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<document id="5235879B1F9264BF317D53A0758E0DBF" ID-CLB-Dataset="26462" ID-DOI="10.11646/zootaxa.4619.1.1" ID-GBIF-Dataset="9aaf6711-748d-4067-a759-26d37802b7fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="3248363" ID-ZooBank="CA6AEB18-2F97-449C-AE34-E1509DFFC841" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1560832679272" checkinUser="plazi" docAuthor="Lee, Bee Yan, Forges, Bertrand Richer De &amp; Ng, Peter K. L." docDate="2019" docId="03C9AC691E78FFB2649FFD53B7B5EA58" docLanguage="en" docName="zootaxa.4619.1.1.pdf" docOrigin="Zootaxa 4619 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Crocydocinus Lee &amp; Forges &amp; Ng 2019, n. gen." docType="treatment" docVersion="17" lastPageNumber="25" masterDocId="FFF0D4111E6EFFAA6408FFAEB427EC58" masterDocTitle="Deep-sea spider crabs of the family Epialtidae MacLeay, 1838, from Papua New Guinea, with a redefinition of Tunepugettia Ng, Komai &amp; Sato, 2017, and descriptions of two new genera (Crustacea: Decapoda: Brachyura: Majoidea)" masterLastPageNumber="44" masterPageNumber="1" pageNumber="23" updateTime="1698735225898" updateUser="ExternalLinkService">
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<mods:title id="2ED82611D2820A9A71D4D33824A88F79">Deep-sea spider crabs of the family Epialtidae MacLeay, 1838, from Papua New Guinea, with a redefinition of Tunepugettia Ng, Komai &amp; Sato, 2017, and descriptions of two new genera (Crustacea: Decapoda: Brachyura: Majoidea)</mods:title>
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<mods:namePart id="BE56669226B20F02A10D3A368D7DB614">Lee, Bee Yan</mods:namePart>
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<mods:namePart id="CBED145727E942EC53039B879CA2AB35">Forges, Bertrand Richer De</mods:namePart>
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<mods:namePart id="8EBCCA1CFECF18C91D1D1E02E6B39B46">Ng, Peter K. L.</mods:namePart>
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<treatment id="03C9AC691E78FFB2649FFD53B7B5EA58" ID-DOI="http://doi.org/10.5281/zenodo.5945520" ID-GBIF-Taxon="158290175" ID-Zenodo-Dep="5945520" LSID="urn:lsid:plazi:treatment:03C9AC691E78FFB2649FFD53B7B5EA58" httpUri="http://treatment.plazi.org/id/03C9AC691E78FFB2649FFD53B7B5EA58" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
<subSubSection id="C37A4EF41E78FFBC649FFD53B5C9EF4F" box="[151,494,765,792]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8BDF1D7F1E78FFBC649FFD53B5C9EF4F" blockId="22.[151,494,765,792]" box="[151,494,765,792]" pageId="22" pageNumber="23">
<heading id="D097AA131E78FFBC649FFD53B5C9EF4F" bold="true" box="[151,494,765,792]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="B914C16D1E78FFBC649FFD53B5C9EF4F" bold="true" box="[151,494,765,792]" pageId="22" pageNumber="23">
Genus
<taxonomicName id="4C6066FC1E78FFBC64E6FD53B5B3EF40" ID-CoL="847QD" authority="Lee &amp; Forges &amp; Ng, 2019" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" box="[238,404,765,792]" class="Malacostraca" family="Epialtidae" genus="Crocydocinus" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B914C16D1E78FFBC64E6FD53B5B3EF40" bold="true" box="[238,404,765,792]" italics="true" pageId="22" pageNumber="23">Crocydocinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2277C161E78FFBC6594FD53B5C9EF4F" box="[412,494,765,791]" pageId="22" pageNumber="23" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37A4EF41E78FFBC649FFCEBB018EF07" box="[151,1087,836,863]" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8BDF1D7F1E78FFBC649FFCEBB018EF07" blockId="22.[151,1437,836,2015]" box="[151,1087,836,863]" pageId="22" pageNumber="23">
<emphasis id="B914C16D1E78FFBC649FFCEBB515EF07" bold="true" box="[151,306,836,863]" pageId="22" pageNumber="23">
<typeStatus id="54DBA3DD1E78FFBC649FFCEBB4F5EF07" box="[151,210,837,863]" pageId="22" pageNumber="23">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C6066FC1E78FFBC6531FCEBB632EF07" authority="Lee &amp; Forges &amp; Ng, 2019" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" box="[313,533,837,863]" class="Malacostraca" family="Epialtidae" genus="Crocydocinus" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="ewok" status="sp. nov.">
<emphasis id="B914C16D1E78FFBC6531FCEBB632EF07" box="[313,533,837,863]" italics="true" pageId="22" pageNumber="23">Crocydocinus ewok</emphasis>
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<taxonomicNameLabel id="A2277C161E78FFBC6613FCEBB67DEF07" box="[539,602,837,863]" pageId="22" pageNumber="23" rank="species">
<emphasis id="B914C16D1E78FFBC6613FCEBB67DEF07" bold="true" box="[539,602,837,863]" pageId="22" pageNumber="23">n. sp.</emphasis>
</taxonomicNameLabel>
, by present designation, gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C37A4EF41E78FFBC64CFFCC6B00AE89F" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8BDF1D7F1E78FFBC64CFFCC6B00AE89F" blockId="22.[151,1437,836,2015]" pageId="22" pageNumber="23">
<emphasis id="B914C16D1E78FFBC64CFFCC6B565EFDB" bold="true" box="[199,322,872,899]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
Carapace pyriform. Carapace region well-defined. Pseudorostral spines short, bifid. Carapace, legs covered in thick layer of setae. Antennae almost as long or longer than pseudorostral spines. Pre-orbital spine roundish to sharp; hiatus between supraorbital eave and post-orbital lobe wide, U-shaped; post-orbital lobe terminally blunt. Basal antennal article longer than broad, outer margin slightly curved to straight margin. Third maxilliped with ischium subquadrate; anterolateral margin of merus weakly produced. Cheliped with margins of merus, carpus, propodus, and palm of chelae carinate. Ambulatory legs slender and cylindrical, merus, carpus and propodus smooth, not carinate, covered with setae, smooth when denuded; P2 longest. Male thoracic sternites 3 and 4 fused without median sutures; lateral margins of sternites 3 and 4 slightly constricted, surface concave. Male pleon triangular, all somites and telson free. Adult female pleon rounded, with all somites and telson free (
<figureCitation id="135B01FA1E78FFBC60D1FB27B143E8FB" box="[1241,1380,1161,1187]" captionStart="FIGURE 16" captionStartId="32.[151,250,1334,1359]" captionTargetBox="[151,1436,186,1315]" captionTargetId="figure@32.[151,1436,186,1315]" captionTargetPageId="32" captionText="FIGURE 16. Overall ventral view. A, Crocydocinus panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; B, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; C, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248395/files/figure.png" pageId="22" pageNumber="23">Fig. 16A, B</figureCitation>
). G1 with single distal angle; G2 shorter than G1, with distal tip rounded (
<figureCitation id="135B01FA1E78FFBC6792FB03B7CDE89F" box="[922,1002,1197,1223]" captionStart="FIGURE 18" captionStartId="35.[151,250,1888,1913]" captionTargetBox="[197,1377,179,1863]" captionTargetId="figure@35.[192,1396,166,1868]" captionTargetPageId="35" captionText="FIGURE 18. AF, Crocydocinus ewok n. sp., holotype, male (22.5 × 15.0 mm) (MNHN-IU-2011-2328), Papua New Guinea, left G1 and G2.A, ventral view; B, ventral view of distal portion; C, dorsal view; D, dorsal view of distal portion; E, ventral view of G2; F, dorsal view of G2. GJ, C. crosnieri (Griffin &amp; Tranter, 1986) comb. nov., male (30.3 × 21.7 mm) (MNHN-IU-2016- 9524), left G1. G, ventral view; H, ventral view of distal portion; I, dorsal view; J, dorsal view of distal portion." httpUri="https://zenodo.org/record/3248399/files/figure.png" pageId="22" pageNumber="23">Fig. 18</figureCitation>
AF).
