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<document id="44403A3BE6A6D4B6F114607F2C91147E" ID-DOI="10.18590/euscorpius.2010.vol2010.iss103.1" ID-ISSN="1536-9307" ID-Zenodo-Dep="12782242" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721369394532" checkinUser="felipe" docAuthor="Lowe, Graeme" docDate="2010" docId="039787C18605FFA2BE3DFE109B65F995" docLanguage="en" docName="Euscorpius.103.1-23.pdf" docOrigin="Euscorpius 103 (103)" docSource="https://mds.marshall.edu/euscorpius/vol2010/iss103/1/" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Hottentotta pellucidus Lowe, 2010, sp.nov." docType="treatment" docUuid="18B24A3E-CBA1-41C2-8295-EEA20A22B79C" docUuidSource="ZooBank" docVersion="3" lastPageNumber="8" masterDocId="FFAEFFB98606FFABBF04FFBA9E09FFEA" masterDocTitle="Two new species of Hottentotta Birula, 1908 (Scorpiones: Buthidae) from northern Oman" masterLastPageNumber="23" masterPageNumber="1" pageNumber="2" updateTime="1721382510913" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="EA5044FCBA7925026ABA708458C9E58D">Two new species of Hottentotta Birula, 1908 (Scorpiones: Buthidae) from northern Oman</mods:title>
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<mods:namePart id="D3E40D2EDE1A292D921A54EBFE48469A">Lowe, Graeme</mods:namePart>
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<mods:identifier id="9F72E1143B539A03D8C779A8AA8B5A69" type="ZooBank">44503249-44BC-4E7B-9EA8-1FDAAC84B559</mods:identifier>
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<treatment id="039787C18605FFA2BE3DFE109B65F995" ID-DOI="http://doi.org/10.5281/zenodo.12782111" ID-Zenodo-Dep="12782111" LSID="urn:lsid:zoobank.org:act:18B24A3E-CBA1-41C2-8295-EEA20A22B79C" httpUri="http://treatment.plazi.org/id/039787C18605FFA2BE3DFE109B65F995" lastPageId="9" lastPageNumber="8" pageId="3" pageNumber="2">
<subSubSection id="C324655C8605FFA8BE3DFE109CA0FE2F" box="[313,681,425,453]" pageId="3" pageNumber="2" type="nomenclature">
<paragraph id="8B8136D78605FFA8BE3DFE109CA0FE2F" blockId="3.[280,715,425,482]" box="[313,681,425,453]" pageId="3" pageNumber="2">
<heading id="D0C981BB8605FFA8BE3DFE109CA0FE2F" box="[313,681,425,453]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="2" reason="2">
<taxonomicName id="4C3E4D548605FFA8BE3DFE109C4CFE2E" ID-CoL="3MQY3" authorityName="Lowe" authorityYear="2010" box="[313,581,425,452]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="pellucidus" status="sp. nov.">
<emphasis id="B94AEAC58605FFA8BE3DFE109C4CFE2E" bold="true" box="[313,581,425,452]" italics="true" pageId="3" pageNumber="2">Hottentotta pellucidus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27957BE8605FFA8BD50FE119CA0FE2F" box="[596,681,427,453]" pageId="3" pageNumber="2" rank="species">sp.nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C324655C8605FFA8BE1CFE719C1DFAF4" pageId="3" pageNumber="2" type="description">
<paragraph id="8B8136D78605FFA8BE1CFE719CC2FE08" blockId="3.[280,715,425,482]" box="[280,715,459,482]" pageId="3" pageNumber="2">
<figureCitation id="13052A528605FFA8BE1CFE719F83FE08" box="[280,394,459,482]" captionStart-0="Figures 1–4" captionStart-1="Figures 5–11" captionStart-2="Figures 12–20" captionStart-3="Figures 21–26" captionStartId-0="4.[192,278,1910,1934]" captionStartId-1="6.[192,278,1662,1685]" captionStartId-2="7.[192,278,1626,1649]" captionStartId-3="8.[192,278,991,1014]" captionTargetBox-0="[233,1392,156,1881]" captionTargetBox-1="[228,1382,170,1606]" captionTargetBox-2="[237,1379,217,1541]" captionTargetBox-3="[199,774,194,958]" captionTargetId-0="figure-12@4.[221,1404,156,1881]" captionTargetId-1="figure-10@6.[192,1441,147,1632]" captionTargetId-2="figure-6@7.[192,1442,168,1594]" captionTargetId-3="figure-10@8.[180,804,154,984]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="Figures 1–4: H. pellucidus, sp. nov., habitus, 1–2, holotype male; 3–4, paratype female. 1, 3. Dorsal aspect. 2, 4. Ventral aspect. Scale bar: 10 mm." captionText-1="Figures 5–11: H. pellucidus, sp. nov. 5, 8–11, holotype male; 6–7, paratype male (Jabal Bani Jabir). 5. Carapace and tergites I– II, dorsal aspect. 6–7, right chelicera. 6. Dorsal aspect. 7. Ventral aspect. 8–9, right basitarsus and telotarsus III. 8. Ventral aspect. 9. Retrolateral aspect. 10–11, metasoma and telson. 10. Right lateral aspect. 11. Ventral aspect. Scale bar: 5: 3.19 mm; 6–7: 2.125 mm; 8–9: 1.5 mm; 10–11: 6.22 mm." captionText-2="Figures 12–20: H. pellucidus, sp.nov., 12–16, 19, holotype male; 17–18, paratype male; 20, paratype female. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17–18. Right hemispermatophore, lobes at base of flagellum. 17. Convex aspect. 18. Inner aspect. 19. Left male pectine, ventral aspect. 20. Left female pectine ventral aspect. Scale bar: 12–16, 19–20: 4.1 mm; 17–18: 1.06 mm." captionText-3="Figures 21–26: H. pellucidus, sp. nov., trichobothrial map, holotype male. 21. Right pedipalp femur, dorsal aspect. 22. Right pedipalp femur, proximal internal aspect. 23. Right pedipalp patella, dorsal aspect. 24. Right pedipalp patella, external aspect. 25. Right pedipalp chela, external aspect. 26. Right pedipalp chela, ventral aspect. Trichobothrial notation after Vachon (1974)." figureDoi-0="http://doi.org/10.5281/zenodo.12782244" figureDoi-1="http://doi.org/10.5281/zenodo.12782246" figureDoi-2="http://doi.org/10.5281/zenodo.12782248" figureDoi-3="http://doi.org/10.5281/zenodo.12782250" httpUri-0="https://zenodo.org/record/12782244/files/figure.png" httpUri-1="https://zenodo.org/record/12782246/files/figure.png" httpUri-2="https://zenodo.org/record/12782248/files/figure.png" httpUri-3="https://zenodo.org/record/12782250/files/figure.png" pageId="3" pageNumber="2">Figs. 1–26</figureCitation>
,
