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<document id="3B3AF5D0CFDFE489643A644CCC580602" ID-CLB-Dataset="28030" ID-DOI="10.11646/zootaxa.4508.4.4" ID-GBIF-Dataset="8b0798f0-9aea-4774-bad9-c5aa8113af96" ID-ISSN="1175-5326" ID-Zenodo-Dep="2607538" ID-ZooBank="E502D036-A6D6-42A0-9064-B68912443F6B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553586398885" checkinUser="plazi" docAuthor="Mashimo, Yuta, Müller, Patrick, Pohl, Hans &amp; Beutel, Rolf G." docDate="2018" docId="03AA7D65FFF88703FF7AFB81FBA4A5DF" docLanguage="en" docName="zootaxa.4508.4.4.pdf" docOrigin="Zootaxa 4508 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zorotypus (Octozoros) hirsutus Mashimo &amp; Müller &amp; Pohl &amp; Beutel 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="566" masterDocId="FF93051DFFF98707FFEDFFD5FFE5A659" masterDocTitle="The “ hairy beast ” - Zorotypus hirsutus sp. n., an unusual new species of Zoraptera (Insecta) from Burmese amber" masterLastPageNumber="568" masterPageNumber="562" pageNumber="563" updateTime="1698713127992" updateUser="ExternalLinkService">
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<mods:title id="2CA097D607751BEDAF6069599DB39BBC">The “ hairy beast ” - Zorotypus hirsutus sp. n., an unusual new species of Zoraptera (Insecta) from Burmese amber</mods:title>
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<mods:namePart id="A8DF313285E1EB5D215426C57F0456B0">Mashimo, Yuta</mods:namePart>
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<mods:namePart id="34C6B7B6715CC0701E82CB5378A9DAF0">Pohl, Hans</mods:namePart>
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<mods:namePart id="629866F3F1C0F2583763C9B22E76AFE6">Beutel, Rolf G.</mods:namePart>
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<mods:title id="1895E5C6C1EA96BBA0289F496C5D4D1C">Zootaxa</mods:title>
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<treatment id="03AA7D65FFF88703FF7AFB81FBA4A5DF" ID-DOI="http://doi.org/10.5281/zenodo.5974548" ID-GBIF-Taxon="154496061" ID-Zenodo-Dep="5974548" LSID="urn:lsid:plazi:treatment:03AA7D65FFF88703FF7AFB81FBA4A5DF" httpUri="http://treatment.plazi.org/id/03AA7D65FFF88703FF7AFB81FBA4A5DF" lastPageId="4" lastPageNumber="566" pageId="1" pageNumber="563">
<subSubSection id="C3199FF8FFF88706FF7AFB81FD39A236" box="[151,732,1108,1135]" pageId="1" pageNumber="563" type="nomenclature">
<paragraph id="8BBCCC73FFF88706FF7AFB81FD39A236" blockId="1.[151,732,1108,1168]" box="[151,732,1108,1135]" pageId="1" pageNumber="563">
<heading id="D0F47B1FFFF88706FF7AFB81FD39A236" bold="true" box="[151,732,1108,1135]" fontSize="11" level="1" pageId="1" pageNumber="563" reason="1">
<emphasis id="B9771061FFF88706FF7AFB81FD39A236" bold="true" box="[151,732,1108,1135]" pageId="1" pageNumber="563">
<taxonomicName id="4C03B7F0FFF88706FF7AFB81FD68A236" ID-CoL="8NFXZ" authority="Mashimo" authorityName="Mashimo &amp; Müller &amp; Pohl &amp; Beutel" authorityYear="2018" box="[151,653,1108,1135]" class="Insecta" family="Zorotypidae" genus="Zorotypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zoraptera" pageId="1" pageNumber="563" phylum="Arthropoda" rank="subGenus" status="sp. nov." subGenus="Octozoros">
<emphasis id="B9771061FFF88706FF7AFB81FEF4A237" bold="true" box="[151,273,1108,1134]" italics="true" pageId="1" pageNumber="563">Zorotypus</emphasis>
(
<emphasis id="B9771061FFF88706FECFFB81FE7FA237" bold="true" box="[290,410,1108,1134]" italics="true" pageId="1" pageNumber="563">Octozoros</emphasis>
)
<emphasis id="B9771061FFF88706FE41FB81FDEAA237" bold="true" box="[428,527,1108,1134]" italics="true" pageId="1" pageNumber="563">hirsutus</emphasis>
Mashimo
</taxonomicName>
,
