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<document ID-DOI="http://dx.doi.org/10.3897/vz.71.e60921" ID-GBIF-Dataset="542339bb-d944-41a5-b45b-664a07c655b6" ID-GBIF-Taxon="191980068" ID-Pensoft-Pub="2625-8498-71-75" ID-Pensoft-UUID="C87DE9A5FDFA5B5FA5603914E35A1446" ID-ZooBank="1C61F5FE1FA44083B91313FD42157FCA" ModsDocID="2625-8498-71-75" checkinTime="1641857443470" checkinUser="pensoft" docAuthor="Quah, Evan S. H., Grismer, L. Lee, Wood Jr., Perry L., Lim, Kelvin K. P., Imbun, Paul Y. &amp; Anuar, M. S. Shahrul" docDate="2021" docId="9C848621A46657E1B4BEE464EAF328BE" docLanguage="en" docName="VertZool 71: 75-99" docOrigin="Vertebrate Zoology 71" docPubDate="2021-03-11" docSource="http://dx.doi.org/10.3897/vz.71.e60921" docTitle="Abavorana decorata comb. nov. (Mocquard 1890" docType="treatment" docVersion="4" id="C87DE9A5FDFA5B5FA5603914E35A1446" lastPageNumber="75" masterDocId="C87DE9A5FDFA5B5FA5603914E35A1446" masterDocTitle="An investigation into the taxonomy of Abavorana luctuosa (Peters, 1871) (Anura, Ranidae) and the resurrection of Rana decorata Mocquard, 1890 from Borneo" masterLastPageNumber="99" masterPageNumber="75" pageId="0" pageNumber="75" updateTime="1641863460135" updateUser="ExternalLinkService">
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<mods:title>An investigation into the taxonomy of Abavorana luctuosa (Peters, 1871) (Anura, Ranidae) and the resurrection of Rana decorata Mocquard, 1890 from Borneo</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Quah, Evan S. H.</mods:namePart>
<mods:affiliation>Institute of Tropical Biodiversity and Sustainable Development, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia, https: // orcid. org / 0000 - 0002 - 5357 - 1953 &amp; Herpetology Laboratory, Department of Biology, La Sierra University, Riverside, California, USA, https: // orcid. org / 0000 - 0001 - 8422 - 3698 &amp; Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, 117377 Singapore, https: // orcid. org / 0000 - 0002 - 0638 - 9198</mods:affiliation>
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<mods:namePart>Grismer, L. Lee</mods:namePart>
<mods:affiliation>Herpetology Laboratory, Department of Biology, La Sierra University, Riverside, California, USA, https: // orcid. org / 0000 - 0001 - 8422 - 3698</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wood Jr., Perry L.</mods:namePart>
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<mods:affiliation>Department of Biological Sciences &amp; Museum of Natural History, Auburn University, Auburn, Alabama 36849, USA, https: // orcid. org / 0000 - 0003 - 3767 - 5274</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Lim, Kelvin K. P.</mods:namePart>
<mods:affiliation>Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, 117377 Singapore, https: // orcid. org / 0000 - 0002 - 0638 - 9198</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Imbun, Paul Y.</mods:namePart>
<mods:affiliation>Museum of Zoology, Research &amp; Education Division, Sabah Parks, 10626 Kota Kinabalu, Sabah, Malaysia</mods:affiliation>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Anuar, M. S. Shahrul</mods:namePart>
<mods:affiliation>School of Biological Sciences, Universiti Sains Malaysia, 11800 Minden, Penang, Malaysia, https: // orcid. org / 0000 - 0003 - 0648 - 7318 &amp; Center for Marine and Coastal Studies, Universiti Sains Malaysia, 11800 USM, Pulau Pinang, Malaysia</mods:affiliation>
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<mods:title>Vertebrate Zoology</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-03-11</mods:number>
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<treatment ID-GBIF-Taxon="191980068" LSID="urn:lsid:plazi:treatment:9C848621A46657E1B4BEE464EAF328BE" httpUri="http://treatment.plazi.org/id/9C848621A46657E1B4BEE464EAF328BE" lastPageNumber="75" pageId="0" pageNumber="75">
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<taxonomicName LSID="9C848621-A466-57E1-B4BE-E464EAF328BE" authority="comb. nov. (Mocquard, 1890)" authorityName="comb. nov. (Mocquard" authorityYear="1890" class="Amphibia" family="Ranidae" genus="Abavorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abavorana decorata" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="decorata">Abavorana decorata comb. nov. (Mocquard, 1890)</taxonomicName>
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<paragraph pageId="0" pageNumber="75">
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Syntypes of Abavorana decorata; A &amp; D: Dorsum and venter of MNHN-RA- 1889.226. B &amp; E: Dorsum and venter of MNHN-RA- 1889.227. C &amp; F: Dorsum and venter of MNHN-RA- 1889.228 (Photographs by Christine Reinhardt). G: Original illustration of the syntypes of Abavorana decorata from Mocquard (1890). Illustrations assignable to specimen MNHN-RA- 1889.226 (left) and MNHN-RA- 1889.228 (right)." figureDoi="10.3897/vz.71.e60921.figure5" httpUri="https://binary.pensoft.net/fig/518978" pageId="0" pageNumber="75">Decorated Mahogany Frog Figs 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Examined Abavorana decorata specimens; Dorsal (A &amp; C) and ventral (B &amp; D) pattern of Abavorana decorata from Borneo. A &amp; B (from left to right): FMNH 109935 from Kiau, Mount Kinabalu, Sabah (left), FMNH 109936 from Kiau, Mount Kinabalu, Sabah (middle), FMNH 231052 from Danum Valley Research Center, Lahad Datu District, Sabah (top right), FMNH 128127 from Bario, Sarawak (bottom right). C &amp; D: SP 26873 from Tawau Hills, Sabah (left) and SP 02810 from Maliau Basin, Sabah." figureDoi="10.3897/vz.71.e60921.figure6" httpUri="https://binary.pensoft.net/fig/518979" pageId="0" pageNumber="75">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Living specimens of Abavorana decorata and Bornean