316 lines
45 KiB
XML
316 lines
45 KiB
XML
<document id="B2532486BB8023B220A1F86B5576CC0B" ID-DOI="10.11646/zootaxa.5514.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="13849826" ID-ZooBank="D745608E-E51C-401E-A1A1-3573A24C51BD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727441634877" checkinUser="plazi" docAuthor="Ramos-Pastrana, Yardany, Merchan-Vargas, Geraldine Lucia & Soares, Matheus M. M." docDate="2024" docId="03A71175FFC83C68CB96689183D4FD91" docLanguage="en" docName="zootaxa.5514.3.3.pdf" docOrigin="Zootaxa 5514 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5514.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systenus bickeli Ramos-Pastrana, Merchan-Vargas & Soares, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="261" masterDocId="FF9E690DFFCA3C6CCB01695C832BFF88" masterDocTitle="First record of the genus Systenus Loew (Diptera: Dolichopodidae) from Colombia, with description of four new species" masterLastPageNumber="272" masterPageNumber="257" pageNumber="259" updateTime="1727443446106" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="5A6885DCDADEB022976B89DEBD9BFE32">
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<mods:title id="2481B7C24332C08C87F730546CD533A8">First record of the genus Systenus Loew (Diptera: Dolichopodidae) from Colombia, with description of four new species</mods:title>
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<mods:name id="F47523C64E66A1FCF0296AD26B769C9B" type="personal">
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<mods:roleTerm id="49FAEF5C85475F96BC163605B0318F2B">Author</mods:roleTerm>
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<mods:namePart id="0E2DC16F006D75D08EDB4151CFD8C95C">Ramos-Pastrana, Yardany</mods:namePart>
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<mods:affiliation id="8D1AAF7730FCB76570154EC66D818127">Universidad de la Amazonia, Grupo de Investigación en Entomología Universidad de la Amazonia - GIEUA-, Laboratorio de Entomología - LEUA-, Av. 11 5 - 69 Juan XXIII, Florencia, Caquetá, Colombia</mods:affiliation>
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<mods:namePart id="20E92FF797D238AC7DD7BD938A86120F">Merchan-Vargas, Geraldine Lucia</mods:namePart>
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<mods:affiliation id="5FF02990FBF90688AC4591BD61B2AD98">Universidad de la Amazonia, Grupo de Investigación en Entomología Universidad de la Amazonia - GIEUA-, Laboratorio de Entomología - LEUA-, Av. 11 5 - 69 Juan XXIII, Florencia, Caquetá, Colombia</mods:affiliation>
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<mods:roleTerm id="417CF6B6E54EB97191816D6C156DD25F">Author</mods:roleTerm>
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<mods:namePart id="67C8A749E8F32CA392A76FC478FC0029">Soares, Matheus M. M.</mods:namePart>
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<mods:affiliation id="094AE6BBD6A55661FCB5FF1DACEAE9FE">Museu de Zoologia da Universidade de São Paulo, Laboratório de Diptera, Av. Nazaré, 481, 04263 - 000 - Ipiranga, São Paulo, São Paulo, Brazil</mods:affiliation>
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<mods:typeOfResource id="136580302CE2EA687EF0D6444ADBBE96">text</mods:typeOfResource>
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<mods:title id="5E8F05430E872FCB5D27622C4918559B">Zootaxa</mods:title>
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<mods:part id="0728116B6E005A0083061B2131730A35">
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<mods:date id="5BFD8581FEC4203C4EA390DA40F4625A">2024</mods:date>
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<mods:number id="28879FC0E165835F7BD2187189DC26B3">2024-09-27</mods:number>
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<mods:number id="A73FBC7A45E1E635AEDDB1B8AD9D265D">5514</mods:number>
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<mods:number id="660306B0F0C945445E79F70BF6C7F864">3</mods:number>
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<mods:url id="24D237DE8EF78C0C2982F94B0F4A221C">http://dx.doi.org/10.11646/zootaxa.5514.3.3</mods:url>
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<mods:classification id="FF99172C77424464E1A957E5A7067336">journal article</mods:classification>
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<mods:identifier id="D0CCED55193C5F0B6199E5E9CD6EE084" type="DOI">10.11646/zootaxa.5514.3.3</mods:identifier>
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<mods:identifier id="48360A49EAEF2119FFF2AA07AA9D4FAC" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A71175FFC83C68CB96689183D4FD91" LSID="urn:lsid:plazi:treatment:03A71175FFC83C68CB96689183D4FD91" httpUri="http://treatment.plazi.org/id/03A71175FFC83C68CB96689183D4FD91" lastPageId="4" lastPageNumber="261" pageId="2" pageNumber="259">
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<subSubSection id="C314F3E8FFC83C6ECB9668918296FE6F" box="[151,445,461,488]" pageId="2" pageNumber="259" type="nomenclature">
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<paragraph id="8BB1A063FFC83C6ECB9668918296FE6F" blockId="2.[151,445,461,523]" box="[151,445,461,488]" pageId="2" pageNumber="259">
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<heading id="D0F9170FFFC83C6ECB9668918296FE6F" bold="true" box="[151,445,461,488]" fontSize="11" level="1" pageId="2" pageNumber="259" reason="1">
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<emphasis id="B97A7C71FFC83C6ECB9668918296FE6F" bold="true" box="[151,445,461,488]" pageId="2" pageNumber="259">
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<taxonomicName id="4C0EDBE0FFC83C6ECB9668918273FE60" authorityName="Ramos-Pastrana & Merchan-Vargas & Soares" authorityYear="2024" box="[151,344,461,488]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="bickeli" status="sp. nov.">
