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<mods:title id="19F4F31A8A4428EBA2AF03A90C608D6C">The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea</mods:title>
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<mods:namePart id="6D64563552CB7A88A2E91ED6A45CFA6C">Landau, Bernard M.</mods:namePart>
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<taxonomicName id="4C675589FFCD0C10FF65FD3BED8EFD77" ID-CoL="86ZT4" authority="Kovacs, 2018" authorityName="Kovacs" authorityYear="2018" box="[151,507,726,753]" class="Gastropoda" family="Tudiclidae" genus="Euthria" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="viciani">
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<emphasis id="B913F218FFCD0C10FF65FD3BED3BFD76" bold="true" box="[151,334,726,753]" italics="true" pageId="87" pageNumber="88">Euthria viciani</emphasis>
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*
<taxonomicName id="4C675589FFCD0C10FF58FCD2ED36FCD0" ID-CoL="86ZT4" authority="Harzhauser &amp; Landau, 2024" authorityName="Harzhauser &amp; Landau" authorityYear="2024" box="[170,323,831,855]" class="Gastropoda" family="Tudiclidae" genus="Euthria" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="viciani" status="sp. nov.">Euthria viciani</taxonomicName>
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, figs 29.
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<emphasis id="B913F218FFCD0C10FF65FC68ED33FC18" bold="true" box="[151,326,901,927]" pageId="87" pageNumber="88">Type material.</emphasis>
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and
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: figs 23),
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(
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<figureCitation id="135C328FFFCD0C10FCB7FC44EFD9FC44" box="[837,940,937,963]" captionStart="FIGURE 46" captionStartId="88.[151,250,1201,1225]" captionTargetBox="[151,1435,181,1177]" captionTargetId="figure-30@88.[151,1435,181,1177]" captionTargetPageId="88" captionText="FIGURE 46. Euthria viciani Kovács, 2018. A1A2. PAL 2018.1.1. HNHM, holotype, Bánd, (Hungary). B1B. PAL 2018.2.1. HNHM, paratype, Bánd (Hungary). C. Coll.V.2017.0203, Collection Vicián, paratype, Bánd (Hungary). Euthria zachosi nov. sp. D1D2. NHMW 2023/0350/0003, holotype, Bad Vöslau (Austria). E1E2. NHMW 2023/0350/0003, paratype, Bad Vöslau (Austria). F. NHMW 2023/0347/0002, paratype, Niederleis (Austria)." figureDoi="http://doi.org/10.5281/zenodo.10841165" httpUri="https://zenodo.org/record/10841165/files/figure.png" pageId="87" pageNumber="88">Figs 46A</figureCitation>
<figureCitation id="135C328FFFCD0C10FC5FFC51EFA2FC43" box="[941,983,938,970]" captionStart="FIGURE 1" captionStartId="5.[151,250,1616,1640]" captionTargetBox="[169,1418,867,1575]" captionTargetId="figure-259@5.[151,1436,849,1593]" captionTargetPageId="5" captionText="FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width." figureDoi="http://doi.org/10.5281/zenodo.10841063" httpUri="https://zenodo.org/record/10841063/files/figure.png" pageId="87" pageNumber="88">
<subScript id="17E32C4FFFCD0C10FC5FFC51EFC0FC4D" attach="right" box="[941,949,956,970]" fontSize="6" pageId="87" pageNumber="88">1</subScript>
A
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<figureCitation id="135C328FFFCD0C10FC25FC51EFAAFC4D" box="[983,991,956,970]" captionStart="FIGURE 2" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[182,1406,181,1850]" captionTargetId="figure-26@7.[182,1406,181,1850]" captionTargetPageId="7" captionText="FIGURE 2. Colubraria subobscura (Hoernes &amp; Auinger, 1884). A1A3. NHMW 1854/0035/0170, paralectotype, Lăpugiu de Sus (Romania). B1B2. NHMW 1859/0037/0018a, paralectotype, Lăpugiu de Sus (Romania). C1C3. NHMW 1865/0001/0183, paralectotype, Lăpugiu de Sus (Romania). D1D2. NHMW 1859/0037/0018b, paralectotype, Lăpugiu de Sus (Romania). E1E3. NHMW 1874/0025/0021, lectotype, Lăpugiu de Sus (Romania). F1F2. NHMW 2023/0338/0001, paralectotype, Lăpugiu de Sus (Romania)." figureDoi="http://doi.org/10.5281/zenodo.10841065" httpUri="https://zenodo.org/record/10841065/files/figure.png" pageId="87" pageNumber="88">
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.
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<typeStatus id="54DC90A8FFCD0C10FC1FFC47E82BFC43" box="[1005,1118,938,964]" pageId="87" pageNumber="88" type="paratype">Paratypes</typeStatus>
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.
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, SL:
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,
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, illustrated in
<bibRefCitation id="EFF653FBFFCD0C10FDDEFC20EEBBFC60" author="Kovacs, Z." box="[556,718,973,999]" pageId="87" pageNumber="88" pagination="179 - 182" refId="ref86477" refString="Kovacs, Z. (2018) New records of the genus Euthria (Mollusca, Buccinidae) in the Miocene Paratethys. Foldtani Kozlony, 148 (2), 179 - 182. https: // doi. org / 10.23928 / foldt. kozl. 2018.148.2.179" type="journal article" year="2018">Kovács (2018</bibRefCitation>
: figs 89), Bánd (
<collectingCountry id="F3706E9AFFCD0C10FC6CFC20E871FC60" box="[926,1028,973,999]" name="Hungary" pageId="87" pageNumber="88">Hungary</collectingCountry>
),
<figureCitation id="135C328FFFCD0C10FBEBFC20E8F4FC60" box="[1049,1153,973,999]" captionStart="FIGURE 46" captionStartId="88.[151,250,1201,1225]" captionTargetBox="[151,1435,181,1177]" captionTargetId="figure-30@88.[151,1435,181,1177]" captionTargetPageId="88" captionText="FIGURE 46. Euthria viciani Kovács, 2018. A1A2. PAL 2018.1.1. HNHM, holotype, Bánd, (Hungary). B1B. PAL 2018.2.1. HNHM, paratype, Bánd (Hungary). C. Coll.V.2017.0203, Collection Vicián, paratype, Bánd (Hungary). Euthria zachosi nov. sp. D1D2. NHMW 2023/0350/0003, holotype, Bad Vöslau (Austria). E1E2. NHMW 2023/0350/0003, paratype, Bad Vöslau (Austria). F. NHMW 2023/0347/0002, paratype, Niederleis (Austria)." figureDoi="http://doi.org/10.5281/zenodo.10841165" httpUri="https://zenodo.org/record/10841165/files/figure.png" pageId="87" pageNumber="88">Figs 46B</figureCitation>
<figureCitation id="135C328FFFCD0C10FB73FC0DE8DCFC6F" box="[1153,1193,974,1006]" captionStart="FIGURE 1" captionStartId="5.[151,250,1616,1640]" captionTargetBox="[169,1418,867,1575]" captionTargetId="figure-259@5.[151,1436,849,1593]" captionTargetPageId="5" captionText="FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width." figureDoi="http://doi.org/10.5281/zenodo.10841063" httpUri="https://zenodo.org/record/10841063/files/figure.png" pageId="87" pageNumber="88">
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B
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<figureCitation id="135C328FFFCD0C10FB5BFC0DE8C4FC69" box="[1193,1201,992,1006]" captionStart="FIGURE 2" captionStartId="7.[151,250,1874,1898]" captionTargetBox="[182,1406,181,1850]" captionTargetId="figure-26@7.[182,1406,181,1850]" captionTargetPageId="7" captionText="FIGURE 2. Colubraria subobscura (Hoernes &amp; Auinger, 1884). A1A3. NHMW 1854/0035/0170, paralectotype, Lăpugiu de Sus (Romania). B1B2. NHMW 1859/0037/0018a, paralectotype, Lăpugiu de Sus (Romania). C1C3. NHMW 1865/0001/0183, paralectotype, Lăpugiu de Sus (Romania). D1D2. NHMW 1859/0037/0018b, paralectotype, Lăpugiu de Sus (Romania). E1E3. NHMW 1874/0025/0021, lectotype, Lăpugiu de Sus (Romania). F1F2. NHMW 2023/0338/0001, paralectotype, Lăpugiu de Sus (Romania)." figureDoi="http://doi.org/10.5281/zenodo.10841065" httpUri="https://zenodo.org/record/10841065/files/figure.png" pageId="87" pageNumber="88">
<subScript id="17E32C4FFFCD0C10FB5BFC0DE8C4FC69" attach="left" box="[1193,1201,992,1006]" fontSize="6" pageId="87" pageNumber="88">2</subScript>
</figureCitation>
.
<collectorName id="26924BDCFFCD0C10FB33FC20E887FC60" box="[1217,1266,973,999]" pageId="87" pageNumber="88">Coll</collectorName>
</materialsCitation>
.
<collectionCode id="ED76B6CFFFCD0C10FB04FC20E97FFC60" box="[1270,1290,973,999]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="87" pageNumber="88" type="Museum">V</collectionCode>
.2017.0203, Collection Vicián, SL:
<quantity id="4C9F83EFFFCD0C10FE6BFC1CED9FFB8B" box="[409,490,1009,1036]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="87" pageNumber="88" unit="mm" value="44.0">44 mm</quantity>
,
<materialsCitation id="3B0F2457FFCD0C10FE07FC1FE91CFB8C" box="[501,1385,1009,1036]" collectionCode="MD" collectorName="Band" country="Hungary" location="Hungary" pageId="87" pageNumber="88" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED76B6CFFFCD0C10FE07FC1FEE53FB8B" box="[501,550,1010,1036]" country="Germany" name="Museum Donaueschingen" pageId="87" pageNumber="88">MD</collectionCode>
:
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, illustrated in
<bibRefCitation id="EFF653FBFFCD0C10FCD0FC1CEFB7FB8C" author="Kovacs, Z." box="[802,962,1009,1035]" pageId="87" pageNumber="88" pagination="179 - 182" refId="ref86477" refString="Kovacs, Z. (2018) New records of the genus Euthria (Mollusca, Buccinidae) in the Miocene Paratethys. Foldtani Kozlony, 148 (2), 179 - 182. https: // doi. org / 10.23928 / foldt. kozl. 2018.148.2.179" type="journal article" year="2018">Kovács (2018</bibRefCitation>
: figs 45),
<collectorName id="26924BDCFFCD0C10FBCDFC1CE80FFB8C" box="[1087,1146,1009,1035]" pageId="87" pageNumber="88">Bánd</collectorName>
(
<collectingCountry id="F3706E9AFFCD0C10FB7BFC1CE884FB8C" box="[1161,1265,1009,1035]" name="Hungary" pageId="87" pageNumber="88">Hungary</collectingCountry>
),
<figureCitation id="135C328FFFCD0C10FAF6FC1CE91CFB8C" box="[1284,1385,1009,1035]" captionStart="FIGURE 46" captionStartId="88.[151,250,1201,1225]" captionTargetBox="[151,1435,181,1177]" captionTargetId="figure-30@88.[151,1435,181,1177]" captionTargetPageId="88" captionText="FIGURE 46. Euthria viciani Kovács, 2018. A1A2. PAL 2018.1.1. HNHM, holotype, Bánd, (Hungary). B1B. PAL 2018.2.1. HNHM, paratype, Bánd (Hungary). C. Coll.V.2017.0203, Collection Vicián, paratype, Bánd (Hungary). Euthria zachosi nov. sp. D1D2. NHMW 2023/0350/0003, holotype, Bad Vöslau (Austria). E1E2. NHMW 2023/0350/0003, paratype, Bad Vöslau (Austria). F. NHMW 2023/0347/0002, paratype, Niederleis (Austria)." figureDoi="http://doi.org/10.5281/zenodo.10841165" httpUri="https://zenodo.org/record/10841165/files/figure.png" pageId="87" pageNumber="88">Fig. 46C</figureCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD82E0AFFCD0C10FF35FBF8EFB0FA3B" blockId="87.[151,1437,901,1827]" pageId="87" pageNumber="88">
<emphasis id="B913F218FFCD0C10FF35FBF8EDC9FBA8" bold="true" box="[199,444,1045,1071]" pageId="87" pageNumber="88">Revised description.</emphasis>
Medium-sized, moderately broad, ovate shell of up to seven teleoconch whorls with slightly coeloconoid early spire passing to cyrtoconoid on later whorls; apical angle ~60°. Protoconch paucispiral of 1.25 smooth whorls. First two teleoconch whorls weakly convex with weak axial ribs and delicate spiral cords. Later teleoconch whorls flat-sided, smooth, separated by indistinct suture. Last whorl smooth, ovate, broad, very high, attaining ~80% of total height, subsutural ramp slightly concave, hardly delimited, broadly convex below; base moderately constricted bearing weak spiral cords; fasciole slightly swollen. Aperture moderately small, ovate. Columella broadly and weakly excavated, weakly angled at transition to siphonal canal. Columellar callus extending as distinct rim along siphonal canal, weakly delimited from base in parietal area. Anal canal incised, accentuated by prominent parietal denticle. Indistinct swelling in area of anal denticle. Outer lip strongly thickened adapically, thinning below, with about ten short lirae starting some distance behind peristome. Siphonal canal moderately long, relatively wide, deflected to the left, shallowly notched. Color pattern of numerous bands of subquadrate brown and white blotches, resulting in a somewhat blurred, chessboard-like pattern.
