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<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.96.81737" ID-GBIF-Dataset="d148b8fa-4f4c-4069-80e5-f04f98561bfe" ID-Pensoft-Pub="2747-8106-96-51" ID-Pensoft-UUID="FCEF2A912BC258BBB022681E86486B06" ID-ZooBank="CD8C70BA025345408EFCD430A12F6B35" ModsDocID="2747-8106-96-51" checkinTime="1649795137967" checkinUser="pensoft" docAuthor="Schwarzhans, Werner &amp; Keupp, Helmut" docDate="2022" docId="16BE13E2884855EE84E8E13A9EAFC2E0" docLanguage="en" docName="Zitteliana 96: 51-67" docOrigin="Zitteliana 96" docPubDate="2022-04-12" docSource="http://dx.doi.org/10.3897/zitteliana.96.81737" docTitle="Leptolepis inaequalis" docType="treatment" docVersion="4" id="FCEF2A912BC258BBB022681E86486B06" lastPageNumber="51" masterDocId="FCEF2A912BC258BBB022681E86486B06" masterDocTitle="Early teleost otolith morphogenesis observed in the Jurassic of Franconia, Bavaria, southern Germany" masterLastPageNumber="67" masterPageNumber="51" pageNumber="51" updateTime="1732488386774" updateUser="ExternalLinkService">
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<mods:title id="00787213B69CC6C0E709D896BD491E03">Early teleost otolith morphogenesis observed in the Jurassic of Franconia, Bavaria, southern Germany</mods:title>
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<mods:namePart id="4482E91D54BEDAE083427FD55E934256">Schwarzhans, Werner</mods:namePart>
<mods:nameIdentifier id="1801EB15867798E652C56C809E440FC9" type="ORCID">https://orcid.org/0000-0003-4842-7989</mods:nameIdentifier>
<mods:affiliation id="A423939610EE99FA16951DB6DE5A64AB">Zoological Museum, Natural History Museum of Denmark, Universitetsparken 15, 2100 Kobenhavn, Denmark &amp; Ahrensburger Weg 103, 22359 Hamburg, Germany</mods:affiliation>
<mods:nameIdentifier id="2FBBD65F71393071304E8E9C18239F08" type="email">wwschwarz@aol.com</mods:nameIdentifier>
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<mods:namePart id="E46008EFB811BA4F25CA724B19D6FC9B">Keupp, Helmut</mods:namePart>
<mods:affiliation id="7E18A4CE465191EB643E49F3BF39B8EC">Freie Universitaet Berlin, Institut fuer Geologische Wissenschaften, Fachrichtung Palaeontologie, Malteser Strasse 74 - 100, Haus D, 12249 Berlin, Germany</mods:affiliation>
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<mods:title id="D8A1C91CF31AD64CBE55CF99EB779C23">Zitteliana</mods:title>
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<mods:date id="846705F26BFA31C69990FB4088F5EF58">2022</mods:date>
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<mods:number id="2F03165F7E1C98D05B5D96A17B43571A">96</mods:number>
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<treatment id="16BE13E2884855EE84E8E13A9EAFC2E0" ID-GBIF-Taxon="194588024" LSID="urn:lsid:plazi:treatment:16BE13E2884855EE84E8E13A9EAFC2E0" httpUri="http://treatment.plazi.org/id/16BE13E2884855EE84E8E13A9EAFC2E0" lastPageNumber="51" pageId="0" pageNumber="51" scope_phylum="Teleostei">
<subSubSection id="9CE8E3E7BAF63AF4A8D51169F3961081" pageId="0" pageNumber="51" type="nomenclature">
<paragraph id="304EE58CFB9C8C8F2B52849554203145" pageId="0" pageNumber="51">
<taxonomicName id="11011D894A270CCC95887A55A2C09022" LSID="16BE13E2-8848-55EE-84E8-E13A9EAFC2E0" authority="(Weiler, 1954)" baseAuthorityName="Weiler" baseAuthorityYear="1954" class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptolepis inaequalis" order="Leptolepidiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="inaequalis">Leptolepis inaequalis (Weiler, 1954)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="171C2EE85AE0F52A32F7D5690489B72A" pageId="0" pageNumber="51" type="description">