A. luctuosa; A: Abavorana decorata from Tawau Hills, Sabah, East Malaysia. B: A. decorata from Maliau Basin, Sabah, East Malaysia. C: A. decorata from Kubah National Park, Sarawak, East Malaysia. D: A. decorata from Melawi Regency, West Kalimantan Province, Indonesia. E &amp; F: A. luctuosa from Kubah National Park, Sarawak, East Malaysia. G: A. luctuosa (FMNH 238720) from Sipitang, Sabah, East Malaysia. H: A. luctuosa (FMNH 242883) from Sipitang, Sabah, East Malaysia. (Photographs by Alexander Haas [A, C &amp; E], Paul Bertner [B], Muhammad Aini [D], Chien C. Lee [F], and Robert F. Inger [G &amp; H], Figs G &amp; H © Field Museum of Natural History. Created by Field Museum of Natural History, Amphibian and Reptile Collection and licensed under CC-BY-SA 4.0)." figureDoi="10.3897/vz.71.e60921.figure7" httpUri="https://binary.pensoft.net/fig/518980" pageId="0" pageNumber="75">, 7A-D</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Colour pattern comparison among species of the genus Abavorana; Dorsum (A, C &amp; E) and venter (B, D &amp; F) in life of A. luctuosa (USMHC 2576) (A &amp; B) from Penang Hill, Penang Island, Malaysia, A. decorata from Maliau Basin, Sabah, East Malaysia, Borneo (C &amp; D) and A. nazgul (USMHC 2233) (E &amp; F) from Gunung Jerai, Kedah, West Malaysia. (Photographs by Evan Quah [A, B, E &amp; F] and Stefan T. Hertwig [C &amp; D])." figureDoi="10.3897/vz.71.e60921.figure9" httpUri="https://binary.pensoft.net/fig/518982" pageId="0" pageNumber="75">, 9C-D</figureCitation>
</paragraph>
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<subSubSection pageId="0" pageNumber="75" type="reference_group">
<paragraph pageId="0" pageNumber="75">
<taxonomicName authorityName="Mocquard" authorityYear="1890" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rana decorata" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="decorata">Rana decorata</taxonomicName>
:
<bibRefCitation author="Mocquard, F" journalOrPublisher="Nouvelles Archives du Museum d'Histoire Naturelle Paris" pageId="0" pageNumber="75" pagination="115 - 168" refId="B70" refString="Mocquard, F, 1890. Recherches sur la faune herpetologique des iles de Borneo et de Palawan. Nouvelles Archives du Museum d'Histoire Naturelle Paris 3: 115 - 168" title="Recherches sur la faune herpetologique des iles de Borneo et de Palawan." volume="3" year="1890">Mocquard 1890</bibRefCitation>
:145-146;
<bibRefCitation author="Guibe, J" journalOrPublisher="Imprimerie nationale, Paris" pageId="0" pageNumber="75" refId="B35" refString="Guibe, J, 1950. Catalogue des types d'amphibiens du Museum national d'Histoire naturelle. Imprimerie nationale, Paris" title="Catalogue des types d'amphibiens du Museum national d'Histoire naturelle." year="1950">
<normalizedToken originalValue="Guibé">Guibe</normalizedToken>
1950
</bibRefCitation>
:41
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1871" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rana luctuosa" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="luctuosa">Rana luctuosa</taxonomicName>
(in part):
<bibRefCitation DOI="https://doi.org/10.1080/00222939109460620" author="Boulenger, GA" journalOrPublisher="Annals and Magazine of Natural History" pageId="0" pageNumber="75" pagination="341 - 346" refId="B4" refString="Boulenger, GA, 1891. Remarks on the herpetological fauna of Mount Kina Baloo, North Borneo. Annals and Magazine of Natural History 6: 341 - 346, DOI: https://doi.org/10.1080/00222939109460620" title="Remarks on the herpetological fauna of Mount Kina Baloo, North Borneo." url="https://doi.org/10.1080/00222939109460620" volume="6" year="1891">Boulenger 1891</bibRefCitation>
:341 &amp; 342, 1912:238, 1920:183 &amp; 184;
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1896.tb03085.x" author="Flower, SS" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="75" pagination="856 - 914" refId="B29" refString="Flower, SS, 1896. Notes on a second collection of batrachians made in the Malay Peninsula 1895-96, with a list of the species recorded from that region. Proceedings of the Zoological Society of London 1896: 856 - 914, DOI: https://doi.org/10.1111/j.1096-3642.1896.tb03085.x" title="Notes on a second collection of batrachians made in the Malay Peninsula 1895 - 96, with a list of the species recorded from that region." url="https://doi.org/10.1111/j.1096-3642.1896.tb03085.x" volume="1896" year="1896">Flower 1896</bibRefCitation>
:904, 1899:896 &amp; 897;
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.115429" author="Hanitsch, R" journalOrPublisher="Journal of the Straits Branch of the Royal Asiatic Society" pageId="0" pageNumber="75" pagination="48 - 88" refId="B37" refString="Hanitsch, R, 1900. An expedition to Mount Kina Balu, British North Borneo. Journal of the Straits Branch of the Royal Asiatic Society 34: 48 - 88, DOI: https://doi.org/10.5962/bhl.title.115429" title="An expedition to Mount Kina Balu, British North Borneo." url="https://doi.org/10.5962/bhl.title.115429" volume="34" year="1900">Hanitsch 1900</bibRefCitation>
:73;
<bibRefCitation author="Butler, AL" journalOrPublisher="Journal of the Bombay Natural History Society" pageId="0" pageNumber="75" pagination="193 - 205" refId="B10" refString="Butler, AL, 1904. A list of the batrachians known to inhabit the Malay Peninsula, with some remarks on their habits and distribution. Journal of the Bombay Natural History Society 15: 193 - 205" title="A list of the batrachians known to inhabit the Malay Peninsula, with some remarks on their habits and distribution." volume="15" year="1904">Butler 1904</bibRefCitation>
:199;
