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<document id="A1083EC9788EDE98CEA11780BBCBCC9A" ID-DOI="10.11646/zootaxa.3931.1.4" ID-GBIF-Dataset="091b43a3-d592-433e-ad7f-6de13b4e9da7" ID-ISSN="1175-5326" ID-Zenodo-Dep="288140" ID-ZooBank="1E897D84-6030-4C39-A215-E55DB18ACB87" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461009762838" checkinUser="plazi" docAuthor="Grismer, L. Lee &amp; Quah, Evan S. H." docDate="2015" docId="03E5A21DFFE93247FF16FF2BFD3DFE69" docLanguage="en" docName="zt03931p070.pdf" docOrigin="Zootaxa 3931 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sphenomorphus senja Grismer &amp; Quah, 2015, sp. nov." docType="treatment" docVersion="6" lastPageNumber="67" masterDocId="FFDCDA65FFEB3242FF81FFF7FFC8FF9C" masterDocTitle="The Rediscovery of Sphenomorphus malayanus Doria, 1888 (Squamata: Scincidae) from the Titiwangsa Mountain Range of Peninsular Malaysia and its re-description as S. senja sp. nov." masterLastPageNumber="70" masterPageNumber="63" pageNumber="65" updateTime="1698566368044" updateUser="plazi">
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<mods:title id="C666D4C5B9C4902EEE44C10C1685A6B6">The Rediscovery of Sphenomorphus malayanus Doria, 1888 (Squamata: Scincidae) from the Titiwangsa Mountain Range of Peninsular Malaysia and its re-description as S. senja sp. nov.</mods:title>
</mods:titleInfo>
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<mods:namePart id="2B1431801454EB9EFE441E4A8B696628">Grismer, L. Lee</mods:namePart>
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<mods:namePart id="93385BFCD9F4E73E38CB09D4450A1FC1">Quah, Evan S. H.</mods:namePart>
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<mods:title id="8111300E183B01D16643479FAAB10226">Zootaxa</mods:title>
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<mods:date id="E545BB4F355F38C90EF1339959676A47">2015</mods:date>
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<mods:number id="21008B3F853900CC9BB721066AD553A6">3931</mods:number>
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<mods:identifier id="6A0B04DB13B303B03EEFC186FFB1F2B3" type="DOI">10.11646/zootaxa.3931.1.4</mods:identifier>
<mods:identifier id="5A58AF005B51BB7944A389075F514274" type="GBIF-Dataset">091b43a3-d592-433e-ad7f-6de13b4e9da7</mods:identifier>
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<treatment id="03E5A21DFFE93247FF16FF2BFD3DFE69" ID-DOI="http://doi.org/10.5281/zenodo.5693929" ID-GBIF-Taxon="127686426" ID-Zenodo-Dep="5693929" LSID="urn:lsid:plazi:treatment:03E5A21DFFE93247FF16FF2BFD3DFE69" httpUri="http://treatment.plazi.org/id/03E5A21DFFE93247FF16FF2BFD3DFE69" lastPageId="5" lastPageNumber="67" pageId="2" pageNumber="65">
<subSubSection id="C3564080FFE93240FF16FF2BFF3DFEA0" pageId="2" pageNumber="65" type="nomenclature">
<paragraph id="8BF3130BFFE93240FF16FF2BFDC1FF6B" blockId="2.[151,521,220,317]" box="[151,521,220,247]" pageId="2" pageNumber="65">
<heading id="D0BBA467FFE93240FF16FF2BFDC1FF6B" bold="true" box="[151,521,220,247]" fontSize="11" level="1" pageId="2" pageNumber="65" reason="1">
<emphasis id="B938CF19FFE93240FF16FF2BFDC1FF6B" bold="true" box="[151,521,220,247]" pageId="2" pageNumber="65">
<taxonomicName id="4C4C6888FFE93240FF16FF2BFE6CFF6A" box="[151,420,220,246]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="65" phylum="Chordata" rank="species" species="senja" status="sp. nov.">
<emphasis id="B938CF19FFE93240FF16FF2BFE6CFF6A" bold="true" box="[151,420,220,246]" italics="true" pageId="2" pageNumber="65">Sphenomorphus senja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B7262FFE93240FE2DFF2AFDC1FF6B" box="[428,521,221,247]" pageId="2" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF3130BFFE93240FF16FF08FF3DFEA0" blockId="2.[151,521,220,317]" pageId="2" pageNumber="65">
Titiwangsa Forest Skink
<figureCitation id="13770F8EFFE93240FF16FED3FF3DFEA0" box="[151,245,292,317]" captionStart="FIGURE 2" captionStartId="3.[151,250,932,954]" captionTargetBox="[150,1436,193,910]" captionTargetId="figure@3.[150,1436,193,911]" captionTargetPageId="3" captionText="FIGURE 2. Upper left and right and lower right: holotype of Sphenomorphus senja sp. nov. (LSUHC 11688) from Gunung Brinchang, Cameron Highlands, Pahang. Lower left: paratype of S. senja sp. nov. (BM 1911.12.5.10) from Gunung Gerah, Perak." httpUri="https://zenodo.org/record/288142/files/figure.png" pageId="2" pageNumber="65">Figure 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3564080FFE93240FF16FE9BFAB4FE6C" pageId="2" pageNumber="65" type="materials_examined">
<paragraph id="8BF3130BFFE93240FF16FE9BFBBCFE34" blockId="2.[151,1436,364,640]" pageId="2" pageNumber="65">
<emphasis id="B938CF19FFE93240FF16FE9BFEC3FE18" bold="true" box="[151,267,364,388]" pageId="2" pageNumber="65">
<typeStatus id="54F7ADA9FFE93240FF16FE9BFECEFE18" box="[151,262,364,388]" pageId="2" pageNumber="65" type="holotype">Holotype</typeStatus>
.
</emphasis>
Adult male (
<collectionCode id="ED5D8BCEFFE93240FE2BFE9BFDCFFE18" box="[426,519,364,388]" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="2" pageNumber="65">LSUHC</collectionCode>
11688) collected from Gunung Brinchang,
<collectingCountry id="F35B539BFFE93240FB88FE9BFBB8FE18" box="[1033,1136,364,388]" name="Cameroon" pageId="2" pageNumber="65">Cameron</collectingCountry>
Highlands, Pahang, West
<collectingCountry id="F35B539BFFE93240FF16FE78FF36FE34" box="[151,254,399,424]" name="Malaysia" pageId="2" pageNumber="65">Malaysia</collectingCountry>
(04°
<geoCoordinate id="EE7875CCFFE93240FEB3FE78FE55FE3B" box="[306,413,399,424]" direction="north" orientation="latitude" pageId="2" pageNumber="65" precision="55" value="31.105">31.105 N</geoCoordinate>
101°
<geoCoordinate id="EE7875CCFFE93240FE54FE78FD8AFE3B" box="[469,578,399,424]" direction="east" orientation="longitude" pageId="2" pageNumber="65" precision="55" value="22.571">22.571 E</geoCoordinate>
; at
<quantity id="4CB4BEEEFFE93240FDE9FE67FD7EFE34" box="[616,694,400,424]" metricMagnitude="3" metricUnit="m" metricValue="1.811" pageId="2" pageNumber="65" unit="m" value="1811.0">1811m</quantity>
) on
<date id="FFF235CBFFE93240FD69FE78FC4EFE34" box="[744,902,399,424]" pageId="2" pageNumber="65" value="2014-04-26">26 April 2014</date>
by Evan S. H. Quah.