<figureCitation id="13052A528605FFA8BE91FE719FD2FE08" box="[405,475,459,482]" captionStart="Figures 53–56" captionStartId="17.[192,278,1091,1114]" captionTargetBox="[206,1424,171,1059]" captionTargetId="figure-6@17.[192,1439,168,1059]" captionTargetPageId="17" captionText="Figures 53–56: Pedipalp finger dentition of new Hottentotta species. 53–54, H. pellucidus, sp. nov., holotype male. 55–56, H. saxinatans, sp. nov., holotype male. 53, 55: right fixed finger, ventral aspect. 54, 56: right movable finger, dorsal aspect. Scale bar: 1 mm. Images acquired under UV fluorescence." figureDoi="http://doi.org/10.5281/zenodo.12782262" httpUri="https://zenodo.org/record/12782262/files/figure.png" pageId="3" pageNumber="2">53–54</figureCitation>
,
<figureCitation id="13052A528605FFA8BEE2FE719C23FE08" box="[486,554,459,482]" captionStart="Figures 57–64" captionStartId="19.[192,278,1836,1860]" captionTargetBox="[201,1429,158,1805]" captionTargetId="figure-6@19.[192,1439,154,1805]" captionTargetPageId="19" captionText="Figures 57–64: In vivo habitus and habitats of new Hottentotta species in Oman. 57–59. H. pellucidus, sp. nov. 57–58. Paratype male. 59. Paratype female. 60. H. saxinatans, sp. nov., paratype female. 61–63. Shir Plateau, Jabal Bani Jabir, type locality of H. pellucidus. 61–62. Terrain photos showing extensive fields of loose rock. 63. Prehistoric tower tomb constructed from broken rock.64. Jabal Shams, type locality of H. saxinatans." figureDoi="http://doi.org/10.5281/zenodo.12782264" httpUri="https://zenodo.org/record/12782264/files/figure.png" pageId="3" pageNumber="2">57–59</figureCitation>
,
<figureCitation id="13052A528605FFA8BD32FE719C72FE08" box="[566,635,459,482]" captionStart="Figures 65–66" captionStartId="20.[192,278,726,749]" captionTargetBox="[200,1434,148,689]" captionTargetId="figure-10@20.[192,1440,144,695]" captionTargetPageId="20" captionText="Figures 65–66: Maps showing collection sites of new species of Hottentotta in northern Oman. 65. Records plotted on larger scale terrain and elevation map of southeast Arabia with geopolitical borders. 66. Records plotted on color topographic map of northern Oman showing locations in the Al Hajar mountain ranges. Magenta circles: H. pellucidus, sp. nov.; red circles: H. saxinatans, sp. nov. The isolated record of H. pellucidus at Muscat may be spurious." figureDoi="http://doi.org/10.5281/zenodo.12782266" httpUri="https://zenodo.org/record/12782266/files/figure.png" pageId="3" pageNumber="2">65–66</figureCitation>
,
<tableCitation id="C6BC036C8605FFA8BD82FE719CC2FE08" box="[646,715,459,482]" captionStart="Table 1" captionStartId="16.[192,257,1818,1841]" captionText="Table 1: Variation in meristics and selected morphometric ratios for Hottentotta pellucidus, sp.nov. and Hottentotta saxinatans, sp.nov. Indicated are ranges, mean ± standard deviation with sample sizes in parentheses. All samples are from adults except pedipalp denticle subrow counts and pectinal tooth counts, which also include measurements from immatures and from both left and right appendages. Values of variables significantly different between the two sexes (P &lt;0.01 for H. pellucidus, P &lt;0.05 for H. saxinatans; Mann Whitney test) are cited separately for males and females, and those not significantly different are pooled for both sexes." pageId="3" pageNumber="2">Tab. 1</tableCitation>
</paragraph>
<paragraph id="8B8136D78605FFA8BFF4FDB39C54FD70" blockId="3.[192,802,521,1310]" pageId="3" pageNumber="2">
<materialsCitation id="3B563C8A8605FFA8BFF4FDB39C54FD70" collectingDate="1995-09-14" collectionCode="UV, NHMB" collectorName="G. Lowe &amp; Lowe, M. D &amp; J. Dundon" country="Oman" elevation="1620" latitude="59.0245" location="Jabal Bani Jabir" pageId="3" pageNumber="2" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B94AEAC58605FFA8BFF4FDB39F5BFDCA" box="[240,338,521,544]" italics="true" pageId="3" pageNumber="2">
<typeStatus id="548588758605FFA8BFF4FDB39F5BFDCA" box="[240,338,521,544]" pageId="3" pageNumber="2" type="holotype">Holotype</typeStatus>
</emphasis>
: adult
<specimenCount id="9D38FD5E8605FFA8BEB1FDB39FC0FDCA" box="[437,457,521,544]" count="1" pageId="3" pageNumber="2" type="male">♂</specimenCount>
,
<collectingCountry id="F32976478605FFA8BED8FDB39C29FDCA" box="[476,544,521,544]" name="Oman" pageId="3" pageNumber="2">Oman</collectingCountry>
,
<location id="8EE1600C8605FFA8BD35FDB39CE2FDCA" LSID="urn:lsid:plazi:treatment:039787C18605FFA2BE3DFE109B65F995:8EE1600C8605FFA8BD35FDB39CE2FDCA" box="[561,747,521,544]" country="Oman" latitude="59.0245" name="Jabal Bani Jabir" pageId="3" pageNumber="2">Jabal Bani Jabir</location>
,
<collectionCode id="ED2FAE128605FFA8BDFFFDB39D28FDCA" box="[763,801,521,544]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="3" pageNumber="2">UV</collectionCode>
detection on rocky terrain, rock strewn plateau with small wadis, on bare rock, 22°49.76'
<geoCoordinate id="EE0A50108605FFA8BD4FFDFC9CD9FDB7" box="[587,720,582,605]" degrees="59" direction="north" minutes="01.47" orientation="latitude" pageId="3" pageNumber="2" precision="9" value="59.0245">N 59°01.47'</geoCoordinate>
E,
<elevation id="0013D1E48605FFA8BDE8FDFC9F1DFD96" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="3" pageNumber="2" unit="m" value="1620.0">
<quantity id="4CC69B328605FFA8BDE8FDFC9EDCFD96" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="3" pageNumber="2" unit="m" value="1620.0">1620 m</quantity>
a.s.l.
</elevation>
,
<date id="FF8010178605FFA8BE23FDDF9C0FFD96" box="[295,518,613,636]" pageId="3" pageNumber="2" value="1995-09-14">
<collectingDate id="EFC4E9FF8605FFA8BE23FDDF9C0FFD96" box="[295,518,613,636]" pageId="3" pageNumber="2" value="1995-09-14">14 September 1995</collectingDate>
</date>
, 23:10 h, leg.
