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<document id="0170D814F46C8A7F399D5F97E2A28B2F" ID-DOI="10.1649/0010-065X-70.2.375" ID-Zenodo-Dep="10106903" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630179577464" checkinUser="diego" docAuthor="Nunes, Rafael V." docDate="2016" docId="03CE7762952CFFF5FD00432340515411" docLanguage="en" docName="0010-065X-70.2.375.pdf" docOrigin="The Coleopterists Bulletin 70 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-70.2.375" docStyle="DocumentStyle:A0A70CFE30726088A421CCB226FF86C7.4:TheColeopBull.2010-2016.journal_article.1cover" docStyleId="A0A70CFE30726088A421CCB226FF86C7" docStyleName="TheColeopBull.2010-2016.journal_article.1cover" docStyleVersion="4" docTitle="Dichotomius (Selenocopris) aielloae Nunes 2016, new species" docType="treatment" docVersion="3" lastPageNumber="379" masterDocId="FFF70F1A952DFFF1FFD94534422A551D" masterDocTitle="A New Species in the SubgenusSelenocoprisBurmeister, 1846 (Coleoptera: Scarabaeidae: Scarabaeinae: DichotomiusHope) from Panama" masterLastPageNumber="379" masterPageNumber="375" pageNumber="375" updateTime="1699638787551" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9BE4753862F753997B91C724AF708F6A">A New Species in the SubgenusSelenocoprisBurmeister, 1846 (Coleoptera: Scarabaeidae: Scarabaeinae: DichotomiusHope) from Panama</mods:title>
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<mods:name id="6F9FB91743B5CF9072523FD24F441AC0" type="personal">
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<mods:namePart id="961B386C2C2BCC069FD744D7D6618534">Nunes, Rafael V.</mods:namePart>
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<mods:title id="E0A45D3A5EB8F1B986A001154AF8AA03">The Coleopterists Bulletin</mods:title>
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<mods:part id="CF29443106E721DFFC2015FEF0881DE4">
<mods:date id="EE72F8FA1992E24354A57AD4DFFEDBA6">2016</mods:date>
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<mods:number id="0C3368D44DF5C3407DB6C2A9F2214A85">2016-06-30</mods:number>
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<mods:number id="58FD8C6F466AC74FF20B78202F749025">70</mods:number>
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<mods:identifier id="8A2E4598FAD9D2154B086FA42C895258" type="DOI">10.1649/0010-065X-70.2.375</mods:identifier>
<mods:identifier id="ACDEE705AB960C003970EF3AC6E3C6CC" type="Zenodo-Dep">10106903</mods:identifier>
</mods:mods>
<treatment id="03CE7762952CFFF5FD00432340515411" LSID="urn:lsid:plazi:treatment:03CE7762952CFFF5FD00432340515411" httpUri="http://treatment.plazi.org/id/03CE7762952CFFF5FD00432340515411" lastPageId="4" pageId="1">
<subSubSection id="C37D95FF952CFFF0FD00432341DA535A" pageId="1" type="nomenclature">
<paragraph id="8BD8C674952CFFF0FD00432341DA535A" blockId="1.[729,1171,1559,1634]" pageId="1">
<heading id="D0907118952CFFF0FD00432341DA535A" centered="true" fontSize="10" level="2" pageId="1" reason="2">
<emphasis id="B9131A66952CFFF0FD00432341DA535A" bold="true" pageId="1">
<taxonomicName id="4C67BDF7952CFFF0FD00432346A75330" ID-CoL="35N9H" authority="Nunes" authorityName="Nunes" authorityYear="2016" box="[729,1165,1559,1581]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="aielloae" status="sp. nov." subGenus="Selenocopris">