<bibRefCitation author="Smith, MA" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="75" pagination="8 - 32" refId="B84" refString="Smith, MA, 1931. The herpetology of Mt. Kinabalu, North Borneo, 13,455 ft. Bulletin of the Raffles Museum 5: 8 - 32" title="The herpetology of Mt. Kinabalu, North Borneo, 13,455 ft." volume="5" year="1931">Smith 1931</bibRefCitation>
:16;
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12187" author="Van Kampen, PN" journalOrPublisher="Brill Ltd., Leiden" pageId="0" pageNumber="75" refId="B92" refString="Van Kampen, PN, 1923. The Amphibia of the Indo-Australian Archipelago. E.J. Brill Ltd., Leiden, DOI: https://doi.org/10.5962/bhl.title.12187" title="The Amphibia of the Indo-Australian Archipelago. E. J." url="https://doi.org/10.5962/bhl.title.12187" year="1923">Van Kampen 1923</bibRefCitation>
:196 &amp; 197;
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.13347" author="Taylor, EH" journalOrPublisher="University of Kansas Science Bulletin" pageId="0" pageNumber="75" pagination="265 - 599" refId="B88" refString="Taylor, EH, 1962. The amphibian fauna of Thailand. University of Kansas Science Bulletin 43: 265 - 599, DOI: https://doi.org/10.5962/bhl.part.13347" title="The amphibian fauna of Thailand." url="https://doi.org/10.5962/bhl.part.13347" volume="43" year="1962">Taylor 1962</bibRefCitation>
:451-453;
<bibRefCitation author="Inger, RF" journalOrPublisher="Journal of the Washington Academy of Sciences" pageId="0" pageNumber="75" pagination="250 - 251" refId="B42" refString="Inger, RF, 1954. On a collection of amphibians from Mount Kina Balu, North Borneo. Journal of the Washington Academy of Sciences 44: 250 - 251" title="On a collection of amphibians from Mount Kina Balu, North Borneo." volume="44" year="1954">Inger 1954</bibRefCitation>
:250, 1966:206-208, 1978:312,
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.3403" author="Inger, RF" journalOrPublisher="Fieldiana: Zoology" pageId="0" pageNumber="75" pagination="1 - 89" refId="B45" refString="Inger, RF, 1985. Tadpoles of the forested regions of Borneo. Fieldiana: Zoology 26: 1 - 89, DOI: https://doi.org/10.5962/bhl.title.3403" title="Tadpoles of the forested regions of Borneo." url="https://doi.org/10.5962/bhl.title.3403" volume="26" year="1985">1985</bibRefCitation>
:iii, 5, 38, 40, 56, 57, 76;
<bibRefCitation author="Grandison, AGC" journalOrPublisher="Bulletin of the British Museum of Natural History (Zoology)" pageId="0" pageNumber="75" pagination="45 - 101" refId="B32" refString="Grandison, AGC, 1972. The Gunong Benom Expedition 1967. 5. Reptiles and amphibians of Gunong Benom with a description of a new species of Macrocalamus. Bulletin of the British Museum of Natural History (Zoology) 23: 45 - 101" title="The Gunong Benom Expedition 1967. 5. Reptiles and amphibians of Gunong Benom with a description of a new species of Macrocalamus." volume="23" year="1972">Grandison 1972</bibRefCitation>
:66 &amp; 67;
<bibRefCitation author="Berry, PY" journalOrPublisher="Tropical Press, Kuala Lumpur" pageId="0" pageNumber="75" refId="B2" refString="Berry, PY, 1975. The amphibian fauna of Peninsular Malaysia. Tropical Press, Kuala Lumpur" title="The amphibian fauna of Peninsular Malaysia." year="1975">Berry 1975</bibRefCitation>
:74 &amp; 75, Fig. 50 (right);
<bibRefCitation author="Matsui, M" journalOrPublisher="Contributions from the Biological Laboratory, Kyoto University" pageId="0" pageNumber="75" pagination="303 - 346" refId="B64" refString="Matsui, M, 1979. Amphibians from Sabah I. Systematic and natural history notes. Contributions from the Biological Laboratory, Kyoto University 25: 303 - 346" title="Amphibians from Sabah I. Systematic and natural history notes." volume="25" year="1979">Matsui 1979</bibRefCitation>
:333; Inger &amp; Stuebing 1992:42, 2005:19, 27, 33, 149 &amp; 150; Inger &amp; Tan 1996:563;
<bibRefCitation author="Inger, RF" editor="Wong, KM" journalOrPublisher="Sabah Society; Sabah Parks, Kota Kinabalu, Sabah" pageId="0" pageNumber="75" pagination="352 - 367" refId="B49" refString="Inger, RF, Stuebing, RB, Tan, FL, 1996. Frogs &amp; toads. In: Wong, KM, Phillipps, A, Eds., Kinabalu: Summit of Borneo. Sabah Society; Sabah Parks, Kota Kinabalu, Sabah: 352 - 367" title="Frogs &amp; toads." volumeTitle="Kinabalu: Summit of Borneo." year="1996">Inger et al. 1996</bibRefCitation>
:363;
<bibRefCitation DOI="https://doi.org/10.1002/mmnz.19940700203" author="Malkmus, R" journalOrPublisher="Mitteilungen aus dem Zoologischen Museum in Berlin" pageId="0" pageNumber="75" pagination="217 - 249" refId="B60" refString="Malkmus, R, 1994. Herpetologische Beobachtungen am Mount Kinabalu, Nord-Borneo. IV. Mitteilungen aus dem Zoologischen Museum in Berlin 70: 217 - 249, DOI: https://doi.org/10.1002/mmnz.19940700203" title="Herpetologische Beobachtungen am Mount Kinabalu, Nord-Borneo. IV." url="https://doi.org/10.1002/mmnz.19940700203" volume="70" year="1994">Malkmus 1994</bibRefCitation>
:244; Manthey &amp; Grossmann 1997:111 &amp; 112; Khonsue &amp; Thirakhupt 2001:73;
<bibRefCitation author="Malkmus, R" journalOrPublisher="Ganter Verlag, Ruggel" pageId="0" pageNumber="75" refId="B61" refString="Malkmus, R, Manthey, U, Vogel, G, Hoffmann, P, Kosuch, J, 2002. Amphibians and Reptiles of Mount Kinabalu (North Borneo). A.R.G. Ganter Verlag, Ruggel" title="Amphibians and Reptiles of Mount Kinabalu (North Borneo). A. R. G." year="2002">Malkmus et al. 2002</bibRefCitation>
: 163-165, figs 159, 160;
<bibRefCitation author="Das, I" journalOrPublisher="Amphibian and Reptile Conservation" pageId="0" pageNumber="75" pagination="3 - 11" refId="B20" refString="Das, I, 2006a. Crocker Range National Park, Sabah, as a refuge for Borneo's montane herpetofauna. Amphibian and Reptile Conservation 4: 3 - 11" title="Crocker Range National Park, Sabah, as a refuge for Borneo's montane herpetofauna." volume="4" year="2006 a">Das 2006a</bibRefCitation>
:5.