</paragraph>
<paragraph id="8BF3130BFFE93240FF46FE43FAB4FE6C" blockId="2.[151,1436,364,640]" pageId="2" pageNumber="65">
<emphasis id="B938CF19FFE93240FF46FE43FEF3FE50" bold="true" box="[199,315,436,460]" pageId="2" pageNumber="65">
<typeStatus id="54F7ADA9FFE93240FF46FE43FEFEFE50" box="[199,310,436,460]" pageId="2" pageNumber="65" type="paratype">Paratype</typeStatus>
.
</emphasis>
BM 1911.12.5.10.
<collectingCountry id="F35B539BFFE93240FD95FE43FDFBFE50" box="[532,563,436,460]" name="American Samoa" pageId="2" pageNumber="65">As</collectingCountry>
noted by
<bibRefCitation id="EFDD6EFAFFE93240FD22FE43FCA1FE50" author="Boulenger" box="[675,873,436,461]" pageId="2" pageNumber="65" refString="Boulenger, G. A. (1912) A Vertebrate Fauna of the Malay Peninsula from the Isthmus of Kra to Singapore Including the Adjacent Islands. Reptilia and Batrachia. Taylor &amp; Francis, London, 294 pp." type="book" year="1912">Boulenger (1912)</bibRefCitation>
the specimen was collected by Mr. J. N. Sheffield at
<quantity id="4CB4BEEEFFE93240FF33FE20FEEAFE6C" box="[178,290,471,496]" metricMagnitude="3" metricUnit="m" metricValue="2.1031199999999997" pageId="2" pageNumber="65" unit="ft" value="6900.0">6,900 feet</quantity>
on Gunong Grah [=Gunung Gerah] in the Titiwangsa Mountain Range, Perak during
<date id="FFF235CBFFE93240FB65FE2FFABFFE6C" box="[1252,1399,471,496]" pageId="2" pageNumber="65" value="1905-08">August 1905</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3564080FFE93240FF46FE0CFBC1FD1C" pageId="2" pageNumber="65" type="diagnosis">
<paragraph id="8BF3130BFFE93240FF46FE0CFBC1FD1C" blockId="2.[151,1436,364,640]" pageId="2" pageNumber="65">
<emphasis id="B938CF19FFE93240FF46FE0CFE89FD88" bold="true" box="[199,321,507,532]" pageId="2" pageNumber="65">Diagnosis.</emphasis>
Adults reaching 65.0 mm SVL; body slender, elongate; limbs separated when adpressed; 32 smooth, midbody scale rows; 72 or 73 paravertebral scales; 68 ventral scale rows; four supraoculars; prefrontals in narrow contact or slightly separated; two loreal scales; 13–17 smooth, lamellae beneath Toe IV; enlarged, preanal scales; dorsal body bands absent; numerous dark markings on back and flanks.
</paragraph>
</subSubSection>
<caption id="DF334383FFE93240FF16F8A0FA93F810" httpUri="https://zenodo.org/record/288141/files/figure.png" pageId="2" pageNumber="65" targetBox="[168,1436,721,1835]" targetPageId="2">
<paragraph id="8BF3130BFFE93240FF16F8A0FA93F810" blockId="2.[151,1436,1879,1932]" pageId="2" pageNumber="65">
<emphasis id="B938CF19FFE93240FF16F8A0FEDFF8F0" bold="true" box="[151,279,1879,1901]" pageId="2" pageNumber="65">FIGURE 1.</emphasis>
Distribution of
<taxonomicName id="4C4C6888FFE93240FE49F8A0FD63F8F1" box="[456,683,1879,1901]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="65" phylum="Chordata" rank="species" species="senja" status="sp. nov.">
<emphasis id="B938CF19FFE93240FE49F8A0FD63F8F1" box="[456,683,1879,1901]" italics="true" pageId="2" pageNumber="65">Sphenomorphus senja</emphasis>
</taxonomicName>
<emphasis id="B938CF19FFE93240FD37F8A0FCC4F8F0" bold="true" box="[694,780,1879,1900]" pageId="2" pageNumber="65">
<taxonomicNameLabel id="A20B7262FFE93240FD37F8A0FCC4F8F0" box="[694,780,1879,1900]" pageId="2" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<emphasis id="B938CF19FFE93240FCC6F8AFFC12F8F1" box="[839,986,1879,1901]" italics="true" pageId="2" pageNumber="65">
<taxonomicName id="4C4C6888FFE93240FCC6F8AFFC1EF8F1" box="[839,982,1879,1901]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="65" phylum="Chordata" rank="species" species="malayanus">S. malayanus</taxonomicName>
.
</emphasis>
Stars denote the type localities of Gunug Singgalang, West Sumatra, Indonesia (yellow) and Gunung Brinchang, Cameron Highlands, Pahang, Malaysia (orange).
</paragraph>
</caption>
<caption id="DF334383FFE83241FF16FC53FF10FC64" httpUri="https://zenodo.org/record/288142/files/figure.png" pageId="3" pageNumber="66" targetBox="[150,1436,193,910]" targetPageId="3">
<paragraph id="8BF3130BFFE83241FF16FC53FF10FC64" blockId="3.[151,1436,932,1016]" pageId="3" pageNumber="66">
<emphasis id="B938CF19FFE83241FF16FC53FEDCFC25" bold="true" box="[151,276,932,954]" pageId="3" pageNumber="66">FIGURE 2.</emphasis>
Upper left and right and lower right: holotype of
<taxonomicName id="4C4C6888FFE83241FC9DFC53FC34FC26" box="[796,1020,932,954]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="66" phylum="Chordata" rank="species" species="senja" status="sp. nov.">
<emphasis id="B938CF19FFE83241FC9DFC53FC34FC26" box="[796,1020,932,954]" italics="true" pageId="3" pageNumber="66">Sphenomorphus senja</emphasis>
</taxonomicName>
<emphasis id="B938CF19FFE83241FB85FC52FB9FFC26" bold="true" box="[1028,1111,933,954]" pageId="3" pageNumber="66">
<taxonomicNameLabel id="A20B7262FFE83241FB85FC52FB9FFC26" box="[1028,1111,933,954]" pageId="3" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(LSUHC 11688) from Gunung Brinchang, Cameron Highlands, Pahang. Lower left: paratype of
<taxonomicName id="4C4C6888FFE83241FCC1FC33FC59FC45" box="[832,913,964,985]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="66" phylum="Chordata" rank="species" species="senja" status="sp. nov.">
<emphasis id="B938CF19FFE83241FCC1FC33FC59FC45" box="[832,913,964,985]" italics="true" pageId="3" pageNumber="66">S. senja</emphasis>
</taxonomicName>
<emphasis id="B938CF19FFE83241FC1BFC34FC25FC44" bold="true" box="[922,1005,963,984]" pageId="3" pageNumber="66">
<taxonomicNameLabel id="A20B7262FFE83241FC1BFC34FC25FC44" box="[922,1005,963,984]" pageId="3" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(BM 1911.12.5.10) from Gunung Gerah, Perak.