<emphasis id="B9131A66952CFFF0FD004323417C5331" bold="true" box="[729,854,1559,1580]" italics="true" pageId="1">Dichotomius</emphasis>
(
<emphasis id="B9131A66952CFFF0FCB1432341CF5331" bold="true" box="[872,997,1559,1580]" italics="true" pageId="1">Selenocopris</emphasis>
)
<emphasis id="B9131A66952CFFF0FC2E432346695331" bold="true" box="[1015,1091,1559,1580]" italics="true" pageId="1">aielloae</emphasis>
Nunes
</taxonomicName>
,
<taxonomicNameLabel id="A220A71D952CFFF0FCA2430641DA535A" box="[891,1008,1586,1607]" pageId="1" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37D95FF952CFFF0FCA2437941DB537F" box="[891,1009,1612,1634]" pageId="1" type="description">
<paragraph id="8BD8C674952CFFF0FCA2437941DB537F" blockId="1.[729,1171,1559,1634]" box="[891,1009,1612,1634]" pageId="1">
(
<figureCitation id="135CDAF1952CFFF0FC5B437941C0537F" box="[898,1002,1612,1634]" captionStart="Fig" captionStartId="2.[175,207,1638,1657]" captionTargetBox="[172,1152,193,1616]" captionTargetId="figure-1@2.[169,1155,189,1621]" captionTargetPageId="2" captionText="Fig. 1. Dichotomius aielloae. A) Holotype male STOCKWELL STRI-ENT 0046987, dorsal habitus, B) Paratype female STOCKWELL STRI-ENT 0046985, dorsal habitus, C) Same paratype female, ventral habitus, D) Frontal view of male cephalic carina with a central blade-like horn. E) Detail of female sixth sternite with white arrow indicating the central pair of tubercles." figureDoi="http://doi.org/10.5281/zenodo.10106907" httpUri="https://zenodo.org/record/10106907/files/figure.png" pageId="1">Fig. 1AE</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37D95FF952CFFF2FD024343436A545C" lastPageId="3" pageId="1" type="materials_examined">
<paragraph id="8BD8C674952CFFF0FD024343468353DC" blockId="1.[707,1194,1655,1729]" pageId="1">
<emphasis id="B9131A66952CFFF0FD02434341585391" bold="true" box="[731,882,1655,1676]" pageId="1">Type Material.</emphasis>
<materialsCitation id="3B0FCC29952CFFF0FCA64343468353DC" collectionCode="T, R, STOCKWELL, STRI-ENT" collectorName="Pr. Cana &amp; Pirre Trail. El." country="Panama" elevation="1250" location="Darien" pageId="1" specimenCount="1" stateProvince="Darien" typeStatus="holotype">
<typeStatus id="54DC78D6952CFFF0FCA6434341C85391" box="[895,994,1655,1676]" pageId="1" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="F37086E4952CFFF0FBED4343468C5391" box="[1076,1190,1655,1676]" name="Panama" pageId="1">PANAMA</collectingCountry>
:
<collectingRegion id="49A30896952CFFF0FD1A43A6412B53BA" box="[707,769,1682,1703]" country="Panama" name="Darien" pageId="1">Darien</collectingRegion>
<collectorName id="2692A3A2952CFFF0FCD143A6417153BA" box="[776,859,1682,1703]" pageId="1">Pr. Cana</collectorName>
,
<collectorName id="2692A3A2952CFFF0FCBF43A641C653BA" box="[870,1004,1682,1703]" pageId="1">Pirre Trail. El.</collectorName>
<quantity id="4C9F6B91952CFFF0FC2A43A6461253BA" box="[1011,1080,1682,1703]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="1" unit="m" value="1250.0">
<elevation id="004A2147952CFFF0FC2A43A6461253BA" box="[1011,1080,1682,1703]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="1" unit="m" value="1250.0">1250m</elevation>
</quantity>
., 4-7-VI-96. F.I.
<collectionCode id="ED765EB1952CFFF0FD3B439840D953DC" box="[738,755,1708,1729]" name="Tavera, Department of Geology and Geophysics" pageId="1">T</collectionCode>
. A.