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1871" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rana (Hylorana) luctuosa" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="luctuosa" subGenus="Hylorana">Rana (Hylorana) luctuosa</taxonomicName>
(in part):
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12471" author="Boulenger, GA" journalOrPublisher="Records of the Indian Museum" pageId="0" pageNumber="75" pagination="1 - 226" refId="B6" refString="Boulenger, GA, 1920. A monograph of the South Asian, Papuan, Melanesian and Australian frogs of the genus Rana. Records of the Indian Museum 20: 1 - 226, DOI: https://doi.org/10.5962/bhl.title.12471" title="A monograph of the South Asian, Papuan, Melanesian and Australian frogs of the genus Rana." url="https://doi.org/10.5962/bhl.title.12471" volume="20" year="1920">Boulenger 1920</bibRefCitation>
:126.
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1871" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rana (Hylarana) luctuosa" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="luctuosa" subGenus="Hylarana">Rana (Hylarana) luctuosa</taxonomicName>
(in part):
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12187" author="Van Kampen, PN" journalOrPublisher="Brill Ltd., Leiden" pageId="0" pageNumber="75" refId="B92" refString="Van Kampen, PN, 1923. The Amphibia of the Indo-Australian Archipelago. E.J. Brill Ltd., Leiden, DOI: https://doi.org/10.5962/bhl.title.12187" title="The Amphibia of the Indo-Australian Archipelago. E. J." url="https://doi.org/10.5962/bhl.title.12187" year="1923">Van Kampen 1923</bibRefCitation>
:196 &amp; 197;
<bibRefCitation author="Bourret, R" journalOrPublisher="lnstitut Oceanographique de l'lndochine, Hanoi" pageId="0" pageNumber="75" refId="B9" refString="Bourret, R, 1942. Les Batraciens de l'lndochine. lnstitut Oceanographique de l'lndochine, Hanoi" title="Les Batraciens de l'lndochine." year="1942">Bourret 1942</bibRefCitation>
:360; Dubois, 1987: 42.
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1871" family="Ranidae" genus="Hylorana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hylorana luctuosa" order="Anura" pageId="0" pageNumber="75" rank="species" species="luctuosa">Hylorana luctuosa</taxonomicName>
(in part):
<bibRefCitation author="Deckert, K" journalOrPublisher="Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin" pageId="0" pageNumber="75" pagination="127 - 184" refId="B24" refString="Deckert, K, 1938. Beitraege zur Osteologie und Systematik ranider Froschlurche. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1938: 127 - 184" title="Beitraege zur Osteologie und Systematik ranider Froschlurche." volume="1938" year="1938">Deckert 1938</bibRefCitation>
:144
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1871" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rana (Pulchrana) luctuosa" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="luctuosa" subGenus="Pulchrana">Rana (Pulchrana) luctuosa</taxonomicName>
(in part):
<bibRefCitation author="Dubois, A" journalOrPublisher="Bulletin Mensuel de la Societe Linneenne de Lyon" pageId="0" pageNumber="75" pagination="305 - 352" refId="B26" refString="Dubois, A, 1992. Notes sur la classification des Ranidae (Amphibiens anoures). Bulletin Mensuel de la Societe Linneenne de Lyon 61: 305 - 352" title="Notes sur la classification des Ranidae (Amphibiens anoures)." volume="61" year="1992">Dubois 1992</bibRefCitation>
:326
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1871" class="Amphibia" family="Ranidae" genus="Pulchrana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchrana luctuosa" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="luctuosa">Pulchrana luctuosa</taxonomicName>
(in part):
<bibRefCitation DOI="https://doi.org/10.1206/0003-0090(2006)297[0001:tatol]2.0.co;2" author="Frost, DR" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="75" refId="B31" refString="Frost, DR, Grant, T, Faivovich, J, Bain, RH, Haas, A, Haddad, CFB, de Sa, RO, Channing, A, Wilkinson, M, Donnellan, SC, Raxworthy, CJ, Campbell, JA, Blotto, BL, Moler, PE, Drewes, RC, Nussbaum, RA, Lynch, JD, Green, DM, Wheeler, WC, 2006. The amphibian tree of life. Bulletin of the American Museum of Natural History 297: 1-291. https://doi.org/10.1206/0003-0090(2006)297[0001:tatol]2.0.co;2" title="The amphibian tree of life. Bulletin of the American Museum of Natural History 297: 1 - 291." url="https://doi.org/10.1206/0003-0090(2006)297[0001:tatol]2.0.co;2" year="2006">Frost et al. 2006</bibRefCitation>
:369;
<bibRefCitation author="Fei, L" journalOrPublisher="Herpetologica Sinica" pageId="0" pageNumber="75" pagination="1 - 43" refId="B28" refString="Fei, L, Ye, C, Jiang, J, 2010. Phylogenetic systematics of Ranidae. Herpetologica Sinica 12: 1 - 43" title="Phylogenetic systematics of Ranidae." volume="12" year="2010">Fei et al. 2010</bibRefCitation>
:33.