</paragraph>
</caption>
<subSubSection id="C3564080FFE83246FF46FBE9FAADF96E" lastPageId="4" lastPageNumber="67" pageId="3" pageNumber="66" type="description">
<paragraph id="8BF3130BFFE83241FF46FBE9FC7DF900" blockId="3.[151,1437,1054,2016]" pageId="3" pageNumber="66">
<emphasis id="B938CF19FFE83241FF46FBE9FE25FBAB" bold="true" box="[199,493,1054,1079]" pageId="3" pageNumber="66">
Description of
<typeStatus id="54F7ADA9FFE83241FEFEFBE8FE20FBAB" box="[383,488,1055,1079]" pageId="3" pageNumber="66" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male, SVL 60.0 mm; head moderate, snout rounded in dorsal and lateral profile, indistinct from neck; rostral wider than long, in broad contact with frontonasal; frontonasal wider than long, not divided anteriorly; prefrontals large, separate or in narrow contact; frontal elongate, diamond-shaped, in contact with first two supraoculars; four supraoculars; frontoparietals in contact posterior to frontal, contacting posterior portion of second, all of third, anterior one-half of fourth supraocular, parietals, and interparietal posteriorly; interparietal diamond-shaped, large, slightly projecting posteriorly; parietal eyespot absent; parietals large, in contact posterior to interparietal, contact posterior corner of fourth supraocular anteriorly; nuchal scales not enlarged; nasals large, widely separated, trapezoidal, contacting rostral anteriorly, frontonasal dorsally, first loreal posteriorly, first supralabial ventrally; nostril in center of nasal; supranasals absent; two loreals, posterior broader than anterior, anterior tallest; two preoculars in contact with posterior margin of second loreal, ventral preocular largest; eight supraciliaries, posteriormost elongate, projecting dorsomedially; two pretemporals; discontinuous row of five suboculars contacting supralabials; seven supralabials, fifth below center of eye and interrupting subocular row; two postsupralabials; one primary temporals; two secondary temporals, uppermost contacting parietals; small, granular scales at posterior corner of eye; lower eyelid transparent, scaly, lacking an enlarged central disc; mental twice as wide as long, rectangular; postmental large, single, contacting first and second infralabials; two pairs of enlarged sublabials following postmental, anterior pair contacting medially, posterior pair separated by a single scale, both pairs contacting infralabials; six infralabials, first smallest; external ear opening large, lacking anterior lobules; tympanum deeply recessed.
</paragraph>
<paragraph id="8BF3130BFFE83241FF46F950FC56F804" blockId="3.[151,1437,1054,2016]" pageId="3" pageNumber="66">Body scales smooth, cycloid, imbricate; ventral scales slightly larger than dorsal scales; 32 longitudinal, scale rows around midbody; 72 paravertebral scale rows; 68 ventral scale rows; two enlarged, medial, preanal scales; limbs small, separated when adpressed; scales of dorsal surface slightly larger than those of ventral surface; palmar and plantar scales rounded; supradigital scales on hand in multiple rows proximally, in a single, enlarged, row distally; supradigital scales on foot in multiple rows throughout digit; 17 smooth, subdigital lamellae on fourth toe; first digit of hand not vestigial; tail robust, not constricted at base, approximately 1.3 (not original) SVL, cylindrical; subcaudal scales slightly larger than dorsal caudal scales.</paragraph>
<paragraph id="8BF3130BFFE83246FF46F855FEC3FE14" blockId="3.[151,1437,1054,2016]" lastBlockId="4.[151,1437,151,789]" lastPageId="4" lastPageNumber="67" pageId="3" pageNumber="66">
<emphasis id="B938CF19FFE83241FF46F855FE3EF827" bold="true" box="[199,502,1954,1979]" pageId="3" pageNumber="66">
Coloration in life (
<figureCitation id="13770F8EFFE83241FE23F855FE20F827" box="[418,488,1954,1979]" captionStart="FIGURE 2" captionStartId="3.[151,250,932,954]" captionTargetBox="[150,1436,193,910]" captionTargetId="figure@3.[150,1436,193,911]" captionTargetPageId="3" captionText="FIGURE 2. Upper left and right and lower right: holotype of Sphenomorphus senja sp. nov. (LSUHC 11688) from Gunung Brinchang, Cameron Highlands, Pahang. Lower left: paratype of S. senja sp. nov. (BM 1911.12.5.10) from Gunung Gerah, Perak." httpUri="https://zenodo.org/record/288142/files/figure.png" pageId="3" pageNumber="66">Fig. 2</figureCitation>
).
</emphasis>
Ground color of dorsal surfaces of head, body, limbs, and tail brown, overlain with numerous darker flecks giving it a speckled appearance; a wide, diffuse, black, lateral stripe extends from posterior margin of eye to where it is most prominent on flanks thence becoming more diffuse and terminating on anterior portion of tail; lateral stripe bordered dorsally by a faint, lighter colored stripe extending from eye to axilla after which it transforms into a series of whitish flecks on trunk; lateral stripe is bordered ventrally by numerous, irregularly shaped, brown and white spots on neck; small whitish spots occur within the stripe; the lower portion of flanks below lateral stripe vermillion orange; gular region, throat and neck beige; pectoral region, abdomen, and undersides of limbs yellow; anterior two-thirds of subcaudal region orange, posterior one-third mottled in orange and black.
</paragraph>
<paragraph id="8BF3130BFFEF3246FF46FE64FB91FC88" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="67">
<emphasis id="B938CF19FFEF3246FF46FE64FEF3FE30" bold="true" box="[199,315,403,428]" pageId="4" pageNumber="67">Variation.</emphasis>
The
<typeStatus id="54F7ADA9FFEF3246FEF9FE62FE10FE31" box="[376,472,405,429]" pageId="4" pageNumber="67" type="paratype">paratype</typeStatus>
BM 1911.12.5.10 closely approaches the
<typeStatus id="54F7ADA9FFEF3246FC2FFE63FBD8FE31" box="[942,1040,404,429]" pageId="4" pageNumber="67" type="holotype">holotype</typeStatus>
in coloration except that the lateral stripe is bordered ventrally by numerous, irregularly shaped, white spots just within the lower region of the stripe, giving it a poorly defined and broken ventral margin and the lower flanks are yellow instead of vermillion (
<bibRefCitation id="EFDD6EFAFFEF3246FAEDFE2BFF1BFD84" author="Lim" pageId="4" pageNumber="67" refString="Lim, L. J. (1998) The taxonomy of West Malaysian and Singapore Scincidae (Reptilia: Sauria). Unpublished M. S. Thesis, National University of Singapore, Kent Ridge, Singapore, 234 pp." type="book" year="1998">Lim 1998</bibRefCitation>
). This may be because the
<typeStatus id="54F7ADA9FFEF3246FD84FE08FDAFFD84" box="[517,615,511,536]" pageId="4" pageNumber="67" type="holotype">holotype</typeStatus>
was a reproductively active male (enlarged testes examined) bearing the breeding coloration. The frontonasal scale of the
<typeStatus id="54F7ADA9FFEF3246FD6BFDD3FC84FDA1" box="[746,844,548,573]" pageId="4" pageNumber="67" type="holotype">holotype</typeStatus>
is not divided whereas it is in the
<typeStatus id="54F7ADA9FFEF3246FA85FDD2FAA0FDA1" box="[1284,1384,549,573]" pageId="4" pageNumber="67" type="paratype">paratype</typeStatus>
; the prefrontals are separate in the
<typeStatus id="54F7ADA9FFEF3246FE77FDB0FD90FDFC" box="[502,600,583,608]" pageId="4" pageNumber="67" type="holotype">holotype</typeStatus>
as opposed to being in narrow contact in the
<typeStatus id="54F7ADA9FFEF3246FBEFFDBFFB1AFDFC" box="[1134,1234,584,608]" pageId="4" pageNumber="67" type="paratype">paratype</typeStatus>
; the
<typeStatus id="54F7ADA9FFEF3246FA8AFDB0FAA5FDFC" box="[1291,1389,583,608]" pageId="4" pageNumber="67" type="holotype">holotype</typeStatus>
has eight as opposed to nine superciliaries; one as opposed to two primary temporals; and 17 as opposed to 13 subdigital lamellae. A discrepancy of four subdigital scales could be considered significant, however, neither
<bibRefCitation id="EFDD6EFAFFEF3246FF16FD43FE94FD50" author="Boulenger" box="[151,348,692,717]" pageId="4" pageNumber="67" refString="Boulenger, G. A. (1912) A Vertebrate Fauna of the Malay Peninsula from the Isthmus of Kra to Singapore Including the Adjacent Islands. Reptilia and Batrachia. Taylor &amp; Francis, London, 294 pp." type="book" year="1912">Boulenger (1912)</bibRefCitation>
or
<bibRefCitation id="EFDD6EFAFFEF3246FE00FD43FDC9FD50" author="Lim" box="[385,513,692,717]" pageId="4" pageNumber="67" refString="Lim, L. J. (1998) The taxonomy of West Malaysian and Singapore Scincidae (Reptilia: Sauria). Unpublished M. S. Thesis, National University of Singapore, Kent Ridge, Singapore, 234 pp." type="book" year="1998">Lim (1998)</bibRefCitation>
explained the method by which this count was taken. We can adjust our method of counting to bring our count down to 15 but elect not to so as to remain consistent with those counts of other Malaysian skinks (
<bibRefCitation id="EFDD6EFAFFEF3246FEE9FD0BFDC0FC88" author="Grismer" box="[360,520,764,789]" pageId="4" pageNumber="67" refString="Grismer, L. L. (2011) Lizards of Peninsular Malaysia, Singapore and Their Adjacent Archipelagos. Edition Chimaira, Frankfurt am Main, 728 pp." type="book" year="2011">Grismer 2011</bibRefCitation>
). Other meristic differences are presented in
<tableCitation id="C6CE26B0FFEF3246FC7EFD0BFB9AFC88" box="[1023,1106,764,789]" captionStart="TABLE 1" captionStartId="4.[151,239,830,853]" captionTargetBox="[159,1328,986,1504]" captionTargetPageId="4" captionText="TABLE 1. Diagnostic character states separating Sphenomorphus malayanus from S. senja. sp. nov. Character states of S. malayanus are from Doria (1888) and of the paratype (BM 1911.12.5.10) from Boulenger (1912) and / or Lim (1998). / — not reported by describing authors." httpUri="http://table.plazi.org/id/DF334383FFEF3246FF16FCC9FDF3FC09" pageId="4" pageNumber="67" tableUuid="DF334383FFEF3246FF16FCC9FDF3FC09">Table 1</tableCitation>
.
</paragraph>
<caption id="DF334383FFEF3246FF16FCC9FDF3FC09" ID-Table-UUID="DF334383FFEF3246FF16FCC9FDF3FC09" httpUri="http://table.plazi.org/id/DF334383FFEF3246FF16FCC9FDF3FC09" pageId="4" pageNumber="67" targetBox="[159,1328,986,1504]" targetIsTable="true" targetPageId="4">
<paragraph id="8BF3130BFFEF3246FF16FCC9FDF3FC09" blockId="4.[151,1436,829,917]" pageId="4" pageNumber="67">
<emphasis id="B938CF19FFEF3246FF16FCC9FEC2FCC9" bold="true" box="[151,266,830,853]" pageId="4" pageNumber="67">TABLE 1.</emphasis>
Diagnostic character states separating
<taxonomicName id="4C4C6888FFEF3246FD2CFCCAFC1BFCC9" box="[685,979,829,853]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="malayanus">
<emphasis id="B938CF19FFEF3246FD2CFCCAFC1BFCC9" box="[685,979,829,853]" italics="true" pageId="4" pageNumber="67">Sphenomorphus malayanus</emphasis>
</taxonomicName>
from
<emphasis id="B938CF19FFEF3246FB97FCC9FBB9FCC9" box="[1046,1137,830,853]" italics="true" pageId="4" pageNumber="67">
<taxonomicName id="4C4C6888FFEF3246FB97FCC9FBA5FCC9" box="[1046,1133,830,853]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="senja">S. senja</taxonomicName>
.
</emphasis>
<emphasis id="B938CF19FFEF3246FBF6FCC9FB06FCC9" bold="true" box="[1143,1230,830,853]" pageId="4" pageNumber="67">sp. nov.</emphasis>
Character states of
<taxonomicName id="4C4C6888FFEF3246FF16FCA9FEEDFCE9" box="[151,293,861,885]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="malayanus">
<emphasis id="B938CF19FFEF3246FF16FCA9FEEDFCE9" box="[151,293,861,885]" italics="true" pageId="4" pageNumber="67">S. malayanus</emphasis>
</taxonomicName>
are from Doria (1888) and of the paratype (BM 1911.12.5.10) from Boulenger (1912) and/or Lim (1998). / — not reported by describing authors.
</paragraph>
</caption>
<paragraph id="8BF3130BFFEF3246FDB2FC46FDFAF91A" blockId="4.[563,902,945,967]" lastBlockId="4.[151,1437,1572,2030]" pageId="4" pageNumber="67">
<emphasis id="B938CF19FFEF3246FDB2FC46FC4EFC5B" box="[563,902,945,967]" italics="true" pageId="4" pageNumber="67">
<taxonomicName id="4C4C6888FFEF3246FDB2FC46FD69FC5B" box="[563,673,945,967]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="malayanus">malayanus</taxonomicName>
<taxonomicName id="4C4C6888FFEF3246FCD0FC45FC4EFC5B" box="[849,902,946,967]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="senja">senja</taxonomicName>
</emphasis>
<emphasis id="B938CF19FFEF3246FF46F9D3FEA8F9A1" bold="true" box="[199,352,1572,1597]" pageId="4" pageNumber="67">Distribution.</emphasis>
<taxonomicName id="4C4C6888FFEF3246FEF1F9D2FDA7F9A1" box="[368,623,1573,1597]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="senja" status="sp. nov.">
<emphasis id="B938CF19FFEF3246FEF1F9D2FDA7F9A1" box="[368,623,1573,1597]" italics="true" pageId="4" pageNumber="67">Sphenomorphus senja</emphasis>
</taxonomicName>
<emphasis id="B938CF19FFEF3246FDFFF9D2FD28F9A1" bold="true" box="[638,736,1573,1597]" pageId="4" pageNumber="67">
<taxonomicNameLabel id="A20B7262FFEF3246FDFFF9D2FD28F9A1" box="[638,736,1573,1597]" pageId="4" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known only from Gunung Gerah, Perak and Gunung Brinchang, Pahang in the Titiwangsa Mountain Range (
<figureCitation id="13770F8EFFEF3246FC92F9BEFC9EF9FE" box="[787,854,1609,1634]" captionStart="FIGURE 1" captionStartId="2.[151,250,1879,1901]" captionTargetBox="[168,1436,721,1835]" captionTargetId="figure@2.[150,1436,721,1835]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of Sphenomorphus senja sp. nov. and S. malayanus. Stars denote the type localities of Gunug Singgalang, West Sumatra, Indonesia (yellow) and Gunung Brinchang, Cameron Highlands, Pahang, Malaysia (orange)." httpUri="https://zenodo.org/record/288141/files/figure.png" pageId="4" pageNumber="67">Fig. 1</figureCitation>
), however, it is likely to range further to the north and south along this upland corridor.