<collectorName id="26CB53018605FFA8BDB3FDDF9D14FD96" box="[695,797,613,636]" pageId="3" pageNumber="2">G. Lowe</collectorName>
, M.D. Gallagher &amp;
<collectorName id="26CB53018605FFA8BE8EFD399FF2FD70" box="[394,507,643,666]" pageId="3" pageNumber="2">J. Dundon</collectorName>
,
<collectionCode id="ED2FAE128605FFA8BD01FD399C50FD70" box="[517,601,643,666]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="3" pageNumber="2">NHMB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8136D78605FFA8BFF4FD189C1DFAF4" blockId="3.[192,802,521,1310]" pageId="3" pageNumber="2">
<materialsCitation id="3B563C8A8605FFA8BFF4FD189D00FD1C" collectingDate="1992-11-06" collectionCode="NHMB" collectorName="A. S. Gardner" country="Oman" elevation="1650" latitude="59.033333" location="Jabal Bani Jabir" pageId="3" pageNumber="2" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B94AEAC58605FFA8BFF4FD189F55FD53" box="[240,348,674,697]" italics="true" pageId="3" pageNumber="2">
<typeStatus id="548588758605FFA8BFF4FD189F55FD53" box="[240,348,674,697]" pageId="3" pageNumber="2" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="F32976478605FFA8BE77FD189FB1FD53" box="[371,440,674,697]" name="Oman" pageId="3" pageNumber="2">Oman</collectingCountry>
:
<specimenCount id="9D38FD5E8605FFA8BECFFD189FF5FD53" box="[459,508,674,697]" count="1" pageId="3" pageNumber="2" type="male">1 ♂</specimenCount>
,
<location id="8EE1600C8605FFA8BD16FD189CDBFD53" LSID="urn:lsid:plazi:treatment:039787C18605FFA2BE3DFE109B65F995:8EE1600C8605FFA8BD16FD189CDBFD53" box="[530,722,674,697]" country="Oman" latitude="59.033333" name="Jabal Bani Jabir" pageId="3" pageNumber="2">Jabal Bani Jabir</location>
, under stone, limestone plateau, 22°50'
<geoCoordinate id="EE0A50108605FFA8BD17FD7B9C7CFD32" box="[531,629,705,728]" degrees="59" direction="north" minutes="02" orientation="latitude" pageId="3" pageNumber="2" precision="925" value="59.033333">N 59°02'</geoCoordinate>
E,
<elevation id="0013D1E48605FFA8BD95FD7B9D15FD32" box="[657,796,705,728]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="3" pageNumber="2" unit="m" value="1650.0">
<quantity id="4CC69B328605FFA8BD95FD7B9CEBFD32" box="[657,738,705,728]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="3" pageNumber="2" unit="m" value="1650.0">1650 m</quantity>
a.s.l.
</elevation>
,
<date id="FF8010178605FFA8BFC4FD659F8DFD1C" box="[192,388,735,758]" pageId="3" pageNumber="2" value="1992-11-06">
<collectingDate id="EFC4E9FF8605FFA8BFC4FD659F8DFD1C" box="[192,388,735,758]" pageId="3" pageNumber="2" value="1992-11-06">6 November 1992</collectingDate>
</date>
, leg.
<collectorName id="26CB53018605FFA8BEB9FD659C46FD1C" box="[445,591,735,758]" pageId="3" pageNumber="2">A.S. Gardner</collectorName>
, SQU41,
<collectionCode id="ED2FAE128605FFA8BDB2FD659D00FD1C" box="[694,777,735,758]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="3" pageNumber="2">NHMB</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B563C8A8605FFA8BC11FD659FD4FCFF" collectionCode="UV" collectorName="Jabal Bani Jabir" country="Oman" pageId="3" pageNumber="2" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D38FD5E8605FFA8BC11FD659EDDFCFF" count="2" pageId="3" pageNumber="2" type="male">2 ♂</specimenCount>
,
<collectorName id="26CB53018605FFA8BFECFD449FADFCFF" box="[232,420,766,789]" pageId="3" pageNumber="2">Jabal Bani Jabir</collectorName>
,
<collectionCode id="ED2FAE128605FFA8BEB2FD449FD4FCFF" box="[438,477,766,789]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="3" pageNumber="2">UV</collectionCode>
</materialsCitation>
<materialsCitation id="3B563C8A8605FFA8BEEEFD449F6EFCB8" collectingDate="1995-09" collectionCode="ONHM" collectorName="A. Francois &amp; C. Delise" country="Oman" elevation="1800" latitude="59.033333" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
detection, 22°49'
<geoCoordinate id="EE0A50108605FFA8BDA0FD449D04FCFF" box="[676,781,766,789]" degrees="59" direction="north" minutes="02" orientation="latitude" pageId="3" pageNumber="2" precision="925" value="59.033333">N 59°02'</geoCoordinate>
E,
<elevation id="0013D1E48605FFA8BFC4FCA79F5AFCDE" box="[192,339,797,820]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="2" unit="m" value="1800.0">
<quantity id="4CC69B328605FFA8BFC4FCA79F1FFCDE" box="[192,278,797,820]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="2" unit="m" value="1800.0">1800 m</quantity>
a.s.l.
</elevation>
,
<date id="FF8010178605FFA8BE61FCA79C13FCDE" box="[357,538,797,820]" pageId="3" pageNumber="2" value="1995-09">
<collectingDate id="EFC4E9FF8605FFA8BE61FCA79C13FCDE" box="[357,538,797,820]" pageId="3" pageNumber="2" value="1995-09">September 1995</collectingDate>
</date>
, leg.