<paragraph id="90C032235BC9F60080057E89D20E4D78" pageId="0" pageNumber="51">
<figureCitation id="50F80C39568D034CB90FDD3E0C5AA4A5" captionStart="Plate 3" captionStartId="F5" captionText="Plate 3. Otoliths of Leptolepis kremmeldorfensis, L. inaequalis, L. macrocephalus, and Leptoelops rhenanus. figs 1 - 6: Leptolepis kremmeldorfensis (Schroeder, 1956); 1: holotype of Ot. kremmeldorfensis Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 35); 2: paratype of Ot. kremmeldorfensis Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 36); 3: holotype of Ot. amygdalinus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 14); 4: paratype of Ot. bambergensis Schroeder, 1956 from Freiahorn (SNSB-BSPG 2022 III 10); 5: holotype of Ot. franconicus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 24); 6: paratype of Ot. franconicus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 27). figs 7, 8: Leptolepis cf. kremmeldorfensis (Schroeder, 1956), from the late Pliensbachian of Buttenheim, SNSB-BSPG 2022 IV 15. figs 9 - 11: Leptolepis inaequalis (Weiler, 1954); 9: holotype of Ot. puenzendorfensis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 39); 10: holotype of Ot. calloviensis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 8); 11: holotype of Ot. inaequalis Weiler, 1954 from the Callovian to Oxfordian of Kandern, Baden-Wuertemberg (SMF P. 3067). figs 12 - 13: Leptolepis macrocephalus (Schroeder, 1956); 12: holotype of Ot. guttaeformis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 6); 13: holotype of Ot. macrocephalus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 1). figs 14 - 16: Leptoelops rhenanus (Weiler, 1954); 14: holotype of Ot. rhenanus Weiler, 1954 from the Callovian of well Bruchsal D 205 (core from 482 to 487 m) near Weingarten, Baden-Wuerttemberg (SMF P. 2953); 15: holotype of Ot. rectisulcatus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 37); 16: holotype of Ot. scissus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 40)." figureDoi="10.3897/zitteliana.96.81737.plate3" httpUri="https://binary.pensoft.net/fig/671633" pageId="0" pageNumber="51">Plate 3, figs 9-11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="679BEF45F842D0D54F5F0A25491FDC0E" pageId="0" pageNumber="51" type="reference_group">
<paragraph id="9EFF093ADC0E94D7C72AB9D960537682" pageId="0" pageNumber="51">
<taxonomicName id="03B51813B4DEDD95C98580665D7A4B3B" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Otolithus inaequalis" order="Leptolepidiformes" pageId="0" pageNumber="51" rank="species" species="inaequalis">Otolithus inaequalis</taxonomicName>
1954
<emphasis id="E76ABA607C6EE1F8303FAAF04A59811E" italics="true" pageId="0" pageNumber="51">Otolithus</emphasis>
(inc.sed.)
<emphasis id="87C6675D79EC2B8D58356BD28CC8C8EB" italics="true" pageId="0" pageNumber="51">inaequalis</emphasis>
- Weiler: pl. 4, fig. 170.
</paragraph>
<paragraph id="E1BC955110447894F43512CA881D3522" pageId="0" pageNumber="51">
<taxonomicName id="EA3308C763E9099708B660AE667720FE" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Otolithus calloviensis" order="Leptolepidiformes" pageId="0" pageNumber="51" rank="species" species="calloviensis">Otolithus calloviensis</taxonomicName>
1956
<taxonomicName id="BFA7F7C476140B378A173ED9AA8F89F1" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Otolithus calloviensis" order="Leptolepidiformes" pageId="0" pageNumber="51" rank="species" species="calloviensis">Otolithus calloviensis</taxonomicName>
-
<normalizedToken id="EE879CB0A7D37B3AED0D6CC8B00A154B" originalValue="Schröder">Schroeder</normalizedToken>
: pl. 6, figs 17-18.