</paragraph>
<paragraph id="8BF3130BFFEF3246FDB2FC2CFB3DFA7C" pageId="4" pageNumber="67">
<table id="F94CE1ABFFEFCDBDFF1EFC2DFAF8FA7C" box="[159,1328,986,1504]" gridcols="4" gridrows="13" pageId="4" pageNumber="67">
<tr id="357C1149FFEFCDBDFF1EFC2DFAF8FC6C" box="[159,1328,986,1008]" gridrow="0" pageId="4" pageNumber="67" rowspan-0="1">
<th id="76AD7835FFEFCDBDFDB3FC2DFD1FFC6C" box="[562,727,986,1008]" gridcol="1" gridrow="0" pageId="4" pageNumber="67">BM 1946.8.3.11</th>
<th id="76AD7835FFEFCDBDFCD0FC2DFBEBFC6C" box="[849,1059,986,1008]" gridcol="2" gridrow="0" pageId="4" pageNumber="67">BM 1911.12.5.10</th>
<th id="76AD7835FFEFCDBDFB0AFC2DFAF8FC6C" box="[1163,1328,986,1008]" gridcol="3" gridrow="0" pageId="4" pageNumber="67">LSUHC 11688</th>
</tr>
<tr id="357C1149FFEFCDBDFF1EFBF3FAF8FB86" box="[159,1328,1028,1050]" gridrow="1" pageId="4" pageNumber="67">
<th id="76AD7835FFEFCDBDFF1EFBF3FE2BFB86" box="[159,483,1028,1050]" gridcol="0" gridrow="1" pageId="4" pageNumber="67">SVL (mm)</th>
<td id="76AD7835FFEFCDBDFDB3FBF3FD1FFB86" box="[562,727,1028,1050]" gridcol="1" gridrow="1" pageId="4" pageNumber="67">53</td>
<td id="76AD7835FFEFCDBDFCD0FBF3FBEBFB86" box="[849,1059,1028,1050]" gridcol="2" gridrow="1" pageId="4" pageNumber="67">65 (Boulenger 1912)</td>
<td id="76AD7835FFEFCDBDFB0AFBF3FAF8FB86" box="[1163,1328,1028,1050]" gridcol="3" gridrow="1" pageId="4" pageNumber="67">60.0</td>
</tr>
<tr id="357C1149FFEFCDBDFF1EFBDAFAF8FBDF" box="[159,1328,1069,1091]" gridrow="2" pageId="4" pageNumber="67">
<th id="76AD7835FFEFCDBDFF1EFBDAFE2BFBDF" box="[159,483,1069,1091]" gridcol="0" gridrow="2" pageId="4" pageNumber="67">Tympanum</th>
<td id="76AD7835FFEFCDBDFDB3FBDAFD1FFBDF" box="[562,727,1069,1091]" gridcol="1" gridrow="2" pageId="4" pageNumber="67">Shallow</td>
<td id="76AD7835FFEFCDBDFCD0FBDAFBEBFBDF" box="[849,1059,1069,1091]" gridcol="2" gridrow="2" pageId="4" pageNumber="67">Deeply recessed</td>
<td id="76AD7835FFEFCDBDFB0AFBDAFAF8FBDF" box="[1163,1328,1069,1091]" gridcol="3" gridrow="2" pageId="4" pageNumber="67">Deeply recessed</td>
</tr>
<tr id="357C1149FFEFCDBDFF1EFBA1FAF8FBF0" box="[159,1328,1110,1132]" gridrow="3" pageId="4" pageNumber="67">
<th id="76AD7835FFEFCDBDFF1EFBA1FE2BFBF0" box="[159,483,1110,1132]" gridcol="0" gridrow="3" pageId="4" pageNumber="67">Midbody scales</th>
<td id="76AD7835FFEFCDBDFDB3FBA1FD1FFBF0" box="[562,727,1110,1132]" gridcol="1" gridrow="3" pageId="4" pageNumber="67">32</td>
<td id="76AD7835FFEFCDBDFCD0FBA1FBEBFBF0" box="[849,1059,1110,1132]" gridcol="2" gridrow="3" pageId="4" pageNumber="67">32</td>
<td id="76AD7835FFEFCDBDFB0AFBA1FAF8FBF0" box="[1163,1328,1110,1132]" gridcol="3" gridrow="3" pageId="4" pageNumber="67">32</td>
</tr>
<tr id="357C1149FFEFCDBDFF1EFB77FAF8FB0A" box="[159,1328,1152,1174]" gridrow="4" pageId="4" pageNumber="67">
<th id="76AD7835FFEFCDBDFF1EFB77FE2BFB0A" box="[159,483,1152,1174]" gridcol="0" gridrow="4" pageId="4" pageNumber="67">Paravertebral scales</th>
<td id="76AD7835FFEFCDBDFDB3FB77FD1FFB0A" box="[562,727,1152,1174]" gridcol="1" gridrow="4" pageId="4" pageNumber="67">76</td>
<td id="76AD7835FFEFCDBDFCD0FB77FBEBFB0A" box="[849,1059,1152,1174]" gridcol="2" gridrow="4" pageId="4" pageNumber="67">73</td>
<td id="76AD7835FFEFCDBDFB0AFB77FAF8FB0A" box="[1163,1328,1152,1174]" gridcol="3" gridrow="4" pageId="4" pageNumber="67">72</td>
</tr>
<tr id="357C1149FFEFCDBDFF1EFB5EFAF8FB23" box="[159,1328,1193,1215]" gridrow="5" pageId="4" pageNumber="67">
<th id="76AD7835FFEFCDBDFF1EFB5EFE2BFB23" box="[159,483,1193,1215]" gridcol="0" gridrow="5" pageId="4" pageNumber="67">Ventral scales</th>
<td id="76AD7835FFEFCDBDFDB3FB5EFD1FFB23" box="[562,727,1193,1215]" gridcol="1" gridrow="5" pageId="4" pageNumber="67">/</td>
<td id="76AD7835FFEFCDBDFCD0FB5EFBEBFB23" box="[849,1059,1193,1215]" gridcol="2" gridrow="5" pageId="4" pageNumber="67">68</td>
<td id="76AD7835FFEFCDBDFB0AFB5EFAF8FB23" box="[1163,1328,1193,1215]" gridcol="3" gridrow="5" pageId="4" pageNumber="67">68</td>
</tr>
<tr id="357C1149FFEFCDBDFF1EFB25FAF8FB74" box="[159,1328,1234,1256]" gridrow="6" pageId="4" pageNumber="67">
<th id="76AD7835FFEFCDBDFF1EFB25FE2BFB74" box="[159,483,1234,1256]" gridcol="0" gridrow="6" pageId="4" pageNumber="67">Superciliary scales</th>
<td id="76AD7835FFEFCDBDFDB3FB25FD1FFB74" box="[562,727,1234,1256]" gridcol="1" gridrow="6" pageId="4" pageNumber="67">10</td>
<td id="76AD7835FFEFCDBDFCD0FB25FBEBFB74" box="[849,1059,1234,1256]" gridcol="2" gridrow="6" pageId="4" pageNumber="67">9</td>
<td id="76AD7835FFEFCDBDFB0AFB25FAF8FB74" box="[1163,1328,1234,1256]" gridcol="3" gridrow="6" pageId="4" pageNumber="67">8</td>
</tr>
<tr id="357C1149FFEFCDBDFF1EFB0BFAF8FA8E" box="[159,1328,1276,1298]" gridrow="7" pageId="4" pageNumber="67">
<th id="76AD7835FFEFCDBDFF1EFB0BFE2BFA8E" box="[159,483,1276,1298]" gridcol="0" gridrow="7" pageId="4" pageNumber="67">Posteriormost superciliary scale</th>
<td id="76AD7835FFEFCDBDFDB3FB0BFD1FFA8E" box="[562,727,1276,1298]" gridcol="1" gridrow="7" pageId="4" pageNumber="67">small</td>
<td id="76AD7835FFEFCDBDFCD0FB0BFBEBFA8E" box="[849,1059,1276,1298]" gridcol="2" gridrow="7" pageId="4" pageNumber="67">large</td>
<td id="76AD7835FFEFCDBDFB0AFB0BFAF8FA8E" box="[1163,1328,1276,1298]" gridcol="3" gridrow="7" pageId="4" pageNumber="67">large</td>
</tr>