<taxonomicNameLabel id="A244AD1AFFF88706FD77FB80FD39A236" box="[666,732,1109,1135]" pageId="1" pageNumber="563" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3199FF8FFF88706FF7AFBA2FEF7A2C9" box="[151,274,1143,1168]" pageId="1" pageNumber="563" type="description">
<paragraph id="8BBCCC73FFF88706FF7AFBA2FEF7A2C9" blockId="1.[151,732,1108,1168]" box="[151,274,1143,1168]" pageId="1" pageNumber="563">
(
<figureCitation id="1338D0F6FFF88706FF72FBA2FF09A2C9" box="[159,236,1143,1168]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="1" pageNumber="563">Figs. 1</figureCitation>
,
<figureCitation id="1338D0F6FFF88706FF17FBADFEECA2C9" box="[250,265,1144,1168]" captionStart="FIGURE 2" captionStartId="3.[151,250,1555,1577]" captionTargetBox="[160,1429,200,1532]" captionTargetId="figure@3.[151,1436,193,1533]" captionTargetPageId="3" captionText="FIGURE 2. Zorotypus hirsutus sp. n.. A: Head in frontal view. Left labial palp behind bubble and barely visible. B: Abdomen in dorsal view. C: Abdomen and metalegs in ventral view. ce, cercus; e, compound eye; ga, galea; lbp, labial palp; la, lacinia; lr, labrum; md, mandible; mxp, maxillary palp; T7, seventh abdominal tergum. Asterisk: lobe-like structure. Scales = 0.2 mm." httpUri="https://zenodo.org/record/2607542/files/figure.png" pageId="1" pageNumber="563">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3199FF8FFF88706FF7AFB15FD97A379" pageId="1" pageNumber="563" type="materials_examined">
<paragraph id="8BBCCC73FFF88706FF7AFB15FD97A379" blockId="1.[151,1437,1215,1997]" pageId="1" pageNumber="563">
<emphasis id="B9771061FFF88706FF7AFB15FEE8A281" bold="true" box="[151,269,1216,1240]" pageId="1" pageNumber="563">
<typeStatus id="54B872D1FFF88706FF7AFB15FEEDA281" box="[151,264,1216,1240]" pageId="1" pageNumber="563" type="holotype">Holotype</typeStatus>
:
</emphasis>
Alate female;
<collectingCountry id="F3148CE3FFF88706FE56FB15FDCFA281" box="[443,554,1216,1240]" name="Myanmar" pageId="1" pageNumber="563">Myanmar</collectingCountry>
,
<collectingRegion id="49C70291FFF88706FDDAFB6AFD6DA281" box="[567,648,1215,1240]" country="Myanmar" name="Kachin" pageId="1" pageNumber="563">Kachin</collectingRegion>
State, Hukawng Valley (
<bibRefCitation id="EF92B182FFF88706FC47FB6AFB9CA281" author="Kania, I. &amp; Wang, B. &amp; Szwedo, J." box="[938,1145,1215,1240]" pageId="1" pageNumber="563" pagination="522 - 530" refId="ref3446" refString="Kania, I., Wang, B. &amp; Szwedo, J. (2015) Dicranoptycha Osten Sacken, 1860 (Diptera, Limoniidae) from the earliest Cenomanian Burmese amber. Cretaceous Research, 52, 522 - 530. https: // doi. org / 10.1016 / j. cretres. 2014.03.002" type="journal article" year="2015">
Kania
<emphasis id="B9771061FFF88706FC15FB14FBCEA281" box="[1016,1067,1216,1240]" italics="true" pageId="1" pageNumber="563">et al</emphasis>
. 2015
</bibRefCitation>
: fig. 1;
<bibRefCitation id="EF92B182FFF88706FB3BFB6AFF31A2A5" author="Jaloszynski, P. &amp; Yamamoto, S. &amp; Takahashi, Y." pageId="1" pageNumber="563" pagination="95 - 102" refId="ref3344" refString="Jaloszynski, P., Yamamoto, S. &amp; Takahashi, Y. (2017) Discovery of a new Mesozoic species of the ancient genus Lepicerus (Coleoptera: Myxophaga: Lepiceridae), with implications for the systematic placement of all previously described extinct ' lepiceroids'. Cretaceous Research, 78, 95 - 102. https: // doi. org / 10.1016 / j. cretres. 2017.06.001" type="journal article" year="2017">
Jałoszyński
<emphasis id="B9771061FFF88706FA8CFB14FA70A281" box="[1377,1429,1216,1240]" italics="true" pageId="1" pageNumber="563">et al</emphasis>
. 2017
</bibRefCitation>
: fig. 1); Albian-Cenomanian boundary, mid Cretaceous. The
<typeStatus id="54B872D1FFF88706FC6DFB31FC07A2A4" box="[896,994,1252,1277]" pageId="1" pageNumber="563" type="holotype">holotype</typeStatus>
is deposited in Patrick Müllers private collection (depository number BUB2785).