</paragraph>
<paragraph id="F315EB38945ABA3F26F527BBD5E0DEB7" pageId="0" pageNumber="51">
<taxonomicName id="6779D0A06FB0006DCA21A816472D0AD8" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Otolithus pünzendorfensis" order="Leptolepidiformes" pageId="0" pageNumber="51" rank="species" species="pünzendorfensis">
Otolithus
<normalizedToken id="AFB921B60DD242616A3784828F359C1E" originalValue="pünzendorfensis">puenzendorfensis</normalizedToken>
</taxonomicName>
1956
<taxonomicName id="EE2A29705AD268C519869C5A773AED00" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Otolithus" order="Leptolepidiformes" pageId="0" pageNumber="51" rank="genus">Otolithus</taxonomicName>
<emphasis id="1427943ABBDBD1C34B2E1992E7A09F87" italics="true" pageId="0" pageNumber="51">
<normalizedToken id="788E4FA19CE3D76D4F66FAEC0F725673" originalValue="pünzendorfensis">puenzendorfensis</normalizedToken>
</emphasis>
-
<normalizedToken id="9A33224710E785171B66DC67E05AE403" originalValue="Schröder">Schroeder</normalizedToken>
: pl. 7, fig. 39.
</paragraph>
<paragraph id="68DAECDB41FB78549A41AE8A86FE6608" pageId="0" pageNumber="51">
<taxonomicName id="A8697ACB98316B46E57E9AA09DF6BD55" baseAuthorityName="Weiler" baseAuthorityYear="1954" class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptolepis inaequalis" order="Leptolepidiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="inaequalis">Leptolepis inaequalis</taxonomicName>
2018
<taxonomicName id="ED8624D6A1CD10D49E0ECE8CB66FA6C0" baseAuthorityName="Weiler" baseAuthorityYear="1954" class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptolepis inaequalis" order="Leptolepidiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="inaequalis">Leptolepis inaequalis</taxonomicName>
(Weiler, 1954) - Schwarzhans: fig. 2E.
</paragraph>
</subSubSection>
<subSubSection id="AFCC0284EB932D0A806D2CBD2DABECEA" pageId="0" pageNumber="51" type="materials_examined">
<paragraph id="C81291A2DCDF0361BD247EBAD81976F6" pageId="0" pageNumber="51">Material.</paragraph>
<paragraph id="4BD8A57F50B8F6841BF5756B2C3251DA" pageId="0" pageNumber="51">
<materialsCitation id="3CAB3B402DF22565C2A0389C04481ACB" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" county="Baden-Wuertemberg" location="Callovian" municipality="Oxfordian of Kandern" specimenCount="4" typeStatus="holotype">
<specimenCount id="08619C750A8E68F2F5D485795129EAE9" type="generic">4 specimens</specimenCount>
:
<normalizedToken id="3AB66E4B064D6F3F7D03A7E73B157D94" originalValue="Weilers">Weiler's</normalizedToken>
<typeStatus id="97811BE9FEA56862DBA616CC98AF2BC6">holotype</typeStatus>
of
<taxonomicName id="954BD07E2141C446B8E94A89428614D5" lsidName="Ot. inaequalis" pageId="0" pageNumber="51" rank="species" species="inaequalis">
<emphasis id="C5B0B5800E6B48B573140FD7E2B9C7A6" italics="true" pageId="0" pageNumber="51">Ot. inaequalis</emphasis>
</taxonomicName>
from the
<location id="380A9461953CD25BF750AB2D08D3BC4A" LSID="urn:lsid:plazi:treatment:16BE13E2884855EE84E8E13A9EAFC2E0:380A9461953CD25BF750AB2D08D3BC4A" county="Baden-Wuertemberg" municipality="Oxfordian of Kandern" name="Callovian">Callovian</location>
to
<collectingMunicipality id="EF13190D48AD0EE786D8DE0AA643B540">Oxfordian of Kandern</collectingMunicipality>
,
<normalizedToken id="BEBD1894CE4E481F6F56BB366A596D5A" originalValue="Baden-Würtemberg">
<collectingCounty id="F6020537A1E8AC5786C3FD5DBC8193C9">Baden-Wuertemberg</collectingCounty>
</normalizedToken>
(SMF P.3067, refigured in Plate