<collectorName id="26CB53018605FFA8BD58FCA79CD7FCDE" box="[604,734,797,820]" pageId="3" pageNumber="2">A. François</collectorName>
&amp;
<collectorName id="26CB53018605FFA8BC0DFCA79F0EFCB8" pageId="3" pageNumber="2">C. Delise</collectorName>
,
<collectionCode id="ED2FAE128605FFA8BE16FC819F6EFCB8" box="[274,359,827,850]" pageId="3" pageNumber="2">ONHM</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B563C8A8605FFA8BE76FC819EEEFC9B" collectionCode="UV" collectorName="Jabal Bani Jabir" country="Oman" pageId="3" pageNumber="2" specimenCount="26" specimenCount-female="10" specimenCount-juvenile="7" specimenCount-male="9" typeStatus="paratype">
<specimenCount id="9D38FD5E8605FFA8BE76FC819F92FCB8" box="[370,411,827,850]" count="9" pageId="3" pageNumber="2" type="male">9 ♂</specimenCount>
,
<specimenCount id="9D38FD5E8605FFA8BEADFC819FD6FCB8" box="[425,479,827,850]" count="10" pageId="3" pageNumber="2" type="female">10 ♀</specimenCount>
,
<specimenCount id="9D38FD5E8605FFA8BEE9FC819C6DFCB8" box="[493,612,827,850]" count="7" pageId="3" pageNumber="2" type="juvenile">7 juveniles</specimenCount>
,
<collectorName id="26CB53018605FFA8BD6BFC819D17FCB8" box="[623,798,827,850]" pageId="3" pageNumber="2">Jabal Bani Jabir</collectorName>
,
<collectionCode id="ED2FAE128605FFA8BFC4FCE09EEEFC9B" box="[192,231,858,881]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="3" pageNumber="2">UV</collectionCode>
</materialsCitation>
<materialsCitation id="3B563C8A8605FFA8BFF1FCE09FC9FBE0" collectingDate="1995-09-14" collectionCode="NHMB" collectorName="G. Lowe &amp; M. D. Gallagher &amp; J. Dundon" country="Oman" elevation="1640" latitude="59.0265" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
detection, stationary on rocks and ground under rocks, loose rock strewn plateau with small wadis, sparse vegetation, few bushes, grass, region of burial towers, 22°49.6'
<geoCoordinate id="EE0A50108605FFA8BE79FC0C9C1BFC27" box="[381,530,950,973]" degrees="59" direction="north" minutes="01.59" orientation="latitude" pageId="3" pageNumber="2" precision="9" value="59.0265">N 59°01.59'</geoCoordinate>
E,
<elevation id="0013D1E48605FFA8BD3AFC0C9CE0FC27" box="[574,745,950,973]" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="3" pageNumber="2" unit="m" value="1640.0">
<quantity id="4CC69B328605FFA8BD3AFC0C9CA9FC27" box="[574,672,950,973]" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="3" pageNumber="2" unit="m" value="1640.0">1640 m</quantity>
a.s.l.
</elevation>
,
<date id="FF8010178605FFA8BC03FC0C9F7BFC06" pageId="3" pageNumber="2" value="1995-09-14">
<collectingDate id="EFC4E9FF8605FFA8BC03FC0C9F7BFC06" pageId="3" pageNumber="2" value="1995-09-14">14 September 1995</collectingDate>
</date>
, 19:40 h, leg.
<collectorName id="26CB53018605FFA8BD08FC6F9C64FC06" box="[524,621,981,1004]" pageId="3" pageNumber="2">G. Lowe</collectorName>
,
<collectorName id="26CB53018605FFA8BD7DFC6F9D28FC06" box="[633,801,981,1004]" pageId="3" pageNumber="2">M.D. Gallagher</collectorName>
&amp;
<collectorName id="26CB53018605FFA8BFE7FC499F52FBE0" box="[227,347,1011,1034]" pageId="3" pageNumber="2">J. Dundon</collectorName>
,
<collectionCode id="ED2FAE128605FFA8BE69FC499FC9FBE0" box="[365,448,1011,1034]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="3" pageNumber="2">NHMB</collectionCode>
</materialsCitation>
, GL, ONHM;
<materialsCitation id="3B563C8A8605FFA8BD75FC499F6EFBA2" collectingDate="1995-09-14" collectionCode="NHMB, MNHN, BMNH, FKCP" collectorName="G. Lowe &amp; Lowe, M. D &amp; J. Dundon" country="Oman" elevation="1620" latitude="59.0245" location="Jabal Bani Jabir" pageId="3" pageNumber="2" specimenCount="32" specimenCount-female="7" specimenCount-juvenile="10" specimenCount-male="15" typeStatus="paratype">
<specimenCount id="9D38FD5E8605FFA8BD75FC499CA7FBE0" box="[625,686,1011,1034]" count="15" pageId="3" pageNumber="2" type="male">15 ♂</specimenCount>
,
<specimenCount id="9D38FD5E8605FFA8BDC7FC499CFBFBE0" box="[707,754,1011,1034]" count="7" pageId="3" pageNumber="2" type="female">7 ♀</specimenCount>
,
<specimenCount id="9D38FD5E8605FFA8BC03FC499F2AFBC3" count="10" pageId="3" pageNumber="2" type="juvenile">10 juveniles</specimenCount>
, same locality as holotype,
<collectionCode id="ED2FAE128605FFA8BD62FBA89CB0FBC3" box="[614,697,1042,1065]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="3" pageNumber="2">NHMB</collectionCode>
,
<collectionCode id="ED2FAE128605FFA8BDCDFBA89D14FBC3" box="[713,797,1042,1065]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="2">MNHN</collectionCode>
,
<collectionCode id="ED2FAE128605FFA8BFC4FB8B9F1AFBA2" box="[192,275,1073,1096]" pageId="3" pageNumber="2">BMNH</collectionCode>
,
<collectionCode id="ED2FAE128605FFA8BE25FB8B9F6EFBA2" box="[289,359,1073,1096]" pageId="3" pageNumber="2">FKCP</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B563C8A8605FFA8BE72FB8B9C2EFB8C" collectingDate="1998-05" collectionCode="NHMB" collectorName="C. Rose" country="Oman" latitude="58.5" location="Ras Al Hamra" pageId="3" pageNumber="2" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D38FD5E8605FFA8BE72FB8B9FA8FBA2" box="[374,417,1073,1096]" count="1" pageId="3" pageNumber="2" type="female">1 ♀</specimenCount>
,
<location id="8EE1600C8605FFA8BEB7FB8B9C5CFBA2" LSID="urn:lsid:plazi:treatment:039787C18605FFA2BE3DFE109B65F995:8EE1600C8605FFA8BEB7FB8B9C5CFBA2" box="[435,597,1073,1096]" country="Oman" latitude="58.5" name="Ras Al Hamra" pageId="3" pageNumber="2">Ras Al Hamra</location>
, 23°38'
<geoCoordinate id="EE0A50108605FFA8BDA2FB8B9D05FBA2" box="[678,780,1073,1096]" degrees="58" direction="north" minutes="30" orientation="latitude" pageId="3" pageNumber="2" precision="925" value="58.5">N 58°30'</geoCoordinate>
E,
<date id="FF8010178605FFA8BFC4FBF59F38FB8C" box="[192,305,1103,1126]" pageId="3" pageNumber="2" value="1998-05">
<collectingDate id="EFC4E9FF8605FFA8BFC4FBF59F38FB8C" box="[192,305,1103,1126]" pageId="3" pageNumber="2" value="1998-05">May 1998</collectingDate>
</date>
, leg.