<tr id="357C1149FFEFCDBDFF1EFAD2FAF8FAA7" box="[159,1328,1317,1339]" gridrow="8" pageId="4" pageNumber="67">
<th id="76AD7835FFEFCDBDFF1EFAD2FE2BFAA7" box="[159,483,1317,1339]" gridcol="0" gridrow="8" pageId="4" pageNumber="67">Toe IV lamellae</th>
<td id="76AD7835FFEFCDBDFDB3FAD2FD1FFAA7" box="[562,727,1317,1339]" gridcol="1" gridrow="8" pageId="4" pageNumber="67">15</td>
<td id="76AD7835FFEFCDBDFCD0FAD2FBEBFAA7" box="[849,1059,1317,1339]" gridcol="2" gridrow="8" pageId="4" pageNumber="67">13</td>
<td id="76AD7835FFEFCDBDFB0AFAD2FAF8FAA7" box="[1163,1328,1317,1339]" gridcol="3" gridrow="8" pageId="4" pageNumber="67">17</td>
</tr>
<tr id="357C1149FFEFCDBDFF1EFAB9FAF8FAF8" box="[159,1328,1358,1380]" gridrow="9" pageId="4" pageNumber="67">
<th id="76AD7835FFEFCDBDFF1EFAB9FE2BFAF8" box="[159,483,1358,1380]" gridcol="0" gridrow="9" pageId="4" pageNumber="67">Lower flanks</th>
<td id="76AD7835FFEFCDBDFDB3FAB9FD1FFAF8" box="[562,727,1358,1380]" gridcol="1" gridrow="9" pageId="4" pageNumber="67">/</td>
<td id="76AD7835FFEFCDBDFCD0FAB9FBEBFAF8" box="[849,1059,1358,1380]" gridcol="2" gridrow="9" pageId="4" pageNumber="67">yellowish</td>
<td id="76AD7835FFEFCDBDFB0AFAB9FAF8FAF8" box="[1163,1328,1358,1380]" gridcol="3" gridrow="9" pageId="4" pageNumber="67">vermillion</td>
</tr>
<tr id="357C1149FFEFCDBDFF1EFA8FFAF8FA12" box="[159,1328,1400,1422]" gridrow="10" pageId="4" pageNumber="67">
<th id="76AD7835FFEFCDBDFF1EFA8FFE2BFA12" box="[159,483,1400,1422]" gridcol="0" gridrow="10" pageId="4" pageNumber="67">Abdomen</th>
<td id="76AD7835FFEFCDBDFDB3FA8FFD1FFA12" box="[562,727,1400,1422]" gridcol="1" gridrow="10" pageId="4" pageNumber="67">yellowish gray</td>
<td id="76AD7835FFEFCDBDFCD0FA8FFBEBFA12" box="[849,1059,1400,1422]" gridcol="2" gridrow="10" pageId="4" pageNumber="67">pale yellow</td>
<td id="76AD7835FFEFCDBDFB0AFA8FFAF8FA12" box="[1163,1328,1400,1422]" gridcol="3" gridrow="10" pageId="4" pageNumber="67">yellow</td>
</tr>
<tr id="357C1149FFEFCDBDFF1EFA56FAF8FA2B" box="[159,1328,1441,1463]" gridrow="11" pageId="4" pageNumber="67">
<th id="76AD7835FFEFCDBDFF1EFA56FE2BFA2B" box="[159,483,1441,1463]" gridcol="0" gridrow="11" pageId="4" pageNumber="67">Pectoral region</th>
<td id="76AD7835FFEFCDBDFDB3FA56FD1FFA2B" box="[562,727,1441,1463]" gridcol="1" gridrow="11" pageId="4" pageNumber="67">yellowish gray</td>
<td id="76AD7835FFEFCDBDFCD0FA56FBEBFA2B" box="[849,1059,1441,1463]" gridcol="2" gridrow="11" pageId="4" pageNumber="67">pale yellow</td>
<td id="76AD7835FFEFCDBDFB0AFA56FAF8FA2B" box="[1163,1328,1441,1463]" gridcol="3" gridrow="11" pageId="4" pageNumber="67">yellow</td>
</tr>
<tr id="357C1149FFEFCDBDFF1EFA3DFAF8FA7C" box="[159,1328,1482,1504]" gridrow="12" pageId="4" pageNumber="67">
<th id="76AD7835FFEFCDBDFF1EFA3DFE2BFA7C" box="[159,483,1482,1504]" gridcol="0" gridrow="12" pageId="4" pageNumber="67">Anterior subcaudal region</th>
<td id="76AD7835FFEFCDBDFDB3FA3DFD1FFA7C" box="[562,727,1482,1504]" gridcol="1" gridrow="12" pageId="4" pageNumber="67">yellowish gray</td>
<td id="76AD7835FFEFCDBDFCD0FA3DFBEBFA7C" box="[849,1059,1482,1504]" gridcol="2" gridrow="12" pageId="4" pageNumber="67">/</td>
<td id="76AD7835FFEFCDBDFB0AFA3DFAF8FA7C" box="[1163,1328,1482,1504]" gridcol="3" gridrow="12" pageId="4" pageNumber="67">vermillion</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF3130BFFEF3246FF46F966FAADF96E" blockId="4.[151,1437,1572,2030]" pageId="4" pageNumber="67">
<emphasis id="B938CF19FFEF3246FF46F966FE4CF936" bold="true" box="[199,388,1681,1706]" pageId="4" pageNumber="67">Natural history.</emphasis>
<taxonomicName id="4C4C6888FFEF3246FE0AF966FD49F935" box="[395,641,1681,1705]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="senja" status="sp. nov.">
<emphasis id="B938CF19FFEF3246FE0AF966FD49F935" box="[395,641,1681,1705]" italics="true" pageId="4" pageNumber="67">Sphenomorphus senja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B7262FFEF3246FD09F965FD10F935" box="[648,728,1682,1705]" pageId="4" pageNumber="67" rank="species">sp. nov</taxonomicNameLabel>
is a upland mossy, cloud forest species (
<figureCitation id="13770F8EFFEF3246FB1CF966FB16F936" box="[1181,1246,1681,1706]" captionStart="FIGURE 3" captionStartId="5.[151,250,1430,1452]" captionTargetBox="[152,1434,555,1408]" captionTargetId="figure@5.[152,1435,555,1408]" captionTargetPageId="5" captionText="FIGURE 3. Mossy cloud forest habitat of Sphenomorphus senja sp. nov. at Gunung Brinchang, Cameron Highlands, Pahang." httpUri="https://zenodo.org/record/288143/files/figure.png" pageId="4" pageNumber="67">Fig. 3</figureCitation>
) ranging from at least
<quantity id="4CB4BEEEFFEF3246FF52F941FEE1F951" box="[211,297,1718,1742]" metricMagnitude="3" metricUnit="m" metricValue="1.811" pageId="4" pageNumber="67" unit="m" value="1811.0">1811 m</quantity>
in elevation at Gunung Brinchang to
<quantity id="4CB4BEEEFFEF3246FD5CF941FCFCF951" box="[733,820,1718,1742]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="67" unit="m" value="2200.0">2200 m</quantity>
in elevation at Gunung Gerah (
<bibRefCitation id="EFDD6EFAFFEF3246FB21F942FA93F952" author="Boulenger" box="[1184,1371,1717,1742]" pageId="4" pageNumber="67" refString="Boulenger, G. A. (1912) A Vertebrate Fauna of the Malay Peninsula from the Isthmus of Kra to Singapore Including the Adjacent Islands. Reptilia and Batrachia. Taylor &amp; Francis, London, 294 pp." type="book" year="1912">Boulenger 1912</bibRefCitation>
). The
<typeStatus id="54F7ADA9FFEF3246FF16F92EFF31F96E" box="[151,249,1753,1778]" pageId="4" pageNumber="67" type="holotype">holotype</typeStatus>
was collected at night beneath leaf-litter along an earthen bank following several days of heavy rain.
</paragraph>
</subSubSection>
<subSubSection id="C3564080FFEF3247FF46F90AFD3DFE69" lastPageId="5" lastPageNumber="68" pageId="4" pageNumber="67" type="etymology">
<paragraph id="8BF3130BFFEF3246FF46F90AFAA6F8A5" blockId="4.[151,1437,1572,2030]" pageId="4" pageNumber="67">
<emphasis id="B938CF19FFEF3246FF46F90AFE86F889" bold="true" box="[199,334,1789,1813]" pageId="4" pageNumber="67">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C4C6888FFEF3246FDB8F909FDBCF889" box="[569,628,1790,1813]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="senja">
<emphasis id="B938CF19FFEF3246FDB8F909FDBCF889" box="[569,628,1790,1813]" italics="true" pageId="4" pageNumber="67">senja</emphasis>
</taxonomicName>
is the Malay word for “twilight” or “dusk” at which time the sky above Gunung Brinchang turns vermillion orange, matching the color on the flanks of
<taxonomicName id="4C4C6888FFEF3246FB96F8D6FAC5F8A5" box="[1047,1293,1825,1849]" class="Reptilia" family="Scincidae" genus="Sphenomoprhus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="senja" status="sp. nov.">
<emphasis id="B938CF19FFEF3246FB96F8D6FAC5F8A5" box="[1047,1293,1825,1849]" italics="true" pageId="4" pageNumber="67">Sphenomoprhus senja</emphasis>
</taxonomicName>
<emphasis id="B938CF19FFEF3246FA95F8D6FAA6F8A5" bold="true" box="[1300,1390,1825,1849]" pageId="4" pageNumber="67">
<taxonomicNameLabel id="A20B7262FFEF3246FA95F8D6FAA6F8A5" box="[1300,1390,1825,1849]" pageId="4" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BF3130BFFEF3247FF46F8B3FD3DFE69" blockId="4.[151,1437,1572,2030]" lastBlockId="5.[151,1436,151,501]" lastPageId="5" lastPageNumber="68" pageId="4" pageNumber="67">
<emphasis id="B938CF19FFEF3246FF46F8B3FEA5F8C1" bold="true" box="[199,365,1860,1885]" pageId="4" pageNumber="67">Comparisons.</emphasis>
<taxonomicName id="4C4C6888FFEF3246FEFBF8B2FDBFF8C1" box="[378,631,1861,1885]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="senja" status="sp. nov.">
<emphasis id="B938CF19FFEF3246FEFBF8B2FDBFF8C1" box="[378,631,1861,1885]" italics="true" pageId="4" pageNumber="67">Sphenomorphus senja</emphasis>
</taxonomicName>
<emphasis id="B938CF19FFEF3246FD05F8B2FD2CF8C1" bold="true" box="[644,740,1861,1885]" pageId="4" pageNumber="67">
<taxonomicNameLabel id="A20B7262FFEF3246FD05F8B2FD2CF8C1" box="[644,740,1861,1885]" pageId="4" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="4C4C6888FFEF3246FC55F8B2FAC7F8C1" box="[980,1295,1861,1885]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="malayanus">
<emphasis id="B938CF19FFEF3246FC55F8B2FAC7F8C1" box="[980,1295,1861,1885]" italics="true" pageId="4" pageNumber="67">Sphenomorphus malayanus</emphasis>
</taxonomicName>
of Gunung Singgalang in West Sumatra but can be separated from it by having a larger SVL (60.0–
<quantity id="4CB4BEEEFFEF3246FB18F89EFB21F81D" box="[1177,1257,1897,1922]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="4" pageNumber="67" unit="mm" value="65.0">65 mm</quantity>
versus
<quantity id="4CB4BEEEFFEF3246FAC5F89EFA5FF81D" box="[1348,1431,1897,1922]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="4" pageNumber="67" unit="mm" value="53.0">53 mm</quantity>
, assuming the
<taxonomicName id="4C4C6888FFEF3246FEB2F879FE01F839" box="[307,457,1933,1957]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="malayanus">
<emphasis id="B938CF19FFEF3246FEB2F879FE01F839" box="[307,457,1933,1957]" italics="true" pageId="4" pageNumber="67">S. malayanus</emphasis>
</taxonomicName>
specimen is an adult); a deep, as opposed to a shallow tympanum; 72 or 73 versus 76 paravertebral scales; eight or nine superciliary scales as opposed to 10; and the posteriromost superciliary scale being large as opposed to small. The anterior subcaudal region is orange in
<taxonomicName id="4C4C6888FFEF3246FC62F821FBF2F871" box="[995,1082,2006,2029]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="senja" status="sp. nov.">
<emphasis id="B938CF19FFEF3246FC62F821FBF2F871" box="[995,1082,2006,2029]" italics="true" pageId="4" pageNumber="67">S. senja</emphasis>
</taxonomicName>
<emphasis id="B938CF19FFEF3246FBC0F822FB52F871" bold="true" box="[1089,1178,2005,2029]" pageId="4" pageNumber="67">
<taxonomicNameLabel id="A20B7262FFEF3246FBC0F822FB52F871" box="[1089,1178,2005,2029]" pageId="4" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and in
<taxonomicName id="4C4C6888FFEF3246FB6CF821FA4AF871" box="[1261,1410,2005,2029]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="67" phylum="Chordata" rank="species" species="malayanus">
<emphasis id="B938CF19FFEF3246FB6CF821FA4AF871" box="[1261,1410,2005,2029]" italics="true" pageId="4" pageNumber="67">S. malayanus</emphasis>
</taxonomicName>
is reported to be yellowish gray (“giallo suducio”,
<bibRefCitation id="EFDD6EFAFFEE3247FD46FF60FC94FF2C" author="Doria" box="[711,860,151,176]" pageId="5" pageNumber="68" refString="Doria, G. (1888) Note Erpetologiche Alcuni nuovi sauri raccolti in Sumatra dal Dr. O. Beccari. Annali del Museo Civico di Storia Natruale di Genova, 2, 646 - 652." type="journal article" year="1888">Doria [1888]</bibRefCitation>
) or “whitish” (de
<bibRefCitation id="EFDD6EFAFFEE3247FBB3FF60FB75FF2C" author="Rooij" box="[1074,1213,151,176]" pageId="5" pageNumber="68" refString="De Rooij, N. (1915) The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. E. J. Brill Ltd., Leiden, 384 pp." type="book" year="1915">Rooij, 1915</bibRefCitation>
). Unfortunately the
<typeStatus id="54F7ADA9FFEE3247FF16FF4BFF31FF49" box="[151,249,188,213]" pageId="5" pageNumber="68" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C4C6888FFEE3247FEA1FF4BFE7EFF48" box="[288,438,188,212]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="68" phylum="Chordata" rank="species" species="malayanus">
<emphasis id="B938CF19FFEE3247FEA1FF4BFE7EFF48" box="[288,438,188,212]" italics="true" pageId="5" pageNumber="68">S. malayanus</emphasis>
</taxonomicName>
could not be located for direct examination (David Gower
<emphasis id="B938CF19FFEE3247FBD8FF4AFB55FF48" box="[1113,1181,188,212]" italics="true" pageId="5" pageNumber="68">in litt.</emphasis>
2014). The only other species in Peninsular
<collectingCountry id="F35B539BFFEE3247FE13FF28FE30FF64" box="[402,504,223,248]" name="Malaysia" pageId="5" pageNumber="68">Malaysia</collectingCountry>
with which
<taxonomicName id="4C4C6888FFEE3247FD0CFF17FD2FFF6B" box="[653,743,224,247]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="68" phylum="Chordata" rank="species" species="senja" status="sp. nov.">
<emphasis id="B938CF19FFEE3247FD0CFF17FD2FFF6B" box="[653,743,224,247]" italics="true" pageId="5" pageNumber="68">S. senja</emphasis>
</taxonomicName>
<emphasis id="B938CF19FFEE3247FD70FF17FC86FF64" bold="true" box="[753,846,224,248]" pageId="5" pageNumber="68">
<taxonomicNameLabel id="A20B7262FFEE3247FD70FF17FC86FF64" box="[753,846,224,248]" pageId="5" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
might be confused with is
<taxonomicName id="4C4C6888FFEE3247FB0EFF17FA54FF64" authority="Smith" authorityName="Smith" box="[1167,1436,223,248]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="68" phylum="Chordata" rank="species" species="cameronicus">
<emphasis id="B938CF19FFEE3247FB0EFF17FAF4FF6B" box="[1167,1340,224,247]" italics="true" pageId="5" pageNumber="68">S. cameronicus</emphasis>
(Smith)
</taxonomicName>
which occurs at lower elevations (
<quantity id="4CB4BEEEFFEE3247FD9BFEF3FDBAFE80" box="[538,626,260,285]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="68" unit="m" value="1500.0">1500 m</quantity>
) in
<collectingCountry id="F35B539BFFEE3247FD21FEF3FCCEFE80" box="[672,774,260,284]" name="Cameroon" pageId="5" pageNumber="68">Cameron</collectingCountry>
Highlands (
<bibRefCitation id="EFDD6EFAFFEE3247FC10FEF3FBE7FE80" author="Grismer" box="[913,1071,260,285]" pageId="5" pageNumber="68" refString="Grismer, L. L. (2011) Lizards of Peninsular Malaysia, Singapore and Their Adjacent Archipelagos. Edition Chimaira, Frankfurt am Main, 728 pp." type="book" year="2011">Grismer 2011</bibRefCitation>
). We examined three specimens of
<taxonomicName id="4C4C6888FFEE3247FF34FEDFFE97FEA3" box="[181,351,296,319]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="68" phylum="Chordata" rank="species" species="cameronicus">
<emphasis id="B938CF19FFEE3247FF34FEDFFE97FEA3" box="[181,351,296,319]" italics="true" pageId="5" pageNumber="68">S. cameronicus</emphasis>
</taxonomicName>
(LSUHC 9738, 9819–20) from its
<typeStatus id="54F7ADA9FFEE3247FD6AFEDFFCD3FEDC" box="[747,795,296,320]" pageId="5" pageNumber="68">type</typeStatus>
locality at Tanah Rata,
<collectingCountry id="F35B539BFFEE3247FBA7FEDFFB45FEDC" box="[1062,1165,296,320]" name="Cameroon" pageId="5" pageNumber="68">Cameron</collectingCountry>
Highlands and find that
<taxonomicName id="4C4C6888FFEE3247FF16FEBBFF39FEF8" box="[151,241,332,356]" class="Reptilia" family="Scincidae" genus="Sphenomorphus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="68" phylum="Chordata" rank="species" species="senja" status="sp. nov.">
<emphasis id="B938CF19FFEE3247FF16FEBBFF39FEF8" box="[151,241,332,356]" italics="true" pageId="5" pageNumber="68">S. senja</emphasis>
</taxonomicName>
<emphasis id="B938CF19FFEE3247FF7DFEBBFE90FEF8" bold="true" box="[252,344,332,356]" pageId="5" pageNumber="68">
<taxonomicNameLabel id="A20B7262FFEE3247FF7DFEBBFE90FEF8" box="[252,344,332,356]" pageId="5" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from it in that the adpressed limbs do not overlap as opposed to overlapping slightly; having 32 midbody scale rows as opposed to 34–38; having 13–17 as opposed to 20 or 21 subdigital lamellae on the fourth toe; having vermillion colored flanks in breeding males as opposed to beige flanks in breeding males (
<bibRefCitation id="EFDD6EFAFFEE3247FF1FFE40FEF6FE4C" author="Grismer" box="[158,318,439,464]" pageId="5" pageNumber="68" refString="Grismer, L. L. (2011) Lizards of Peninsular Malaysia, Singapore and Their Adjacent Archipelagos. Edition Chimaira, Frankfurt am Main, 728 pp." type="book" year="2011">Grismer 2011</bibRefCitation>
); having yellow as opposed to beige pectoral and abdomen regions; and that the anterior two-thirds of the subcaudal region is orange as opposed to beige.
</paragraph>
</subSubSection>
</treatment>
</document>
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