<collectionCode id="ED765EB1952CFFF0FCCE4398410053DC" box="[791,810,1708,1729]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1">R</collectionCode>
. Gillogly /
<collectionCode id="ED765EB1952CFFF0FC7B4398468353DC" box="[930,1193,1708,1729]" pageId="1">STOCKWELL STRI-ENT</collectionCode>
</materialsCitation>
</paragraph>
<paragraph id="8BD8C674952CFFF0FD4243D2409153E4" blockId="1.[667,699,1766,1785]" box="[667,699,1766,1785]" pageId="1">375</paragraph>
<caption id="DF1896FC952FFFF3FF76435243A153DC" ID-DOI="http://doi.org/10.5281/zenodo.10106907" ID-Zenodo-Dep="10106907" httpUri="https://zenodo.org/record/10106907/files/figure.png" pageId="2" startId="2.[175,207,1638,1657]" targetBox="[172,1152,193,1616]" targetPageId="2">
<paragraph id="8BD8C674952FFFF3FF76435243A153DC" blockId="2.[151,1172,1638,1729]" pageId="2">
<emphasis id="B9131A66952FFFF3FF76435242C15364" bold="true" box="[175,235,1638,1657]" pageId="2">Fig. 1.</emphasis>
<taxonomicName id="4C67BDF7952FFFF3FED8435243925364" authorityName="Nunes" authorityYear="2016" box="[257,440,1638,1657]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="aielloae">
<emphasis id="B9131A66952FFFF3FED8435243925364" box="[257,440,1638,1657]" italics="true" pageId="2">Dichotomius aielloae</emphasis>
</taxonomicName>
. A) Holotype male STOCKWELL STRI-ENT 0046987, dorsal habitus, B) Paratype female STOCKWELL STRI-ENT 0046985, dorsal habitus, C) Same paratype female, ventral habitus, D) Frontal view of male cephalic carina with a central blade-like horn. E) Detail of female sixth sternite with white arrow indicating the central pair of tubercles.
</paragraph>
</caption>
<paragraph id="8BD8C674952EFFF2FF7445F6436A545C" blockId="3.[173,659,194,375]" pageId="3">
<materialsCitation id="3B0FCC29952EFFF2FF7445F6402E55CA" box="[173,516,194,215]" collectionCode="STRI-ENT" pageId="3" specimenCount="1" typeStatus="paratype">
0046987 [
<collectionCode id="ED765EB1952EFFF2FECB45F643AC55CA" box="[274,390,194,215]" pageId="3">STRI-ENT</collectionCode>
].
<typeStatus id="54DC78D6952EFFF2FE4145F6402E55CA" box="[408,516,194,215]" pageId="3" type="paratype">
<emphasis id="B9131A66952EFFF2FE4145F6402E55CA" bold="true" box="[408,516,194,215]" pageId="3">Paratypes</emphasis>
</typeStatus>
</materialsCitation>
<emphasis id="B9131A66952EFFF2FDDD45F6402255CA" bold="true" box="[516,520,194,215]" pageId="3">:</emphasis>
<materialsCitation id="3B0FCC29952EFFF2FDCB45F643BF5411" collectionCode="STOCKWELL, STRI-ENT" collectorName="Pr. Cana &amp; Pirre Trail. El." country="Panama" elevation="1250" location="Darien" pageId="3" specimenCode="STRI-ENT 0046986" specimenCount="1" specimenCount-male="1" stateProvince="Darien" typeStatus="paratype">
same data as holotype /
<collectionCode id="ED765EB1952EFFF2FEF845E9439D55EF" box="[289,439,221,242]" pageId="3">STOCKWELL</collectionCode>
<specimenCode id="DBC16E0F952EFFF2FE1845E940B955EF" box="[449,659,221,242]" collectionCode="STRI-ENT" pageId="3">STRI-ENT 0046986</specimenCode>
[
<specimenCount id="9D610DFD952EFFF2FF6C45C342D35411" box="[181,249,247,268]" pageId="3" type="male">1 male</specimenCount>
at
<collectionCode id="ED765EB1952EFFF2FEC445C343A55411" box="[285,399,247,268]" pageId="3">STRI-ENT</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B0FCC29952EFFF2FE7B45C34316545C" collectionCode="STOCKWELL, STRI-ENT" collectorName="Pr. Cana &amp; Pirre Trail. El." country="Panama" elevation="1250" location="Darien" pageId="3" specimenCode="STRI-ENT 0046985" specimenCount="1" specimenCount-female="1" stateProvince="Darien" typeStatus="paratype">
same data as holotype /
<collectionCode id="ED765EB1952EFFF2FF744426436F543A" box="[173,325,274,295]" pageId="3">STOCKWELL</collectionCode>
<specimenCode id="DBC16E0F952EFFF2FE964426400E543A" box="[335,548,274,295]" collectionCode="STRI-ENT" pageId="3">STRI-ENT 0046985</specimenCode>
[
<specimenCount id="9D610DFD952EFFF2FDEE442640B9543A" box="[567,659,274,295]" pageId="3" type="female">1 female</specimenCount>
at
<collectionCode id="ED765EB1952EFFF2FF1F4418431D545C" box="[198,311,300,321]" pageId="3">STRI-ENT</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37D95FF952EFFF5FF1C447340515411" lastPageId="4" pageId="3" type="diagnosis">
<paragraph id="8BD8C674952EFFF2FF1C447340B9546A" blockId="3.[173,659,194,375]" pageId="3">
<emphasis id="B9131A66952EFFF2FF1C447343135441" bold="true" box="[197,313,327,348]" pageId="3">Diagnosis.</emphasis>
The angulation of the clypeogenal junction (straight on males, obtuse female)
</paragraph>
<paragraph id="8BD8C674952EFFF2FD1A45F646B8546A" blockId="3.[707,1194,194,375]" pageId="3">
(
<figureCitation id="135CDAF1952EFFF2FD1345F6411E55CA" box="[714,820,194,216]" captionStart="Fig" captionStartId="2.[175,207,1638,1657]" captionTargetBox="[172,1152,193,1616]" captionTargetId="figure-1@2.[169,1155,189,1621]" captionTargetPageId="2" captionText="Fig. 1. Dichotomius aielloae. A) Holotype male STOCKWELL STRI-ENT 0046987, dorsal habitus, B) Paratype female STOCKWELL STRI-ENT 0046985, dorsal habitus, C) Same paratype female, ventral habitus, D) Frontal view of male cephalic carina with a central blade-like horn. E) Detail of female sixth sternite with white arrow indicating the central pair of tubercles." figureDoi="http://doi.org/10.5281/zenodo.10106907" httpUri="https://zenodo.org/record/10106907/files/figure.png" pageId="3">Fig. 1AB</figureCitation>
) and pair of tubercles on the female sixth abdominal sternite (
<figureCitation id="135CDAF1952EFFF2FC6845E941D255EF" box="[945,1016,221,242]" captionStart="Fig" captionStartId="2.[175,207,1638,1657]" captionTargetBox="[172,1152,193,1616]" captionTargetId="figure-1@2.[169,1155,189,1621]" captionTargetPageId="2" captionText="Fig. 1. Dichotomius aielloae. A) Holotype male STOCKWELL STRI-ENT 0046987, dorsal habitus, B) Paratype female STOCKWELL STRI-ENT 0046985, dorsal habitus, C) Same paratype female, ventral habitus, D) Frontal view of male cephalic carina with a central blade-like horn. E) Detail of female sixth sternite with white arrow indicating the central pair of tubercles." figureDoi="http://doi.org/10.5281/zenodo.10106907" httpUri="https://zenodo.org/record/10106907/files/figure.png" pageId="3">Fig. 1E</figureCitation>
) place this species in the
<taxonomicName id="4C67BDF7952EFFF2FCDF45C3463E5411" baseAuthorityName="Felsche" baseAuthorityYear="1901" box="[774,1044,247,268]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="quadraticeps">
<emphasis id="B9131A66952EFFF2FCDF45C3463E5411" box="[774,1044,247,268]" italics="true" pageId="3">Dichotomius quadraticeps</emphasis>
</taxonomicName>
species-group (Nunes and Vaz-de-Mello, unpublished data).
<taxonomicName id="4C67BDF7952EFFF2FD1A441841AD545C" authorityName="Nunes" authorityYear="2016" box="[707,903,300,321]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="aielloae">
<emphasis id="B9131A66952EFFF2FD1A441841AD545C" box="[707,903,300,321]" italics="true" pageId="3">Dichotomius aielloae</emphasis>
</taxonomicName>
is separated from other species in this group by the male cephalic carina, which bears a central blade-like horn (ventral view) (
<figureCitation id="135CDAF1952EFFF2FBE2445646AC546A" box="[1083,1158,354,375]" captionStart="Fig" captionStartId="2.[175,207,1638,1657]" captionTargetBox="[172,1152,193,1616]" captionTargetId="figure-1@2.[169,1155,189,1621]" captionTargetPageId="2" captionText="Fig. 1. Dichotomius aielloae. A) Holotype male STOCKWELL STRI-ENT 0046987, dorsal habitus, B) Paratype female STOCKWELL STRI-ENT 0046985, dorsal habitus, C) Same paratype female, ventral habitus, D) Frontal view of male cephalic carina with a central blade-like horn. E) Detail of female sixth sternite with white arrow indicating the central pair of tubercles." figureDoi="http://doi.org/10.5281/zenodo.10106907" httpUri="https://zenodo.org/record/10106907/files/figure.png" pageId="3">Fig. 1D</figureCitation>
).
</paragraph>
<caption id="DF1896FC952EFFF2FF1C43A2466653DC" ID-DOI="http://doi.org/10.5281/zenodo.10106909" ID-Zenodo-Dep="10106909" httpUri="https://zenodo.org/record/10106909/files/figure.png" pageId="3" startId="3.[197,229,1686,1705]" targetBox="[173,1179,437,1663]" targetPageId="3">
<paragraph id="8BD8C674952EFFF2FF1C43A2466653DC" blockId="3.[173,1194,1686,1729]" pageId="3">
<emphasis id="B9131A66952EFFF2FF1C43A2432B53B4" bold="true" box="[197,257,1686,1705]" pageId="3">Fig. 2.</emphasis>
A) Type locality of
<taxonomicName id="4C67BDF7952EFFF2FE6443A2405A53B4" authorityName="Nunes" authorityYear="2016" box="[445,624,1686,1705]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="aielloae">
<emphasis id="B9131A66952EFFF2FE6443A2405A53B4" box="[445,624,1686,1705]" italics="true" pageId="3">Dichotomius aielloae</emphasis>
</taxonomicName>
(open circle in square) and locality (star) for
<taxonomicName id="4C67BDF7952EFFF2FC2843A2468053B4" box="[1009,1194,1686,1705]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="fonsecae">
<emphasis id="B9131A66952EFFF2FC2843A2468053B4" box="[1009,1194,1686,1705]" italics="true" pageId="3">Dichotomius fonsecae</emphasis>
</taxonomicName>
in Ecuador, B) Satellite image of type locality (open circle) showing the main mountain ranges in the region.
</paragraph>
</caption>
<paragraph id="8BD8C6749529FFF5FF7645F640515411" blockId="4.[151,638,194,269]" pageId="4">
This species can be separated from other Panamanian
<taxonomicName id="4C67BDF79529FFF5FEC345E843C255EF" box="[282,488,220,242]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9131A669529FFF5FEC345E843BC55EC" box="[282,406,220,241]" italics="true" pageId="4">Dichotomius</emphasis>
species
</taxonomicName>
by modifying couplet 6 of Howden and Young s (1981) key:
</paragraph>
</subSubSection>
</treatment>
</document>