122 lines
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122 lines
15 KiB
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<document id="405A3D7F9261DA83FD9CB7B6E51CC3F0" ID-CLB-Dataset="203546" ID-DOI="10.11646/zootaxa.5219.1.3" ID-GBIF-Dataset="0523df85-98fc-4cc3-baff-61c49020a01b" ID-ISSN="1175-5326" ID-Zenodo-Dep="7408120" ID-ZooBank="C39DD77A-45E5-475C-B214-85DEA9D75AA9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1670399898510" checkinUser="plazi" docAuthor="Thandar, A. S. & Arumugam, P." docDate="2022" docId="039A87C2267CB863A0E8FE4AFD989AD5" docLanguage="en" docName="zootaxa.5219.1.3.pdf" docOrigin="Zootaxa 5219 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sclerothyone Thandar 1989" docType="treatment" docVersion="6" lastPageNumber="66" masterDocId="FFA3FFBA267DB862A07FFF6CFFE29955" masterDocTitle="Referral of Thyone neofusus Deichmann, 1941, Thyone adinopoda (Pawson & Miller, 1981 and Havelockia obunca (Lampert, 1885) to the genus Sclerothyone Thandar, 1990, and a replacement name for the preoccupied genus Neothyone Deichmann, 1941 (Echinodermata: Holothuroidea: Dendrochirotida)" masterLastPageNumber="71" masterPageNumber="65" pageNumber="66" updateTime="1699413116561" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3C9773BFC4D409AA62FF12CCB2E98F91">Referral of Thyone neofusus Deichmann, 1941, Thyone adinopoda (Pawson & Miller, 1981 and Havelockia obunca (Lampert, 1885) to the genus Sclerothyone Thandar, 1990, and a replacement name for the preoccupied genus Neothyone Deichmann, 1941 (Echinodermata: Holothuroidea: Dendrochirotida)</mods:title>
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<mods:namePart id="A95AF64AF71B046350CE2D6049D45067">Thandar, A. S.</mods:namePart>
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<mods:affiliation id="6942AC9910D43F14D2687B0ED280594F">University of KwaZulu-Natal, Westville Campus, P / Bag X 54001, Durban 4000</mods:affiliation>
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<mods:namePart id="ED2B817018D33DBBDE87E253EE1E8459">Arumugam, P.</mods:namePart>
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<mods:affiliation id="CE0F120A2B18D5CC1E83720CC2F0F7EA">University of KwaZulu-Natal, Westville Campus, P / Bag X 54001, Durban 4000 & 205500752 @ stu. ukzn. ac. za; https: // orcid. org 0000 - 0002 - 1945 - 7268</mods:affiliation>
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<mods:nameIdentifier id="F230FEB712915BB0F9D64750FC9B1D40" type="email">205500752@stu.ukzn.ac.za</mods:nameIdentifier>
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<mods:typeOfResource id="4172C0641B6204724F7A1488724D90FF">text</mods:typeOfResource>
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<mods:title id="DCDD52C862B5062C69DA14B7BF0A425D">Zootaxa</mods:title>
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<mods:date id="1FA4F522B6D58EDFA207C4F78241123F">2022</mods:date>
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<treatment id="039A87C2267CB863A0E8FE4AFD989AD5" ID-DOI="http://doi.org/10.5281/zenodo.7408122" ID-GBIF-Taxon="205875619" ID-Zenodo-Dep="7408122" LSID="urn:lsid:plazi:treatment:039A87C2267CB863A0E8FE4AFD989AD5" httpUri="http://treatment.plazi.org/id/039A87C2267CB863A0E8FE4AFD989AD5" lastPageNumber="66" pageId="1" pageNumber="66">
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<subSubSection id="C329655F267CB863A0E8FE4AFDA89815" box="[151,586,294,321]" pageId="1" pageNumber="66" type="nomenclature">
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<paragraph id="8B8C36D4267CB863A0E8FE4AFDA89815" blockId="1.[151,586,294,321]" box="[151,586,294,321]" pageId="1" pageNumber="66">
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<heading id="D0C481B8267CB863A0E8FE4AFDA89815" bold="true" box="[151,586,294,321]" fontSize="11" level="1" pageId="1" pageNumber="66" reason="1">
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<emphasis id="B947EAC6267CB863A0E8FE4AFDA89815" bold="true" box="[151,586,294,321]" pageId="1" pageNumber="66">
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Genus
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<taxonomicName id="4C334D57267CB863A091FE4AFDA89815" ID-CoL="7PZ4F" authority="Thandar, 1989" authorityName="Thandar" authorityYear="1989" box="[238,586,294,321]" class="Holothuroidea" family="Sclerodactylidae" genus="Sclerothyone" kingdom="Animalia" order="Dendrochirotida" pageId="1" pageNumber="66" phylum="Echinodermata" rank="genus">
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<emphasis id="B947EAC6267CB863A091FE4AFE6E9814" bold="true" box="[238,396,294,321]" italics="true" pageId="1" pageNumber="66">Sclerothyone</emphasis>
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<bibRefCitation id="EFA24B25267CB863A1EBFE4AFDA89815" author="Thandar, A. S." box="[404,586,294,320]" pageId="1" pageNumber="66" pagination="290 - 304" refId="ref5553" refString="Thandar, A. S. (1989) The sclerodactylid holothurians of southern Africa, with the erection of one new subfamily and two new genera. South African Journal of Zoology, 24 (4), 290 - 304. https: // doi. org / 10.1080 / 02541858.1989.11448167" type="journal article" year="1989">Thandar, 1989</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C329655F267CB863A0E8FE02FD989AD5" pageId="1" pageNumber="66" type="diagnosis">
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<paragraph id="8B8C36D4267CB863A0E8FE02FDA69B69" blockId="1.[151,1437,366,897]" pageId="1" pageNumber="66">
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<emphasis id="B947EAC6267CB863A0E8FE02FEE898DD" bold="true" box="[151,266,366,392]" pageId="1" pageNumber="66">Diagnosis</emphasis>
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(from
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<bibRefCitation id="EFA24B25267CB863A120FE02FD9998DD" author="Martins, L. & Tavares, M." box="[351,635,366,393]" pageId="1" pageNumber="66" pagination="375 - 382" refId="ref5355" refString="Martins, L. & Tavares, M. (2019) Two new species of Sclerothyone from the Southwestern Atlantic Ocean, with a key to the genera and species of Sclerothyoninae (Holothuroidea: Dendrochirotida: Sclerodactylidae). Zootaxa, 4658 (2), 375 - 382. https: // doi. org / 10.11646 / zootaxa. 4658.2.1" type="journal article" year="2019">Martins & Tavares 2019</bibRefCitation>
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, amended herein). Tube feet in double rows, restricted to ambulacra, interambulacra mostly naked, or tube feet scattered over body. Calcareous ring compact, non-tubular, posterior processes of radial plates long, entire or subdivided. Body wall tables with regular, oval, four-holed disc, with or without a handle, or large, irregular multilocular disc, also with or without a handle. Spire with pillars separate or fused. Introvert with plates, rods, rosettes and/or tables. Tentacles with rods, plates, rosettes or tables. Tube feet with endplate, supporting tables and plates.
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</paragraph>
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<paragraph id="8B8C36D4267CB863A0B8FD2AFD989AD5" blockId="1.[151,1437,366,897]" pageId="1" pageNumber="66">
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<emphasis id="B947EAC6267CB863A0B8FD2AFE719B35" bold="true" box="[199,403,582,608]" pageId="1" pageNumber="66">Included species.</emphasis>
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All genera and species included, except those here dealt with, are tabulated by Martins & Tavares (1998), together with their distribution and ossicle morphology and there is no need to repeat this here, except that
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<taxonomicName id="4C334D57267CB863A164FDE3FD229BFD" authority="Mitsukuri, 2012" authorityName="Mitsukuri" authorityYear="2012" box="[283,704,654,681]" class="Holothuroidea" family="Cucumariidae" genus="Cucumaria" kingdom="Animalia" order="Dendrochirotida" pageId="1" pageNumber="66" phylum="Echinodermata" rank="species" species="nozawai">
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<emphasis id="B947EAC6267CB863A164FDE3FE1C9BFD" box="[283,510,655,681]" italics="true" pageId="1" pageNumber="66">Cucumaria nozawai</emphasis>
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Mitsukuri, 2012
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</taxonomicName>
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, from
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<collectingCountry id="F3247644267CB863A374FDE3FCAC9BFC" box="[779,846,655,681]" name="Japan" pageId="1" pageNumber="66">Japan</collectingCountry>
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, was transferred to
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<taxonomicName id="4C334D57267CB863A451FDE2FB5D9BFD" authorityName="Thandar" authorityYear="1989" box="[1070,1215,654,680]" class="Holothuroidea" family="Sclerodactylidae" genus="Sclerothyone" kingdom="Animalia" order="Dendrochirotida" pageId="1" pageNumber="66" phylum="Echinodermata" rank="genus">
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<emphasis id="B947EAC6267CB863A451FDE2FB5D9BFD" box="[1070,1215,654,680]" italics="true" pageId="1" pageNumber="66">Sclerothyone</emphasis>
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</taxonomicName>
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by Thandar (2021) and, in the current paper,
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<taxonomicName id="4C334D57267CB863A1C1FDDEFCAC9B99" authority="Deichmann, 1941" authorityName="Deichmann" authorityYear="1941" box="[446,846,690,716]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="1" pageNumber="66" phylum="Echinodermata" rank="species" species="neofusus">
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<emphasis id="B947EAC6267CB863A1C1FDDEFD9E9B99" box="[446,636,690,716]" italics="true" pageId="1" pageNumber="66">Thyone neofusus</emphasis>
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<bibRefCitation id="EFA24B25267CB863A2FAFDDEFCAC9B99" author="Deichmann, E." box="[645,846,690,716]" pageId="1" pageNumber="66" pagination="61 - 153" refId="ref5136" refString="Deichmann, E. (1941) The Holothurioidea collected by the Velero III during the years 1932 - 38. Part 1. Dendrochirota. Allan Hancock Paciic Expedition, 8, 61 - 153, 6 text-figs, pls. 10 - 30." type="journal article" year="1941">Deichmann, 1941</bibRefCitation>
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</taxonomicName>
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from the West Atlantic,
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<taxonomicName id="4C334D57267CB863A416FDDEFEA79BA5" authority="Pawson & Miller. 1981" authorityName="Pawson & Miller." authorityYear="1981" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="1" pageNumber="66" phylum="Echinodermata" rank="species" species="adinopoda">
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<emphasis id="B947EAC6267CB863A416FDDEFADE9B99" box="[1129,1340,690,716]" italics="true" pageId="1" pageNumber="66">Thyone adinopoda</emphasis>
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Pawson & Miller. 1981
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</taxonomicName>
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also from the West Atlantic and
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<taxonomicName id="4C334D57267CB863A2B4FDBAFB8E9BA4" authority="(Lampert, 1885)" baseAuthorityName="Lampert" baseAuthorityYear="1885" box="[715,1132,726,753]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="1" pageNumber="66" phylum="Echinodermata" rank="species" species="obunca">
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<emphasis id="B947EAC6267CB863A2B4FDBAFC459BA5" box="[715,935,726,752]" italics="true" pageId="1" pageNumber="66">Havelockia obunca</emphasis>
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(
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<bibRefCitation id="EFA24B25267CB863A3C6FDBAFB869BA4" author="Lampert, K." box="[953,1124,726,753]" pageId="1" pageNumber="66" pagination="1 - 312" refId="ref5239" refString="Lampert, K. (1885) Die Seewalzen (Holothroidea). In: Semper, C., Philippinen Reisen im Archipel der, Wiesbaden, Series 2, 4 (3), 1 - 312, 1 pl." type="journal article" year="1885">Lampert, 1885</bibRefCitation>
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)
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</taxonomicName>
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from
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<collectingCountry id="F3247644267CB863A4C8FDBBFB189BA4" box="[1207,1274,727,753]" name="Japan" pageId="1" pageNumber="66">Japan</collectingCountry>
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, are also now transferred to this genus as suggested by
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<bibRefCitation id="EFA24B25267CB863A219FD96FCD89A41" author="Arumugam, P." box="[614,826,762,788]" pageId="1" pageNumber="66" refId="ref5043" refString="Arumugam, P. (2011) A critical assessment of the dendrochirotid subfamilies, Sclerodactylinae and Thyoninae, with the taxonomic management of the ' supergenus' Thyone (Echinodermata: Holothuroidea). M. Sc. dissertation, unpublished, University of KwaZulu-Natal, Berea, Durban, xvi + 229 pp." type="book" year="2011">Arumugam (2011)</bibRefCitation>
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. The inclusion of
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<taxonomicName id="4C334D57267CB863A47AFD97FB7B9A41" box="[1029,1177,762,788]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="1" pageNumber="66" phylum="Echinodermata" rank="species" species="adinopoda">
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<emphasis id="B947EAC6267CB863A47AFD97FB7B9A41" box="[1029,1177,762,788]" italics="true" pageId="1" pageNumber="66">T. adinopoda</emphasis>
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</taxonomicName>
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was not questioned by
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<bibRefCitation id="EFA24B25267CB863A0E8FC72FE5F9A6D" author="Martins, L. & Tavares, M." box="[151,445,798,825]" pageId="1" pageNumber="66" pagination="375 - 382" refId="ref5355" refString="Martins, L. & Tavares, M. (2019) Two new species of Sclerothyone from the Southwestern Atlantic Ocean, with a key to the genera and species of Sclerothyoninae (Holothuroidea: Dendrochirotida: Sclerodactylidae). Zootaxa, 4658 (2), 375 - 382. https: // doi. org / 10.11646 / zootaxa. 4658.2.1" type="journal article" year="2019">Martins & Tavares (2019)</bibRefCitation>
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but they excluded it from their table and the key. Hence altogether 13 species currently stand in
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<taxonomicName id="4C334D57267CB863A086FC2EFE689A09" authorityName="Thandar" authorityYear="1989" box="[249,394,834,860]" class="Holothuroidea" family="Sclerodactylidae" genus="Sclerothyone" kingdom="Animalia" order="Dendrochirotida" pageId="1" pageNumber="66" phylum="Echinodermata" rank="genus">
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<emphasis id="B947EAC6267CB863A086FC2EFE689A09" box="[249,394,834,860]" italics="true" pageId="1" pageNumber="66">Sclerothyone</emphasis>
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</taxonomicName>
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. The key provided by
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<bibRefCitation id="EFA24B25267CB863A2F2FC2EFC569A09" author="Martins, L. & Tavares, M." box="[653,948,834,861]" pageId="1" pageNumber="66" pagination="375 - 382" refId="ref5355" refString="Martins, L. & Tavares, M. (2019) Two new species of Sclerothyone from the Southwestern Atlantic Ocean, with a key to the genera and species of Sclerothyoninae (Holothuroidea: Dendrochirotida: Sclerodactylidae). Zootaxa, 4658 (2), 375 - 382. https: // doi. org / 10.11646 / zootaxa. 4658.2.1" type="journal article" year="2019">Martins & Tavares (2019)</bibRefCitation>
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is here modified to include all the species. The debatable species are discussed below.
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</paragraph>
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</subSubSection>
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</treatment>
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