</paragraph>
</subSubSection>
<subSubSection id="C3199FF8FFF88706FF2AFAF9FBC0A33E" pageId="1" pageNumber="563" type="etymology">
<paragraph id="8BBCCC73FFF88706FF2AFAF9FBC0A33E" blockId="1.[151,1437,1215,1997]" pageId="1" pageNumber="563">
<emphasis id="B9771061FFF88706FF2AFAF9FEABA31D" bold="true" box="[199,334,1324,1348]" pageId="1" pageNumber="563">Etymology.</emphasis>
The specific name is taken from a Latin adjective
<taxonomicName id="4C03B7F0FFF88706FC64FAF9FC06A31D" authorityName="Mashimo &amp; Müller &amp; Pohl &amp; Beutel" authorityYear="2018" box="[905,995,1324,1348]" class="Insecta" family="Zorotypidae" genus="Zorotypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zoraptera" pageId="1" pageNumber="563" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B9771061FFF88706FC64FAF9FC06A31D" box="[905,995,1324,1348]" italics="true" pageId="1" pageNumber="563">hirsutus</emphasis>
</taxonomicName>
meaning “hairy, shaggy”, and refers to the bristly or hirsute appearance, with a dense vestiture of long and slender setae.
</paragraph>
</subSubSection>
<subSubSection id="C3199FF8FFF88706FF2AFAA6FC72A066" pageId="1" pageNumber="563" type="diagnosis">
<paragraph id="8BBCCC73FFF88706FF2AFAA6FC72A066" blockId="1.[151,1437,1215,1997]" pageId="1" pageNumber="563">
<emphasis id="B9771061FFF88706FF2AFAA6FEA4A3D5" bold="true" box="[199,321,1395,1420]" pageId="1" pageNumber="563">Diagnosis.</emphasis>
<taxonomicName id="4C03B7F0FFF88706FEA6FAA0FDC5A3D5" authority="Mashimo &amp; Müller &amp; Pohl &amp; Beutel, 2018" authorityName="Mashimo &amp; Müller &amp; Pohl &amp; Beutel" authorityYear="2018" box="[331,544,1396,1420]" class="Insecta" family="Zorotypidae" genus="Zorotypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zoraptera" pageId="1" pageNumber="563" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B9771061FFF88706FEA6FAA0FDC5A3D5" box="[331,544,1396,1420]" italics="true" pageId="1" pageNumber="563">Zorotypus hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A244AD1AFFF88706FDC6FAA1FD8BA3D5" box="[555,622,1396,1420]" pageId="1" pageNumber="563" rank="species">
<emphasis id="B9771061FFF88706FDC6FAA1FD8BA3D5" bold="true" box="[555,622,1396,1420]" pageId="1" pageNumber="563">sp. n.</emphasis>
</taxonomicNameLabel>
is tentatively assigned to the subgenus
<taxonomicName id="4C03B7F0FFF88706FBACFAA1FB56A3D2" box="[1089,1203,1396,1419]" class="Insecta" family="Zorotypidae" genus="Zorotypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zoraptera" pageId="1" pageNumber="563" phylum="Arthropoda" rank="subGenus" subGenus="Octozoros">
<emphasis id="B9771061FFF88706FBACFAA1FB56A3D2" box="[1089,1203,1396,1419]" italics="true" pageId="1" pageNumber="563">Octozoros</emphasis>
</taxonomicName>
based on the eightsegmented antenna. This species is readily distinguished from the other species of
<taxonomicName id="4C03B7F0FFF88706FBC3FA4DFB45A3F6" box="[1070,1184,1432,1455]" class="Insecta" family="Zorotypidae" genus="Zorotypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zoraptera" pageId="1" pageNumber="566" phylum="Arthropoda" rank="subGenus" subGenus="Octozoros">
<emphasis id="B9771061FFF88706FBC3FA4DFB45A3F6" box="[1070,1184,1432,1455]" italics="true" pageId="1" pageNumber="563">Octozoros</emphasis>
</taxonomicName>
by the combination of the following characters: strongly developed vestiture of bristles on the entire body; very slender, elongate antennomeres; elongated head with concave genal region; absence of thorn-like protuberances on mesonotal anterolateral corners; absence of jugate setae along posterior margin of forewings; relatively slender tibiae; empodium of meta-pretarsus reduced to a slender hair-like structure.