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<emphasis id="B97A7C71FFC83C6ECB9668918273FE60" bold="true" box="[151,344,461,488]" italics="true" pageId="2" pageNumber="259">Systenus bickeli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A249C10AFFC83C6ECA6168918296FE6F" box="[352,445,461,487]" pageId="2" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C314F3E8FFC83C6ECB9668AD8220FD83" box="[151,267,497,523]" pageId="2" pageNumber="259" type="description">
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<paragraph id="8BB1A063FFC83C6ECB9668AD8220FD83" blockId="2.[151,445,461,523]" box="[151,267,497,523]" pageId="2" pageNumber="259">
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(
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<figureCitation id="1335BCE6FFC83C6ECB9E68AD83CDFD83" box="[159,230,497,523]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="2" pageNumber="259">Figs 1</figureCitation>
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,
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<figureCitation id="1335BCE6FFC83C6ECBF568AD8228FD83" box="[244,259,497,523]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[301,1286,1429,1917]" captionTargetId="figure-56@13.[299,1288,1427,1919]" captionTargetPageId="13" captionText="FIGURE 7. Geographical records of Systenus Loew in Colombia. S. beatae Naglis, 2000; S. bickeli sp. nov.; S. colombiensis sp. nov.; S. flaviatus Naglis, 2000; S. fuscus sp. nov.; S. grossus sp. nov.; S. maculipennis Bickel, 2015; S. naranjensis Bickel, 2015 and S. parkeri Bickel, 2015." figureDoi="http://doi.org/10.5281/zenodo.13849845" httpUri="https://zenodo.org/record/13849845/files/figure.png" pageId="2" pageNumber="259">7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C314F3E8FFC83C6ECB966B6583EEFCA3" pageId="2" pageNumber="259" type="diagnosis">
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<paragraph id="8BB1A063FFC83C6ECB966B6583EEFCA3" blockId="2.[151,1437,569,2036]" pageId="2" pageNumber="259">
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<emphasis id="B97A7C71FFC83C6ECB966B658239FDDB" bold="true" box="[151,274,569,595]" pageId="2" pageNumber="259">Diagnosis.</emphasis>
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Antenna dark brown, except ventral surface of scape, pedicel, and basoventral subtriangular area of postpedicel orange-yellow; postpedicel abruptly narrowing at apical 1/2, 4X longer than high as highest point (
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<figureCitation id="1335BCE6FFC83C6ECE6E6B0183F6FD13" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="2" pageNumber="259">Fig. 1A, B</figureCitation>
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). Metepimeron grayish brown (
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<figureCitation id="1335BCE6FFC83C6EC9426BDD81B6FD13" box="[579,669,641,667]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="2" pageNumber="259">Fig. 1A</figureCitation>
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). Legs yellow, except apical 1/4 of coxae II and III, tarsus II and III from apex of basitarsus dark brown (
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<figureCitation id="1335BCE6FFC83C6EC9306BF981A1FD37" box="[561,650,677,703]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="2" pageNumber="259">Fig. 1A</figureCitation>
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). Wing light brown (
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<figureCitation id="1335BCE6FFC83C6EC8736BF980E0FD37" box="[882,971,677,703]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="2" pageNumber="259">Fig. 1D</figureCitation>
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). Surstylus wide at basal 1/3 and abruptly narrowing at apical 2/3, slightly curved ventrally at apex, with 1 long and acute basodorsal digitiform process, about as long as the width of surstylus at base (
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<figureCitation id="1335BCE6FFC83C6EC96A6BB181EBFC8F" box="[619,704,749,775]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="2" pageNumber="259">Fig. 1F</figureCitation>
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). Cercus long, digitiform, about 1.5X longer than surstylus (
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<figureCitation id="1335BCE6FFC83C6ECE6E6BB1839CFCA3" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="2" pageNumber="259">Fig. 1F</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C314F3E8FFC83C6FCBC66A698289F82D" lastPageId="3" lastPageNumber="260" pageId="2" pageNumber="259" type="description">
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<paragraph id="8BB1A063FFC83C6FCBC66A698192FF5D" blockId="2.[151,1437,569,2036]" lastBlockId="3.[151,1437,150,213]" lastPageId="3" lastPageNumber="260" pageId="2" pageNumber="259">
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<emphasis id="B97A7C71FFC83C6ECBC66A6982B4FCC7" bold="true" box="[199,415,821,847]" pageId="2" pageNumber="259">Description. Male</emphasis>
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(
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<figureCitation id="1335BCE6FFC83C6ECAB16A6982D8FCC7" box="[432,499,821,847]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="2" pageNumber="259">Fig. 1</figureCitation>
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). Body length:
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<quantity id="4CF60D86FFC83C6EC9AB6A69802EFCD8" box="[682,773,821,848]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="259" unit="mm" value="3.2">3.2 mm</quantity>
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; wing
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<quantity id="4CF60D86FFC83C6EC8556A698084FCD8" box="[852,943,821,848]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="259" unit="mm" value="2.8">2.8 mm</quantity>
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.