</paragraph>
</subSubSection>
<subSubSection id="C37A4EF41E78FFBC64CFFB7FB58FE90F" pageId="22" pageNumber="23" type="etymology">
<paragraph id="8BDF1D7F1E78FFBC64CFFB7FB58FE90F" blockId="22.[151,1437,836,2015]" pageId="22" pageNumber="23">
<emphasis id="B914C16D1E78FFBC64CFFB7FB569E8B3" bold="true" box="[199,334,1233,1259]" pageId="22" pageNumber="23">Etymology.</emphasis>
The new genus is named after the characteristic thick layer of setae covering the entire carapace and legs of the crab—“
<emphasis id="B914C16D1E78FFBC6590FB5BB5D6E957" box="[408,497,1269,1295]" italics="true" pageId="22" pageNumber="23">crocydo</emphasis>
” meaning downy fibers on woollen cloth in Latin, which is originally a Greek word, “
<emphasis id="B914C16D1E78FFBC64ACFAB7B52FE96B" box="[164,264,1305,1331]" italics="true" pageId="22" pageNumber="23">krokydos</emphasis>
”. It is used in arbitrary combination with the Latin word “
<emphasis id="B914C16D1E78FFBC679EFAB4B7D2E96B" box="[918,1013,1306,1331]" italics="true" pageId="22" pageNumber="23">carcinus</emphasis>
” from the Greek word “
<emphasis id="B914C16D1E78FFBC6100FAB7B14FE96B" box="[1288,1384,1305,1331]" italics="true" pageId="22" pageNumber="23">karkinus</emphasis>
” for crab. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C37A4EF41E78FFB264CFFACFB7B5EA58" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8BDF1D7F1E78FFBC64CFFACFB17DEB17" blockId="22.[151,1437,836,2015]" pageId="22" pageNumber="23">
<emphasis id="B914C16D1E78FFBC64CFFACFB51CE923" bold="true" box="[199,315,1377,1403]" pageId="22" pageNumber="23">Remarks.</emphasis>
This new genus is morphologically similar to
<taxonomicName id="4C6066FC1E78FFBC6731FACCB0C0E923" authority="Ng, Komai &amp; Sato, 2017" authorityName="Ng, Komai &amp; Sato" authorityYear="2017" box="[825,1255,1377,1403]" class="Malacostraca" family="Epialtidae" genus="Tunepugettia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B914C16D1E78FFBC6731FACCB7ECE923" box="[825,971,1378,1403]" italics="true" pageId="22" pageNumber="23">Tunepugettia</emphasis>
<bibRefCitation id="EFF1608E1E78FFBC67D9FACFB0C0E923" author="Ng, P. K. L. &amp; Komai, T. &amp; Sato, T." box="[977,1255,1377,1403]" pageId="22" pageNumber="23" pagination="133 - 152" refId="ref28986" refString="Ng, P. K. L., Komai, T. &amp; Sato, T. (2017) On the trail of a Japanese &quot; ghost species &quot; the identify of Goniopugettia tanakae Sakai, 1986, and the establishment of a new genus for Pugettia sagamiensis Gordon, 1930 (Decapoda, Brahyura, Epialtidae). Crustacean Research, 46, 133 - 152." type="journal article" year="2017">Ng, Komai &amp; Sato, 2017</bibRefCitation>
</taxonomicName>
, in general carapace form and having relatively wide male thoracic sternites 3 and 4 with slightly constricted lateral margins and a concave surface.