</paragraph>
</subSubSection>
<subSubSection id="C3199FF8FFF88703FF2AF99EFB71A567" lastPageId="4" lastPageNumber="566" pageId="1" pageNumber="563" type="description">
<paragraph id="8BBCCC73FFF88706FF2AF99EFBBCA195" blockId="1.[151,1437,1215,1997]" pageId="1" pageNumber="563">
<emphasis id="B9771061FFF88706FF2AF99EFEBCA03D" bold="true" box="[199,345,1611,1636]" pageId="1" pageNumber="563">Description.</emphasis>
Alate female. Integument blackish brown except antennomeres VII and VIII (
<figureCitation id="1338D0F6FFF88706FB3DF999FAAEA03D" box="[1232,1355,1612,1637]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="1" pageNumber="563">Fig. 1A, B</figureCitation>
). Head subtriangular, moderately elongated, with concave genal region (
<figureCitation id="1338D0F6FFF88706FC9AF9BAFC3FA0D1" box="[887,986,1647,1672]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="1" pageNumber="563">Figs. 1C</figureCitation>
,
<figureCitation id="1338D0F6FFF88706FC05F9A5FBEBA0D1" box="[1000,1038,1648,1672]" captionStart="FIGURE 2" captionStartId="3.[151,250,1555,1577]" captionTargetBox="[160,1429,200,1532]" captionTargetId="figure@3.[151,1436,193,1533]" captionTargetPageId="3" captionText="FIGURE 2. Zorotypus hirsutus sp. n.. A: Head in frontal view. Left labial palp behind bubble and barely visible. B: Abdomen in dorsal view. C: Abdomen and metalegs in ventral view. ce, cercus; e, compound eye; ga, galea; lbp, labial palp; la, lacinia; lr, labrum; md, mandible; mxp, maxillary palp; T7, seventh abdominal tergum. Asterisk: lobe-like structure. Scales = 0.2 mm." httpUri="https://zenodo.org/record/2607542/files/figure.png" pageId="1" pageNumber="563">2A</figureCitation>
). Surface covered with vestiture of long setae (
<figureCitation id="1338D0F6FFF88706FEF8F941FE93A0F5" box="[277,374,1684,1709]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="1" pageNumber="563">Figs. 1C</figureCitation>
,
<figureCitation id="1338D0F6FFF88706FE6FF941FE4DA0F5" box="[386,424,1684,1708]" captionStart="FIGURE 2" captionStartId="3.[151,250,1555,1577]" captionTargetBox="[160,1429,200,1532]" captionTargetId="figure@3.[151,1436,193,1533]" captionTargetPageId="3" captionText="FIGURE 2. Zorotypus hirsutus sp. n.. A: Head in frontal view. Left labial palp behind bubble and barely visible. B: Abdomen in dorsal view. C: Abdomen and metalegs in ventral view. ce, cercus; e, compound eye; ga, galea; lbp, labial palp; la, lacinia; lr, labrum; md, mandible; mxp, maxillary palp; T7, seventh abdominal tergum. Asterisk: lobe-like structure. Scales = 0.2 mm." httpUri="https://zenodo.org/record/2607542/files/figure.png" pageId="1" pageNumber="563">2A</figureCitation>
). Compound eyes prominent; three ocelli present. Antennae 8-segmented, with vestiture of setae of moderate length (
<figureCitation id="1338D0F6FFF88706FE39F962FDBEA089" box="[468,603,1719,1744]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="1" pageNumber="563">Fig. 1A, B</figureCitation>
); antennomere I elongate, approximately 4.5 times longer than wide; antennomere II relatively short, about one-third as long as antennomere I; antennomere III elongate, approximately twice as long as antennomere II; antennomeres IVVI distinctly elongated, slender, approximately six times as long as wide; antennomeres VII-VIII very slender, approximately four times as long as wide. Maxilla only partly visible; galea with comb of mesally directed setae on apical region (
<figureCitation id="1338D0F6FFF88706FC71F892FC1AA139" box="[924,1023,1863,1888]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="1" pageNumber="563">Figs. 1C</figureCitation>
,
<figureCitation id="1338D0F6FFF88706FBE1F89DFBD7A139" box="[1036,1074,1864,1888]" captionStart="FIGURE 2" captionStartId="3.[151,250,1555,1577]" captionTargetBox="[160,1429,200,1532]" captionTargetId="figure@3.[151,1436,193,1533]" captionTargetPageId="3" captionText="FIGURE 2. Zorotypus hirsutus sp. n.. A: Head in frontal view. Left labial palp behind bubble and barely visible. B: Abdomen in dorsal view. C: Abdomen and metalegs in ventral view. ce, cercus; e, compound eye; ga, galea; lbp, labial palp; la, lacinia; lr, labrum; md, mandible; mxp, maxillary palp; T7, seventh abdominal tergum. Asterisk: lobe-like structure. Scales = 0.2 mm." httpUri="https://zenodo.org/record/2607542/files/figure.png" pageId="1" pageNumber="563">2A</figureCitation>
); slender lacinia with two small mesally directed teeth on distal part (
<figureCitation id="1338D0F6FFF88706FDDFF8B9FD69A1DD" box="[562,652,1900,1925]" captionStart="FIGURE 2" captionStartId="3.[151,250,1555,1577]" captionTargetBox="[160,1429,200,1532]" captionTargetId="figure@3.[151,1436,193,1533]" captionTargetPageId="3" captionText="FIGURE 2. Zorotypus hirsutus sp. n.. A: Head in frontal view. Left labial palp behind bubble and barely visible. B: Abdomen in dorsal view. C: Abdomen and metalegs in ventral view. ce, cercus; e, compound eye; ga, galea; lbp, labial palp; la, lacinia; lr, labrum; md, mandible; mxp, maxillary palp; T7, seventh abdominal tergum. Asterisk: lobe-like structure. Scales = 0.2 mm." httpUri="https://zenodo.org/record/2607542/files/figure.png" pageId="1" pageNumber="563">Fig. 2A</figureCitation>
); maxillary palpus with palpomere I not visible; palpomeres II, III and V distinctly elongated; palpomere IV slightly longer than wide. Labial palpus with palpomere I partly visible; palpomere II slightly elongated and palpomere III distinctly elongated (
<figureCitation id="1338D0F6FFF88706FC55F861FBFCA195" box="[952,1049,1972,1997]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="1" pageNumber="563">Figs. 1C</figureCitation>
,
<figureCitation id="1338D0F6FFF88706FBCBF861FBA9A195" box="[1062,1100,1972,1996]" captionStart="FIGURE 2" captionStartId="3.[151,250,1555,1577]" captionTargetBox="[160,1429,200,1532]" captionTargetId="figure@3.[151,1436,193,1533]" captionTargetPageId="3" captionText="FIGURE 2. Zorotypus hirsutus sp. n.. A: Head in frontal view. Left labial palp behind bubble and barely visible. B: Abdomen in dorsal view. C: Abdomen and metalegs in ventral view. ce, cercus; e, compound eye; ga, galea; lbp, labial palp; la, lacinia; lr, labrum; md, mandible; mxp, maxillary palp; T7, seventh abdominal tergum. Asterisk: lobe-like structure. Scales = 0.2 mm." httpUri="https://zenodo.org/record/2607542/files/figure.png" pageId="1" pageNumber="563">2A</figureCitation>
).