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<emphasis id="B97A7C71FFC83C6EC8BD6A6980D7FCC7" bold="true" box="[956,1020,821,847]" pageId="2" pageNumber="259">Head</emphasis>
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(
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<figureCitation id="1335BCE6FFC83C6ECF0C6A6987A0FCC7" box="[1037,1163,821,847]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="2" pageNumber="259">Fig. 1A, B</figureCitation>
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). Face narrowed below, narrower than ocellar tubercle; clypeus short, rectangular, ending above lower eye margin; face, clypeus and frons metallic green and covered by silvery pruinosity. Palpus pale brown with short black setae at apical 1/2 and 1 strong, black apical seta (shorter than palpus). Proboscis brown. Antenna dark brown, except ventral surface of scape, pedicel and basoventral subtriangular area of postpedicel; postpedicel abruptly narrowed at apical 1/2, 4X longer than high at widest point; arista-like stylus broken off. Ocellar setae lost, 1 pair of tiny postocellar setae; vertical setae lost; 1 pair of short paravertical setae; one row of postocular, uppermost 6 black, remaining setae pale brown; postcranium metallic green and covered by silvery pruinosity.
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<emphasis id="B97A7C71FFC83C6EC8646D6D8095FBC3" bold="true" box="[869,958,1073,1099]" pageId="2" pageNumber="259">Thorax</emphasis>
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(
|
||
<figureCitation id="1335BCE6FFC83C6EC8CE6D6D8764FBC3" box="[975,1103,1073,1099]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="2" pageNumber="259">Fig. 1A, C</figureCitation>
|
||
). Mostly metallic light green with weak bluish reflections and coppery acrostichal stripe ending close to scutellum, thorax wholly covered by silvery pruinosity. Pleura grayish brown, with weak greenish and bluish reflections. Metepimeron grayish brown.
|
||
<emphasis id="B97A7C71FFC83C6ECB966DC1823CFB3F" box="[151,279,1181,1207]" italics="true" pageId="2" pageNumber="259">Chaetotaxy</emphasis>
|
||
. Pronotum with row of strong, yellow setae; lower surface of proepisternum with 1 light brown strong seta and upper surface with 1 thin, light brown seta; 2 regular rows of acrostichals setae ending between fourth and fifth posterior dorsocentrals; 6 strong dorsocentrals; 1 pre-, 1 sutural and 1 postsutural intra-alars; 1 pre-, and 2 postsutural supra-alars; 1 strong and 1 tiny postpronotal; 2 notopleurals; 1 postalar; 1 pair of strong medial and 1 pair of short lateral scutellar setae.
|
||
<emphasis id="B97A7C71FFC83C6EC9456C718157FACF" bold="true" box="[580,636,1325,1351]" pageId="2" pageNumber="259">Legs</emphasis>
|
||
(
|
||
<figureCitation id="1335BCE6FFC83C6EC98C6C7181C2FACF" box="[653,745,1325,1351]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="2" pageNumber="259">Fig. 1A</figureCitation>
|
||
). Yellow, except apical 1/4 of coxae II and III, tarsus II and III from apex of basitarsus dark brown.
|
||
<emphasis id="B97A7C71FFC83C6EC9666C0D8184FAE3" bold="true" box="[615,687,1361,1387]" pageId="2" pageNumber="259">Leg I.</emphasis>
|
||
Podomere ratios: 80, 78, 37/25/17/?/? (It
|
||
<subScript id="178AA226FFC83C6ECF8D6C3F878FFAF9" attach="left" box="[1164,1188,1379,1393]" fontSize="6" pageId="2" pageNumber="259">4–5</subScript>
|
||
broken off). Anterior surface of coxa I covered by short vestiture of yellow setae, 4 long yellow setae at apical edge. Femur I clothed by short vestiture of brown setae, except ventral surface mostly bare. It
|
||
<subScript id="178AA226FFC83C6EC8CD6CF780CFFA31" attach="left" box="[972,996,1451,1465]" fontSize="6" pageId="2" pageNumber="259">4–5</subScript>
|
||
lost.
|
||
<emphasis id="B97A7C71FFC83C6ECF276CC58752FA3B" bold="true" box="[1062,1145,1433,1459]" pageId="2" pageNumber="259">Leg II.</emphasis>
|
||
Podomere ratios: 90, 91, 50/45/30/20/10. Anterior surface of coxa II covered by short vestiture of yellow setae, longer at apical edge. Femur II mostly clothed by short vestiture of black setae, except ventral surface with pale setae. Tibia II with 1 dorsal and 1 posterior seta at basal 1/4, and crown of setae at apex: 1 anteroventral, 1 ventral, 1 posterodorsal and 1 dorsal seta.