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1871" class="Amphibia" family="Ranidae" genus="Hylarana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hylarana luctuosa" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="luctuosa">Hylarana luctuosa</taxonomicName>
(in part):
<bibRefCitation author="Manthey, U" journalOrPublisher="- herpetofauna, Weinstadt" pageId="0" pageNumber="75" pagination="20 - 31" refId="B62" refString="Manthey, U, 1983. Exkursion am Mt. Kinabalu 4100 m, Nord - Borneo. Teil 3: Checkliste der Herpetofauna oberhalb 600m ueNN. - herpetofauna, Weinstadt 5: 20 - 31" title="Exkursion am Mt. Kinabalu 4100 m, Nord - Borneo. Teil 3: Checkliste der Herpetofauna oberhalb 600 m ueNN." volume="5" year="1983">Manthey 1983</bibRefCitation>
:22;
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2006.11.032" author="Che, J" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="75" pagination="1 - 13" refId="B17" refString="Che, J, Pang, J, Zhao, H, Wu, GF, Zhao, EM, Zhang, YP, 2007. Phylogeny of Raninae (Anura: Ranidae) inferred from mitochondrial and nuclear sequences. Molecular Phylogenetics and Evolution 43: 1 - 13, DOI: https://doi.org/10.1016/j.ympev.2006.11.032" title="Phylogeny of Raninae (Anura: Ranidae) inferred from mitochondrial and nuclear sequences." url="https://doi.org/10.1016/j.ympev.2006.11.032" volume="43" year="2007">Che et al. 2007</bibRefCitation>
: 1-13 (by implication);
<bibRefCitation author="Das, I" journalOrPublisher="Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut" pageId="0" pageNumber="75" pagination="141 - 174" refId="B23" refString="Das, I, Jankowski, A, Makmor, MIB, Haas, A, 2007. Species diversity, elevational distribution and reproductive modes in an amphibian community at the Matang Range, Sarawak (Borneo). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 104: 141 - 174" title="Species diversity, elevational distribution and reproductive modes in an amphibian community at the Matang Range, Sarawak (Borneo)." volume="104" year="2007">Das et al. 2007</bibRefCitation>
:159, 160, 167, 170;
<bibRefCitation author="Imbun, PY" journalOrPublisher="Dewan Bahasa dan Pustaka, Kuala Lumpur" pageId="0" pageNumber="75" refId="B41" refString="Imbun, PY, 2014. Amfibia Sabah. Dewan Bahasa dan Pustaka, Kuala Lumpur" title="Amfibia Sabah." year="2014">Imbun 2014</bibRefCitation>
:99.
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1871" class="Amphibia" family="Ranidae" genus="Abavorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abavorana luctuosa" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="luctuosa">Abavorana luctuosa</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2015.05.001" author="Oliver, LA" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="75" pagination="176 - 192" refId="B71" refString="Oliver, LA, Prendini, E, Kraus, F, Raxworthy, CJ, 2015. Systematics and biogeography of the Hylarana frog (Anura: Ranidae) radiation across tropical Australasia, Southeast Asia, and Africa. Molecular Phylogenetics and Evolution 90: 176 - 192, DOI: https://doi.org/10.1016/j.ympev.2015.05.001" title="Systematics and biogeography of the Hylarana frog (Anura: Ranidae) radiation across tropical Australasia, Southeast Asia, and Africa." url="https://doi.org/10.1016/j.ympev.2015.05.001" volume="90" year="2015">Oliver et al. 2015</bibRefCitation>
:186 (in part);
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4320.2.4" author="Quah, ESH" journalOrPublisher="Zootaxa" pageId="0" pageNumber="75" pagination="272 - 288" refId="B75" refString="Quah, ESH, Shahrul, AMS, Grismer, LL, Wood, JrPL, Siti, AMN, Muin, MA, 2017. A new species of frog of the genus Abavorana Oliver, Prendini, Kraus &amp; Raxworthy 2015 (Anura: Ranidae) from Gunung Jerai, Kedah, northwestern Peninsular Malaysia. Zootaxa 4320: 272 - 288, DOI: https://doi.org/10.11646/zootaxa.4320.2.4" title="A new species of frog of the genus Abavorana Oliver, Prendini, Kraus &amp; Raxworthy 2015 (Anura: Ranidae) from Gunung Jerai, Kedah, northwestern Peninsular Malaysia." url="https://doi.org/10.11646/zootaxa.4320.2.4" volume="4320" year="2017">Quah et al. 2017</bibRefCitation>
: 272-288 (in part);
<bibRefCitation author="Zainudin, R" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="75" pagination="876 - 891" refId="B93" refString="Zainudin, R, Zain, BMM, Ahmad, N, Nor, SM, 2017. Microhabitat partitioning of closely related Sarawak (Malaysian Borneo) frog species previously assigned to the genus Hylarana (Amphibia: Anura). Turkish Journal of Zoology 41: 876 - 891" title="Microhabitat partitioning of closely related Sarawak (Malaysian Borneo) frog species previously assigned to the genus Hylarana (Amphibia: Anura)." volume="41" year="2017">Zainudin et al. 2017</bibRefCitation>
:876-891 (in part);
<bibRefCitation DOI="https://doi.org/10.3897/evolsyst.2.27020" author="Haas, A" journalOrPublisher="Evolutionary Systematics" pageId="0" pageNumber="75" pagination="89 - 114" refId="B36" refString="Haas, A, Kueh, BH, Joseph, A, Asri, Mb, Das, I, Hagmann, R, Schwander, L, Hertwig, ST, 2018. An updated checklist of the amphibian diversity of Maliau Basin Conservation Area, Sabah, Malaysia. Evolutionary Systematics 2: 89 - 114, DOI: https://doi.org/10.3897/evolsyst.2.27020" title="An updated checklist of the amphibian diversity of Maliau Basin Conservation Area, Sabah, Malaysia." url="https://doi.org/10.3897/evolsyst.2.27020" volume="2" year="2018">Haas et al. 2018</bibRefCitation>
:92, 93, 100, fig. 4b.