<collectorName id="26CB53018605FFA8BE6AFBF59FCEFB8C" box="[366,455,1103,1126]" pageId="3" pageNumber="2">C. Rose</collectorName>
,
<collectionCode id="ED2FAE128605FFA8BED0FBF59C2EFB8C" box="[468,551,1103,1126]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="3" pageNumber="2">NHMB</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B563C8A8605FFA8BD30FBF59EEEFB6F" collectionCode="UV" collectorName="Jabal Bani Jabir" country="Oman" pageId="3" pageNumber="2" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D38FD5E8605FFA8BD30FBF59C57FB8C" box="[564,606,1103,1126]" count="1" pageId="3" pageNumber="2" type="female">1 ♀</specimenCount>
,
<collectorName id="26CB53018605FFA8BD69FBF59D17FB8C" box="[621,798,1103,1126]" pageId="3" pageNumber="2">Jabal Bani Jabir</collectorName>
,
<collectionCode id="ED2FAE128605FFA8BFC4FBD49EEEFB6F" box="[192,231,1134,1157]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="3" pageNumber="2">UV</collectionCode>
</materialsCitation>
<materialsCitation id="3B563C8A8605FFA8BFEAFBD49F73FB28" collectingDate="2001-12-14" collectionCode="NHMB" collectorName="A. Winkler" country="Oman" elevation="1686" latitude="59.022667" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
detection, rocky slope, 22°49.86'
<geoCoordinate id="EE0A50108605FFA8BD49FBD49CD8FB6F" box="[589,721,1134,1157]" degrees="59" direction="north" minutes="01.36" orientation="latitude" pageId="3" pageNumber="2" precision="9" value="59.022667">N 59°01.36'</geoCoordinate>
E,
<elevation id="0013D1E48605FFA8BDE8FBD49F1FFB4E" metricMagnitude="3" metricUnit="m" metricValue="1.686" pageId="3" pageNumber="2" unit="m" value="1686.0">
<quantity id="4CC69B328605FFA8BDE8FBD49EDCFB4E" metricMagnitude="3" metricUnit="m" metricValue="1.686" pageId="3" pageNumber="2" unit="m" value="1686.0">1686 m</quantity>
a.s.l.
</elevation>
,
<date id="FF8010178605FFA8BE2EFB379C0EFB4E" box="[298,519,1165,1188]" pageId="3" pageNumber="2" value="2001-12-14">
<collectingDate id="EFC4E9FF8605FFA8BE2EFB379C0EFB4E" box="[298,519,1165,1188]" pageId="3" pageNumber="2" value="2001-12-14">14 December 2001</collectingDate>
</date>
, 15:30–17:30 h, leg.
<collectorName id="26CB53018605FFA8BC0CFB379F13FB28" pageId="3" pageNumber="2">A. Winkler</collectorName>
,
<collectionCode id="ED2FAE128605FFA8BE23FB119F73FB28" box="[295,378,1195,1218]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="3" pageNumber="2">NHMB</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B563C8A8605FFA8BE83FB119C65FB28" box="[391,620,1195,1218]" collectionCode="UV" country="Oman" location="Jabal Bani Jabir" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
<location id="8EE1600C8605FFA8BE83FB119C30FB28" LSID="urn:lsid:plazi:treatment:039787C18605FFA2BE3DFE109B65F995:8EE1600C8605FFA8BE83FB119C30FB28" box="[391,569,1195,1218]" country="Oman" name="Jabal Bani Jabir" pageId="3" pageNumber="2">Jabal Bani Jabir</location>
,
<collectionCode id="ED2FAE128605FFA8BD41FB119C65FB28" box="[581,620,1195,1218]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="3" pageNumber="2">UV</collectionCode>
</materialsCitation>
<materialsCitation id="3B563C8A8605FFA8BD70FB119C19FAF4" collectingDate="2001-12-12" collectionCode="NHMB" collectorName="A. Winkler &amp; B. Winkler" country="Oman" elevation="1695" latitude="59.022667" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
detection, rocky slope, between rocks, 22°49.6'
<geoCoordinate id="EE0A50108605FFA8BD1FFB709CAEFB0B" box="[539,679,1226,1249]" degrees="59" direction="north" minutes="01.36" orientation="latitude" pageId="3" pageNumber="2" precision="9" value="59.022667">N 59°01.36'</geoCoordinate>
E,
<elevation id="0013D1E48605FFA8BDCDFB709EFBFAEA" metricMagnitude="3" metricUnit="m" metricValue="1.6949999999999998" pageId="3" pageNumber="2" unit="m" value="1695.0">
<quantity id="4CC69B328605FFA8BDCDFB709D2BFB0B" box="[713,802,1226,1249]" metricMagnitude="3" metricUnit="m" metricValue="1.6949999999999998" pageId="3" pageNumber="2" unit="m" value="1695.0">1695 m</quantity>
a.s.l.
</elevation>
,
<date id="FF8010178605FFA8BE08FB539FFAFAEA" box="[268,499,1257,1280]" pageId="3" pageNumber="2" value="2001-12-12">
<collectingDate id="EFC4E9FF8605FFA8BE08FB539FFAFAEA" box="[268,499,1257,1280]" pageId="3" pageNumber="2" value="2001-12-12">12 December 2001</collectingDate>
</date>
, 18:30–21:30 h, leg.