|
||
<emphasis id="B97A7C71FFC83C6ECBD56F758211F9CB" bold="true" box="[212,314,1577,1603]" pageId="2" pageNumber="259">Leg. III.</emphasis>
|
||
Podomere ratios: 98, 110, 25/55/33/23/12. Lateral surface of coxa III with 1 strong, yellow seta near middle. Femur III clothed by short vestiture of brown setae, except ventral surface mostly bare. Tibia III with dorsal row of slightly longer, thicker setae, starting at basal 1/4 and ending before near apex.
|
||
<emphasis id="B97A7C71FFC83C6ECFA26F2D87CFF903" bold="true" box="[1187,1252,1649,1675]" pageId="2" pageNumber="259">Wing</emphasis>
|
||
(
|
||
<figureCitation id="1335BCE6FFC83C6ECFF26F2D8660F903" box="[1267,1355,1649,1675]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="2" pageNumber="259">Fig. 1D</figureCitation>
|
||
). Light brown, veins brown. Costal ending at M
|
||
<subScript id="178AA226FFC83C6EC9606FF48142F93E" attach="left" box="[609,617,1704,1718]" fontSize="6" pageId="2" pageNumber="259">1</subScript>
|
||
; R
|
||
<subScript id="178AA226FFC83C6EC98D6FF481BFF93E" attach="left" box="[652,660,1704,1718]" fontSize="6" pageId="2" pageNumber="259">1</subScript>
|
||
ending at basal 3.5/8; dm ending at basal 5.5/8; R
|
||
<subScript id="178AA226FFC83C6ECFD06FF487C1F93E" attach="left" box="[1233,1258,1704,1718]" fontSize="6" pageId="2" pageNumber="259">2+3</subScript>
|
||
nearly straight; R
|
||
<subScript id="178AA226FFC83C6ECBAB6F9783EFF951" attach="left" box="[170,196,1739,1753]" fontSize="6" pageId="2" pageNumber="259">4+5</subScript>
|
||
and M
|
||
<subScript id="178AA226FFC83C6ECA136F908231F952" attach="left" box="[274,282,1740,1754]" fontSize="6" pageId="2" pageNumber="259">1</subScript>
|
||
slightly bowed at apical 6.5/8, ending nearly parallel sided at apex; CuA+CuP as indistinct fold, length of crossvein dm-m/last part of M
|
||
<subScript id="178AA226FFC83C6EC9096FAC813BF976" attach="left" box="[520,528,1776,1790]" fontSize="6" pageId="2" pageNumber="259">4</subScript>
|
||
(“CuAx ratio”): 0.7. Halter whitish yellow.
|
||
<emphasis id="B97A7C71FFC83C6EC8FC6F818759F97F" bold="true" box="[1021,1138,1757,1783]" pageId="2" pageNumber="259">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="1335BCE6FFC83C6ECF806F8187FDF97F" box="[1153,1238,1757,1783]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="2" pageNumber="259">Fig. 1E</figureCitation>
|
||
). Mostly metallic green, covered by short vestiture of black setae, tergite 1 with long setae at posterior margin, tergites 2–5 with dark brown dorsal band, tergites 6–7 entirely dark brown, tergite 7 long and setose, forming long peduncle with sternite 7, sternite 8 brown.
|
||
<emphasis id="B97A7C71FFC83C6ECA816E158137F8EB" bold="true" box="[384,540,1865,1891]" pageId="2" pageNumber="259">Hypopygium</emphasis>
|
||
(
|
||
<figureCitation id="1335BCE6FFC83C6EC92F6E1581AFF8EB" box="[558,644,1865,1891]" captionStart="FIGURE 1" captionStartId="3.[151,250,1689,1713]" captionTargetBox="[162,1426,238,1658]" captionTargetId="figure-44@3.[158,1430,234,1662]" captionTargetPageId="3" captionText="FIGURE 1. Systenus bickeli sp. nov., holotype male (before dissection). A. Habitus, lateral view; B. Head, lateral view; C. Thorax, dorsal view; D. Right wing; E. Abdomen, dorsal view; F. Hypopygium, left lateral view. Abbreviations: CuA+CuP = anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M 1 = first branch of media; M 4 = fourth branch of media; R 1 = anterior branch of radius; R 2+3 = second branch of radius; R = third 4+5 branch of radius." figureDoi="http://doi.org/10.5281/zenodo.13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="2" pageNumber="259">Fig. 1F</figureCitation>
|
||
). Surstylus about as long as epandrial capsule, wide at basal 1/3 and abruptly narrowed at apical 2/3, slightly curved ventrally at apex, 1 long, acute basodorsal digitiform process, about as long as width of surstylus at base, 2 short and 1 long setae at basal 1/3 of ventral edge, a few short setae at dorsal and ventral edges before apex, and 1 ribbon-like seta at apex. Epandrial lobe as wide as surstylus at widest point, gradually narrowed at apex, nearly straight, only slightly curved ventrally at apex, with 1 short basal and 1 long apical seta. Cercus yellow, long and narrow, about 1.5X longer than surstylus and epandrial capsule, with rounded swollen base, and covered by short yellow setae.