</paragraph>
<paragraph pageId="0" pageNumber="75">
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1871" class="Amphibia" family="Ranidae" genus="Hylarana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hylarana (Abavorana) luctuosa" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="luctuosa" subGenus="Abavorana">Hylarana (Abavorana) luctuosa</taxonomicName>
(in part):
<bibRefCitation author="Inger, RF" journalOrPublisher=", Kota Kinabalu, Sabah" pageId="0" pageNumber="75" refId="B48" refString="Inger, RF, Stuebing, RB, Grafe, TU, Dehling, JM, 2017. A Field Guide to the Frogs of Borneo. 3rd Edition. Natural History Publications (Borneo) Sdn Bhd., Kota Kinabalu, Sabah" title="A Field Guide to the Frogs of Borneo. 3 rd Edition. Natural History Publications (Borneo) Sdn Bhd." year="2017">Inger et al. 2017</bibRefCitation>
:147 &amp; 148.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="75" type="syntypes">
<paragraph pageId="0" pageNumber="75">Syntypes.</paragraph>
<paragraph pageId="0" pageNumber="75">
MNHNP 1889.226-28 from
<normalizedToken originalValue="“Bornéo">&quot;Borneo</normalizedToken>
Kina Balu&quot;, Sabah (Borneo), Malaysia. (
<bibRefCitation author="Guibe, J" journalOrPublisher="Imprimerie nationale, Paris" pageId="0" pageNumber="75" refId="B35" refString="Guibe, J, 1950. Catalogue des types d'amphibiens du Museum national d'Histoire naturelle. Imprimerie nationale, Paris" title="Catalogue des types d'amphibiens du Museum national d'Histoire naturelle." year="1950">
<normalizedToken originalValue="Guibé">Guibe</normalizedToken>
1950
</bibRefCitation>
) (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Syntypes of Abavorana decorata; A &amp; D: Dorsum and venter of MNHN-RA- 1889.226. B &amp; E: Dorsum and venter of MNHN-RA- 1889.227. C &amp; F: Dorsum and venter of MNHN-RA- 1889.228 (Photographs by Christine Reinhardt). G: Original illustration of the syntypes of Abavorana decorata from Mocquard (1890). Illustrations assignable to specimen MNHN-RA- 1889.226 (left) and MNHN-RA- 1889.228 (right)." figureDoi="10.3897/vz.71.e60921.figure5" httpUri="https://binary.pensoft.net/fig/518978" pageId="0" pageNumber="75">5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="75" type="diagnosis">
<paragraph pageId="0" pageNumber="75">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="75">
In addition to its phylogenetic placement (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogenetic tree of the genus Abavorana; Maximum likelihood phylogram for species of the genus Abavorana. Nodal support is indicated by Bayesian posterior probability (BPP) / Ultrafast bootstrap (UFboot) respectively." figureDoi="10.3897/vz.71.e60921.figure1" httpUri="https://binary.pensoft.net/fig/518974" pageId="0" pageNumber="75">1</figureCitation>
),
<taxonomicName authorityName="Mocquard" authorityYear="1890" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rana decorata" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="decorata">
<emphasis italics="true" pageId="0" pageNumber="75">Rana decorata</emphasis>
</taxonomicName>
is reassigned as a member of the genus
<taxonomicName authorityName="Oliver, Prendini, Kraus &amp; Raxworthy" authorityYear="2015" class="Amphibia" family="Ranidae" genus="Abavorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abavorana" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="75">Abavorana</emphasis>
</taxonomicName>
based on the combination of having a robust, medium-sized body; no vocal sacs in males; length of 1st finger greater than 2nd finger; disc width to finger width ratios of finger 3 and toe 4 is 1-1.5; dorsolateral fold indistinct or absent; the colour of the dorsolateral line being white or yellow; the humeral gland in males is prominent, raised and centrally positioned on the ventral surface of the upper arm; a weak or absent rictal ridge; outer metatarsal tubercle weak or absent; skin of dorsum smooth or finely shagreened; throat, abdomen and flanks smooth; posterior section of venter and back of the thigh rugose; no pale colouration on the margins of the tympanum; flanks dark-brown or black below the dorsal fold grading into a pale venter (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.3147" author="Inger, RF" journalOrPublisher="Fieldiana: Zoology" pageId="0" pageNumber="75" pagination="1 - 402" refId="B43" refString="Inger, RF, 1966. The systematics and zoogeography of the Amphibia of Borneo. Fieldiana: Zoology 52: 1 - 402, DOI: https://doi.org/10.5962/bhl.title.3147" title="The systematics and zoogeography of the Amphibia of Borneo." url="https://doi.org/10.5962/bhl.title.3147" volume="52" year="1966">Inger 1966</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2015.05.001" author="Oliver, LA" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="75" pagination="176 - 192" refId="B71" refString="Oliver, LA, Prendini, E, Kraus, F, Raxworthy, CJ, 2015. Systematics and biogeography of the Hylarana frog (Anura: Ranidae) radiation across tropical Australasia, Southeast Asia, and Africa. Molecular Phylogenetics and Evolution 90: 176 - 192, DOI: https://doi.org/10.1016/j.ympev.2015.05.001" title="Systematics and biogeography of the Hylarana frog (Anura: Ranidae) radiation across tropical Australasia, Southeast Asia, and Africa." url="https://doi.org/10.1016/j.ympev.2015.05.001" volume="90" year="2015">Oliver et al. 2015</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4320.2.4" author="Quah, ESH" journalOrPublisher="Zootaxa" pageId="0" pageNumber="75" pagination="272 - 288" refId="B75" refString="Quah, ESH, Shahrul, AMS, Grismer, LL, Wood, JrPL, Siti, AMN, Muin, MA, 2017. A new species of frog of the genus Abavorana Oliver, Prendini, Kraus &amp; Raxworthy 2015 (Anura: Ranidae) from Gunung Jerai, Kedah, northwestern Peninsular Malaysia. Zootaxa 4320: 272 - 288, DOI: https://doi.org/10.11646/zootaxa.4320.2.4" title="A new species of frog of the genus Abavorana Oliver, Prendini, Kraus &amp; Raxworthy 2015 (Anura: Ranidae) from Gunung Jerai, Kedah, northwestern Peninsular Malaysia." url="https://doi.org/10.11646/zootaxa.4320.2.4" volume="4320" year="2017">Quah et al. 2017</bibRefCitation>
). Dorsum reddish-orange to rust-brown, encircled by a white or cream coloured dorsolateral line that encircles the snout, canthus rostralis, outer edge of the upper eyelids, and dorsum along the dorsolateral fold to the vent; lower flanks dark-brown or black below the dorsolateral line grading into a paler venter; dorsal colouration of the limbs light-grey or brown with speckling and prominent dark-brown or black bands.