<collectorName id="26CB53018605FFA8BC0CFB539F1EFAF4" pageId="3" pageNumber="2">A. Winkler</collectorName>
&amp;
<collectorName id="26CB53018605FFA8BE3EFABD9FBBFAF4" box="[314,434,1287,1310]" pageId="3" pageNumber="2">B. Winkler</collectorName>
,
<collectionCode id="ED2FAE128605FFA8BEB9FABD9C19FAF4" box="[445,528,1287,1310]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="3" pageNumber="2">NHMB</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C324655C8605FFA8BFC4FAFF9D69FE18" pageId="3" pageNumber="2" type="diagnosis">
<paragraph id="8B8136D78605FFA8BFC4FAFF9D69FE18" blockId="3.[192,802,1349,1893]" lastBlockId="3.[830,1440,199,498]" pageId="3" pageNumber="2">
<emphasis id="B94AEAC58605FFA8BFC4FAFF9F3CFAB6" bold="true" box="[192,309,1349,1372]" pageId="3" pageNumber="2">Diagnosis.</emphasis>
<materialsCitation id="3B563C8A8605FFA8BE47FAFF9D55FE18" collectionCode="VI, III" collectorName="Medium &amp; Sissom &amp; Kovarik &amp; Sun &amp; Zhu &amp; Lourenco" longitude="-5.9" pageId="3" pageNumber="2" specimenCode="III 1.29, 1.48, V 2.02, 2.40" specimenCount="1">
<collectorName id="26CB53018605FFA8BE47FAFF9F94FAB6" box="[323,413,1349,1372]" pageId="3" pageNumber="2">Medium</collectorName>
sized
<taxonomicName id="4C3E4D548605FFA8BEF5FAFF9C62FAB6" authorityName="Birula" authorityYear="1908" box="[497,619,1349,1372]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAC58605FFA8BEF5FAFF9C62FAB6" box="[497,619,1349,1372]" italics="true" pageId="3" pageNumber="2">Hottentotta</emphasis>
</taxonomicName>
(
<collectorName id="26CB53018605FFA8BD85FAFF9CDBFAB6" box="[641,722,1349,1372]" pageId="3" pageNumber="2">Sissom</collectorName>
, 1990;
<collectorName id="26CB53018605FFA8BFC4FAD99F13FA90" box="[192,282,1379,1402]" pageId="3" pageNumber="2">Kovařík</collectorName>
, 2007;
<collectorName id="26CB53018605FFA8BE6EFAD99F9FFA90" box="[362,406,1379,1402]" pageId="3" pageNumber="2">Sun</collectorName>
,
<collectorName id="26CB53018605FFA8BEA6FAD99FC4FA90" box="[418,461,1379,1402]" pageId="3" pageNumber="2">Zhu</collectorName>
&amp;
<collectorName id="26CB53018605FFA8BEF5FAD99C53FA90" box="[497,602,1379,1402]" pageId="3" pageNumber="2">Lourenço</collectorName>
, 2010), adults
<quantity id="4CC69B328605FFA8BDFEFAD99F1AFA73" metricMagnitude="-2" metricUnit="m" metricValue="6.3" metricValueMax="7.0" metricValueMin="5.6" pageId="3" pageNumber="2" unit="mm" value="63.0" valueMax="70.0" valueMin="56.0">56– 70 mm</quantity>
; color uniform pale yellow without contrasting markings; carapace and tergites with strong, granular carinae; anterior margin of carapace with coarse granules; tergites tricarinate, posterior margins with carinae protruding in short spiniform processes; pedipalp femur, pedipalp patella, legs, metasoma and telson conspicuously hirsute, with numerous long and short macrosetae; metasoma with numerous intercarinal macrosetae; genital opercula densely setose; pedipalp chela finely hirsute with mostly short macrosetae; posterior margin of tergites I–
<collectionCode id="ED2FAE128605FFA8BD13F90F9C3AF926" box="[535,563,1717,1740]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="2">VI</collectionCode>
mostly bare, usually with just one pair of short macrosetae internal to lateral carinae; pedipalps slender, femur L/
<geoCoordinate id="EE0A50108605FFA8BD50F9489C92F8E3" box="[596,667,1778,1801]" degrees="3.6" direction="west" orientation="longitude" pageId="3" pageNumber="2" precision="5555" value="-3.6">W 3.6</geoCoordinate>
–4.4, patella L/
<geoCoordinate id="EE0A50108605FFA8BFDCF8AB9F12F8C2" box="[216,283,1809,1832]" degrees="3.0" direction="west" orientation="longitude" pageId="3" pageNumber="2" precision="5555" value="-3.0">W 3.0</geoCoordinate>
–3.5, chela L/
<geoCoordinate id="EE0A50108605FFA8BEAAF8AB9FF8F8C2" box="[430,497,1809,1832]" degrees="5.9" direction="west" orientation="longitude" pageId="3" pageNumber="2" precision="5555" value="-5.9">W 5.9</geoCoordinate>
–6.5; pedipalp chela without carinae; pedipalp fixed finger primary denticles divided into 13–14 subrows (98 % of cases), movable finger primary denticles divided into 14–15 subrows (98 % of cases); male pedipalp fingers without strong basal scalloping; metasoma moderately robust, segment L/W ratios: metasoma I 1.01–1.15, metasoma II 1.24–1.40, metasoma
<specimenCode id="DB989EAC8605FFA8BCC5FEF89A28FEB3" box="[961,1057,322,345]" collectionCode="III" pageId="3" pageNumber="2">III 1.29</specimenCode>
–
<specimenCode id="DB989EAC8605FFA8BB34FEF89A56FEB3" box="[1072,1119,322,345]" collectionCode="III" pageId="3" pageNumber="2">1.48</specimenCode>
, metasoma IV 1.50–1.85, metasoma
<specimenCode id="DB989EAC8605FFA8BCB6FEDA9DF4FE9D" box="[946,1021,352,375]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="2" type="Museum">V 2.02</specimenCode>
–
<specimenCode id="DB989EAC8605FFA8BB08FEDA9A35FE9D" box="[1036,1084,352,375]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="2" type="Museum">2.40</specimenCode>
; metasoma I–
<collectionCode id="ED2FAE128605FFA8BBD5FEDA9AE5FE9D" box="[1233,1260,352,375]" pageId="3" pageNumber="2">III</collectionCode>
with 10 carinae, median lateral carinae complete but weak on segment
<collectionCode id="ED2FAE128605FFA8BC3AFE249D55FE5F" box="[830,860,414,437]" pageId="3" pageNumber="2">III</collectionCode>
; lateral surfaces of metasoma I–IV coarsely granulated; pectine teeth in males 29–35, in females 25– 33
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C324655C8605FFA8BC3AFDA29B49F9C9" pageId="3" pageNumber="2" type="etymology">
<paragraph id="8B8136D78605FFA8BC3AFDA29D8DFD61" blockId="3.[830,1440,536,651]" pageId="3" pageNumber="2">
<emphasis id="B94AEAC58605FFA8BC3AFDA29DB0FDC5" bold="true" box="[830,953,536,559]" pageId="3" pageNumber="2">Etymology</emphasis>
. The specific epithet refers to the uniform, pale yellow, semi-translucent body, which contrasts with the varied dark coloration of many other members of the genus.
</paragraph>
<paragraph id="8B8136D78605FFA8BC3AFD089A4EFB38" blockId="3.[830,1441,690,1571]" pageId="3" pageNumber="2">
<emphasis id="B94AEAC58605FFA8BC3AFD089DD4FD23" bold="true" box="[830,989,690,713]" pageId="3" pageNumber="2">Comparisons.</emphasis>
Other species of
<taxonomicName id="4C3E4D548605FFA8BBDBFD089B51FD23" authorityName="Birula" authorityYear="1908" box="[1247,1368,690,713]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAC58605FFA8BBDBFD089B51FD23" box="[1247,1368,690,713]" italics="true" pageId="3" pageNumber="2">Hottentotta</emphasis>
</taxonomicName>
with pedipalps bearing numerous long macrosetae are differentiated as follows:
<taxonomicName id="4C3E4D548605FFA8BB28FD559B73FCEC" authority="(Birula, 1914)" baseAuthorityName="Birula" baseAuthorityYear="1914" box="[1068,1402,751,774]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="franzwerneri">
<emphasis id="B94AEAC58605FFA8BB28FD559A36FCEC" box="[1068,1087,751,774]" italics="true" pageId="3" pageNumber="2">H</emphasis>
.