|
||
</paragraph>
|
||
<caption id="DF71F0EBFFC93C6FCB966FC58268F8C9" ID-DOI="http://doi.org/10.5281/zenodo.13849830" ID-Zenodo-Dep="13849830" httpUri="https://zenodo.org/record/13849830/files/figure.png" pageId="3" pageNumber="260" startId="3.[151,250,1689,1713]" targetBox="[162,1426,238,1658]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BB1A063FFC93C6FCB966FC586B7F939" blockId="3.[151,1436,1689,1857]" box="[151,1436,1689,1713]" pageId="3" pageNumber="260">
|
||
<emphasis id="B97A7C71FFC93C6FCB966FC5823EF939" bold="true" box="[151,277,1689,1713]" pageId="3" pageNumber="260">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C0EDBE0FFC93C6FCA1C6FC582EAF939" authorityName="Ramos-Pastrana & Merchan-Vargas & Soares" authorityYear="2024" box="[285,449,1689,1713]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="260" phylum="Arthropoda" rank="species" species="bickeli" status="sp. nov.">
|
||
<emphasis id="B97A7C71FFC93C6FCA1C6FC582EAF939" box="[285,449,1689,1713]" italics="true" pageId="3" pageNumber="260">Systenus bickeli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97A7C71FFC93C6FCAC96FC58130F939" bold="true" box="[456,539,1689,1713]" pageId="3" pageNumber="260">
|
||
<taxonomicNameLabel id="A249C10AFFC93C6FCAC96FC58130F939" box="[456,539,1689,1713]" pageId="3" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (before dissection).
|
||
<emphasis id="B97A7C71FFC93C6FC8936FC58080F939" bold="true" box="[914,939,1689,1713]" pageId="3" pageNumber="260">A.</emphasis>
|
||
Habitus, lateral view;
|
||
<emphasis id="B97A7C71FFC93C6FCF996FC58784F939" bold="true" box="[1176,1199,1689,1713]" pageId="3" pageNumber="260">B.</emphasis>
|
||
Head, lateral view;
|
||
<emphasis id="B97A7C71FFC93C6FCE856FC586B7F939" bold="true" box="[1412,1436,1689,1713]" pageId="3" pageNumber="260">C.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB1A063FFC93C6FCB966FE186B7F95C" blockId="3.[151,1436,1689,1857]" box="[151,1436,1725,1749]" pageId="3" pageNumber="260">
|
||
Thorax, dorsal view;
|
||
<emphasis id="B97A7C71FFC93C6FCA726FE182A0F95D" bold="true" box="[371,395,1725,1749]" pageId="3" pageNumber="260">D.</emphasis>
|
||
Right wing;
|
||
<emphasis id="B97A7C71FFC93C6FC9126FE18101F95D" bold="true" box="[531,554,1725,1749]" pageId="3" pageNumber="260">E.</emphasis>
|
||
Abdomen, dorsal view;
|
||
<emphasis id="B97A7C71FFC93C6FC8266FE18011F95D" bold="true" box="[807,826,1725,1749]" pageId="3" pageNumber="260">F.</emphasis>
|
||
Hypopygium, left lateral view. Abbreviations: CuA+CuP =
|
||
</paragraph>
|
||
<paragraph id="8BB1A063FFC93C6FCB966FBD8268F8C9" blockId="3.[151,1436,1689,1857]" pageId="3" pageNumber="260">
|
||
anterior branch of cubital vein + posterior branch of cubital vein; dm = discal medial cell; dm-m = discal medial crossvein; M
|
||
<subScript id="178AA226FFC93C6FCE946FAD86B7F977" attach="left" box="[1429,1436,1777,1791]" fontSize="6" pageId="3" pageNumber="260">1</subScript>
|
||
= first branch of media; M
|
||
<subScript id="178AA226FFC93C6FCAA06E498283F8AB" attach="left" box="[417,424,1813,1827]" fontSize="6" pageId="3" pageNumber="260">4</subScript>
|
||
= fourth branch of media; R
|
||
<subScript id="178AA226FFC93C6FC9C66E4981E5F8AB" attach="left" box="[711,718,1813,1827]" fontSize="6" pageId="3" pageNumber="260">1</subScript>
|
||
= anterior branch of radius; R
|
||
<subScript id="178AA226FFC93C6FC8FC6E49873EF8AB" attach="left" box="[1021,1045,1813,1827]" fontSize="6" pageId="3" pageNumber="260">2+3</subScript>
|
||
= second branch of radius; R = third
|
||
<subScript id="178AA226FFC93C6FCE3A6E4A8678F8AC" attach="left" box="[1339,1363,1814,1828]" fontSize="6" pageId="3" pageNumber="260">4+5</subScript>
|
||
branch of radius.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB1A063FFC93C6FCBC66ED78289F82D" blockId="3.[151,1437,1931,2030]" box="[199,418,1931,1957]" pageId="3" pageNumber="260">
|
||
<emphasis id="B97A7C71FFC93C6FCBC66ED7820EF82D" bold="true" box="[199,293,1931,1957]" pageId="3" pageNumber="260">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C314F3E8FFC93C68CBC66EF38125FF5D" lastPageId="4" lastPageNumber="261" pageId="3" pageNumber="260" type="materials_examined">
|
||
<paragraph id="8BB1A063FFC93C6FCBC66EF386B6F865" blockId="3.[151,1437,1931,2030]" pageId="3" pageNumber="260">