<taxonomicName id="4C6066FC1E78FFBC6566FA07B62EE99B" authority="Lee &amp; Forges &amp; Ng, 2019" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" box="[366,521,1449,1475]" class="Malacostraca" family="Epialtidae" genus="Crocydocinus" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B914C16D1E78FFBC6566FA07B62EE99B" box="[366,521,1449,1475]" italics="true" pageId="22" pageNumber="23">Crocydocinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2277C161E78FFBC6618FA07B678E99B" box="[528,607,1449,1475]" pageId="22" pageNumber="23" rank="genus">
<emphasis id="B914C16D1E78FFBC6618FA07B678E99B" bold="true" box="[528,607,1449,1475]" pageId="22" pageNumber="23">n. gen.</emphasis>
</taxonomicNameLabel>
, however, differs from
<taxonomicName id="4C6066FC1E78FFBC6762FA04B7DBE99B" authorityName="Ng, Komai &amp; Sato" authorityYear="2017" box="[874,1020,1450,1475]" class="Malacostraca" family="Epialtidae" genus="Tunepugettia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B914C16D1E78FFBC6762FA04B7DBE99B" box="[874,1020,1450,1475]" italics="true" pageId="22" pageNumber="23">Tunepugettia</emphasis>
</taxonomicName>
in having the carapace regions more clearly defined, often with granules on each region (
<figureCitation id="135B01FA1E78FFBC66DFFA63B714E9BF" box="[727,819,1485,1511]" captionStart="FIGURE 13" captionStartId="25.[151,250,1863,1888]" captionTargetBox="[151,1436,690,1844]" captionTargetId="figure@25.[151,1436,690,1844]" captionTargetPageId="25" captionText="FIGURE 13. Overall dorsal view. A, Crocydocinus ewok n. sp., holotype, male (22.5 × 15.0 mm) (MNHN-IU-2011-2328), Papua New Guinea; B, C. brevirostris (Doflein, 1904) comb. nov., holotype, female (22.4 × 15.5 mm) (ZMB 13647), Sumatra; C, C. crosnieri (Griffin &amp; Tranter, 1986) comb. nov., holotype, female (20.1 × 13.9 mm) (MNHN-B16073), Réunion; D, C. decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam." httpUri="https://zenodo.org/record/3248389/files/figure.png" pageId="22" pageNumber="23">Figs. 13</figureCitation>
,
<figureCitation id="135B01FA1E78FFBC6736FA63B77CE9BF" box="[830,859,1485,1511]" captionStart="FIGURE 14" captionStartId="30.[151,250,1652,1677]" captionTargetBox="[151,1436,474,1633]" captionTargetId="figure@30.[151,1436,474,1633]" captionTargetPageId="30" captionText="FIGURE 14. Overall dorsal view. A, Crocydocinus panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; B, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; C, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248391/files/figure.png" pageId="22" pageNumber="23">14</figureCitation>
) (versus carapace regions defined by strongly swollen regions and usually with no or few granules in
<taxonomicName id="4C6066FC1E78FFBC66CCFA5CB771EA53" authorityName="Ng, Komai &amp; Sato" authorityYear="2017" box="[708,854,1522,1547]" class="Malacostraca" family="Epialtidae" genus="Tunepugettia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B914C16D1E78FFBC66CCFA5CB771EA53" box="[708,854,1522,1547]" italics="true" pageId="22" pageNumber="23">Tunepugettia</emphasis>
</taxonomicName>
;
<figureCitation id="135B01FA1E78FFBC676CFA5FB7CEEA53" box="[868,1001,1521,1547]" captionStart="FIGURE 11" captionStartId="23.[151,250,1902,1927]" captionTargetBox="[151,1436,166,1882]" captionTargetId="figure@23.[151,1436,166,1882]" captionTargetPageId="23" captionText="FIGURE 11. AC, Tunepugettia sagamiensis (Gordon, 1930), male (46.0 × 31.0 mm) (ZRC 2016.0183), Japan. A, overall dorsal view; B, overall ventral view; C, lateral view of carapace. DF, T. corbariae n. sp., holotype, male (31.4 × 21.2 mm) (MNHN-IU-2014-9773), Papua New Guinea. D, overall dorsal view; E, overall ventral view; F, lateral view of carapace." httpUri="https://zenodo.org/record/3248385/files/figure.png" pageId="22" pageNumber="23">Fig. 11A, D</figureCitation>
); the pseudorostral spines are short and straight, cylindrical and diverging to form a distinct V-shaped structure (
<figureCitation id="135B01FA1E78FFBC67CFF9BBB002EA77" box="[967,1061,1557,1583]" captionStart="FIGURE 13" captionStartId="25.[151,250,1863,1888]" captionTargetBox="[151,1436,690,1844]" captionTargetId="figure@25.[151,1436,690,1844]" captionTargetPageId="25" captionText="FIGURE 13. Overall dorsal view. A, Crocydocinus ewok n. sp., holotype, male (22.5 × 15.0 mm) (MNHN-IU-2011-2328), Papua New Guinea; B, C. brevirostris (Doflein, 1904) comb. nov., holotype, female (22.4 × 15.5 mm) (ZMB 13647), Sumatra; C, C. crosnieri (Griffin &amp; Tranter, 1986) comb. nov., holotype, female (20.1 × 13.9 mm) (MNHN-B16073), Réunion; D, C. decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam." httpUri="https://zenodo.org/record/3248389/files/figure.png" pageId="22" pageNumber="23">Figs. 13</figureCitation>
,
<figureCitation id="135B01FA1E78FFBC603AF9BBB068EA77" box="[1074,1103,1557,1583]" captionStart="FIGURE 14" captionStartId="30.[151,250,1652,1677]" captionTargetBox="[151,1436,474,1633]" captionTargetId="figure@30.[151,1436,474,1633]" captionTargetPageId="30" captionText="FIGURE 14. Overall dorsal view. A, Crocydocinus panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; B, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; C, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248391/files/figure.png" pageId="22" pageNumber="23">14</figureCitation>
) (versus pseudorostral spines short, straight, slightly dorso-ventrally flattened with the outer margin slightly curved in
<taxonomicName id="4C6066FC1E78FFBC6086F994B107EA0B" authorityName="Ng, Komai &amp; Sato" authorityYear="2017" box="[1166,1312,1594,1619]" class="Malacostraca" family="Epialtidae" genus="Tunepugettia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B914C16D1E78FFBC6086F994B107EA0B" box="[1166,1312,1594,1619]" italics="true" pageId="22" pageNumber="23">Tunepugettia</emphasis>