</paragraph>
</subSubSection>
<subSubSection id="C37D7D81FFCD0C10FF35FA28ED98F95C" pageId="87" pageNumber="88" type="discussion">
<paragraph id="8BD82E0AFFCD0C10FF35FA28EDC0F9CB" blockId="87.[151,1437,901,1827]" pageId="87" pageNumber="88">
<emphasis id="B913F218FFCD0C10FF35FA28ED39FA58" bold="true" box="[199,332,1477,1503]" pageId="87" pageNumber="88">Discussion.</emphasis>
<taxonomicName id="4C675589FFCD0C10FEA4FA28EE77FA58" authorityName="Harzhauser &amp; Landau" authorityYear="2024" box="[342,514,1477,1503]" class="Gastropoda" family="Tudiclidae" genus="Euthria" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="viciani">
<emphasis id="B913F218FFCD0C10FEA4FA28EE77FA58" box="[342,514,1477,1503]" italics="true" pageId="87" pageNumber="88">Euthria viciani</emphasis>
</taxonomicName>
is unique within the Circum-Mediterranean
<taxonomicName id="4C675589FFCD0C10FBFBFA28E8C8FA58" box="[1033,1213,1477,1503]" pageId="87" pageNumber="88">
<emphasis id="B913F218FFCD0C10FBFBFA28E82AFA58" box="[1033,1119,1477,1503]" italics="true" pageId="87" pageNumber="88">Euthria</emphasis>
species
</taxonomicName>
, due to its peculiar color pattern and due to the change from coeloconoid to cyrtoconoid spire. Its shape is slightly reminiscent of
<taxonomicName id="4C675589FFCD0C10FF65F9E0EE94F9A0" authority="(Hoernes &amp; Auinger, 1890)" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1890" box="[151,737,1549,1576]" class="Gastropoda" family="Tudiclidae" genus="Euthria" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="fuscocingulatus">
<emphasis id="B913F218FFCD0C10FF65F9E0EDD1F9A0" box="[151,420,1549,1575]" italics="true" pageId="87" pageNumber="88">Euthria fuscocingulatus</emphasis>
(Hoernes &amp; Auinger, 1890)
</taxonomicName>
and
<taxonomicName id="4C675589FFCD0C10FCEAF9E0E8C7F9A0" authority="Bandat, 1943" authorityName="Bandat" authorityYear="1943" box="[792,1202,1549,1575]" class="Gastropoda" family="Tudiclidae" genus="Euthria" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="depressospira">
<emphasis id="B913F218FFCD0C10FCEAF9E0E867F9A0" box="[792,1042,1549,1575]" italics="true" pageId="87" pageNumber="88">Euthria depressospira</emphasis>
<bibRefCitation id="EFF653FBFFCD0C10FBEBF9E0E8C7F9A0" author="Bandat, H." box="[1049,1202,1549,1575]" pageId="87" pageNumber="88" pagination="289 - 367" refId="ref77008" refString="Bandat, H. (1943) Ujharmadkori csigak Delalbaniabol (Tertiary gastropods from South Albania). Foldtani Szemle, 1 (6), 289 - 367." type="journal article" year="1943">Bandat, 1943</bibRefCitation>
</taxonomicName>
, which both lack the concave subsutural ramp.