|
||
<emphasis id="B97A7C71FFC93C6FCBC66EF38259F841" bold="true" box="[199,370,1967,1993]" pageId="3" pageNumber="260">Type material.</emphasis>
|
||
<materialsCitation id="3B66AA3EFFC93C6FCA776EF386B6F865" collectingDate="2017-03-29" collectingDateMax="2017-04-14" collectingDateMin="2017-03-29" collectionCode="LEUA" collectorName="Y. Ramos-Pastrana Leg." country="Colombia" county="Florencia" elevation="1719" latitude="1.8511112" location="Trampa Malaise en Bosque" longLatPrecision="21" longitude="-75.66694" municipality="Vereda" pageId="3" pageNumber="260" specimenCount="1" specimenCount-male="1" stateProvince="Caqueta" typeStatus="holotype">
|
||
<emphasis id="B97A7C71FFC93C6FCA776EF3813EF841" bold="true" box="[374,533,1967,1993]" pageId="3" pageNumber="260">
|
||
<typeStatus id="54B51EC1FFC93C6FCA776EF3813EF841" box="[374,533,1967,1993]" pageId="3" pageNumber="260" type="holotype">HOLOTYPE</typeStatus>
|
||
</emphasis>
|
||
<specimenCount id="9D086BEAFFC93C6FC9186EEC811BF842" box="[537,560,1968,1994]" count="1" pageId="3" pageNumber="260" type="male">♂</specimenCount>
|
||
, labelled:“
|
||
<collectingCountry id="F319E0F3FFC93C6FC9A66EF38069F841" box="[679,834,1967,1993]" name="Colombia" pageId="3" pageNumber="260">COLOMBIA</collectingCountry>
|
||
,
|
||
<collectingRegion id="49CA6E81FFC93C6FC84B6EF38082F841" box="[842,937,1967,1993]" country="Colombia" name="Caqueta" pageId="3" pageNumber="260">Caquetá</collectingRegion>
|
||
,
|
||
<collectingCounty id="62D0D8EFFFC93C6FC8B06EF38735F841" box="[945,1054,1967,1993]" pageId="3" pageNumber="260">Florencia</collectingCounty>
|
||
, Vda.[
|
||
<collectingMunicipality id="6BD53A19FFC93C6FCF5E6EF3879EF841" box="[1119,1205,1967,1993]" pageId="3" pageNumber="260">Vereda</collectingMunicipality>
|
||
]
|
||
<collectingRegion id="49CA6E81FFC93C6FCFBE6EF38629F841" box="[1215,1282,1967,1993]" country="Colombia" name="Sucre" pageId="3" pageNumber="260">Sucre</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE3AC6A4FFC93C6FCE0B6EF386B8F842" box="[1290,1427,1967,1994]" degrees="01" direction="north" minutes="51" orientation="latitude" pageId="3" pageNumber="260" precision="15" seconds="04" value="1.8511112">01º51’04’’N</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE3AC6A4FFC93C6FCB966E8F8203F866" box="[151,296,2003,2030]" degrees="75" direction="west" minutes="40" orientation="longitude" pageId="3" pageNumber="260" precision="15" seconds="01" value="-75.66694">75º40’01’’W</geoCoordinate>
|
||
,
|
||
<quantity id="4CF60D86FFC93C6FCA346E8F82A6F866" box="[309,397,2003,2030]" metricMagnitude="3" metricUnit="m" metricValue="1.719" pageId="3" pageNumber="260" unit="m" value="1719.0">
|
||
<elevation id="00234750FFC93C6FCA346E8F82A6F866" box="[309,397,2003,2030]" metricMagnitude="3" metricUnit="m" metricValue="1.719" pageId="3" pageNumber="260" unit="m" value="1719.0">1719 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFB086A3FFC93C6FCA9B6E8F8149F865" box="[410,610,2003,2029]" pageId="3" pageNumber="260" value="2017-03-29" valueMax="2017-04-14" valueMin="2017-03-29">
|
||
<collectingDate id="EFF47F4BFFC93C6FCA9B6E8F8149F865" box="[410,610,2003,2029]" pageId="3" pageNumber="260" value="2017-03-29" valueMax="2017-04-14" valueMin="2017-03-29">29.III–14.IV.2017</collectingDate>
|
||
</date>
|
||
/
|
||
<location id="8ED1F6B8FFC93C6FC97B6E88809BF866" LSID="urn:lsid:plazi:treatment:03A71175FFC83C68CB96689183D4FD91:8ED1F6B8FFC93C6FC97B6E88809BF866" box="[634,944,2003,2030]" country="Colombia" county="Florencia" latitude="1.8511112" longLatPrecision="21" longitude="-75.66694" municipality="Vereda" name="Trampa Malaise en Bosque" pageId="3" pageNumber="260" stateProvince="Caqueta">Trampa Malaise en Bosque</location>
|
||
(Dosel),
|
||
<collectorName id="26FBC5B5FFC93C6FCF1B6E888607F865" box="[1050,1324,2003,2030]" pageId="3" pageNumber="260">Y. Ramos-Pastrana Leg.</collectorName>
|
||
” (
|
||
<collectionCode id="ED1F38A6FFC93C6FCE476E8F86BFF865" box="[1350,1428,2003,2029]" pageId="3" pageNumber="260">LEUA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BB1A063FFCE3C68CB9669CB8125FF5D" blockId="4.[151,1436,151,537]" pageId="4" pageNumber="261">
|
||
(photographed specimen).