</taxonomicName>
;
<figureCitation id="135B01FA1E78FFBC6139F997B489EA2F" captionStart="FIGURE 11" captionStartId="23.[151,250,1902,1927]" captionTargetBox="[151,1436,166,1882]" captionTargetId="figure@23.[151,1436,166,1882]" captionTargetPageId="23" captionText="FIGURE 11. AC, Tunepugettia sagamiensis (Gordon, 1930), male (46.0 × 31.0 mm) (ZRC 2016.0183), Japan. A, overall dorsal view; B, overall ventral view; C, lateral view of carapace. DF, T. corbariae n. sp., holotype, male (31.4 × 21.2 mm) (MNHN-IU-2014-9773), Papua New Guinea. D, overall dorsal view; E, overall ventral view; F, lateral view of carapace." httpUri="https://zenodo.org/record/3248385/files/figure.png" pageId="22" pageNumber="23">Fig. 11A, D</figureCitation>
); the postorbital lobe is slender with a rounded distal margin (
<figureCitation id="135B01FA1E78FFBC6764F9F3B799EA2F" box="[876,958,1629,1655]" captionStart="FIGURE 17" captionStartId="33.[151,250,1804,1829]" captionTargetBox="[184,1405,624,1785]" captionTargetId="figure@33.[184,1405,624,1786]" captionTargetPageId="33" captionText="FIGURE 17. Lateral view of carapace.A, Crocydocinus ewok n. sp., holotype, male (22.5 × 15.0 mm) (MNHN-IU-2011-2328), Papua New Guinea; B, C. brevirostris (Doflein, 1904) comb. nov., holotype, female (22.4 × 15.5 mm) (ZMB 13647), Sumatra; C, C. crosnieri (Griffin &amp; Tranter, 1986) comb. nov., holotype, female (20.1 × 13.9 mm) (MNHN-B16073), Ile de la Réunion; D, C. decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam; E, C. panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; F, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; G, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248397/files/figure.png" pageId="22" pageNumber="23">Fig. 17</figureCitation>
) (versus cup-like postorbital lobe with the side slightly flatten in
<taxonomicName id="4C6066FC1E78FFBC659AF92CB603EAC3" authorityName="Ng, Komai &amp; Sato" authorityYear="2017" box="[402,548,1666,1691]" class="Malacostraca" family="Epialtidae" genus="Tunepugettia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B914C16D1E78FFBC659AF92CB603EAC3" box="[402,548,1666,1691]" italics="true" pageId="22" pageNumber="23">Tunepugettia</emphasis>
</taxonomicName>
;
<figureCitation id="135B01FA1E78FFBC663BF92FB692EAC3" box="[563,693,1665,1691]" captionStart="FIGURE 11" captionStartId="23.[151,250,1902,1927]" captionTargetBox="[151,1436,166,1882]" captionTargetId="figure@23.[151,1436,166,1882]" captionTargetPageId="23" captionText="FIGURE 11. AC, Tunepugettia sagamiensis (Gordon, 1930), male (46.0 × 31.0 mm) (ZRC 2016.0183), Japan. A, overall dorsal view; B, overall ventral view; C, lateral view of carapace. DF, T. corbariae n. sp., holotype, male (31.4 × 21.2 mm) (MNHN-IU-2014-9773), Papua New Guinea. D, overall dorsal view; E, overall ventral view; F, lateral view of carapace." httpUri="https://zenodo.org/record/3248385/files/figure.png" pageId="22" pageNumber="23">Fig. 11C, F</figureCitation>
); the basal antennal article has a relatively straight to slightly convex outer margin with distal angle distinct (
<figureCitation id="135B01FA1E78FFBC6676F90BB6FBEAE7" box="[638,732,1700,1727]" captionStart="FIGURE 15" captionStartId="31.[151,250,1906,1931]" captionTargetBox="[151,1436,751,1887]" captionTargetId="figure@31.[151,1436,751,1887]" captionTargetPageId="31" captionText="FIGURE 15. Overall ventral view. A, Crocydocinus ewok n. sp., holotype, male (22.5 × 15.0 mm) (MNHN-IU-2011-2328), Papua New Guinea; B, C. brevirostris (Doflein, 1904) comb. nov., holotype, female (22.4 × 15.5 mm) (ZMB 13647), Sumatra; C, C. crosnieri (Griffin &amp; Tranter, 1986) comb. nov., holotype, female (20.1 × 13.9 mm) (MNHN-B16073), Réunion; D, C. decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam." httpUri="https://zenodo.org/record/3248393/files/figure.png" pageId="22" pageNumber="23">Figs. 15</figureCitation>
,
<figureCitation id="135B01FA1E78FFBC66E0F90BB722EA98" box="[744,773,1701,1728]" captionStart="FIGURE 16" captionStartId="32.[151,250,1334,1359]" captionTargetBox="[151,1436,186,1315]" captionTargetId="figure@32.[151,1436,186,1315]" captionTargetPageId="32" captionText="FIGURE 16. Overall ventral view. A, Crocydocinus panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; B, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; C, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248395/files/figure.png" pageId="22" pageNumber="23">16</figureCitation>
) (versus outer margin of basal antennal article prominently convex with distal angle less pronounced in
<taxonomicName id="4C6066FC1E78FFBC669BF964B702EABB" authorityName="Ng, Komai &amp; Sato" authorityYear="2017" box="[659,805,1738,1763]" class="Malacostraca" family="Epialtidae" genus="Tunepugettia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B914C16D1E78FFBC669BF964B702EABB" box="[659,805,1738,1763]" italics="true" pageId="22" pageNumber="23">Tunepugettia</emphasis>
</taxonomicName>
;
<figureCitation id="135B01FA1E78FFBC673DF967B79BEABB" box="[821,956,1737,1763]" captionStart="FIGURE 11" captionStartId="23.[151,250,1902,1927]" captionTargetBox="[151,1436,166,1882]" captionTargetId="figure@23.[151,1436,166,1882]" captionTargetPageId="23" captionText="FIGURE 11. AC, Tunepugettia sagamiensis (Gordon, 1930), male (46.0 × 31.0 mm) (ZRC 2016.0183), Japan. A, overall dorsal view; B, overall ventral view; C, lateral view of carapace. DF, T. corbariae n. sp., holotype, male (31.4 × 21.2 mm) (MNHN-IU-2014-9773), Papua New Guinea. D, overall dorsal view; E, overall ventral view; F, lateral view of carapace." httpUri="https://zenodo.org/record/3248385/files/figure.png" pageId="22" pageNumber="23">Fig. 11B, E</figureCitation>
