs of left leg II, prolateral view. (D) Ventrally bulged tarsus of left leg II, prolateral view. (E) Scopuliform hair on left tarsus II. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743975" httpUri="https://zenodo.org/record/5743975/files/figure.png" pageId="6" pageNumber="428">Fig. 10D</figureCitation>
). Females distinguished from those of 
<taxonomicName id="22D04D21C62FFF911138FBC357B3FBDA" ID-CoL="98X62" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[328,463,1054,1077]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="428" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D7A4EAB0C62FFF911138FBC357B3FBDA" box="[328,463,1054,1077]" italics="true" pageId="6" pageNumber="428">Malayathele</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC62FFF9111AEFBC35434FBDA" box="[478,584,1054,1077]" pageId="6" pageNumber="428" rank="genus">gen. nov.</taxonomicNameLabel>
by possessing median receptacular stalks with unsclerotised walls (
<figureCitation id="7DEB2A27C62FFF9110EFFB835694FB9A" box="[159,232,1118,1141]" captionStart="Fig" captionStartId="8.[151,186,2027,2048]" captionTargetBox="[159,1431,770,1960]" captionTargetId="figure-262@8.[151,1435,763,1970]" captionTargetPageId="8" captionText="Fig. 4. Leptothele bencha; vulvae of six females, dorsal view. (A) Female from Ko Libong. (B-C) Two females from Ko Tarutao. (D) Female from Ko Lanta. (E-F) Two females from Ko Siray. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743962" httpUri="https://zenodo.org/record/5743962/files/figure.png" pageId="6" pageNumber="428">Figs 4</figureCitation>
, 
<figureCitation id="7DEB2A27C62FFF911087FB835778FB9A" box="[247,260,1118,1141]" captionStart="Fig" captionStartId="12.[151,186,1998,2019]" captionTargetBox="[155,1430,1261,1937]" captionTargetId="figure-579@12.[151,1435,1255,1940]" captionTargetPageId="12" captionText="Fig. 7. Leptothele chang sp. nov.; vulvae of female allotype (A) and of three female paratypes (B-D), dorsal view. (A) Drawn from exuvia of allotype; cuticular gland duct bases visible. (B-D) Drawn from dissected paratypes; gland duct bases not visible but gland pores instead. (C) Right lateral receptacle carrying a tiny secondary receptacle. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743968" httpUri="https://zenodo.org/record/5743968/files/figure.png" pageId="6" pageNumber="428">7</figureCitation>
cf. 
<figureCitation id="7DEB2A27C62FFF911144FB835700FB9A" box="[308,380,1118,1141]" captionStart="Fig" captionStartId="17.[151,186,1715,1736]" captionTargetBox="[154,1431,263,1659]" captionTargetId="figure-71@17.[151,1435,259,1666]" captionTargetPageId="17" captionText="Fig. 9. Malayathele ulu sp. nov.; vulvae of seven females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B-C) Paratypes from the type locality (Ulu Groh). (D-G) Paratypes from the Cameron Highlands: Parit Falls, 26.IV.2009 (D); Tanah Rata, 7.VIII.1972 (E); Gunung Perdah, 2.-14.V.2009 (F); trails 4 &amp; 13, 23.III.1993, heads of median receptacles collapsed and right lateral receptacle torn off (G). Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743972" httpUri="https://zenodo.org/record/5743972/files/figure.png" pageId="6" pageNumber="428">Figs 9</figureCitation>
, 
<figureCitation id="7DEB2A27C62FFF9111FBFB8357C6FB9A" box="[395,442,1118,1141]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="6" pageNumber="428">11K</figureCitation>
, 
<figureCitation id="7DEB2A27C62FFF9111B9FB83579AFB9A" box="[457,486,1118,1141]" captionStart="Fig" captionStartId="25.[151,186,2027,2048]" captionTargetBox="[155,1418,1378,1967]" captionTargetId="figure-608@25.[151,1422,1374,1976]" captionTargetPageId="25" captionText="Fig. 14. Malayathele maculosa sp. nov., vulvae of three females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B) Paratype from Genting Highlands. (C) Juvenile female paratype from Fraser’s Hill. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743985" httpUri="https://zenodo.org/record/5743985/files/figure.png" pageId="6" pageNumber="428">14</figureCitation>
, 
<figureCitation id="7DEB2A27C62FFF911185FB83546FFB9A" box="[501,531,1118,1141]" captionStart="Fig" captionStartId="28.[152,187,2025,2046]" captionTargetBox="[157,1428,1213,1987]" captionTargetId="figure-544@28.[151,1431,1208,1994]" captionTargetPageId="28" captionText="Fig. 16. Malayathele kanching sp. nov.; vulvae of four female paratypes, dorsal (A-B) to increasingly more postero-dorsal view (C- D). (A, C-D) Specimens from the Templer Park. (B) Specimen from Ulu Gombak. Scale lines 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5743991" httpUri="https://zenodo.org/record/5743991/files/figure.png" pageId="6" pageNumber="428">16</figureCitation>
). Revised character: teeth on unpaired leg claws either all sessile (
<figureCitation id="7DEB2A27C62FFF9112F0FBA354C0FB7A" box="[640,700,1150,1173]" captionStart="Fig" captionStartId="2.[151,186,1849,1870]" captionTargetBox="[161,1430,259,1801]" captionTargetId="figure-12@2.[153,1438,254,1809]" captionTargetPageId="2" captionText="Fig. 1. Photos of unpaired leg claws, lateral view (A-J), and of serrula on palpal coxa, anteroventral view (K-L). (A) Masteria cf. cavicola (Simon, 1892), female (in MHNG; 13.IV.1977; leg. P. Strinati &amp; V. Aellen) from a cave near Bagio, Luzon, the Phillipines, leg II. (B) Same, leg IV.(C) Phyxioschema huberi Schwendinger, 2009, exuvia of female paratype, leg I. (D) Same, leg IV. (E) Leptothele bencha, female from Ko Sirey, leg I. (F) Same, leg III. (G) Leptothele chang sp. nov., exuvia of female paratype, leg I. (H) Same, leg III. (I) Malayathele ulu sp. nov., exuvia of female paratype, leg I. (J) Same, leg IV. (K) Leptothele chang sp. nov., exuvia of female paratype. (L) Malayathele kanching sp. nov., exuvia of female paratype. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743954" httpUri="https://zenodo.org/record/5743954/files/figure.png" pageId="6" pageNumber="428">Fig. 1</figureCitation>
F-G) or proximal teeth raised on indistinct common base (
<figureCitation id="7DEB2A27C62FFF9110EFFB62576FFB39" box="[159,275,1215,1238]" captionStart="Fig" captionStartId="2.[151,186,1849,1870]" captionTargetBox="[161,1430,259,1801]" captionTargetId="figure-12@2.[153,1438,254,1809]" captionTargetPageId="2" captionText="Fig. 1. Photos of unpaired leg claws, lateral view (A-J), and of serrula on palpal coxa, anteroventral view (K-L). (A) Masteria cf. cavicola (Simon, 1892), female (in MHNG; 13.IV.1977; leg. P. Strinati &amp; V. Aellen) from a cave near Bagio, Luzon, the Phillipines, leg II. (B) Same, leg IV.(C) Phyxioschema huberi Schwendinger, 2009, exuvia of female paratype, leg I. (D) Same, leg IV. (E) Leptothele bencha, female from Ko Sirey, leg I. (F) Same, leg III. (G) Leptothele chang sp. nov., exuvia of female paratype, leg I. (H) Same, leg III. (I) Malayathele ulu sp. nov., exuvia of female paratype, leg I. (J) Same, leg IV. (K) Leptothele chang sp. nov., exuvia of female paratype. (L) Malayathele kanching sp. nov., exuvia of female paratype. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743954" httpUri="https://zenodo.org/record/5743954/files/figure.png" pageId="6" pageNumber="428">Fig. 1E, H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="ADCA6529C62FFF9110E7FB3356AAFACA" pageId="6" pageNumber="428" type="distribution">
<paragraph id="E56F36A2C62FFF9110E7FB3356AAFACA" blockId="6.[151,756,1262,1317]" pageId="6" pageNumber="428">
<emphasis id="D7A4EAB0C62FFF9110E7FB335752FAEA" bold="true" box="[151,302,1262,1285]" pageId="6" pageNumber="428">Distribution:</emphasis>
Southern 
<collectingCountry id="9DC77632C62FFF9111C3FB33546FFAEA" box="[435,531,1262,1285]" name="Thailand" pageId="6" pageNumber="428">Thailand</collectingCountry>
(
<figureCitation id="7DEB2A27C62FFF91125EFB335407FAEA" box="[558,635,1262,1285]" captionStart="Fig" captionStartId="4.[155,190,1853,1874]" captionTargetBox="[154,1433,260,1820]" captionTargetId="figure-12@4.[151,1437,256,1824]" captionTargetPageId="4" captionText="Fig. 2. Localities of Leptothele spp. and Malayathele spp. on the Thai-Malay Peninsula. The coast of Sumatra is omitted. 1 - Ko Sirey (L. bencha); 2 - Ko Yao Noi (L. bencha); 3 - Khao Phanom Bencha (L. bencha, type locality); 4 - Thab Khaek - Hang Nak Hill and Khlong Jilat (L. bencha); 5 - Ban Chong Phlie (L. bencha); 6 - Ko Lanta (L. bencha); 7 - Ko Muk (L. bencha); 8 - Ko Libong (L. bencha); 9 - Ko Sukon (L. bencha); 10 - Ko Tarutao (L. bencha); 11 - Tham Khao Chang Hai (L. chang sp. nov., type locality); 12 - Ulu Groh (M. ulu sp. nov., type locality); 13 - Chenderiang (Malayathele sp.); 14 - Cameron Highlands (M. cameronensis sp. nov., type locality; M. ulu sp. nov.); 15 - Fraser’s Hill (M. maculosa sp. nov., type locality); 16 - Genting Highlands (M. maculosa sp. nov.); 17 - Templer Park (M. kanching sp. nov.; type locality); 18 - Ulu Gombak (M. kanching sp. nov.); 19 - Jeram Pasu Waterfall (Malayathele sp.); 20 - Singapore (Euagridae/Dipluridae gen. sp.)." figureDoi="http://doi.org/10.5281/zenodo.5743958" httpUri="https://zenodo.org/record/5743958/files/figure.png" pageId="6" pageNumber="428">Fig. 2</figureCitation>
, localities 1-11).
</paragraph>
</subSubSection>
</treatment>
</document>