<emphasis id="B94AEAC58605FFA8BB4AFD559AD1FCEC" box="[1102,1240,751,774]" italics="true" pageId="3" pageNumber="2">franzwerneri</emphasis>
(Birula, 1914)
</taxonomicName>
,
<taxonomicName id="4C3E4D548605FFA8BA82FD559A39FCCF" authority="(Pallary, 1924)" baseAuthorityName="Pallary" baseAuthorityYear="1924" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="gentili">
<emphasis id="B94AEAC58605FFA8BA82FD559B90FCEC" box="[1414,1433,751,774]" italics="true" pageId="3" pageNumber="2">H</emphasis>
.
<emphasis id="B94AEAC58605FFA8BC3AFCB49D8BFCCF" box="[830,898,782,805]" italics="true" pageId="3" pageNumber="2">gentili</emphasis>
(Pallary, 1924)
</taxonomicName>
,
<taxonomicName id="4C3E4D548605FFA8BB38FCB49B46FCCF" authority="(Simon, 1880)" baseAuthorityName="Simon" baseAuthorityYear="1880" box="[1084,1359,782,805]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="saulcyi">
<emphasis id="B94AEAC58605FFA8BB38FCB49A46FCCF" box="[1084,1103,782,805]" italics="true" pageId="3" pageNumber="2">H</emphasis>
.
<emphasis id="B94AEAC58605FFA8BB5BFCB49AA3FCCF" box="[1119,1194,782,805]" italics="true" pageId="3" pageNumber="2">saulcyi</emphasis>
(Simon, 1880)
</taxonomicName>
and
<taxonomicName id="4C3E4D548605FFA8BA82FCB49A3CFCA9" authority="(Birula, 1905)" baseAuthorityName="Birula" baseAuthorityYear="1905" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="schach">
<emphasis id="B94AEAC58605FFA8BA82FCB49B90FCCF" box="[1414,1433,782,805]" italics="true" pageId="3" pageNumber="2">H</emphasis>
.
<emphasis id="B94AEAC58605FFA8BC3AFC969D81FCA9" box="[830,904,812,835]" italics="true" pageId="3" pageNumber="2">schach</emphasis>
(Birula, 1905)
</taxonomicName>
: larger scorpions (
<quantity id="4CC69B328605FFA8BA0DFC969B9CFCA9" box="[1289,1429,812,835]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="1.3" metricValueMin="0.7" pageId="3" pageNumber="2" unit="mm" value="100.0" valueMax="130.0" valueMin="70.0">70–130 mm</quantity>
), dark color on carapace or body, tergites with numerous macrosetae, male with more swollen pedipalp chela manus;
<taxonomicName id="4C3E4D548605FFA8BC90FC329ADEFC75" authority="Kovarik, 1997" authorityName="Kovarik" authorityYear="1997" box="[916,1239,904,927]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="B94AEAC58605FFA8BC90FC329DAEFC75" box="[916,935,904,927]" italics="true" pageId="3" pageNumber="2">H</emphasis>
.
<emphasis id="B94AEAC58605FFA8BCB2FC329A3BFC75" box="[950,1074,904,927]" italics="true" pageId="3" pageNumber="2">zagrosensis</emphasis>
Kovařík, 1997
</taxonomicName>
and
<taxonomicName id="4C3E4D548605FFA8BA0BFC329A4BFC54" authority="Navidpour et al., 2010" authorityName="Navidpour" authorityYear="2010" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="lorestanus">
<emphasis id="B94AEAC58605FFA8BA0BFC329B2BFC75" box="[1295,1314,904,927]" italics="true" pageId="3" pageNumber="2">H</emphasis>
.
<emphasis id="B94AEAC58605FFA8BA35FC329BA9FC75" box="[1329,1440,904,927]" italics="true" pageId="3" pageNumber="2">lorestanus</emphasis>
Navidpour et al., 2010
</taxonomicName>
: larger scorpions (&gt;
<quantity id="4CC69B328605FFA8BA37FC1D9B9DFC54" box="[1331,1428,935,958]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="2" unit="mm" value="100.0">100 mm</quantity>
), body entirely black, tergites with numerous macrosetae;
<taxonomicName id="4C3E4D548605FFA8BC3AFC5E9A9FFC11" authority="Kovarik, 2007" authorityName="Kovarik" authorityYear="2007" box="[830,1174,996,1019]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="jabalpurensis">
<emphasis id="B94AEAC58605FFA8BC3AFC5E9D58FC11" box="[830,849,996,1019]" italics="true" pageId="3" pageNumber="2">H</emphasis>
.
<emphasis id="B94AEAC58605FFA8BC5BFC5E9DF9FC11" box="[863,1008,996,1019]" italics="true" pageId="3" pageNumber="2">jabalpurensis</emphasis>
Kovařík, 2007
</taxonomicName>
: male with more swollen pedipalp chela manus, tergites with numerous macrosetae;
<taxonomicName id="4C3E4D548605FFA8BC87FB989AB7FBD3" authority="(Ehrenberg, 1828)" baseAuthorityName="Ehrenberg" baseAuthorityYear="1828" box="[899,1214,1058,1081]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="scaber">
<emphasis id="B94AEAC58605FFA8BC87FB989D9FFBD3" box="[899,918,1058,1081]" italics="true" pageId="3" pageNumber="2">H</emphasis>
.
<emphasis id="B94AEAC58605FFA8BCA2FB989DE4FBD3" box="[934,1005,1058,1081]" italics="true" pageId="3" pageNumber="2">scaber</emphasis>
(Ehrenberg, 1828)
</taxonomicName>
: light and dark color pattern, metasoma with sparse setation, metasoma I–III more stout;
<taxonomicName id="4C3E4D548605FFA8BCBBFBE59B38FB9C" authority="Teruel et Rein, 2010" authorityName="Teruel et Rein" authorityYear="2010" box="[959,1329,1119,1142]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="flavidulus">
<emphasis id="B94AEAC58605FFA8BCBBFBE59DDBFB9C" box="[959,978,1119,1142]" italics="true" pageId="3" pageNumber="2">H</emphasis>
.
<emphasis id="B94AEAC58605FFA8BCDBFBE59A41FB9C" box="[991,1096,1119,1142]" italics="true" pageId="3" pageNumber="2">flavidulus</emphasis>
Teruel et Rein, 2010
</taxonomicName>
: similar to
<taxonomicName id="4C3E4D548605FFA8BC3AFBC49DD3FB7F" authorityName="Lowe" authorityYear="2010" box="[830,986,1150,1173]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="pellucidus">
<emphasis id="B94AEAC58605FFA8BC3AFBC49D58FB7F" box="[830,849,1150,1173]" italics="true" pageId="3" pageNumber="2">H</emphasis>
.