</paragraph>
<caption id="DF7C9CFBFFFB8705FF7AF90BFED3A1B0" httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="2" pageNumber="564" startId="2.[151,250,1758,1780]" targetBox="[164,1425,195,1738]" targetPageId="2">
<paragraph id="8BBCCC73FFFB8705FF7AF90BFED3A1B0" blockId="2.[151,1436,1758,2025]" pageId="2" pageNumber="564">
<emphasis id="B9771061FFFB8705FF7AF90BFEF0A0AA" bold="true" box="[151,277,1758,1780]" pageId="2" pageNumber="564">FIGURE 1.</emphasis>
<taxonomicName id="4C03B7F0FFFB8705FEF0F90AFE38A0AD" authority="Mashimo &amp; Müller &amp; Pohl &amp; Beutel, 2018" authorityName="Mashimo &amp; Müller &amp; Pohl &amp; Beutel" authorityYear="2018" box="[285,477,1758,1780]" class="Insecta" family="Zorotypidae" genus="Zorotypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zoraptera" pageId="2" pageNumber="564" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B9771061FFFB8705FEF0F90AFE38A0AD" box="[285,477,1758,1780]" italics="true" pageId="2" pageNumber="564">Zorotypus hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A244AD1AFFFB8705FE08F90BFDFAA0AA" box="[485,543,1758,1779]" pageId="2" pageNumber="564" rank="species">
<emphasis id="B9771061FFFB8705FE08F90BFDFAA0AA" bold="true" box="[485,543,1758,1779]" pageId="2" pageNumber="564">sp. n.</emphasis>
</taxonomicNameLabel>
, holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the
<emphasis id="B9771061FFFB8705FC76F861FC11A193" box="[923,1012,1972,1994]" italics="true" pageId="2" pageNumber="564">Remarks</emphasis>
). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G.
</paragraph>
</caption>
<caption id="DF7C9CFBFFFA8704FF7AF9C6FA92A03F" httpUri="https://zenodo.org/record/2607542/files/figure.png" pageId="3" pageNumber="565" startId="3.[151,250,1555,1577]" targetBox="[160,1429,200,1532]" targetPageId="3">
<paragraph id="8BBCCC73FFFA8704FF7AF9C6FA92A03F" blockId="3.[151,1436,1555,1638]" pageId="3" pageNumber="565">
<emphasis id="B9771061FFFA8704FF7AF9C6FEF6A071" bold="true" box="[151,275,1555,1577]" pageId="3" pageNumber="565">FIGURE 2.</emphasis>
<taxonomicName id="4C03B7F0FFFA8704FEF4F9C1FE32A070" authority="Mashimo &amp; Müller &amp; Pohl &amp; Beutel, 2018" authorityName="Mashimo &amp; Müller &amp; Pohl &amp; Beutel" authorityYear="2018" box="[281,471,1555,1577]" class="Insecta" family="Zorotypidae" genus="Zorotypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zoraptera" pageId="3" pageNumber="565" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B9771061FFFA8704FEF4F9C1FE32A070" box="[281,471,1555,1577]" italics="true" pageId="3" pageNumber="565">Zorotypus hirsutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A244AD1AFFFA8704FE30F9C6FDF2A071" box="[477,535,1555,1576]" pageId="3" pageNumber="565" rank="species">
<emphasis id="B9771061FFFA8704FE30F9C6FDF2A071" bold="true" box="[477,535,1555,1576]" pageId="3" pageNumber="565">sp. n.</emphasis>
</taxonomicNameLabel>
. A: Head in frontal view. Left labial palp behind bubble and barely visible. B: Abdomen in dorsal view. C: Abdomen and metalegs in ventral view. ce, cercus; e, compound eye; ga, galea; lbp, labial palp; la, lacinia; lr, labrum; md, mandible; mxp, maxillary palp; T7, seventh abdominal tergum. Asterisk: lobe-like structure. Scales = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BBCCC73FFFA8704FF2AF941FD82A139" blockId="3.[151,1436,1683,2032]" pageId="3" pageNumber="565">
Pronotum subrectangular, longer than wide, slightly narrowing anteriorly; with vestiture of setae of moderate length on surface and also densely covered with long setae along margins (
<figureCitation id="1338D0F6FFFA8704FBF8F962FB97A096" box="[1045,1138,1719,1744]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="3" pageNumber="565">Fig. 1D</figureCitation>
). Mesonotum only partly visible; distinctly broader than long, about half as long as pronotum; thorn-like protuberances on the anterolateral corners absent; with vestiture of setae of moderate length on surface and along margins. Metanotum (and part of wings) covered by bubble; distinctly broader than long, slightly shorter than mesonotum, with setae of moderate length along lateral and posterior margin.
