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<mods:title id="2AB593EB3E8962E330669644BB1B6C42">Two new species of Coccomorpha (Hemiptera: Sternorrhyncha) collected from Japanese silver grass, Miscanthus sinensis (Poaceae) in Okinawa Island, Japan</mods:title>
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<mods:namePart id="1BA92910F0CB339B0547A631B5C4052A">Tanaka, Hirotaka</mods:namePart>
<mods:affiliation id="9A06021D0E92FBD063A0A2051B1465EF">Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, Ehime 790 - 8566, Japan. &amp; The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka, 812 - 8581 Japan.</mods:affiliation>
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<mods:namePart id="D3F47888E4F5E11ECB1054A79A07E9BD">Kamitani, Satoshi</mods:namePart>
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<mods:affiliation id="4604B4E11002655FD32C2F3573FAF7D0">Entomological Laboratory, Faculty of Agriculture, Kyushu University, Motooka 744, Fukuoka, 819 - 0395 Japan. kamitani @ agr. kyushu-u. ac. jp; https: // orcid. org / 0000 - 0003 - 0870 - 5163</mods:affiliation>
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<paragraph id="8BDFAC5432223346FF64F9C4FDEDF9EE" blockId="4.[151,554,1547,1574]" box="[151,554,1547,1574]" pageId="4" pageNumber="573">
<heading id="D0971B3832223346FF64F9C4FDEDF9EE" bold="true" box="[151,554,1547,1574]" fontSize="11" level="1" pageId="4" pageNumber="573" reason="1">
<emphasis id="B914704632223346FF64F9C4FDEDF9EE" bold="true" box="[151,554,1547,1574]" pageId="4" pageNumber="573">
Genus
<taxonomicName id="4C60D7D732223346FF1DF9C3FDEDF9EE" ID-CoL="495R" ID-ENA="38113" authority="Ferris, 1950" authorityName="Ferris" authorityYear="1950" box="[238,554,1547,1574]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="573" phylum="Arthropoda" rank="genus">
<emphasis id="B914704632223346FF1DF9C3FE4BF9EE" bold="true" box="[238,396,1548,1574]" italics="true" pageId="4" pageNumber="573">Dysmicoccus</emphasis>
<bibRefCitation id="EFF1D1A532223346FE67F9C4FDEDF9EE" author="Ferris, G. F." box="[404,554,1547,1574]" pageId="4" pageNumber="573" refId="ref5949" refString="Ferris, G. F. (1950) Atlas of the Scale Insects of North America. Series 5. Vol. 5. Part. I. The Pseudococcidae (Part I). Stanford University Press, Palo Alto, California, 278 pp." type="book" year="1950">Ferris, 1950</bibRefCitation>
</taxonomicName>
</emphasis>
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</paragraph>
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<subSubSection id="C37AFFDF32223346FF64F99CFBDFF81E" pageId="4" pageNumber="573" type="diagnosis">
<paragraph id="8BDFAC5432223346FF64F99CFBDFF81E" blockId="4.[151,1437,1619,2042]" pageId="4" pageNumber="573">
<emphasis id="B914704632223346FF64F99CFECDF9A5" bold="true" box="[151,266,1619,1645]" pageId="4" pageNumber="573">Diagnosis</emphasis>
(adopted and slightly modified from
<bibRefCitation id="EFF1D1A532223346FD55F99BFC88F9A6" author="Williams, D. J." box="[678,847,1620,1646]" pageId="4" pageNumber="573" refId="ref6902" refString="Williams, D. J. (2004) Mealybugs of Southern Asia. The Natural History Museum. Southdene SDN. BHD, Kuala Lumpur, 896 pp." type="book" year="2004">Williams 2004</bibRefCitation>
). Body of adult female elongate to broadly oval. Anal lobes usually developed, either membranous or sclerotised, each lobe bearing a normal apical seta. Ventral margins of abdominal segments anterior to anal lobes always membranous. Antennae each with 68 segments. Legs well developed; translucent pores present or absent; tarsal digitules usually knobbed. Claw without a denticle. Cerarii numbering 617 pairs. Auxiliary setae present, at least in anal lobe cerarii. Anal lobe cerarii each bearing either 2 cerarian setae or as many as 8, these usually conical; sometimes conical setae replaced by flagellate setae but cerarii always recognisable by concentrations of trilocular pores. Anal ring normally situated at apex of abdomen, usually bearing 6 setae. Anterior and posterior ostioles present. Dorsal setae variously shaped. Ventral setae flagellate. Trilocular pores present on both dorsal and ventral surface. Multilocular pores usually present, at least on venter. Quinquelocular pores always absent. Oral collar tubular ducts usually present. Oral rim tubular ducts always absent. Discoidal pores present, sometimes large, occasionally present next to each eye.
</paragraph>
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<paragraph id="8BDFAC5432223347FF34F82FFAAFFF31" blockId="4.[151,1437,1619,2042]" lastBlockId="5.[151,1437,151,250]" lastPageId="5" lastPageNumber="574" pageId="4" pageNumber="573">
<emphasis id="B914704632223346FF34F82FFEFCF832" bold="true" box="[199,315,2016,2042]" pageId="4" pageNumber="573">Remarks.</emphasis>
The molecular phylogenetic study on mealybugs conducted by
<bibRefCitation id="EFF1D1A532223346FC08F82FFAD5F832" author="Downie, D. A. &amp; Gullan, P. J." box="[1019,1298,2016,2042]" pageId="4" pageNumber="573" pagination="238 - 259" refId="ref5844" refString="Downie, D. A. &amp; Gullan, P. J. (2004) Phylogenetic analysis of mealybugs (Hemiptera: Coccoidea: Pseudococcidae) based on DNA sequences from three nuclear genes, and a review of the higher classification. Systematic Entomology, 29, 238 - 259. https: // doi. org / 10.1111 / j. 0307 - 6970.2004.00241. x" type="journal article" year="2004">Downie &amp; Gullan (2004)</bibRefCitation>
showed that the genus
<taxonomicName id="4C60D7D732233347FEF4FF58FE5CFF78" authorityName="Ferris" authorityYear="1950" box="[263,411,151,176]" class="Insecta" family="Pseudococcidae" genus="Dysmicoccus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B914704632233347FEF4FF58FE5CFF78" box="[263,411,151,176]" italics="true" pageId="5" pageNumber="574">Dysmicoccus</emphasis>
</taxonomicName>
is probably not a simple monophletic group but forms a large single clade with several other genera, such as
<taxonomicName id="4C60D7D732233347FEBBFF74FD6EFF1D" authority="Westwood, 1840" authorityName="Westwood" authorityYear="1840" box="[328,681,187,213]" class="Insecta" family="Pseudococcidae" genus="Pseudococcus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B914704632233347FEBBFF74FE20FF1D" box="[328,487,187,213]" italics="true" pageId="5" pageNumber="574">Pseudococcus</emphasis>
<bibRefCitation id="EFF1D1A532233347FE1DFF74FD6EFF1D" author="Westwood, J. O." box="[494,681,187,213]" pageId="5" pageNumber="574" refId="ref6839" refString="Westwood, J. O. (1840) An introduction to the modern classification of insects; founded on the natural habits and corresponding organization of different families. Vol. II. Longman, Orme, Brown, Green and Longmans, London, 587 pp. https: // doi. org / 10.5962 / bhl. title. 12455" type="book" year="1840">Westwood, 1840</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C60D7D732233347FD13FF74FC1BFF1D" authority="Berg, 1899" authorityName="Berg" authorityYear="1899" box="[736,988,187,213]" class="Insecta" family="Pseudococcidae" genus="Trionymus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B914704632233347FD13FF74FC91FF1C" box="[736,854,187,212]" italics="true" pageId="5" pageNumber="574">Trionymus</emphasis>
<bibRefCitation id="EFF1D1A532233347FCAEFF74FC1BFF1D" author="Berg, C." box="[861,988,187,213]" pageId="5" pageNumber="574" pagination="77 - 80" refId="ref5742" refString="Berg, C. (1899) Substitucion de nombres genericos. III. Comunicaciones del Museo Nacional de Buenos Aires, 1, 77 - 80." type="journal article" year="1899">Berg, 1899</bibRefCitation>
</taxonomicName>
, so the current definition of the genus is probably arbitrary (see below). Further molecular and morphological studies on this genus are greatly needed.
</paragraph>
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