<taxonomicName authorityName="comb. nov. (Mocquard" authorityYear="1890" class="Amphibia" family="Ranidae" genus="Abavorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abavorana decorata" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="decorata">
<emphasis italics="true" pageId="0" pageNumber="75">Abavorana decorata</emphasis>
</taxonomicName>
can be differentiated from its congeners on the basis of its ventral colour pattern which is reticulated in black and white especially on the lower flanks, underside of the limbs (especially hind-limbs) are boldly banded in black and white, and a prominent white streak under the eye and/or tympanum to the corner of the jaw (Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Syntypes of Abavorana decorata; A &amp; D: Dorsum and venter of MNHN-RA- 1889.226. B &amp; E: Dorsum and venter of MNHN-RA- 1889.227. C &amp; F: Dorsum and venter of MNHN-RA- 1889.228 (Photographs by Christine Reinhardt). G: Original illustration of the syntypes of Abavorana decorata from Mocquard (1890). Illustrations assignable to specimen MNHN-RA- 1889.226 (left) and MNHN-RA- 1889.228 (right)." figureDoi="10.3897/vz.71.e60921.figure5" httpUri="https://binary.pensoft.net/fig/518978" pageId="0" pageNumber="75">5</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Examined Abavorana decorata specimens; Dorsal (A &amp; C) and ventral (B &amp; D) pattern of Abavorana decorata from Borneo. A &amp; B (from left to right): FMNH 109935 from Kiau, Mount Kinabalu, Sabah (left), FMNH 109936 from Kiau, Mount Kinabalu, Sabah (middle), FMNH 231052 from Danum Valley Research Center, Lahad Datu District, Sabah (top right), FMNH 128127 from Bario, Sarawak (bottom right). C &amp; D: SP 26873 from Tawau Hills, Sabah (left) and SP 02810 from Maliau Basin, Sabah." figureDoi="10.3897/vz.71.e60921.figure6" httpUri="https://binary.pensoft.net/fig/518979" pageId="0" pageNumber="75">6</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Colour pattern comparison among species of the genus Abavorana; Dorsum (A, C &amp; E) and venter (B, D &amp; F) in life of A. luctuosa (USMHC 2576) (A &amp; B) from Penang Hill, Penang Island, Malaysia, A. decorata from Maliau Basin, Sabah, East Malaysia, Borneo (C &amp; D) and A. nazgul (USMHC 2233) (E &amp; F) from Gunung Jerai, Kedah, West Malaysia. (Photographs by Evan Quah [A, B, E &amp; F] and Stefan T. Hertwig [C &amp; D])." figureDoi="10.3897/vz.71.e60921.figure9" httpUri="https://binary.pensoft.net/fig/518982" pageId="0" pageNumber="75">9C-D</figureCitation>
). Adult males with SVL 46.21-56.0 mm, adult females with SVL 29.66-58.42 mm; adult males with SW 9.91-10.71 mm, adult females with SW 6.25-11.68 mm; adult males with IOD 5.59-5.95 mm, adult females with IOD 3.91-6.78 mm; adult males with TD 4.54-4.60 mm, adult females with TD 2.27-5.63 mm; adult males with FL 25.58-26.10 mm, adult females with FL 17.19-31.34 mm (Table
<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Summary statistics and ventral colour pattern among members of the genus Abavorana. Summary statistics and ventral colour pattern for Abavorana decorata, A. nazgul, and A. luctuosa populations from the Malay Peninsula and Borneo, SD = standard deviation, N = sample size, / = no data. Humeral gland only present in males and all measurements in mm." httpUri="http://table.plazi.org/id/B91F734E592A25063FCA3E8C57D388D3" pageId="0" pageNumber="75" tableUuid="B91F734E592A25063FCA3E8C57D388D3">5</tableCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="75">
<emphasis bold="true" pageId="0" pageNumber="75">
Distribution (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution map of Abavorana decorata and A. luctuosa in Borneo; Known distribution of Abavorana decorata (red) and A. luctuosa (blue) in Borneo. Star = type locality." figureDoi="10.3897/vz.71.e60921.figure10" httpUri="https://binary.pensoft.net/fig/518983" pageId="0" pageNumber="75">10</figureCitation>
)
</emphasis>
. Endemic to Borneo. The species is known from Sabah state, Malaysian Borneo: Mount Kinabalu, Kampung Kiau Taburi, Danum Valley, Tawau Hills Park, Maliau Basin, Tambunan and Penampang (CalPhotos ID: 0000 0000 0912 1159); Sarawak state, Malaysia Borneo: Kubah National Park (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Living specimens of Abavorana decorata and Bornean A. luctuosa; A: Abavorana decorata from Tawau Hills, Sabah, East Malaysia. B: A. decorata from Maliau Basin, Sabah, East Malaysia. C: A. decorata from Kubah National Park, Sarawak, East Malaysia. D: A. decorata from Melawi Regency, West Kalimantan Province, Indonesia. E &amp; F: A. luctuosa from Kubah National Park, Sarawak, East Malaysia. G: A. luctuosa (FMNH 238720) from Sipitang, Sabah, East Malaysia. H: A. luctuosa (FMNH 242883) from Sipitang, Sabah, East Malaysia. (Photographs by Alexander Haas [A, C &amp; E], Paul Bertner [B], Muhammad Aini [D], Chien C. Lee [F], and Robert F. Inger [G &amp; H], Figs G &amp; H © Field Museum of Natural History. Created by Field Museum of Natural History, Amphibian and Reptile Collection and licensed under CC-BY-SA 4.0)." figureDoi="10.3897/vz.71.e60921.figure7" httpUri="https://binary.pensoft.net/fig/518980" pageId="0" pageNumber="75">7C</figureCitation>
) and Bario (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Examined Abavorana decorata specimens; Dorsal (A &amp; C) and ventral (B &amp; D) pattern of Abavorana decorata from Borneo. A &amp; B (from left to right): FMNH 109935 from Kiau, Mount Kinabalu, Sabah (left), FMNH 109936 from Kiau, Mount Kinabalu, Sabah (middle), FMNH 231052 from Danum Valley Research Center, Lahad Datu District, Sabah (top right), FMNH 128127 from Bario, Sarawak (bottom right). C &amp; D: SP 26873 from Tawau Hills, Sabah (left) and SP 02810 from Maliau Basin, Sabah." figureDoi="10.3897/vz.71.e60921.figure6" httpUri="https://binary.pensoft.net/fig/518979" pageId="0" pageNumber="75">6A-B</figureCitation>
); and West Kalimantan, Indonesia: Melawi Regency (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Living specimens of Abavorana decorata and Bornean A. luctuosa; A: Abavorana decorata from Tawau Hills, Sabah, East Malaysia. B: A. decorata from Maliau Basin, Sabah, East Malaysia. C: A. decorata from Kubah National Park, Sarawak, East Malaysia. D: A. decorata from Melawi Regency, West Kalimantan Province, Indonesia. E &amp; F: A. luctuosa from Kubah National Park, Sarawak, East Malaysia. G: A. luctuosa (FMNH 238720) from Sipitang, Sabah, East Malaysia. H: A. luctuosa (FMNH 242883) from Sipitang, Sabah, East Malaysia. (Photographs by Alexander Haas [A, C &amp; E], Paul Bertner [B], Muhammad Aini [D], Chien C. Lee [F], and Robert F. Inger [G &amp; H], Figs G &amp; H © Field Museum of Natural History. Created by Field Museum of Natural History, Amphibian and Reptile Collection and licensed under CC-BY-SA 4.0)." figureDoi="10.3897/vz.71.e60921.figure7" httpUri="https://binary.pensoft.net/fig/518980" pageId="0" pageNumber="75">7D</figureCitation>
). The species is expected to be wider ranging on the island.
</paragraph>
<caption doi="10.3897/vz.71.e60921.figure7" httpUri="https://binary.pensoft.net/fig/518980" pageId="0" pageNumber="75" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="75">
<emphasis bold="true" pageId="0" pageNumber="75">Figure 7.</emphasis>
Living specimens of
<taxonomicName authorityName="comb. nov. (Mocquard" authorityYear="1890" class="Amphibia" family="Ranidae" genus="Abavorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abavorana decorata" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="decorata">
<emphasis italics="true" pageId="0" pageNumber="75">Abavorana decorata</emphasis>
</taxonomicName>
and Bornean
<taxonomicName genus="A." lsidName="A. luctuosa" pageId="0" pageNumber="75" rank="species" species="luctuosa">
<emphasis italics="true" pageId="0" pageNumber="75">A. luctuosa</emphasis>
</taxonomicName>
; A:
<taxonomicName authorityName="comb. nov. (Mocquard" authorityYear="1890" class="Amphibia" family="Ranidae" genus="Abavorana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abavorana decorata" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="decorata">
<emphasis italics="true" pageId="0" pageNumber="75">Abavorana decorata</emphasis>
</taxonomicName>
from Tawau Hills, Sabah, East Malaysia. B:
<taxonomicName genus="A." lsidName="A. decorata" pageId="0" pageNumber="75" rank="species" species="decorata">
<emphasis italics="true" pageId="0" pageNumber="75">A. decorata</emphasis>
</taxonomicName>
from Maliau Basin, Sabah, East Malaysia. C:
<taxonomicName genus="A." lsidName="A. decorata" pageId="0" pageNumber="75" rank="species" species="decorata">
<emphasis italics="true" pageId="0" pageNumber="75">A. decorata</emphasis>
</taxonomicName>
from Kubah National Park, Sarawak, East Malaysia. D:
<taxonomicName genus="A." lsidName="A. decorata" pageId="0" pageNumber="75" rank="species" species="decorata">
<emphasis italics="true" pageId="0" pageNumber="75">A. decorata</emphasis>
</taxonomicName>
from Melawi Regency, West Kalimantan Province, Indonesia. E &amp; F:
<taxonomicName genus="A." lsidName="A. luctuosa" pageId="0" pageNumber="75" rank="species" species="luctuosa">
<emphasis italics="true" pageId="0" pageNumber="75">A. luctuosa</emphasis>
</taxonomicName>
from Kubah National Park, Sarawak, East Malaysia. G:
<taxonomicName genus="A." lsidName="A. luctuosa" pageId="0" pageNumber="75" rank="species" species="luctuosa">
<emphasis italics="true" pageId="0" pageNumber="75">A. luctuosa</emphasis>
</taxonomicName>
(FMNH 238720) from Sipitang, Sabah, East Malaysia. H:
<taxonomicName genus="A." lsidName="A. luctuosa" pageId="0" pageNumber="75" rank="species" species="luctuosa">
<emphasis italics="true" pageId="0" pageNumber="75">A. luctuosa</emphasis>
</taxonomicName>
(FMNH 242883) from Sipitang, Sabah, East Malaysia. (Photographs by Alexander Haas [A, C &amp; E], Paul Bertner [B], Muhammad Aini [D], Chien C. Lee [F], and Robert F. Inger [G &amp; H], Figs G &amp; H ©Field Museum of Natural History. Created by Field Museum of Natural History, Amphibian and Reptile Collection and licensed under CC-BY-SA 4.0).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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