|
||
<typeStatus id="54B51EC1FFCE3C68CAC669CB8104FF39" box="[455,559,151,177]" pageId="4" pageNumber="261" type="holotype">Holotype</typeStatus>
|
||
condition: good, right leg I and left It
|
||
<subScript id="178AA226FFCE3C68C8EB69F58729FF3F" attach="left" box="[1002,1026,169,183]" fontSize="6" pageId="4" pageNumber="261">4–5</subScript>
|
||
broken off, abdomen dissected and stored in microvial with glycerin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C314F3E8FFCE3C68CBC66982800DFF71" box="[199,806,222,249]" pageId="4" pageNumber="261" type="distribution">
|
||
<paragraph id="8BB1A063FFCE3C68CBC66982800DFF71" blockId="4.[151,1436,151,537]" box="[199,806,222,249]" pageId="4" pageNumber="261">
|
||
<emphasis id="B97A7C71FFCE3C68CBC66982824AFF70" bold="true" box="[199,353,222,248]" pageId="4" pageNumber="261">Distribution.</emphasis>
|
||
<collectingCountry id="F319E0F3FFCE3C68CA66698382FEFF71" box="[359,469,223,249]" name="Colombia" pageId="4" pageNumber="261">Colombia</collectingCountry>
|
||
(
|
||
<collectingRegion id="49CA6E81FFCE3C68CAE46983816CFF71" box="[485,583,223,249]" country="Colombia" name="Caqueta" pageId="4" pageNumber="261">Caquetá</collectingRegion>
|
||
: Florencia) (
|
||
<figureCitation id="1335BCE6FFCE3C68C9D269838033FF71" box="[723,792,223,249]" captionStart="FIGURE 7" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[301,1286,1429,1917]" captionTargetId="figure-56@13.[299,1288,1427,1919]" captionTargetPageId="13" captionText="FIGURE 7. Geographical records of Systenus Loew in Colombia. S. beatae Naglis, 2000; S. bickeli sp. nov.; S. colombiensis sp. nov.; S. flaviatus Naglis, 2000; S. fuscus sp. nov.; S. grossus sp. nov.; S. maculipennis Bickel, 2015; S. naranjensis Bickel, 2015 and S. parkeri Bickel, 2015." figureDoi="http://doi.org/10.5281/zenodo.13849845" httpUri="https://zenodo.org/record/13849845/files/figure.png" pageId="4" pageNumber="261">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C314F3E8FFCE3C68CBC6685F804DFE59" pageId="4" pageNumber="261" type="discussion">
|
||
<paragraph id="8BB1A063FFCE3C68CBC6685F804DFE59" blockId="4.[151,1436,151,537]" pageId="4" pageNumber="261">
|
||
<emphasis id="B97A7C71FFCE3C68CBC6685F8210FE95" bold="true" box="[199,315,259,285]" pageId="4" pageNumber="261">Remarks.</emphasis>
|
||
The new species runs to
|
||
<taxonomicName id="4C0EDBE0FFCE3C68C959685F8194FE95" authorityName="Naglis" authorityYear="2000" box="[600,703,259,285]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="beatae">
|
||
<emphasis id="B97A7C71FFCE3C68C959685F814DFE95" box="[600,614,259,285]" italics="true" pageId="4" pageNumber="261">S</emphasis>
|
||
.
|
||
<emphasis id="B97A7C71FFCE3C68C975685F8194FE95" box="[628,703,259,285]" italics="true" pageId="4" pageNumber="261">beatae</emphasis>
|
||
</taxonomicName>
|
||
in the key of
|
||
<bibRefCitation id="EF9FDD92FFCE3C68C85A685F80DDFE95" author="Naglis, S." box="[859,1014,259,285]" pageId="4" pageNumber="261" pagination="1 - 11" refId="ref11237" refString="Naglis, S. (2000) Six new species of Systenus (Diptera, Dolichopodidae) from Brazil, with a key to the Neotropical species. Studia dipterologica, 7, 1 - 11. https: // doi. org / 10.11646 / zootaxa. 4020.1.7" type="journal article" year="2000">Naglis (2000)</bibRefCitation>
|
||
and to
|
||
<taxonomicName id="4C0EDBE0FFCE3C68CF48685F8623FE95" authorityName="Bickel" authorityYear="2015" box="[1097,1288,259,285]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="flavifemoratus">
|
||
<emphasis id="B97A7C71FFCE3C68CF48685F877CFE95" box="[1097,1111,259,285]" italics="true" pageId="4" pageNumber="261">S</emphasis>
|
||
.