</paragraph>
<paragraph id="8BD82E0AFFCD0C10FF35F9B8EFBBF914" blockId="87.[151,1437,901,1827]" pageId="87" pageNumber="88">
The extant
<taxonomicName id="4C675589FFCD0C10FEB5F9B8EF78F9E8" authority="Fraussen &amp; Rolan, 2003" authorityName="Fraussen &amp; Rolan" authorityYear="2003" box="[327,781,1621,1648]" class="Gastropoda" family="Tudiclidae" genus="Euthria" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="cecilea">
<emphasis id="B913F218FFCD0C10FEB5F9B8ED86F9E8" box="[327,499,1621,1647]" italics="true" pageId="87" pageNumber="88">Euthria cecilea</emphasis>
<bibRefCitation id="EFF653FBFFCD0C10FE09F9BBEF78F9E8" author="Fraussen, K. &amp; Rolan, E." box="[507,781,1621,1648]" pageId="87" pageNumber="88" pagination="76 - 93" refId="ref81718" refString="Fraussen, K. &amp; Rolan, E. (2003) Four new Euthria (Gastropoda: Buccinidae) from the Cape Verde Archipelago. Gloria Maris, 42 (4 - 5), 76 - 93." type="journal article" year="2003">Fraussen &amp; Rolán, 2003</bibRefCitation>
</taxonomicName>
from the
<collectingCountry id="F3706E9AFFCD0C10FC8FF9B8EF8AF9E8" box="[893,1023,1621,1647]" name="Cape Verde" pageId="87" pageNumber="88">Cape Verde</collectingCountry>
Islands, is reminiscent of
<taxonomicName id="4C675589FFCD0C10FADBF9BBE9E0F9E8" authorityName="Harzhauser &amp; Landau" authorityYear="2024" box="[1321,1429,1622,1647]" class="Gastropoda" family="Tudiclidae" genus="Euthria" kingdom="Animalia" order="Neogastropoda" pageId="87" pageNumber="88" phylum="Mollusca" rank="species" species="viciani">
<emphasis id="B913F218FFCD0C10FADBF9BBE9E0F9E8" box="[1321,1429,1622,1647]" italics="true" pageId="87" pageNumber="88">E. viciani</emphasis>
</taxonomicName>
, but its blotches are less strictly subquadratic and its spire is more conical.
</paragraph>
<paragraph id="8BD82E0AFFCD0C10FF35F970ED98F95C" blockId="87.[151,1437,901,1827]" pageId="87" pageNumber="88">
<emphasis id="B913F218FFCD0C10FF35F970EDDDF930" bold="true" box="[199,424,1693,1719]" pageId="87" pageNumber="88">Paleoenvironment.</emphasis>
The locality Bánd comprises an assemblage indicative for a shallow marine depositional environment (own data M.H.).
</paragraph>
</subSubSection>
<subSubSection id="C37D7D81FFCD0C10FF35F908EFB3F8A4" pageId="87" pageNumber="88" type="distribution">
<paragraph id="8BD82E0AFFCD0C10FF35F908EFB3F8A4" blockId="87.[151,1437,901,1827]" pageId="87" pageNumber="88">
<emphasis id="B913F218FFCD0C10FF35F908EE0CF978" bold="true" box="[199,633,1765,1791]" pageId="87" pageNumber="88">Distribution in Central Paratethys.</emphasis>
Badenian (Middle Miocene):
<emphasis id="B913F218FFCD0C10FC12F90BE8F7F978" box="[992,1154,1766,1791]" italics="true" pageId="87" pageNumber="88">
<collectingRegion id="49A3E0E8FFCD0C10FC12F90BE858F978" box="[992,1069,1766,1791]" country="Austria" name="Wien" pageId="87" pageNumber="88">Vienna</collectingRegion>
Basin:
</emphasis>
Niederleis, Bad Vöslau (
<collectingCountry id="F3706E9AFFCD0C10FF6DF8E4EC80F8A4" box="[159,245,1801,1827]" name="Austria" pageId="87" pageNumber="88">Austria</collectingCountry>
) (hoc opus);
<emphasis id="B913F218FFCD0C10FE7BF8E7EE21F8A4" box="[393,596,1802,1827]" italics="true" pageId="87" pageNumber="88">Pannonian Basin:</emphasis>
Bánd (
<collectingCountry id="F3706E9AFFCD0C10FD54F8E4EF7FF8A4" box="[678,778,1801,1827]" name="Hungary" pageId="87" pageNumber="88">Hungary</collectingCountry>
) (
<bibRefCitation id="EFF653FBFFCD0C10FCD2F8E4EFCFF8A4" author="Kovacs, Z." box="[800,954,1801,1827]" pageId="87" pageNumber="88" pagination="179 - 182" refId="ref86477" refString="Kovacs, Z. (2018) New records of the genus Euthria (Mollusca, Buccinidae) in the Miocene Paratethys. Foldtani Kozlony, 148 (2), 179 - 182. https: // doi. org / 10.23928 / foldt. kozl. 2018.148.2.179" type="journal article" year="2018">Kovács 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>