</paragraph>
<paragraph id="8BBCCC73FFFA8703FF2AF8B9FCC2A73D" blockId="3.[151,1436,1683,2032]" lastBlockId="4.[151,1437,151,902]" lastPageId="4" lastPageNumber="566" pageId="3" pageNumber="565">
Legs densely covered with setae of moderate length; tarsi 2-segmented. Protibia with bristles arranged as comb along distal ventral half and pair of spurs inserted apically (
<figureCitation id="1338D0F6FFFA8704FCADF85AFC5EA1FE" box="[832,955,1935,1960]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="3" pageNumber="565">Fig. 1C, E</figureCitation>
). Mesotibia with apical pair of spurs (
<figureCitation id="1338D0F6FFFA8704FA82F85AFF52A192" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="3" pageNumber="565">Fig. 1F</figureCitation>
). Metafemur proximally expanded, gently tapering towards apex; eight stout spines (sp18) placed on tubercles along posterior border of ventral surface (
<quantity id="4CFB6196FFFA8704FD84F802FD54A1A9" box="[617,689,2007,2032]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.778" metricValueMin="0.254" pageId="3" pageNumber="565" unit="in" value="4.0" valueMax="7.0" valueMin="1.0">17 in</quantity>
<figureCitation id="1338D0F6FFFA8704FD5AF802FCEFA1B6" box="[695,778,2007,2032]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="3" pageNumber="565">Fig. 1F</figureCitation>
); sp5 and sp7 distinctly elongated; sp1 slightly longer than remaining spines (
<figureCitation id="1338D0F6FFFD8703FE89FF42FE5DA6E9" box="[356,440,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="4" pageNumber="566">Fig. 1E</figureCitation>
); right metafemur with one additional small spine between sp6 and sp7 (6+ in the inset in
<figureCitation id="1338D0F6FFFD8703FF7AFF69FF09A68D" box="[151,236,188,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="4" pageNumber="566">Fig. 1F</figureCitation>
); three stiff bristles inserted at preapical region of anterior border of ventral surface (
<figureCitation id="1338D0F6FFFD8703FB2EFF69FAFCA68D" box="[1219,1305,188,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="4" pageNumber="566">Fig. 1F</figureCitation>
). Metatibia slender, with two stout spines at apical one-third and at apex (a, b in
<figureCitation id="1338D0F6FFFD8703FC5AFF0AFBF4A6A1" box="[951,1041,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="4" pageNumber="566">Fig. 1A</figureCitation>
), additionally with tiny spine near most apical spine (white arrow in
<figureCitation id="1338D0F6FFFD8703FDF0FED1FD79A745" box="[541,668,260,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="4" pageNumber="566">Fig. 1A, G</figureCitation>
). Meta-pretarsus with pair of small pulvilli (black arrowhead in the inset in
<figureCitation id="1338D0F6FFFD8703FF1FFEF2FEAEA719" box="[242,331,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="4" pageNumber="566">Fig. 1G</figureCitation>
); empodium reduced to slender hair-like structure (white arrowhead in the inset in
<figureCitation id="1338D0F6FFFD8703FB11FEF2FAB0A719" box="[1276,1365,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="4" pageNumber="566">Fig. 1G</figureCitation>
). Stiff spine present between metacoxae (black arrow in
<figureCitation id="1338D0F6FFFD8703FD2BFE99FCFCA73D" box="[710,793,332,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="4" pageNumber="566">Fig. 1F</figureCitation>
).
</paragraph>
<paragraph id="8BBCCC73FFFD8703FF2AFEBAFC4CA465" blockId="4.[151,1437,151,902]" pageId="4" pageNumber="566">
Abdominal setae dense (
<figureCitation id="1338D0F6FFFD8703FE32FEBAFDDFA7D1" box="[479,570,367,392]" captionStart="FIGURE 2" captionStartId="3.[151,250,1555,1577]" captionTargetBox="[160,1429,200,1532]" captionTargetId="figure@3.[151,1436,193,1533]" captionTargetPageId="3" captionText="FIGURE 2. Zorotypus hirsutus sp. n.. A: Head in frontal view. Left labial palp behind bubble and barely visible. B: Abdomen in dorsal view. C: Abdomen and metalegs in ventral view. ce, cercus; e, compound eye; ga, galea; lbp, labial palp; la, lacinia; lr, labrum; md, mandible; mxp, maxillary palp; T7, seventh abdominal tergum. Asterisk: lobe-like structure. Scales = 0.2 mm." httpUri="https://zenodo.org/record/2607542/files/figure.png" pageId="4" pageNumber="566">Fig. 2C</figureCitation>
). Abdominal tergum I (T1) with transverse row of setae of moderate length along posterior margin (
<figureCitation id="1338D0F6FFFD8703FE44FE41FDE7A7F5" box="[425,514,404,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1555,1577]" captionTargetBox="[160,1429,200,1532]" captionTargetId="figure@3.[151,1436,193,1533]" captionTargetPageId="3" captionText="FIGURE 2. Zorotypus hirsutus sp. n.. A: Head in frontal view. Left labial palp behind bubble and barely visible. B: Abdomen in dorsal view. C: Abdomen and metalegs in ventral view. ce, cercus; e, compound eye; ga, galea; lbp, labial palp; la, lacinia; lr, labrum; md, mandible; mxp, maxillary palp; T7, seventh abdominal tergum. Asterisk: lobe-like structure. Scales = 0.2 mm." httpUri="https://zenodo.org/record/2607542/files/figure.png" pageId="4" pageNumber="566">Fig. 2B</figureCitation>
); T210 with vestiture of setae of moderate length; T27 with pair of long, erect setae on both sides of posterior region; T89 with two pairs of long, erect setae on both sides of posterior region. Median up-curved projections missing (
<figureCitation id="1338D0F6FFFD8703FD8DFE09FD5EA7AD" box="[608,699,476,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1555,1577]" captionTargetBox="[160,1429,200,1532]" captionTargetId="figure@3.[151,1436,193,1533]" captionTargetPageId="3" captionText="FIGURE 2. Zorotypus hirsutus sp. n.. A: Head in frontal view. Left labial palp behind bubble and barely visible. B: Abdomen in dorsal view. C: Abdomen and metalegs in ventral view. ce, cercus; e, compound eye; ga, galea; lbp, labial palp; la, lacinia; lr, labrum; md, mandible; mxp, maxillary palp; T7, seventh abdominal tergum. Asterisk: lobe-like structure. Scales = 0.2 mm." httpUri="https://zenodo.org/record/2607542/files/figure.png" pageId="4" pageNumber="566">Fig. 2C</figureCitation>
). Cercus unsegmented, conical, with four or five long subapical setae almost as long as cercus, proximally with moderate to long, fine setae (
<figureCitation id="1338D0F6FFFD8703FC03FE2AFB94A44E" box="[1006,1137,511,536]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="4" pageNumber="566">Figs. 1A, B</figureCitation>
,
<figureCitation id="1338D0F6FFFD8703FB90FDD5FB24A441" box="[1149,1217,512,536]" captionStart="FIGURE 2" captionStartId="3.[151,250,1555,1577]" captionTargetBox="[160,1429,200,1532]" captionTargetId="figure@3.[151,1436,193,1533]" captionTargetPageId="3" captionText="FIGURE 2. Zorotypus hirsutus sp. n.. A: Head in frontal view. Left labial palp behind bubble and barely visible. B: Abdomen in dorsal view. C: Abdomen and metalegs in ventral view. ce, cercus; e, compound eye; ga, galea; lbp, labial palp; la, lacinia; lr, labrum; md, mandible; mxp, maxillary palp; T7, seventh abdominal tergum. Asterisk: lobe-like structure. Scales = 0.2 mm." httpUri="https://zenodo.org/record/2607542/files/figure.png" pageId="4" pageNumber="566">2B, C</figureCitation>
). Abdominal sterna IIII (S13) not visible; S48 with setae of moderate length (
<figureCitation id="1338D0F6FFFD8703FCA9FDF1FC79A465" box="[836,924,548,573]" captionStart="FIGURE 2" captionStartId="3.[151,250,1555,1577]" captionTargetBox="[160,1429,200,1532]" captionTargetId="figure@3.[151,1436,193,1533]" captionTargetPageId="3" captionText="FIGURE 2. Zorotypus hirsutus sp. n.. A: Head in frontal view. Left labial palp behind bubble and barely visible. B: Abdomen in dorsal view. C: Abdomen and metalegs in ventral view. ce, cercus; e, compound eye; ga, galea; lbp, labial palp; la, lacinia; lr, labrum; md, mandible; mxp, maxillary palp; T7, seventh abdominal tergum. Asterisk: lobe-like structure. Scales = 0.2 mm." httpUri="https://zenodo.org/record/2607542/files/figure.png" pageId="4" pageNumber="566">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="8BBCCC73FFFD8703FF2AFD92FB71A567" blockId="4.[151,1437,151,902]" pageId="4" pageNumber="566">
Wing venation (
<figureCitation id="1338D0F6FFFD8703FE91FD92FE11A439" box="[380,500,583,608]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="4" pageNumber="566">Fig. 1A, B</figureCitation>
) visible as faint, fuscous lines; membrane hyaline except for brownish pterostigma of forewing, covered with minute setae; both fore and hind wings with dense fringes of short setae, slightly longer than those of membrane; posterior margin of forewings lacking stiff, jugate setae. Forewing (
<figureCitation id="1338D0F6FFFD8703FB55FD5AFAEAA4F1" box="[1208,1295,655,680]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="4" pageNumber="566">Fig. 1A</figureCitation>
) R reaching pterostigma base, evanescent distally; Rs separating from radial stem near midpoint of wing, connected with M by short rs-m cross vein; M reaching posterior wing margin, slightly proximal to termination of Rs; CuA
<subScript id="1787CE36FFFD8703FAF0FD33FAC0A4AD" attach="left" box="[1309,1317,742,756]" fontSize="6" pageId="4" pageNumber="566">1</subScript>
extending over third-fifths of wing, terminating on posterior margin; CuA
<subScript id="1787CE36FFFD8703FCB3FCD8FC83A542" attach="left" box="[862,870,781,795]" fontSize="6" pageId="4" pageNumber="566">2</subScript>
present as a very short stub in basal third of wing. Hind wing with R+M furcated near apex, both R and M reaching wing margins; Cu absent.
</paragraph>
</subSubSection>
<subSubSection id="C3199FF8FFFD8703FF2AFC9CFBA4A5DF" pageId="4" pageNumber="566" type="discussion">
<paragraph id="8BBCCC73FFFD8703FF2AFC9CFBA4A5DF" blockId="4.[151,1437,151,902]" pageId="4" pageNumber="566">
<emphasis id="B9771061FFFD8703FF2AFC9CFEDEA538" bold="true" box="[199,315,841,865]" pageId="4" pageNumber="566">Remarks.</emphasis>
A lobe-like structure is partly visible lateral to the left galea (asterisk in
<figureCitation id="1338D0F6FFFD8703FB8EFC9CFB26A53B" box="[1123,1219,841,866]" captionStart="FIGURE 1" captionStartId="2.[151,250,1758,1780]" captionTargetBox="[164,1425,195,1738]" captionTargetId="figure@2.[164,1425,195,1738]" captionTargetPageId="2" captionText="FIGURE 1. Zorotypus hirsutus sp. n., holotype (BUB2785). A, B: Habitus in ventral (A) and dorsal (B) views. C: Head in frontal view. D: Thorax in dorsal view. E: Left foreleg. F: Abdomen and right legs in ventral view; inset: an enlargement of right metafemur with an additional spine. G: Right metatarsus; inset: enlargement of right meta-pretarsus apex. 18, metafemoral spines 18; 6+, the additional metafemoral spine; a, b, metatibial spines a, b; a18, antennomeres IVIII; bac, bristles arranged as comb; ce, cercus; cx1, procoxa; e, compound eye; fe1, profemur; ga, galea; lbp, labial palp; lr, labrum; mxp, maxillary palp; oc, ocelli; S8, eighth abdominal sternum; sp, spurs; th1, prothorax; ti1, protibia. Black and white arrows show the distinct spine between metacoxae and tiny metatibial spine, respectively. Black and white arrowheads show pulvilli and empodium, respectively. Asterisk shows the lobe-like structure (see the Remarks). Scales = 0.5 mm in AB; 0.2 mm in C F; 0.1 mm in G." httpUri="https://zenodo.org/record/2607540/files/figure.png" pageId="4" pageNumber="566">Figs. 1C</figureCitation>
,
<figureCitation id="1338D0F6FFFD8703FB3DFC9FFB13A53B" box="[1232,1270,842,866]" captionStart="FIGURE 2" captionStartId="3.[151,250,1555,1577]" captionTargetBox="[160,1429,200,1532]" captionTargetId="figure@3.[151,1436,193,1533]" captionTargetPageId="3" captionText="FIGURE 2. Zorotypus hirsutus sp. n.. A: Head in frontal view. Left labial palp behind bubble and barely visible. B: Abdomen in dorsal view. C: Abdomen and metalegs in ventral view. ce, cercus; e, compound eye; ga, galea; lbp, labial palp; la, lacinia; lr, labrum; md, mandible; mxp, maxillary palp; T7, seventh abdominal tergum. Asterisk: lobe-like structure. Scales = 0.2 mm." httpUri="https://zenodo.org/record/2607542/files/figure.png" pageId="4" pageNumber="566">2A</figureCitation>
), but could not be unambiguously identified; corresponding part on right side concealed by bubble.
</paragraph>
</subSubSection>
</treatment>
</document>