|
||
<emphasis id="B97A7C71FFCE3C68CF67685F8623FE95" box="[1126,1288,259,285]" italics="true" pageId="4" pageNumber="261">flavifemoratus</emphasis>
|
||
</taxonomicName>
|
||
in the key of
|
||
<bibRefCitation id="EF9FDD92FFCE3C68CB96687B821EFEC9" author="Bickel, D. J." box="[151,309,294,321]" pageId="4" pageNumber="261" pagination="169 - 182" refId="ref10716" refString="Bickel, D. J. (2015) The Costa Rican Systenus Low (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus. Zootaxa, 4020 (1), 169 - 182. https: // doi. org / 10.11646 / zootaxa. 4020.1.7" type="journal article" year="2015">Bickel (2015)</bibRefCitation>
|
||
, but can be differentiated from
|
||
<taxonomicName id="4C0EDBE0FFCE3C68C998687B802AFEC9" authorityName="Naglis" authorityYear="2000" box="[665,769,295,321]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="beatae">
|
||
<emphasis id="B97A7C71FFCE3C68C998687B818CFEC9" box="[665,679,295,321]" italics="true" pageId="4" pageNumber="261">S</emphasis>
|
||
.
|
||
<emphasis id="B97A7C71FFCE3C68C9B7687B802AFEC9" box="[694,769,295,321]" italics="true" pageId="4" pageNumber="261">beatae</emphasis>
|
||
</taxonomicName>
|
||
by the surstylus shorter than cercus and abruptly narrowed at apical 2/3 and by the presence of 1 long, acute basodorsal digitiform process (surstylus longer than cercus, only slightly narrowed at apex, with 1 wide digitiform process at middle of dorsal edge in
|
||
<taxonomicName id="4C0EDBE0FFCE3C68CFB66833860AFE01" authorityName="Naglis" authorityYear="2000" box="[1207,1313,367,393]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="beatae">
|
||
<emphasis id="B97A7C71FFCE3C68CFB6683387EEFE01" box="[1207,1221,367,393]" italics="true" pageId="4" pageNumber="261">S</emphasis>
|
||
.
|
||
<emphasis id="B97A7C71FFCE3C68CFD76833860AFE01" box="[1238,1313,367,393]" italics="true" pageId="4" pageNumber="261">beatae</emphasis>
|
||
</taxonomicName>
|
||
) and from
|
||
<taxonomicName id="4C0EDBE0FFCE3C68CB9668CF8273FE25" authorityName="Bickel" authorityYear="2015" box="[151,344,403,429]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="flavifemoratus">
|
||
<emphasis id="B97A7C71FFCE3C68CB9668CF838EFE25" box="[151,165,403,429]" italics="true" pageId="4" pageNumber="261">S</emphasis>
|
||
.
|
||
<emphasis id="B97A7C71FFCE3C68CBB768CF8273FE25" box="[182,344,403,429]" italics="true" pageId="4" pageNumber="261">flavifemoratus</emphasis>
|
||
</taxonomicName>
|
||
by the wing light brown and the surstylus slightly curved ventrally at apex (wing hyaline and surstylus abruptly curved ventrally at apex in
|
||
<taxonomicName id="4C0EDBE0FFCE3C68C99968EB807DFE59" authorityName="Bickel" authorityYear="2015" box="[664,854,439,465]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="flavifemoratus">
|
||
<emphasis id="B97A7C71FFCE3C68C99968EB818DFE59" box="[664,678,439,465]" italics="true" pageId="4" pageNumber="261">S</emphasis>
|
||
.
|
||
<emphasis id="B97A7C71FFCE3C68C9B568EB807DFE59" box="[692,854,439,465]" italics="true" pageId="4" pageNumber="261">flavifemoratus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C314F3E8FFCE3C68CBC6688783D4FD91" pageId="4" pageNumber="261" type="etymology">
|
||
<paragraph id="8BB1A063FFCE3C68CBC6688783D4FD91" blockId="4.[151,1436,151,537]" pageId="4" pageNumber="261">
|
||
<emphasis id="B97A7C71FFCE3C68CBC668878265FE7D" bold="true" box="[199,334,475,501]" pageId="4" pageNumber="261">Etymology.</emphasis>
|
||
Named after Daniel Bickel (Australian Museum) for his contribution to the knowledge of the genus
|
||
<taxonomicName id="4C0EDBE0FFCE3C68CB9668A383DCFD91" authorityName="Loew" authorityYear="1857" box="[151,247,511,537]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="261" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97A7C71FFCE3C68CB9668A383DCFD91" box="[151,247,511,537]" italics="true" pageId="4" pageNumber="261">Systenus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |