treatments-xml/data/03/97/87/039787C2D23BFFD0166AFE0E5B75EB7E.xml
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<document id="7CFA455DB04140EBBC162FF5038A7181" ID-DOI="10.5281/zenodo.171327" ID-GBIF-Dataset="f27a15be-7b4a-442a-a141-ad361f119be0" ID-ISSN="1175­5326" ID-Zenodo-Dep="171327" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459720545083" checkinUser="plazi" docAuthor="Winterton, Shaun L. &amp; Metz, Mark A." docDate="2005" docId="039787C2D23BFFD0166AFE0E5B75EB7E" docLanguage="en" docName="zt00975.pdf" docOrigin="Zootaxa 975" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Cyrtosathe kirkspriggsi Winterton &amp; Metz" docType="treatment" docVersion="10" lastPageNumber="7" masterDocId="FFAEFFBAD238FFD61762FFFF585AEC20" masterDocTitle="Cyrtosathe gen. n.: the first non­scenopinine window fly from sub­ Saharan Africa (Diptera: Scenopinidae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="4" updateTime="1698209236972" updateUser="plazi">
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<mods:title id="8A65A08C3C394DEA5D242EF6165FEF33">Cyrtosathe gen. n.: the first non­scenopinine window fly from sub­ Saharan Africa (Diptera: Scenopinidae)</mods:title>
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<mods:name id="4C59A13B354502C54258EE7DC6A0526C" type="personal">
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<mods:namePart id="BD1B7142EDA1B1A62372E125A515C419">Winterton, Shaun L.</mods:namePart>
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<mods:namePart id="7B8952DCFE027A1E2F2AF836D623FBD6">Metz, Mark A.</mods:namePart>
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<mods:typeOfResource id="D2082E7B47A950F0F096AD2850833AA9">text</mods:typeOfResource>
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<mods:title id="A74DA2DF0EBFB3D3A587A55C95510D71">Zootaxa</mods:title>
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<mods:date id="896BBFD6EE79C851B24526A2857E15BC">2005</mods:date>
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<mods:number id="DD5261BE01CA1AAC682F7DB1A87ACE08">975</mods:number>
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<treatment id="039787C2D23BFFD0166AFE0E5B75EB7E" ID-DOI="http://doi.org/10.5281/zenodo.5686461" ID-GBIF-Taxon="119335518" ID-Zenodo-Dep="5686461" LSID="urn:lsid:plazi:treatment:039787C2D23BFFD0166AFE0E5B75EB7E" httpUri="http://treatment.plazi.org/id/039787C2D23BFFD0166AFE0E5B75EB7E" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4">
<subSubSection id="C324655FD23BFFD5166AFE0E59DBEE0E" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8B8136D4D23BFFD5166AFE0E5B96EE2B" blockId="3.[264,972,497,558]" box="[264,972,497,523]" pageId="3" pageNumber="4">
<heading id="D0C981B8D23BFFD5166AFE0E5B96EE2B" bold="true" box="[264,972,497,523]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<emphasis id="B94AEAC6D23BFFD5166AFE0E5B96EE2B" bold="true" box="[264,972,497,523]" pageId="3" pageNumber="4">
<taxonomicName id="4C3E4D57D23BFFD5166AFE0E5B4BEE2B" ID-CoL="33NP2" authority="Winterton &amp; Metz" authorityName="Winterton &amp; Metz" box="[264,785,497,523]" class="Insecta" family="Scenopinidae" genus="Cyrtosathe" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="kirkspriggsi">
<emphasis id="B94AEAC6D23BFFD5166AFE0E5A7BEE2B" bold="true" box="[264,545,497,523]" italics="true" pageId="3" pageNumber="4">Cyrtosathe kirkspriggsi</emphasis>
Winterton &amp; Metz
</taxonomicName>
gen.
<emphasis id="B94AEAC6D23BFFD51430FE0E5B3DEE2B" bold="true" box="[850,871,497,523]" italics="true" pageId="3" pageNumber="4">et</emphasis>
sp. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8136D4D23BFFD5166AFDEB59DBEE0E" blockId="3.[264,972,497,558]" box="[264,385,532,558]" pageId="3" pageNumber="4">
(
<figureCitation id="13052A51D23BFFD51672FDEB592DEE0E" box="[272,375,532,558]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="4.[264,367,1766,1790]" captionStartId-1="5.[264,368,899,923]" captionStartId-2="6.[264,368,1452,1476]" captionStartId-3="7.[264,368,1703,1727]" captionTargetBox-0="[276,1317,624,1746]" captionTargetBox-1="[264,1322,319,873]" captionTargetBox-2="[267,1309,397,1413]" captionTargetBox-3="[353,1214,294,1685]" captionTargetId-0="figure@4.[264,1323,614,1756]" captionTargetId-1="figure@5.[207,1379,309,900]" captionTargetId-2="figure@6.[264,1323,386,1425]" captionTargetId-3="figure@7.[340,1222,283,1688]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 1. Cyrtosathe kirkspriggsi gen. et sp. nov.: Male: A, head, lateral; B, same anterior. Female: C, head lateral; D, same anterior; E, antenna, right lateral. F, abdominal tergite 2 sensory setae patch (enlarged at right). Scale lines: 0.1 mm." captionText-1="FIGURE 2. Cyrtosathe kirkspriggsi gen. et sp. nov.: Wing. Scale line: 0.2 mm." captionText-2="FIGURE 3. Cyrtosathe kirkspriggsi gen. et sp. nov.: Male genitalia: A. Epandrium, dorsal; B, aedeagus and gonocoxites (epandrium and vestiture removed), dorso­lateral view; C, same, lateral; D, detail of basal portion of aedeagus, lateral; E, aedeagus and gonocoxites, dorsal; F, detail of gonocoxites. lateral, Scale line: 0.1 mm. Abbreviations: c, cercus; d, distiphallus; da, dorsal apodeme of parameral sheath; e, epandrium; ea, ejaculatory apodeme; g, gonocoxite; ga, gonocoxal apodeme; gs, gonostylus; h, hypandrium; hy, hypoproct." captionText-3="FIGURE 4. Cyrtosathe kirkspriggsi gen. et sp. nov.: Female genitalia: A, Genitalia in contacted position, postero­lateral; B, detail of same with acanthophorites slightly relaxed posteriorly, lateral; C, same, showing distal reproductive system (tergite 8 cut away and vestiture removed), dorsal. Scale lines: 0.1 mm. Abbreviations: A 1, acanthophorite macrosetae; ag, accessory gland duct (gland cut away); f, furca; ss, spermathecal sac; s, spermatheca; sd, spermathecal duct; s 8, sternite 8; s 10, sternite 10; t 8 ­ t 10, tergites 8 to 10." httpUri-0="https://zenodo.org/record/171328/files/figure.png" httpUri-1="https://zenodo.org/record/171329/files/figure.png" httpUri-2="https://zenodo.org/record/171330/files/figure.png" httpUri-3="https://zenodo.org/record/171331/files/figure.png" pageId="3" pageNumber="4">Figs 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324655FD23BFFD5166AFD9C5C64EE5E" box="[264,1086,611,638]" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="8B8136D4D23BFFD5166AFD9C5C64EE5E" blockId="3.[264,1325,611,1881]" box="[264,1086,611,638]" pageId="3" pageNumber="4">
<emphasis id="B94AEAC6D23BFFD5166AFD9C59F0EE5D" bold="true" box="[264,426,611,637]" pageId="3" pageNumber="4">
<typeStatus id="54858876D23BFFD5166AFD9C591FEE5D" box="[264,325,611,637]" pageId="3" pageNumber="4">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C3E4D57D23BFFD516D0FD9B5A9CEE5D" box="[434,710,612,637]" class="Insecta" family="Scenopinidae" genus="Cyrtosathe" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="kirkspriggsi" status="sp. nov.">
<emphasis id="B94AEAC6D23BFFD516D0FD9B5A9CEE5D" box="[434,710,612,637]" italics="true" pageId="3" pageNumber="4">Cyrtosathe kirkspriggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27957BDD23BFFD515AEFD9B5B7DEE5E" box="[716,807,612,638]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C324655FD23BFFD5165AFD735D57EEEE" pageId="3" pageNumber="4" type="etymology">
<paragraph id="8B8136D4D23BFFD5165AFD735D57EEEE" blockId="3.[264,1325,611,1881]" pageId="3" pageNumber="4">
<emphasis id="B94AEAC6D23BFFD5165AFD73599EEE86" bold="true" box="[312,452,652,678]" pageId="3" pageNumber="4">Etymology.</emphasis>
<taxonomicName id="4C3E4D57D23BFFD516ACFD735A15EE85" box="[462,591,652,677]" class="Insecta" family="Scenopinidae" genus="Cyrtosathe" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAC6D23BFFD516ACFD735A15EE85" box="[462,591,652,677]" italics="true" pageId="3" pageNumber="4">Cyrtosathe</emphasis>
</taxonomicName>
is derived from the Greek,
<emphasis id="B94AEAC6D23BFFD514FFFD735B83EE85" box="[925,985,652,677]" italics="true" pageId="3" pageNumber="4">cyrto</emphasis>
, bent;
<emphasis id="B94AEAC6D23BFFD5134FFD735C30EE85" box="[1069,1130,652,677]" italics="true" pageId="3" pageNumber="4">sathe</emphasis>
penis. The specific epithet is named in honour of Ashley Kirk­Spriggs, the collector of the
<typeStatus id="54858876D23BFFD513EBFD4B5CE1EEEE" box="[1161,1211,692,718]" pageId="3" pageNumber="4">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="C324655FD23BFFD5165AFD23593CE87E" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="8B8136D4D23BFFD5165AFD23593CE87E" blockId="3.[264,1325,611,1881]" pageId="3" pageNumber="4">
<emphasis id="B94AEAC6D23BFFD5165AFD2359A9EED6" bold="true" box="[312,499,732,758]" pageId="3" pageNumber="4">
<typeStatus id="54858876D23BFFD5165AFD23592FEED6" box="[312,373,732,758]" pageId="3" pageNumber="4">Type</typeStatus>
material:
</emphasis>
<typeStatus id="54858876D23BFFD5169FFD235A30EED6" box="[509,618,732,758]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F3297644D23BFFD515DDFD235B1EEED6" box="[703,836,732,758]" name="Namibia" pageId="3" pageNumber="4">NAMIBIA</collectingCountry>
: Luderitz District, Ob[i]b Waters, pitfall trap (
<geoCoordinate id="EE0A5013D23BFFD5162EFCFB5980EF3E" box="[332,474,772,798]" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="15" value="-28.002222">28°0008”S</geoCoordinate>
, 16°3646”),
<date id="FF801014D23BFFD515E7FCFB5B19EF3E" box="[645,835,772,798]" pageId="3" pageNumber="4" value="1998-08-10" valueMax="1998-08-26" valueMin="1998-08-10">1026.viii.1998</date>
, Kirk­Spriggs &amp; Marais (MEI#119440) (
<collectionCode id="ED2FAE11D23BFFD51672FCD3592DEF66" box="[272,375,812,838]" httpUri="http://grbio.org/cool/y6s1-e7mm" name="National Museum of Namibia" pageId="3" pageNumber="4">NMNW</collectionCode>
).
<typeStatus id="54858876D23BFFD516E8FCD359A5EF66" box="[394,511,812,838]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
,
<collectingCountry id="F3297644D23BFFD51568FCD35ACAEF66" box="[522,656,812,838]" name="Namibia" pageId="3" pageNumber="4">NAMIBIA</collectingCountry>
:
<specimenCount id="9D38FD5DD23BFFD515F8FCD35AA2EF66" box="[666,760,812,838]" pageId="3" pageNumber="4" type="male">2 males</specimenCount>
,
<specimenCount id="9D38FD5DD23BFFD51461FCD35B36EF66" box="[771,876,812,838]" pageId="3" pageNumber="4" type="female">1 female</specimenCount>
, same data as
<typeStatus id="54858876D23BFFD51370FCD35C23EF66" box="[1042,1145,812,838]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
(MEI#119434, 119442, 119465) (
<collectionCode id="ED2FAE11D23BFFD516BCFCAB5A63EF4E" LSID="urn:lsid:biocol.org:col:34871" box="[478,569,852,878]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="4">USNM</collectionCode>
); male, Obib Waters, Malaise trap (
<geoCoordinate id="EE0A5013D23BFFD514B8FCAB5C32EF4E" box="[986,1128,852,878]" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="15" value="-28.002222">28°0008”S</geoCoordinate>
, 16°3846”)
<date id="FF801014D23BFFD51262FCAB59C2EFB6" pageId="3" pageNumber="4" value="1998-08-25" valueMax="1998-08-26" valueMin="1998-08-25">25 26.viii.1998</date>
, Kirk­Spriggs &amp; Marais (MEI#
<date id="FF801014D23BFFD51469FC835B3EEFB6" box="[779,868,892,918]" pageId="3" pageNumber="4" value="32-11-29">112932</date>
) (
<collectionCode id="ED2FAE11D23BFFD51419FC835B8DEFB6" LSID="urn:lsid:biocol.org:col:34871" box="[891,983,892,918]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="4">USNM</collectionCode>
);
<specimenCount id="9D38FD5DD23BFFD51489FC835C12EFB6" box="[1003,1096,892,918]" pageId="3" pageNumber="4" type="male">6 males</specimenCount>
,
<specimenCount id="9D38FD5DD23BFFD51337FC835CE7EFB6" box="[1109,1213,892,918]" pageId="3" pageNumber="4" type="female">1 female</specimenCount>
, Luderitz District,
<quantity id="4CC69B31D23BFFD51612FC5B59F7EF9E" box="[368,429,932,958]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="4" unit="km" value="8.0">8 km</quantity>
W Rosh Pinah, pitfall trap (
<geoCoordinate id="EE0A5013D23BFFD5159FFC5B5BD1EF9E" box="[765,907,932,958]" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="15" value="-27.991112">27°5928”S</geoCoordinate>
, 16°
<date id="FF801014D23BFFD514A2FC5B5C5CEF9E" box="[960,1030,932,958]" pageId="3" pageNumber="4" value="1939-01-14">39í14</date>
”)
<date id="FF801014D23BFFD51341FC5B5CBAEF9E" box="[1059,1248,932,958]" pageId="3" pageNumber="4" value="1998-08-20" valueMax="1998-08-26" valueMin="1998-08-20">2026.viii.1998</date>
, Kirk­ Spriggs &amp; Marais (MEI#119436, 119438, 119441, 1194456, 119451, 119453) (
<collectionCode id="ED2FAE11D23BFFD513DBFC335D7AEFC6" box="[1209,1312,972,998]" httpUri="http://grbio.org/cool/y6s1-e7mm" name="National Museum of Namibia" pageId="3" pageNumber="4">NMNW</collectionCode>
);
<specimenCount id="9D38FD5DD23BFFD5166AFC0B593AE82E" box="[264,352,1012,1038]" pageId="3" pageNumber="4" type="male">1 male</specimenCount>
,
<specimenCount id="9D38FD5DD23BFFD51610FC0B59BBE82E" box="[370,481,1012,1038]" pageId="3" pageNumber="4" type="female">1 female</specimenCount>
, Luderitz District,
<quantity id="4CC69B31D23BFFD515B3FC0B5B49E82E" box="[721,787,1012,1038]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="4" unit="km" value="8.0">8 km</quantity>
W Rosh Pinah, Malaise trap (
<geoCoordinate id="EE0A5013D23BFFD513F8FC0B5D72E82E" box="[1178,1320,1012,1038]" direction="south" orientation="latitude" pageId="3" pageNumber="4" precision="15" value="-27.991112">27°5928”S</geoCoordinate>
, 16°3914”)
<date id="FF801014D23BFFD516FFFBE35A00E816" box="[413,602,1052,1078]" pageId="3" pageNumber="4" value="1998-08-20" valueMax="1998-08-26" valueMin="1998-08-20">2026.viii.1998</date>
, Kirk­Spriggs &amp; Marais (MEI#119448, 119454) (
<collectionCode id="ED2FAE11D23BFFD513BBFBE35D7EE816" box="[1241,1316,1052,1078]" pageId="3" pageNumber="4">MEIC</collectionCode>
/
<collectionCode id="ED2FAE11D23BFFD5166AFBBB5900E87E" box="[264,346,1092,1118]" pageId="3" pageNumber="4">CASC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C324655FD23BFFD5165AFB935C31E9D1" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8B8136D4D23BFFD5165AFB935C31E9D1" blockId="3.[264,1325,611,1881]" pageId="3" pageNumber="4">
<emphasis id="B94AEAC6D23BFFD5165AFB9359EAE8A6" bold="true" box="[312,432,1132,1158]" pageId="3" pageNumber="4">Diagnosis</emphasis>
. Autapomorphic characters: Female tergite 8 covered with erect, elongate setae arranged in a ring pattern (i.e. glabrous centre); hypandrium bowl­like with posterior processes; dorsal apodeme of parameral sheath enlarged into a ladle­like structure enclosing ejaculatory apodeme; aedeagus folded onto itself posteriorly such that the ejaculatory apodeme is directed posteriorly. Shared characters with other genera: head and thorax mostly glabrous; costal vein ending just past R5; wing vein M2 present; hind coxal knob absent; abdominal tergite 2 sensory patch present as two hemispheres, sensory setae with apices acute (not truncated); male epandrium divided medially; hypandrium separate from gonocoxites; gonocoxal apodeme slightly longer than gonocoxites; distiphallus bifid, greatly elongate and narrow; ventral apodeme of parameral sheath absent.
</paragraph>
</subSubSection>
<subSubSection id="C324655FD23BFFD0165AFA0159A5ED7E" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4" type="description">
<paragraph id="8B8136D4D23BFFD5165AFA015B95EB79" blockId="3.[264,1325,611,1881]" pageId="3" pageNumber="4">
<emphasis id="B94AEAC6D23BFFD5165AFA015993EA38" bold="true" box="[312,457,1534,1560]" pageId="3" pageNumber="4">Description</emphasis>
. Body length: 2.02.5 mm.
<emphasis id="B94AEAC6D23BFFD51477FA005B0EEA38" box="[789,852,1535,1560]" italics="true" pageId="3" pageNumber="4">Head</emphasis>
. Spherical, glossy dark brown to black; frons glabrous, narrow in male (
<figureCitation id="13052A51D23BFFD515F7F9D85ABAEA61" box="[661,736,1575,1601]" captionStart="FIGURE 1" captionStartId="4.[264,367,1766,1790]" captionTargetBox="[276,1317,624,1746]" captionTargetId="figure@4.[264,1323,614,1756]" captionTargetPageId="4" captionText="FIGURE 1. Cyrtosathe kirkspriggsi gen. et sp. nov.: Male: A, head, lateral; B, same anterior. Female: C, head lateral; D, same anterior; E, antenna, right lateral. F, abdominal tergite 2 sensory setae patch (enlarged at right). Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/171328/files/figure.png" pageId="3" pageNumber="4">Figs 1</figureCitation>
A, B) with eyes contiguous along middle third, female (
<figureCitation id="13052A51D23BFFD51605F9B059F4EA49" box="[359,430,1615,1641]" captionStart="FIGURE 1" captionStartId="4.[264,367,1766,1790]" captionTargetBox="[276,1317,624,1746]" captionTargetId="figure@4.[264,1323,614,1756]" captionTargetPageId="4" captionText="FIGURE 1. Cyrtosathe kirkspriggsi gen. et sp. nov.: Male: A, head, lateral; B, same anterior. Female: C, head lateral; D, same anterior; E, antenna, right lateral. F, abdominal tergite 2 sensory setae patch (enlarged at right). Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/171328/files/figure.png" pageId="3" pageNumber="4">Figs 1</figureCitation>
C, D) frons much wider than ocellar tubercle along entire length; male eyes with facets larger in upper half; ocellar tubercle raised, ocelli red­brown; occiput flat to slightly concave laterally; postocular ridge with irregular rows of minute postocular setae; gena with short setae; mouthparts brown, relatively elongate, approximately equal in length to antennae; labellum relatively small and rounded; antenna (
<figureCitation id="13052A51D23BFFD51345F9105C33EB29" box="[1063,1129,1775,1801]" captionStart="FIGURE 1" captionStartId="4.[264,367,1766,1790]" captionTargetBox="[276,1317,624,1746]" captionTargetId="figure@4.[264,1323,614,1756]" captionTargetPageId="4" captionText="FIGURE 1. Cyrtosathe kirkspriggsi gen. et sp. nov.: Male: A, head, lateral; B, same anterior. Female: C, head lateral; D, same anterior; E, antenna, right lateral. F, abdominal tergite 2 sensory setae patch (enlarged at right). Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/171328/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
E) brown, overlain with fine silver pruinescence; scape and pedicel short, broad; flagellum
<date id="FF801014D23BFFD513F5F8E85CEEEB11" box="[1175,1204,1815,1841]" pageId="3" pageNumber="4">2x</date>
length of pedicel, flattened, apex pointed; style apical, single segment.
</paragraph>
<paragraph id="8B8136D4D23CFFD2165AFEE359EDEE23" blockId="4.[264,1323,284,515]" pageId="4" pageNumber="5">
<emphasis id="B94AEAC6D23CFFD2165AFEE359D6ED15" box="[312,396,284,309]" italics="true" pageId="4" pageNumber="5">Thorax</emphasis>
. Glossy black, glabrous except for fine, pale setae on katepisternum and short dark setae on proepimeron and scutellum; macrosetae absent; wing (
<figureCitation id="13052A51D23CFFD21349FEBB5C2BED7E" box="[1067,1137,324,350]" captionStart="FIGURE 2" captionStartId="5.[264,368,899,923]" captionTargetBox="[264,1322,319,873]" captionTargetId="figure@5.[207,1379,309,900]" captionTargetPageId="5" captionText="FIGURE 2. Cyrtosathe kirkspriggsi gen. et sp. nov.: Wing. Scale line: 0.2 mm." httpUri="https://zenodo.org/record/171329/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
) hyaline, venation pale, venation in posterior half of wing poorly sclerotised and difficult to detect; costal vein ending at R5 vein; vein M2 present, extending from middle of discal cell; haltere stem brown, knob pale cream­white; legs dark brown to black, tibiae pale yellow, covered with fine pale setae.
</paragraph>
<caption id="DF41665CD23CFFD2166AF9195B76EB63" httpUri="https://zenodo.org/record/171328/files/figure.png" pageId="4" pageNumber="5" targetBox="[276,1317,624,1746]" targetPageId="4">
<paragraph id="8B8136D4D23CFFD2166AF9195B76EB63" blockId="4.[264,1323,1766,1859]" pageId="4" pageNumber="5">
<emphasis id="B94AEAC6D23CFFD2166AF91959D2EADE" bold="true" box="[264,392,1766,1790]" pageId="4" pageNumber="5">FIGURE 1</emphasis>
.
<taxonomicName id="4C3E4D57D23CFFD216F8F9185AC0EADE" box="[410,666,1767,1790]" class="Insecta" family="Scenopinidae" genus="Cyrtosathe" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kirkspriggsi">
<emphasis id="B94AEAC6D23CFFD216F8F9185AC0EADE" box="[410,666,1767,1790]" italics="true" pageId="4" pageNumber="5">Cyrtosathe kirkspriggsi</emphasis>
</taxonomicName>
gen.
<emphasis id="B94AEAC6D23CFFD215BFF9185AAAEADE" box="[733,752,1767,1790]" italics="true" pageId="4" pageNumber="5">et</emphasis>
sp. nov.: Male: A, head, lateral; B, same anterior. Female: C, head lateral; D, same anterior; E, antenna, right lateral. F, abdominal tergite 2 sensory setae patch (enlarged at right). Scale lines: 0.1 mm.
</paragraph>
</caption>
<caption id="DF41665CD23DFFD3166AFC7C5C0DEFBA" box="[264,1111,899,923]" httpUri="https://zenodo.org/record/171329/files/figure.png" pageId="5" pageNumber="6" targetBox="[264,1322,319,873]" targetPageId="5">
<paragraph id="8B8136D4D23DFFD3166AFC7C5C0DEFBA" blockId="5.[264,1111,899,923]" box="[264,1111,899,923]" pageId="5" pageNumber="6">
<emphasis id="B94AEAC6D23DFFD3166AFC7C59D0EFBB" bold="true" box="[264,394,899,923]" pageId="5" pageNumber="6">FIGURE 2.</emphasis>
<taxonomicName id="4C3E4D57D23DFFD316F3FC7C5AD6EFBA" box="[401,652,899,922]" class="Insecta" family="Scenopinidae" genus="Cyrtosathe" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kirkspriggsi">
<emphasis id="B94AEAC6D23DFFD316F3FC7C5AD6EFBA" box="[401,652,899,922]" italics="true" pageId="5" pageNumber="6">Cyrtosathe kirkspriggsi</emphasis>
</taxonomicName>
gen.
<emphasis id="B94AEAC6D23DFFD315A5FC7C5A80EFBA" box="[711,730,899,922]" italics="true" pageId="5" pageNumber="6">et</emphasis>
sp. nov.: Wing. Scale line: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B8136D4D23DFFD3165AFC115C53E810" blockId="5.[264,1323,1006,1872]" pageId="5" pageNumber="6">
<emphasis id="B94AEAC6D23DFFD3165AFC1159FCE827" box="[312,422,1006,1031]" italics="true" pageId="5" pageNumber="6">Abdomen</emphasis>
. Glossy black, sparsely covered with elongate, pale setae in male; tergite 2 with two medial hemispherical patches of sensory setae (
<figureCitation id="13052A51D23DFFD314C0FBE95BBEE810" box="[930,996,1046,1072]" captionStart="FIGURE 1" captionStartId="4.[264,367,1766,1790]" captionTargetBox="[276,1317,624,1746]" captionTargetId="figure@4.[264,1323,614,1756]" captionTargetPageId="4" captionText="FIGURE 1. Cyrtosathe kirkspriggsi gen. et sp. nov.: Male: A, head, lateral; B, same anterior. Female: C, head lateral; D, same anterior; E, antenna, right lateral. F, abdominal tergite 2 sensory setae patch (enlarged at right). Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/171328/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
F).
</paragraph>
<paragraph id="8B8136D4D23DFFD3165AFBC15CCFEA18" blockId="5.[264,1323,1006,1872]" pageId="5" pageNumber="6">
<emphasis id="B94AEAC6D23DFFD3165AFBC159B1E877" box="[312,491,1086,1111]" italics="true" pageId="5" pageNumber="6">Male Genitalia</emphasis>
. Genitalia aligned with body axis (i.e. epandrium dorsal), not rotated; epandrium (
<figureCitation id="13052A51D23DFFD316F5FB9959B8E8A0" box="[407,482,1126,1152]" captionStart="FIGURE 3" captionStartId="6.[264,368,1452,1476]" captionTargetBox="[267,1309,397,1413]" captionTargetId="figure@6.[264,1323,386,1425]" captionTargetPageId="6" captionText="FIGURE 3. Cyrtosathe kirkspriggsi gen. et sp. nov.: Male genitalia: A. Epandrium, dorsal; B, aedeagus and gonocoxites (epandrium and vestiture removed), dorso­lateral view; C, same, lateral; D, detail of basal portion of aedeagus, lateral; E, aedeagus and gonocoxites, dorsal; F, detail of gonocoxites. lateral, Scale line: 0.1 mm. Abbreviations: c, cercus; d, distiphallus; da, dorsal apodeme of parameral sheath; e, epandrium; ea, ejaculatory apodeme; g, gonocoxite; ga, gonocoxal apodeme; gs, gonostylus; h, hypandrium; hy, hypoproct." httpUri="https://zenodo.org/record/171330/files/figure.png" pageId="5" pageNumber="6">Figs 3</figureCitation>
A, C) quadrangular, arched, covering terminalia, split medially in two sections; cerci short, rounded; gonocoxites (
<figureCitation id="13052A51D23DFFD3146DFB715B02E888" box="[783,856,1166,1192]" captionStart="FIGURE 3" captionStartId="6.[264,368,1452,1476]" captionTargetBox="[267,1309,397,1413]" captionTargetId="figure@6.[264,1323,386,1425]" captionTargetPageId="6" captionText="FIGURE 3. Cyrtosathe kirkspriggsi gen. et sp. nov.: Male genitalia: A. Epandrium, dorsal; B, aedeagus and gonocoxites (epandrium and vestiture removed), dorso­lateral view; C, same, lateral; D, detail of basal portion of aedeagus, lateral; E, aedeagus and gonocoxites, dorsal; F, detail of gonocoxites. lateral, Scale line: 0.1 mm. Abbreviations: c, cercus; d, distiphallus; da, dorsal apodeme of parameral sheath; e, epandrium; ea, ejaculatory apodeme; g, gonocoxite; ga, gonocoxal apodeme; gs, gonostylus; h, hypandrium; hy, hypoproct." httpUri="https://zenodo.org/record/171330/files/figure.png" pageId="5" pageNumber="6">Figs 3</figureCitation>
BC, EF) elongate with spatulate posterior process and a short, strong seta dorsally; gonocoxites widely separated medially by a large rounded bowl­like hypandrium; hypandrium with paired posterior processes and medial process (
<figureCitation id="13052A51D23DFFD316B1FAF95A78E900" box="[467,546,1286,1312]" captionStart="FIGURE 3" captionStartId="6.[264,368,1452,1476]" captionTargetBox="[267,1309,397,1413]" captionTargetId="figure@6.[264,1323,386,1425]" captionTargetPageId="6" captionText="FIGURE 3. Cyrtosathe kirkspriggsi gen. et sp. nov.: Male genitalia: A. Epandrium, dorsal; B, aedeagus and gonocoxites (epandrium and vestiture removed), dorso­lateral view; C, same, lateral; D, detail of basal portion of aedeagus, lateral; E, aedeagus and gonocoxites, dorsal; F, detail of gonocoxites. lateral, Scale line: 0.1 mm. Abbreviations: c, cercus; d, distiphallus; da, dorsal apodeme of parameral sheath; e, epandrium; ea, ejaculatory apodeme; g, gonocoxite; ga, gonocoxal apodeme; gs, gonostylus; h, hypandrium; hy, hypoproct." httpUri="https://zenodo.org/record/171330/files/figure.png" pageId="5" pageNumber="6">Figs 3</figureCitation>
BC, EF); gonostylus elongate, narrow, apical third upturned slightly; gonocoxal apodeme elongate, equal to gonocoxite in length, curved outwards then recurved inwards at apex around dorsal apodeme of parameral sheath, apex spatulate; aedeagus extended anteriorly beyond gonocoxites, folded dorsally on itself, distiphallus bifid, greatly narrowed, elongate, slightly coiled, ribbon­like lateral flanges near base; dorsal apodeme of parameral sheath large, ladle­shaped with lateral wings at base, cradling ejaculatory apodeme (
<figureCitation id="13052A51D23DFFD31569FA095A0EEA30" box="[523,596,1526,1552]" captionStart="FIGURE 3" captionStartId="6.[264,368,1452,1476]" captionTargetBox="[267,1309,397,1413]" captionTargetId="figure@6.[264,1323,386,1425]" captionTargetPageId="6" captionText="FIGURE 3. Cyrtosathe kirkspriggsi gen. et sp. nov.: Male genitalia: A. Epandrium, dorsal; B, aedeagus and gonocoxites (epandrium and vestiture removed), dorso­lateral view; C, same, lateral; D, detail of basal portion of aedeagus, lateral; E, aedeagus and gonocoxites, dorsal; F, detail of gonocoxites. lateral, Scale line: 0.1 mm. Abbreviations: c, cercus; d, distiphallus; da, dorsal apodeme of parameral sheath; e, epandrium; ea, ejaculatory apodeme; g, gonocoxite; ga, gonocoxal apodeme; gs, gonostylus; h, hypandrium; hy, hypoproct." httpUri="https://zenodo.org/record/171330/files/figure.png" pageId="5" pageNumber="6">Figs 3</figureCitation>
BE); lateral ejaculatory apodemes not clearly evident; ejaculatory apodeme narrow with greatly enlarged paddle­shaped apex posteriorly.
</paragraph>
<paragraph id="8B8136D4D23DFFD0165AF9B959A5ED7E" blockId="5.[264,1323,1006,1872]" lastBlockId="6.[264,1323,284,350]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="B94AEAC6D23DFFD3165AF9B95A5CEA7F" box="[312,518,1606,1631]" italics="true" pageId="5" pageNumber="6">Female Genitalia</emphasis>
. Tergite 8 covered with erect, elongate pale setae arranged in a ringlike pattern (i.e. glabrous in the centre) (
<figureCitation id="13052A51D23DFFD315BFF9915B79EAA8" box="[733,803,1646,1672]" captionStart="FIGURE 4" captionStartId="7.[264,368,1703,1727]" captionTargetBox="[353,1214,294,1685]" captionTargetId="figure@7.[340,1222,283,1688]" captionTargetPageId="7" captionText="FIGURE 4. Cyrtosathe kirkspriggsi gen. et sp. nov.: Female genitalia: A, Genitalia in contacted position, postero­lateral; B, detail of same with acanthophorites slightly relaxed posteriorly, lateral; C, same, showing distal reproductive system (tergite 8 cut away and vestiture removed), dorsal. Scale lines: 0.1 mm. Abbreviations: A 1, acanthophorite macrosetae; ag, accessory gland duct (gland cut away); f, furca; ss, spermathecal sac; s, spermatheca; sd, spermathecal duct; s 8, sternite 8; s 10, sternite 10; t 8 ­ t 10, tergites 8 to 10." httpUri="https://zenodo.org/record/171331/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
A); tergite 9+10 joined to furca internally by a bridge formed by tergite 9 (
<figureCitation id="13052A51D23DFFD31500F9695AF4EA90" box="[610,686,1686,1712]" captionStart="FIGURE 4" captionStartId="7.[264,368,1703,1727]" captionTargetBox="[353,1214,294,1685]" captionTargetId="figure@7.[340,1222,283,1688]" captionTargetPageId="7" captionText="FIGURE 4. Cyrtosathe kirkspriggsi gen. et sp. nov.: Female genitalia: A, Genitalia in contacted position, postero­lateral; B, detail of same with acanthophorites slightly relaxed posteriorly, lateral; C, same, showing distal reproductive system (tergite 8 cut away and vestiture removed), dorsal. Scale lines: 0.1 mm. Abbreviations: A 1, acanthophorite macrosetae; ag, accessory gland duct (gland cut away); f, furca; ss, spermathecal sac; s, spermatheca; sd, spermathecal duct; s 8, sternite 8; s 10, sternite 10; t 8 ­ t 10, tergites 8 to 10." httpUri="https://zenodo.org/record/171331/files/figure.png" pageId="5" pageNumber="6">Figs 4</figureCitation>
BC); acanthophorites with four to six stout A1 setae; furca (
<figureCitation id="13052A51D23DFFD3163BF94159FFEAF8" box="[345,421,1726,1752]" captionStart="FIGURE 4" captionStartId="7.[264,368,1703,1727]" captionTargetBox="[353,1214,294,1685]" captionTargetId="figure@7.[340,1222,283,1688]" captionTargetPageId="7" captionText="FIGURE 4. Cyrtosathe kirkspriggsi gen. et sp. nov.: Female genitalia: A, Genitalia in contacted position, postero­lateral; B, detail of same with acanthophorites slightly relaxed posteriorly, lateral; C, same, showing distal reproductive system (tergite 8 cut away and vestiture removed), dorsal. Scale lines: 0.1 mm. Abbreviations: A 1, acanthophorite macrosetae; ag, accessory gland duct (gland cut away); f, furca; ss, spermathecal sac; s, spermatheca; sd, spermathecal duct; s 8, sternite 8; s 10, sternite 10; t 8 ­ t 10, tergites 8 to 10." httpUri="https://zenodo.org/record/171331/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
C) circular, lightly sclerotised; sternite 8 bowl shaped; two spermathecae present; spermathecal ducts thickened basally, narrower distally, with reticulated tissue patterning; spermatheca membranous, truncated distally with thickened marginal band; spermathecal sac very small; spermathecae join to roof of bursa separately from and immediately posterior of spermathecal sac duct; accessory glands join to bursa just posterior to spermathecae.
</paragraph>
</subSubSection>
<caption id="DF41665CD23EFFD0166AFA535B05EA51" httpUri="https://zenodo.org/record/171330/files/figure.png" pageId="6" pageNumber="7" targetBox="[267,1309,397,1413]" targetPageId="6">
<paragraph id="8B8136D4D23EFFD0166AFA535B05EA51" blockId="6.[264,1323,1452,1649]" pageId="6" pageNumber="7">
<emphasis id="B94AEAC6D23EFFD0166AFA5359D4E9E4" bold="true" box="[264,398,1452,1476]" pageId="6" pageNumber="7">FIGURE 3.</emphasis>
<taxonomicName id="4C3E4D57D23EFFD016F8FA535AC2E9E3" box="[410,664,1452,1475]" class="Insecta" family="Scenopinidae" genus="Cyrtosathe" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kirkspriggsi">
<emphasis id="B94AEAC6D23EFFD016F8FA535AC2E9E3" box="[410,664,1452,1475]" italics="true" pageId="6" pageNumber="7">Cyrtosathe kirkspriggsi</emphasis>
</taxonomicName>
gen.
<emphasis id="B94AEAC6D23EFFD015BEFA535AB5E9E3" box="[732,751,1452,1475]" italics="true" pageId="6" pageNumber="7">et</emphasis>
sp. nov.: Male genitalia: A. Epandrium, dorsal; B, aedeagus and gonocoxites (epandrium and vestiture removed), dorso­lateral view; C, same, lateral; D, detail of basal portion of aedeagus, lateral; E, aedeagus and gonocoxites, dorsal; F, detail of gonocoxites. lateral, Scale line: 0.1 mm. Abbreviations:
<emphasis id="B94AEAC6D23EFFD0140DF9EB5B21EA0B" box="[879,891,1556,1579]" italics="true" pageId="6" pageNumber="7">c</emphasis>
, cercus;
<emphasis id="B94AEAC6D23EFFD014BDF9EB5BB6EA0B" box="[991,1004,1556,1579]" italics="true" pageId="6" pageNumber="7">d</emphasis>
, distiphallus;
<emphasis id="B94AEAC6D23EFFD013E5F9EB5CF8EA0B" box="[1159,1186,1556,1579]" italics="true" pageId="6" pageNumber="7">da</emphasis>
, dorsal apodeme of parameral sheath;
<emphasis id="B94AEAC6D23EFFD01556F9C85A1AEA6E" box="[564,576,1591,1614]" italics="true" pageId="6" pageNumber="7">e</emphasis>
, epandrium;
<emphasis id="B94AEAC6D23EFFD015B0F9C85AB1EA6E" box="[722,747,1591,1614]" italics="true" pageId="6" pageNumber="7">ea</emphasis>
, ejaculatory apodeme;
<emphasis id="B94AEAC6D23EFFD0148EF9C85BA3EA6E" box="[1004,1017,1591,1614]" italics="true" pageId="6" pageNumber="7">g</emphasis>
, gonocoxite;
<emphasis id="B94AEAC6D23EFFD013F3F9C85CF6EA6E" box="[1169,1196,1591,1614]" italics="true" pageId="6" pageNumber="7">ga</emphasis>
, gonocoxal apodeme;
<emphasis id="B94AEAC6D23EFFD01614F9A559D4EA51" box="[374,398,1626,1649]" italics="true" pageId="6" pageNumber="7">gs</emphasis>
, gonostylus;
<emphasis id="B94AEAC6D23EFFD0157FF9A55A70EA51" box="[541,554,1626,1649]" italics="true" pageId="6" pageNumber="7">h</emphasis>
, hypandrium;
<emphasis id="B94AEAC6D23EFFD015A5F9A55A84EA51" box="[711,734,1626,1649]" italics="true" pageId="6" pageNumber="7">hy</emphasis>
, hypoproct.
</paragraph>
</caption>
<subSubSection id="C324655FD23EFFD0165AF95B5B75EB7E" pageId="6" pageNumber="7" type="discussion">
<paragraph id="8B8136D4D23EFFD0165AF95B5B75EB7E" blockId="6.[264,1324,1700,1886]" pageId="6" pageNumber="7">
<emphasis id="B94AEAC6D23EFFD0165AF95B59E5EA9E" bold="true" box="[312,447,1700,1726]" pageId="6" pageNumber="7">Comments</emphasis>
.
<taxonomicName id="4C3E4D57D23EFFD016AFF95A5AB8EA9E" box="[461,738,1701,1726]" class="Insecta" family="Scenopinidae" genus="Cyrtosathe" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kirkspriggsi">
<emphasis id="B94AEAC6D23EFFD016AFF95A5AB8EA9E" box="[461,738,1701,1726]" italics="true" pageId="6" pageNumber="7">Cyrtosathe kirkspriggsi</emphasis>
</taxonomicName>
gen.
<emphasis id="B94AEAC6D23EFFD01441F95A5B62EA9E" box="[803,824,1701,1726]" italics="true" pageId="6" pageNumber="7">et</emphasis>
sp. nov. is known from a single collecting locality in the Luderitz District,
<collectingCountry id="F3297644D23EFFD015E7F9335AB4EAC6" box="[645,750,1740,1766]" name="Namibia" pageId="6" pageNumber="7">Namibia</collectingCountry>
. All adults were collected by Malaise and pitfall traps. As with all other non­scenopinine window flies, the immature stages are unknown. The
<typeStatus id="54858876D23EFFD0165EF8E35935EB16" box="[316,367,1820,1846]" pageId="6" pageNumber="7">type</typeStatus>
series is only in fair condition as most specimens are shriveled, presumably when they were removed from alcohol and mounted.
</paragraph>
</subSubSection>
</treatment>
</document>