); the telson of the male pleon is generally more dome shaped (
<figureCitation id="135B01FA1E78FFBC6570F943B62BEB5F" box="[376,524,1772,1799]" captionStart="FIGURE 15" captionStartId="31.[151,250,1906,1931]" captionTargetBox="[151,1436,751,1887]" captionTargetId="figure@31.[151,1436,751,1887]" captionTargetPageId="31" captionText="FIGURE 15. Overall ventral view. A, Crocydocinus ewok n. sp., holotype, male (22.5 × 15.0 mm) (MNHN-IU-2011-2328), Papua New Guinea; B, C. brevirostris (Doflein, 1904) comb. nov., holotype, female (22.4 × 15.5 mm) (ZMB 13647), Sumatra; C, C. crosnieri (Griffin &amp; Tranter, 1986) comb. nov., holotype, female (20.1 × 13.9 mm) (MNHN-B16073), Réunion; D, C. decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam." httpUri="https://zenodo.org/record/3248393/files/figure.png" pageId="22" pageNumber="23">Figs. 15A, D</figureCitation>
,
<figureCitation id="135B01FA1E78FFBC661FF943B660EB50" box="[535,583,1773,1800]" captionStart="FIGURE 16" captionStartId="32.[151,250,1334,1359]" captionTargetBox="[151,1436,186,1315]" captionTargetId="figure@32.[151,1436,186,1315]" captionTargetPageId="32" captionText="FIGURE 16. Overall ventral view. A, Crocydocinus panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; B, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; C, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248395/files/figure.png" pageId="22" pageNumber="23">16C</figureCitation>
) (versus the telson of the male pleon is more triangular in
<taxonomicName id="4C6066FC1E78FFBC60C7F940B146EB5F" authorityName="Ng, Komai &amp; Sato" authorityYear="2017" box="[1231,1377,1774,1799]" class="Malacostraca" family="Epialtidae" genus="Tunepugettia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B914C16D1E78FFBC60C7F940B146EB5F" box="[1231,1377,1774,1799]" italics="true" pageId="22" pageNumber="23">Tunepugettia</emphasis>
</taxonomicName>
;
<figureCitation id="135B01FA1E78FFBC6178F943B4C1EB73" captionStart="FIGURE 11" captionStartId="23.[151,250,1902,1927]" captionTargetBox="[151,1436,166,1882]" captionTargetId="figure@23.[151,1436,166,1882]" captionTargetPageId="23" captionText="FIGURE 11. AC, Tunepugettia sagamiensis (Gordon, 1930), male (46.0 × 31.0 mm) (ZRC 2016.0183), Japan. A, overall dorsal view; B, overall ventral view; C, lateral view of carapace. DF, T. corbariae n. sp., holotype, male (31.4 × 21.2 mm) (MNHN-IU-2014-9773), Papua New Guinea. D, overall dorsal view; E, overall ventral view; F, lateral view of carapace." httpUri="https://zenodo.org/record/3248385/files/figure.png" pageId="22" pageNumber="23">Fig. 11B, E</figureCitation>
); and the ambulatory legs are smooth with the merus, carpus and propodus not carinate (
<figureCitation id="135B01FA1E78FFBC60B1F8BFB132EB73" box="[1209,1301,1809,1835]" captionStart="FIGURE 13" captionStartId="25.[151,250,1863,1888]" captionTargetBox="[151,1436,690,1844]" captionTargetId="figure@25.[151,1436,690,1844]" captionTargetPageId="25" captionText="FIGURE 13. Overall dorsal view. A, Crocydocinus ewok n. sp., holotype, male (22.5 × 15.0 mm) (MNHN-IU-2011-2328), Papua New Guinea; B, C. brevirostris (Doflein, 1904) comb. nov., holotype, female (22.4 × 15.5 mm) (ZMB 13647), Sumatra; C, C. crosnieri (Griffin &amp; Tranter, 1986) comb. nov., holotype, female (20.1 × 13.9 mm) (MNHN-B16073), Réunion; D, C. decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam." httpUri="https://zenodo.org/record/3248389/files/figure.png" pageId="22" pageNumber="23">Figs. 13</figureCitation>
,
<figureCitation id="135B01FA1E78FFBC6128F8BFB11AEB73" box="[1312,1341,1809,1835]" captionStart="FIGURE 14" captionStartId="30.[151,250,1652,1677]" captionTargetBox="[151,1436,474,1633]" captionTargetId="figure@30.[151,1436,474,1633]" captionTargetPageId="30" captionText="FIGURE 14. Overall dorsal view. A, Crocydocinus panglao n. sp., holotype, ovigerous female (32.4 × 22.6 mm) (NMCR 50091), Philippines; B, C. porg n. sp., holotype, female (28.5 × 19.3 mm) (MNHN-IU-2013-3042), Papua New Guinea; C, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu." httpUri="https://zenodo.org/record/3248391/files/figure.png" pageId="22" pageNumber="23">14</figureCitation>
) (versus ambulatory legs with margins of merus, carpus and propodus directly carinate in
<taxonomicName id="4C6066FC1E78FFBC602CF898B091EB17" authorityName="Ng, Komai &amp; Sato" authorityYear="2017" box="[1060,1206,1846,1871]" class="Malacostraca" family="Epialtidae" genus="Tunepugettia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B914C16D1E78FFBC602CF898B091EB17" box="[1060,1206,1846,1871]" italics="true" pageId="22" pageNumber="23">Tunepugettia</emphasis>
</taxonomicName>
;
<figureCitation id="135B01FA1E78FFBC60CDF89BB16AEB17" box="[1221,1357,1845,1871]" captionStart="FIGURE 11" captionStartId="23.[151,250,1902,1927]" captionTargetBox="[151,1436,166,1882]" captionTargetId="figure@23.[151,1436,166,1882]" captionTargetPageId="23" captionText="FIGURE 11. AC, Tunepugettia sagamiensis (Gordon, 1930), male (46.0 × 31.0 mm) (ZRC 2016.0183), Japan. A, overall dorsal view; B, overall ventral view; C, lateral view of carapace. DF, T. corbariae n. sp., holotype, male (31.4 × 21.2 mm) (MNHN-IU-2014-9773), Papua New Guinea. D, overall dorsal view; E, overall ventral view; F, lateral view of carapace." httpUri="https://zenodo.org/record/3248385/files/figure.png" pageId="22" pageNumber="23">Fig. 11A, D</figureCitation>
).
</paragraph>
<paragraph id="8BDF1D7F1E78FFBC64CFF8F7B7EFEB87" blockId="22.[151,1437,836,2015]" pageId="22" pageNumber="23">
The G1s of all the species of
<taxonomicName id="4C6066FC1E78FFBC6603F8F4B6BAEB2B" authorityName="Ng, Komai &amp; Sato" authorityYear="2017" box="[523,669,1882,1907]" class="Malacostraca" family="Epialtidae" genus="Tunepugettia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B914C16D1E78FFBC6603F8F4B6BAEB2B" box="[523,669,1882,1907]" italics="true" pageId="22" pageNumber="23">Tunepugettia</emphasis>
</taxonomicName>
end in a bilobed tip (
<figureCitation id="135B01FA1E78FFBC6780F8F7B7C1EB2B" box="[904,998,1881,1907]" captionStart="FIGURE 12" captionStartId="24.[151,250,1294,1319]" captionTargetBox="[172,1393,190,1256]" captionTargetId="figure@24.[151,1436,166,1275]" captionTargetPageId="24" captionText="FIGURE 12. Tunepugettia corbariae n. sp., holotype, male (31.4 × 21.2 mm) (MNHN-IU-2014-9773), Papua New Guinea, left G1. A, ventral view; B, ventral view of distal portion; C, dorsal view; D, dorsal view of distal portion." httpUri="https://zenodo.org/record/3248387/files/figure.png" pageId="22" pageNumber="23">Figs. 12</figureCitation>
;
<bibRefCitation id="EFF1608E1E78FFBC67F9F8F7B0B3EB2B" author="Ng, P. K. L. &amp; Komai, T. &amp; Sato, T." box="[1009,1172,1881,1907]" pageId="22" pageNumber="23" pagination="133 - 152" refId="ref28986" refString="Ng, P. K. L., Komai, T. &amp; Sato, T. (2017) On the trail of a Japanese &quot; ghost species &quot; the identify of Goniopugettia tanakae Sakai, 1986, and the establishment of a new genus for Pugettia sagamiensis Gordon, 1930 (Decapoda, Brahyura, Epialtidae). Crustacean Research, 46, 133 - 152." type="journal article" year="2017">
Ng
<emphasis id="B914C16D1E78FFBC6012F8F4B06CEB2B" box="[1050,1099,1881,1907]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
. 2017
</bibRefCitation>
: fig. 7); while in almost all species of
<taxonomicName id="4C6066FC1E78FFBC653AF8D3B5EAEBCF" authority="Lee &amp; Forges &amp; Ng, 2019" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" box="[306,461,1917,1943]" class="Malacostraca" family="Epialtidae" genus="Crocydocinus" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B914C16D1E78FFBC653AF8D3B5EAEBCF" box="[306,461,1917,1943]" italics="true" pageId="22" pageNumber="23">Crocydocinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2277C161E78FFBC65DCF8D3B604EBCF" box="[468,547,1917,1943]" pageId="22" pageNumber="23" rank="genus">
<emphasis id="B914C16D1E78FFBC65DCF8D3B604EBCF" bold="true" box="[468,547,1917,1943]" pageId="22" pageNumber="23">n. gen.</emphasis>
</taxonomicNameLabel>
, the G1 distal tip is simple (
<figureCitation id="135B01FA1E78FFBC676CF8D3B7E5EBCF" box="[868,962,1917,1943]" captionStart="FIGURE 18" captionStartId="35.[151,250,1888,1913]" captionTargetBox="[197,1377,179,1863]" captionTargetId="figure@35.[192,1396,166,1868]" captionTargetPageId="35" captionText="FIGURE 18. AF, Crocydocinus ewok n. sp., holotype, male (22.5 × 15.0 mm) (MNHN-IU-2011-2328), Papua New Guinea, left G1 and G2.A, ventral view; B, ventral view of distal portion; C, dorsal view; D, dorsal view of distal portion; E, ventral view of G2; F, dorsal view of G2. GJ, C. crosnieri (Griffin &amp; Tranter, 1986) comb. nov., male (30.3 × 21.7 mm) (MNHN-IU-2016- 9524), left G1. G, ventral view; H, ventral view of distal portion; I, dorsal view; J, dorsal view of distal portion." httpUri="https://zenodo.org/record/3248399/files/figure.png" pageId="22" pageNumber="23">Figs. 18</figureCitation>
,
<figureCitation id="135B01FA1E78FFBC67C6F8D3B7CEEBCF" box="[974,1001,1917,1943]" captionStart="FIGURE 19" captionStartId="36.[151,250,1890,1915]" captionTargetBox="[281,1307,176,1860]" captionTargetId="figure@36.[243,1344,166,1870]" captionTargetPageId="36" captionText="FIGURE 19. AD, Crocydocinus decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam, left G1. A, ventral view; B, ventral view of distal portion; C, dorsal view; D, dorsal view of distal portion. EH, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu, left G1. E, ventral view; F, ventral view of distal portion; G, dorsal view; H, dorsal view of distal portion." httpUri="https://zenodo.org/record/3248401/files/figure.png" pageId="22" pageNumber="23">19</figureCitation>
EH); the exception being
<taxonomicName id="4C6066FC1E78FFBC6113F8D3B5C6EBE3" authority="(Williams &amp; Eldredge, 1994)" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" baseAuthorityName="Williams &amp; Eldredge" baseAuthorityYear="1994" class="Malacostraca" family="Epialtidae" genus="Crocydocinus" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="decipata">
<emphasis id="B914C16D1E78FFBC6113F8D3B109EBCF" box="[1307,1326,1917,1943]" italics="true" pageId="22" pageNumber="23">C</emphasis>
.
<emphasis id="B914C16D1E78FFBC6134F8D3B1BBEBCF" box="[1340,1436,1917,1943]" italics="true" pageId="22" pageNumber="23">decipata</emphasis>
(
<bibRefCitation id="EFF1608E1E78FFBC6497F80FB5FEEBE3" author="Williams, A. B. &amp; Eldredge, L. G." box="[159,473,1953,1979]" pageId="22" pageNumber="23" pagination="1 - 4" refId="ref31137" refString="Williams, A. B. &amp; Eldredge, L. G. (1994) A new species of spider crab from Guam, Rochinia decipiata (Brachyura: Majidae). Crustacean Research, 23, 1 - 4. https: // doi. org / 10.18353 / crustacea. 23.0 _ 1" type="journal article" year="1994">Williams &amp; Eldredge, 1994</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="135B01FA1E78FFBC65F9F80FB665EBE3" box="[497,578,1953,1979]" captionStart="FIGURE 19" captionStartId="36.[151,250,1890,1915]" captionTargetBox="[281,1307,176,1860]" captionTargetId="figure@36.[243,1344,166,1870]" captionTargetPageId="36" captionText="FIGURE 19. AD, Crocydocinus decipata (Williams &amp; Eldredge, 1994) comb. nov., holotype, male (14.2 × 9.1 mm) (USNM251434), Guam, left G1. A, ventral view; B, ventral view of distal portion; C, dorsal view; D, dorsal view of distal portion. EH, C. vanuatu n. sp., holotype, male (16.0 × 11.0 mm) (ZRC 2011.1047), Vanuatu, left G1. E, ventral view; F, ventral view of distal portion; G, dorsal view; H, dorsal view of distal portion." httpUri="https://zenodo.org/record/3248401/files/figure.png" pageId="22" pageNumber="23">Fig. 19</figureCitation>
AD). In the form of the basal antennal article and non-carinate ambulatory legs, however,
<taxonomicName id="4C6066FC1E78FFBC6548F86BB5E7EB87" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" baseAuthorityName="Williams &amp; Eldredge" baseAuthorityYear="1994" box="[320,448,1989,2015]" class="Malacostraca" family="Epialtidae" genus="Crocydocinus" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="decipata">
<emphasis id="B914C16D1E78FFBC6548F86BB574EB87" box="[320,339,1989,2015]" italics="true" pageId="22" pageNumber="23">C</emphasis>
.
<emphasis id="B914C16D1E78FFBC6568F86BB5E7EB87" box="[352,448,1989,2015]" italics="true" pageId="22" pageNumber="23">decipata</emphasis>
</taxonomicName>
should be transferred to
<taxonomicName id="4C6066FC1E78FFBC66D0F86BB754EB87" authority="Lee &amp; Forges &amp; Ng, 2019" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" box="[728,883,1989,2015]" class="Malacostraca" family="Epialtidae" genus="Crocydocinus" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B914C16D1E78FFBC66D0F86BB754EB87" box="[728,883,1989,2015]" italics="true" pageId="22" pageNumber="23">Crocydocinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2277C161E78FFBC6771F86BB7EFEB87" box="[889,968,1989,2015]" pageId="22" pageNumber="23" rank="genus">
<emphasis id="B914C16D1E78FFBC6771F86BB7EFEB87" bold="true" box="[889,968,1989,2015]" pageId="22" pageNumber="23">n. gen.</emphasis>
</taxonomicNameLabel>
</paragraph>
<caption id="DF1F4DF71E79FFBD649FF8C0B147EB96" httpUri="https://zenodo.org/record/3248385/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,1902,1927]" targetBox="[151,1436,166,1882]" targetPageId="23">
<paragraph id="8BDF1D7F1E79FFBD649FF8C0B147EB96" blockId="23.[151,1436,1902,1999]" pageId="23" pageNumber="24">
<emphasis id="B914C16D1E79FFBD649FF8C0B506EBDF" bold="true" box="[151,289,1902,1927]" pageId="23" pageNumber="24">FIGURE 11.</emphasis>
AC,
<taxonomicName id="4C6066FC1E79FFBD6561F8C1B73AEBDF" authority="(Gordon, 1930)" authorityName="Gordon" authorityYear="1930" baseAuthorityName="Gordon" baseAuthorityYear="1930" box="[361,797,1902,1927]" class="Malacostraca" family="Epialtidae" genus="Tunepugettia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="sagamiensis">
<emphasis id="B914C16D1E79FFBD6561F8C1B655EBDE" box="[361,626,1903,1926]" italics="true" pageId="23" pageNumber="24">Tunepugettia sagamiensis</emphasis>
(
<bibRefCitation id="EFF1608E1E79FFBD668AF8C0B731EBDF" author="Gordon I." box="[642,790,1902,1927]" pageId="23" pageNumber="24" pagination="519 - 525" refId="ref27090" refString="Gordon I. (1930) Seven new species of Brachyura from the coasts of China. Annals and Magazine of Natural History, including Zoology, Botany, and Geology, Series 10, 6 (34), 519 - 525. https: // doi. org / 10.1080 / 00222933008673240" type="journal article" year="1930">Gordon, 1930</bibRefCitation>
)
</taxonomicName>
, male (46.0 × 31.0 mm) (ZRC 2016.0183), Japan. A, overall dorsal view; B, overall ventral view; C, lateral view of carapace. DF,
<taxonomicName id="4C6066FC1E79FFBD677DF83DB7D2EBF3" authority="Lee &amp; Forges &amp; Ng, 2019" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" box="[885,1013,1939,1963]" class="Malacostraca" family="Epialtidae" genus="Tunepugettia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="corbariae" status="sp. nov.">
<emphasis id="B914C16D1E79FFBD677DF83DB7A4EBF2" box="[885,899,1939,1962]" italics="true" pageId="23" pageNumber="24">T</emphasis>
.
<emphasis id="B914C16D1E79FFBD6799F83DB7D2EBF3" box="[913,1013,1939,1963]" italics="true" pageId="23" pageNumber="24">corbariae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2277C161E79FFBD67F5F83DB01FEBF3" box="[1021,1080,1939,1963]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B914C16D1E79FFBD67F5F83DB01FEBF3" bold="true" box="[1021,1080,1939,1963]" pageId="23" pageNumber="24">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, male (31.4 × 21.2 mm) (MNHN-IU-2014-9773), Papua New Guinea. D, overall dorsal view; E, overall ventral view; F, lateral view of carapace.
</paragraph>
</caption>
<caption id="DF1F4DF71E76FFB2649FFAA0B0EEE913" httpUri="https://zenodo.org/record/3248387/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,250,1294,1319]" targetBox="[172,1393,190,1256]" targetPageId="24">
<paragraph id="8BDF1D7F1E76FFB2649FFAA0B0EEE913" blockId="24.[151,1436,1294,1355]" pageId="24" pageNumber="25">
<emphasis id="B914C16D1E76FFB2649FFAA0B506E97F" bold="true" box="[151,289,1294,1319]" pageId="24" pageNumber="25">FIGURE 12.</emphasis>
<taxonomicName id="4C6066FC1E76FFB26521FABEB63EE97F" authority="Lee &amp; Forges &amp; Ng, 2019" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" box="[297,537,1295,1319]" class="Malacostraca" family="Epialtidae" genus="Tunepugettia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="corbariae" status="sp. nov.">
<emphasis id="B914C16D1E76FFB26521FABEB63EE97F" box="[297,537,1295,1319]" italics="true" pageId="24" pageNumber="25">Tunepugettia corbariae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2277C161E76FFB26629FAA1B67BE97F" box="[545,604,1295,1319]" pageId="24" pageNumber="25" rank="species">
<emphasis id="B914C16D1E76FFB26629FAA1B67BE97F" bold="true" box="[545,604,1295,1319]" pageId="24" pageNumber="25">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, male (31.4 × 21.2 mm) (MNHN-IU-2014-9773), Papua New Guinea, left G1. A, ventral view; B, ventral view of distal portion; C, dorsal view; D, dorsal view of distal portion.
</paragraph>
</caption>
<paragraph id="8BDF1D7F1E76FFB264CFFAD7B7B5EA58" blockId="24.[151,1437,1401,1536]" pageId="24" pageNumber="25">
With the description of four new species from
<collectingCountry id="F3775DEF1E76FFB266D2FAD4B79FE9CB" box="[730,952,1401,1428]" name="Papua New Guinea" pageId="24" pageNumber="25">Papua New Guinea</collectingCountry>
,
<collectingCountry id="F3775DEF1E76FFB267CCFAD7B062E9CB" box="[964,1093,1401,1427]" name="Philippines" pageId="24" pageNumber="25">Philippines</collectingCountry>
, and
<collectingCountry id="F3775DEF1E76FFB26089FAD4B0C7E9CC" box="[1153,1248,1402,1428]" name="Vanuatu" pageId="24" pageNumber="25">Vanuatu</collectingCountry>
, and the transfer of four species to
<taxonomicName id="4C6066FC1E76FFB2656FFA33B625E9EF" authority="Lee &amp; Forges &amp; Ng, 2019" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" box="[359,514,1437,1463]" class="Malacostraca" family="Epialtidae" genus="Crocydocinus" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B914C16D1E76FFB2656FFA33B625E9EF" box="[359,514,1437,1463]" italics="true" pageId="24" pageNumber="25">Crocydocinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2277C161E76FFB26603FA30B67CE9E0" box="[523,603,1438,1464]" pageId="24" pageNumber="25" rank="genus">
<emphasis id="B914C16D1E76FFB26603FA30B67CE9E0" bold="true" box="[523,603,1438,1464]" pageId="24" pageNumber="25">n. gen.</emphasis>
</taxonomicNameLabel>
:
<taxonomicName id="4C6066FC1E76FFB26664FA30B079E9E0" authority="(Alcock &amp; Anderson, 1894)" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" baseAuthorityName="Alcock &amp; Anderson" baseAuthorityYear="1894" box="[620,1118,1437,1464]" class="Malacostraca" family="Epialtidae" genus="Crocydocinus" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="beauchampi" status="comb. nov.">
<emphasis id="B914C16D1E76FFB26664FA30B658E9E0" box="[620,639,1438,1464]" italics="true" pageId="24" pageNumber="25">C</emphasis>
.
<emphasis id="B914C16D1E76FFB26686FA33B730E9EF" box="[654,791,1437,1463]" italics="true" pageId="24" pageNumber="25">beauchampi</emphasis>
(
<bibRefCitation id="EFF1608E1E76FFB2672FFA33B071E9E0" author="Alcock, A. &amp; Anderson, A. R. S." box="[807,1110,1437,1464]" pageId="24" pageNumber="25" pagination="141 - 185" refId="ref26443" refString="Alcock, A. &amp; Anderson, A. R. S. (1894) Natural History Notes from H. M. Indian Marine Survey Steamer ' Investigator, ' Commander C. F. Oldham, R. N., commanding. Series II., No. 14. An Account of a Recent Collection of Deep Sea Crustacea from the Bay of Bengal and Laccadive Sea. Journal of the Asiatic Society of Bengal, 63 (Part 2, no. 3), 141 - 185, pl. IX." type="journal article" year="1894">Alcock &amp; Anderson, 1894</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A2277C161E76FFB2606FFA30B0CEE9E0" box="[1127,1257,1438,1464]" pageId="24" pageNumber="25" rank="species">
<emphasis id="B914C16D1E76FFB2606FFA30B0CEE9E0" bold="true" box="[1127,1257,1438,1464]" pageId="24" pageNumber="25">comb. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C6066FC1E76FFB260F1FA30B560E984" authority="(Doflein, 1904)" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" baseAuthorityName="Doflein" baseAuthorityYear="1904" class="Malacostraca" family="Epialtidae" genus="Rochinia" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="brevirostris" status="comb. nov.">
<emphasis id="B914C16D1E76FFB260F1FA30B12BE9E0" box="[1273,1292,1438,1464]" italics="true" pageId="24" pageNumber="25">C</emphasis>
.
<emphasis id="B914C16D1E76FFB26114FA33B1BAE9EF" box="[1308,1437,1437,1463]" italics="true" pageId="24" pageNumber="25">brevirostris</emphasis>
(
<bibRefCitation id="EFF1608E1E76FFB26497FA6FB518E984" author="Doflein, F." box="[159,319,1473,1500]" pageId="24" pageNumber="25" pagination="1 - 58" refId="ref27028" refString="Doflein, F. (1904) Brachyura. In: Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899. Vol. 6. Gustav Fischer, Jena, pp. 1 - 6 + i-xiv + 1 - 314 pp., figs. I-LVIII, atlas (pls. 1 - 58)." type="book chapter" year="1904">Doflein, 1904</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A2277C161E76FFB26547FA6CB5F6E984" box="[335,465,1474,1500]" pageId="24" pageNumber="25" rank="species">
<emphasis id="B914C16D1E76FFB26547FA6CB5F6E984" bold="true" box="[335,465,1474,1500]" pageId="24" pageNumber="25">comb. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="4C6066FC1E76FFB265E9FA6CB7A9E984" authority="(Griffin &amp; Tranter, 1986)" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" baseAuthorityName="Griffin &amp; Tranter" baseAuthorityYear="1986" box="[481,910,1473,1500]" class="Malacostraca" family="Epialtidae" genus="Crocydocinus" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="crosnieri" status="comb. nov.">
<emphasis id="B914C16D1E76FFB265E9FA6CB5D3E984" box="[481,500,1474,1500]" italics="true" pageId="24" pageNumber="25">C</emphasis>
.
<emphasis id="B914C16D1E76FFB2660BFA6CB640E983" box="[515,615,1474,1499]" italics="true" pageId="24" pageNumber="25">crosnieri</emphasis>
(Griffin &amp; Tranter, 1986)
</taxonomicName>
<taxonomicNameLabel id="A2277C161E76FFB2679FFA6CB03EE984" box="[919,1049,1474,1500]" pageId="24" pageNumber="25" rank="species">
<emphasis id="B914C16D1E76FFB2679FFA6CB03EE984" bold="true" box="[919,1049,1474,1500]" pageId="24" pageNumber="25">comb. nov.</emphasis>
</taxonomicNameLabel>
, and
<taxonomicName id="4C6066FC1E76FFB26051FA6CB514EA58" authority="(Williams &amp; Eldredge, 1994)" authorityName="Lee &amp; Forges &amp; Ng" authorityYear="2019" baseAuthorityName="Williams &amp; Eldredge" baseAuthorityYear="1994" class="Malacostraca" family="Epialtidae" genus="Crocydocinus" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="decipata" status="comb. nov.">
<emphasis id="B914C16D1E76FFB26051FA6CB04BE984" box="[1113,1132,1474,1500]" italics="true" pageId="24" pageNumber="25">C</emphasis>
.
<emphasis id="B914C16D1E76FFB26073FA6FB0FCE983" box="[1147,1243,1473,1499]" italics="true" pageId="24" pageNumber="25">decipata</emphasis>
(
<bibRefCitation id="EFF1608E1E76FFB260E4FA6FB50DEA58" author="Williams, A. B. &amp; Eldredge, L. G." pageId="24" pageNumber="25" pagination="1 - 4" refId="ref31137" refString="Williams, A. B. &amp; Eldredge, L. G. (1994) A new species of spider crab from Guam, Rochinia decipiata (Brachyura: Majidae). Crustacean Research, 23, 1 - 4. https: // doi. org / 10.18353 / crustacea. 23.0 _ 1" type="journal article" year="1994">Williams &amp; Eldredge, 1994</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A2277C161E76FFB26532FA48B59DEA58" box="[314,442,1510,1536]" pageId="24" pageNumber="25" rank="species">
<emphasis id="B914C16D1E76FFB26532FA48B59DEA58" bold="true" box="[314,442,1510,1536]" pageId="24" pageNumber="25">comb. nov.</emphasis>
</taxonomicNameLabel>
, there are eight species in this new genus.
</paragraph>
</subSubSection>
</treatment>
</document>