<figureCitation id="864B44A4B3400534E4EB7B43FEADDA5D" captionStart="Plate 3" captionStartId="F5" captionText="Plate 3. Otoliths of Leptolepis kremmeldorfensis, L. inaequalis, L. macrocephalus, and Leptoelops rhenanus. figs 1 - 6: Leptolepis kremmeldorfensis (Schroeder, 1956); 1: holotype of Ot. kremmeldorfensis Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 35); 2: paratype of Ot. kremmeldorfensis Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 36); 3: holotype of Ot. amygdalinus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 14); 4: paratype of Ot. bambergensis Schroeder, 1956 from Freiahorn (SNSB-BSPG 2022 III 10); 5: holotype of Ot. franconicus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 24); 6: paratype of Ot. franconicus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 27). figs 7, 8: Leptolepis cf. kremmeldorfensis (Schroeder, 1956), from the late Pliensbachian of Buttenheim, SNSB-BSPG 2022 IV 15. figs 9 - 11: Leptolepis inaequalis (Weiler, 1954); 9: holotype of Ot. puenzendorfensis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 39); 10: holotype of Ot. calloviensis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 8); 11: holotype of Ot. inaequalis Weiler, 1954 from the Callovian to Oxfordian of Kandern, Baden-Wuertemberg (SMF P. 3067). figs 12 - 13: Leptolepis macrocephalus (Schroeder, 1956); 12: holotype of Ot. guttaeformis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 6); 13: holotype of Ot. macrocephalus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 1). figs 14 - 16: Leptoelops rhenanus (Weiler, 1954); 14: holotype of Ot. rhenanus Weiler, 1954 from the Callovian of well Bruchsal D 205 (core from 482 to 487 m) near Weingarten, Baden-Wuerttemberg (SMF P. 2953); 15: holotype of Ot. rectisulcatus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 37); 16: holotype of Ot. scissus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 40)." figureDoi="10.3897/zitteliana.96.81737.plate3" httpUri="https://binary.pensoft.net/fig/671633" pageId="0" pageNumber="51">3</figureCitation>
, fig. 11)
</materialsCitation>
;
<materialsCitation id="EC5F4F032209E9B9B807DEF9AFBC1789" collectingDate="2022-03-08" location="Puenzendorf" specimenCount="3" typeStatus="holotype">
<specimenCount id="18D9B26C74FA9FE65C88B08994D15287" type="generic">3 specimens</specimenCount>
from
<normalizedToken id="1814DBE942541B3CAD51C0E9FF313088" originalValue="Schröders">Schroeder's</normalizedToken>
collection from the early Callovian:
<typeStatus id="D6DC1BB46276BAF4C71630970D4A45B1">holotype</typeStatus>
of
<taxonomicName id="757307026D3935F2820007654BA05E2B" lsidName="Ot. calloviensis" pageId="0" pageNumber="51" rank="species" species="calloviensis">
<emphasis id="FC3345C6A207F979805098C2E53273F1" italics="true" pageId="0" pageNumber="51">Ot. calloviensis</emphasis>
</taxonomicName>
from
<normalizedToken id="EC88171D8AE0E0EEE63EA6035A01F9C2" originalValue="Pünzendorf">
<location id="EC74F3C25675A5DA705D55D2FD53610A" LSID="urn:lsid:plazi:treatment:16BE13E2884855EE84E8E13A9EAFC2E0:EC74F3C25675A5DA705D55D2FD53610A" name="Puenzendorf">Puenzendorf</location>
</normalizedToken>
(SNSB-BSPG
<collectingDate id="49E081FDA43ECD540EC276205616A1C2" value="2022-03-08">2022 III 8</collectingDate>
, Plate
<figureCitation id="250A8536A5669245DD29D6ABA2F27FE6" captionStart="Plate 3" captionStartId="F5" captionText="Plate 3. Otoliths of Leptolepis kremmeldorfensis, L. inaequalis, L. macrocephalus, and Leptoelops rhenanus. figs 1 - 6: Leptolepis kremmeldorfensis (Schroeder, 1956); 1: holotype of Ot. kremmeldorfensis Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 35); 2: paratype of Ot. kremmeldorfensis Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 36); 3: holotype of Ot. amygdalinus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 14); 4: paratype of Ot. bambergensis Schroeder, 1956 from Freiahorn (SNSB-BSPG 2022 III 10); 5: holotype of Ot. franconicus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 24); 6: paratype of Ot. franconicus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 27). figs 7, 8: Leptolepis cf. kremmeldorfensis (Schroeder, 1956), from the late Pliensbachian of Buttenheim, SNSB-BSPG 2022 IV 15. figs 9 - 11: Leptolepis inaequalis (Weiler, 1954); 9: holotype of Ot. puenzendorfensis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 39); 10: holotype of Ot. calloviensis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 8); 11: holotype of Ot. inaequalis Weiler, 1954 from the Callovian to Oxfordian of Kandern, Baden-Wuertemberg (SMF P. 3067). figs 12 - 13: Leptolepis macrocephalus (Schroeder, 1956); 12: holotype of Ot. guttaeformis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 6); 13: holotype of Ot. macrocephalus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 1). figs 14 - 16: Leptoelops rhenanus (Weiler, 1954); 14: holotype of Ot. rhenanus Weiler, 1954 from the Callovian of well Bruchsal D 205 (core from 482 to 487 m) near Weingarten, Baden-Wuerttemberg (SMF P. 2953); 15: holotype of Ot. rectisulcatus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 37); 16: holotype of Ot. scissus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 40)." figureDoi="10.3897/zitteliana.96.81737.plate3" httpUri="https://binary.pensoft.net/fig/671633" pageId="0" pageNumber="51">3</figureCitation>
, fig. 10) and
</materialsCitation>
<specimenCount id="D3484E95AF41F5CD5F07BC97C45FFCBB" type="generic" typeStatus="paratype">1 paratype</specimenCount>
from
<normalizedToken id="B6DA4F57467F40BEEF03A27DEB81F315" originalValue="Pünzendorf">Puenzendorf</normalizedToken>
(SNSB-BSPG
<date id="C1262188B718510964E421ABA2A08450" value="2022-03-09">2022 III 9</date>
),
<typeStatus id="E8B0C2A30EFFF80C2E0EB2CBEBCA0A5A">holotype</typeStatus>
of
<taxonomicName id="F6BCC485A1B8F68A62CD6E7BF2E31C97" lsidName="Ot. puenzendorfensis" pageId="0" pageNumber="51" rank="species" species="puenzendorfensis">
<emphasis id="8F81C8F95C04831AD4EC86E6AF10252B" italics="true" pageId="0" pageNumber="51">Ot. puenzendorfensis</emphasis>
</taxonomicName>
(emended spelling, SNSB-BSPG
<date id="072C074EAD910FD47448AF8698FF1182" value="2022-03">2022 III</date>
39, Plate
<figureCitation id="8D6695B94E3BFCE99134C70DCB3F453E" captionStart="Plate 3" captionStartId="F5" captionText="Plate 3. Otoliths of Leptolepis kremmeldorfensis, L. inaequalis, L. macrocephalus, and Leptoelops rhenanus. figs 1 - 6: Leptolepis kremmeldorfensis (Schroeder, 1956); 1: holotype of Ot. kremmeldorfensis Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 35); 2: paratype of Ot. kremmeldorfensis Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 36); 3: holotype of Ot. amygdalinus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 14); 4: paratype of Ot. bambergensis Schroeder, 1956 from Freiahorn (SNSB-BSPG 2022 III 10); 5: holotype of Ot. franconicus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 24); 6: paratype of Ot. franconicus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 27). figs 7, 8: Leptolepis cf. kremmeldorfensis (Schroeder, 1956), from the late Pliensbachian of Buttenheim, SNSB-BSPG 2022 IV 15. figs 9 - 11: Leptolepis inaequalis (Weiler, 1954); 9: holotype of Ot. puenzendorfensis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 39); 10: holotype of Ot. calloviensis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 8); 11: holotype of Ot. inaequalis Weiler, 1954 from the Callovian to Oxfordian of Kandern, Baden-Wuertemberg (SMF P. 3067). figs 12 - 13: Leptolepis macrocephalus (Schroeder, 1956); 12: holotype of Ot. guttaeformis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 6); 13: holotype of Ot. macrocephalus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 1). figs 14 - 16: Leptoelops rhenanus (Weiler, 1954); 14: holotype of Ot. rhenanus Weiler, 1954 from the Callovian of well Bruchsal D 205 (core from 482 to 487 m) near Weingarten, Baden-Wuerttemberg (SMF P. 2953); 15: holotype of Ot. rectisulcatus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 37); 16: holotype of Ot. scissus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 40)." figureDoi="10.3897/zitteliana.96.81737.plate3" httpUri="https://binary.pensoft.net/fig/671633" pageId="0" pageNumber="51">3</figureCitation>
, fig. 9).
</paragraph>
</subSubSection>
<subSubSection id="567678F9BCBBEFEFB3F4DEFF1A5861FC" pageId="0" pageNumber="51" type="differential diagnosis">
<paragraph id="692907A16024F98D03C2EA27F07FC39C" pageId="0" pageNumber="51">Differential diagnosis.</paragraph>
<paragraph id="35F54D13B620A8F8B744581EBDA0D4E3" pageId="0" pageNumber="51">
<taxonomicName id="1644723C49E7D2C8766BC49F6DF5112C" baseAuthorityName="Weiler" baseAuthorityYear="1954" class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptolepis inaequalis" order="Elopiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="inaequalis">
<emphasis id="FD4D689D133728E8E06314C21F22868B" italics="true" pageId="0" pageNumber="51">Leptolepis inaequalis</emphasis>
</taxonomicName>
is characterized by being very thin with nearly flat inner and outer faces. It shares these features only with
<taxonomicName id="2FA2415245478A68D54575FE25EACC91" class="Actinopterygii" kingdom="Animalia" lsidName="L. macrocephalus" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="macrocephalus">
<emphasis id="0A441275304D179DB9B30DF8C91B1431" italics="true" pageId="0" pageNumber="51">L. macrocephalus</emphasis>
</taxonomicName>
(see below), from which it differs in its slightly slenderer shape (OL:OH = 1.5 vs. 1.35-1.4) and its shallow and rather long postdorsal rim (vs. rounded and expanded).
</paragraph>
</subSubSection>
<subSubSection id="BB7D749C66D9A62E936B34721C27EE95" pageId="0" pageNumber="51" type="discussion">
<paragraph id="DA6F1CDE651982BBF7E27504937BFC71" pageId="0" pageNumber="51">Discussion.</paragraph>
<paragraph id="79FD5958426F93606FCD5AB26F28A4C5" pageId="0" pageNumber="51">
Due to the fragile nature of these thin and delicate otoliths, only one complete specimen of
<taxonomicName id="FC915115AB53DD166D436573B6154AAA" class="Actinopterygii" kingdom="Animalia" lsidName="L. inaequalis" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="inaequalis">
<emphasis id="999B37E29A90038B6ACE21101A334830" italics="true" pageId="0" pageNumber="51">L. inaequalis</emphasis>
</taxonomicName>
is known: the holotype from Weiler. All three specimens from
<normalizedToken id="B9C82B57BD9CC3E706604D2D9592ED9E" originalValue="Schröder">Schroeder</normalizedToken>
, which we synonymized with
<taxonomicName id="EEC42FA40D5FEBC6B3C7B67E96B65E3B" class="Actinopterygii" kingdom="Animalia" lsidName="L. inaequalis" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="inaequalis">
<emphasis id="D5A09BDF78AFD45797005CB604DA3521" italics="true" pageId="0" pageNumber="51">L. inaequalis</emphasis>
</taxonomicName>
, are incomplete because they lack the rostrum, but they do show the typical flat inner face in combination with the shallow and long postdorsal rim.
<normalizedToken id="DB6FADD4D849BCC8247488CBE02A240E" originalValue="Schröders">Schroeder's</normalizedToken>
drawing of the holotype of
<taxonomicName id="0342D7E910E441537CB9370EB80D1937" lsidName="Ot. calloviensis" pageId="0" pageNumber="51" rank="species" species="calloviensis">
<emphasis id="2AADA2B9AEECC4E846F085723D3894D6" italics="true" pageId="0" pageNumber="51">Ot. calloviensis</emphasis>
</taxonomicName>
indicates a complete specimen, but the one found in his collection is slightly damaged, which possibly happened after the drawing was made.
</paragraph>
</subSubSection>
</treatment>
</document>