<emphasis id="B94AEAC58605FFA8BC68FBC49DD3FB7F" box="[876,986,1150,1173]" italics="true" pageId="3" pageNumber="2">pellucidus</emphasis>
</taxonomicName>
in being a smaller
<taxonomicName id="4C3E4D548605FFA8BBE0FBC49B57FB7F" authorityName="Birula" authorityYear="1908" box="[1252,1374,1150,1173]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAC58605FFA8BBE0FBC49B57FB7F" box="[1252,1374,1150,1173]" italics="true" pageId="3" pageNumber="2">Hottentotta</emphasis>
</taxonomicName>
with uniform yellow coloration, but metasoma is more stout and lacks dense setation.
</paragraph>
<paragraph id="8B8136D78605FFA8BC6AFB609B49F9C9" blockId="3.[830,1441,690,1571]" pageId="3" pageNumber="2">
In
<collectingCountry id="F32976478605FFA8BC94FB609DDDFB1B" box="[912,980,1242,1265]" name="Oman" pageId="3" pageNumber="2">Oman</collectingCountry>
, the hirsute species
<taxonomicName id="4C3E4D548605FFA8BBBDFB609D74FAE5" authority="(Pocock, 1895)" baseAuthorityName="Pocock" baseAuthorityYear="1895" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="jayakari">
<emphasis id="B94AEAC58605FFA8BBBDFB609AC5FB1B" box="[1209,1228,1242,1265]" italics="true" pageId="3" pageNumber="2">H</emphasis>
.
<emphasis id="B94AEAC58605FFA8BBDAFB609B3EFB1B" box="[1246,1335,1242,1265]" italics="true" pageId="3" pageNumber="2">jayakari</emphasis>
(Pocock, 1895)
</taxonomicName>
and
<taxonomicName id="4C3E4D548605FFA8BCB9FB429AD8FAE5" authority="(Vachon, 1980)" baseAuthorityName="Vachon" baseAuthorityYear="1980" box="[957,1233,1272,1295]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="salei">
<emphasis id="B94AEAC58605FFA8BCB9FB429DD9FAE5" box="[957,976,1272,1295]" italics="true" pageId="3" pageNumber="2">H</emphasis>
.
<emphasis id="B94AEAC58605FFA8BCE0FB429A1FFAE5" box="[996,1046,1272,1295]" italics="true" pageId="3" pageNumber="2">salei</emphasis>
(Vachon, 1980)
</taxonomicName>
differ as follows: larger species (
<quantity id="4CC69B328605FFA8BCE1FAAD9A69FAC4" box="[997,1120,1303,1326]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="3" pageNumber="2" unit="mm" value="75.0" valueMax="90.0" valueMin="60.0">60–90 mm</quantity>
), dark pigmentation on body and pedipalps; denser pilosity on legs and pedipalps; numerous macrosetae on entire posterior margins of tergites I–VI; metasomal segments of adults more robust, with more sparse intercarinal setation (
<taxonomicName id="4C3E4D548605FFA8BA45FA289B9EFA43" baseAuthorityName="Vachon" baseAuthorityYear="1980" box="[1345,1431,1426,1449]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="salei">
<emphasis id="B94AEAC58605FFA8BA45FA289B5DFA43" box="[1345,1364,1426,1449]" italics="true" pageId="3" pageNumber="2">H</emphasis>
.
<emphasis id="B94AEAC58605FFA8BA61FA289B9EFA43" box="[1381,1431,1426,1449]" italics="true" pageId="3" pageNumber="2">salei</emphasis>
</taxonomicName>
) or bare, lacking macrosetae (
<taxonomicName id="4C3E4D548605FFA8BBABFA0A9B31FA2D" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[1199,1336,1456,1479]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="jayakari">
<emphasis id="B94AEAC58605FFA8BBABFA0A9ACBFA2D" box="[1199,1218,1456,1479]" italics="true" pageId="3" pageNumber="2">H</emphasis>
.
<emphasis id="B94AEAC58605FFA8BBDBFA0A9B31FA2D" box="[1247,1336,1456,1479]" italics="true" pageId="3" pageNumber="2">jayakari</emphasis>
</taxonomicName>
); lateral intercarinal surfaces of metasoma I–IV smooth, not granulated; male pedipalp chelae with more swollen manus and distinct scalloping at base of fingers.
</paragraph>
</subSubSection>
<subSubSection id="C324655C8605FFA2BC3AF9F49B11FDC5" lastPageId="9" lastPageNumber="8" pageId="3" pageNumber="2" type="description">
<paragraph id="8B8136D78605FFA8BC3AF9F49DB5F967" blockId="3.[830,1440,1614,1677]" pageId="3" pageNumber="2">
<emphasis id="B94AEAC58605FFA8BC3AF9F49DB5F967" bold="true" pageId="3" pageNumber="2">
Description (
<typeStatus id="548588758605FFA8BCEAF9F49A52F982" box="[1006,1115,1614,1640]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
male unless otherwise specified).
</emphasis>
</paragraph>
<paragraph id="8B8136D78605FFA8BC3AF90E9A09F88F" blockId="3.[830,1440,1716,1893]" pageId="3" pageNumber="2">
<emphasis id="B94AEAC58605FFA8BC3AF90E9DB0F921" bold="true" box="[830,953,1716,1739]" pageId="3" pageNumber="2">Coloration</emphasis>
(
<figureCitation id="13052A528605FFA8BCD6F90E9A4DF921" box="[978,1092,1716,1739]" captionStart="Figures 1–4" captionStartId="4.[192,278,1910,1934]" captionTargetBox="[233,1392,156,1881]" captionTargetId="figure-12@4.[221,1404,156,1881]" captionTargetPageId="4" captionText="Figures 1–4: H. pellucidus, sp. nov., habitus, 1–2, holotype male; 3–4, paratype female. 1, 3. Dorsal aspect. 2, 4. Ventral aspect. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.12782244" httpUri="https://zenodo.org/record/12782244/files/figure.png" pageId="3" pageNumber="2">Figs. 1–4</figureCitation>
). Base color pale translucent yellow; without contrasting markings; melanic pigmentation confined to median and lateral eyes; denticles of chelicerae and pedipalp fingers, articular condyles of pedipalp movable fingers, tarsal ungues and aculeus of telson castaneous.
</paragraph>
<caption id="DF41665F8602FFAFBFC4F8CC9FBEF84D" ID-DOI="http://doi.org/10.5281/zenodo.12782244" ID-Zenodo-Dep="12782244" httpUri="https://zenodo.org/record/12782244/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,278,1910,1934]" targetBox="[233,1392,156,1881]" targetPageId="4" targetType="figure">
<paragraph id="8B8136D78602FFAFBFC4F8CC9FBEF84D" blockId="4.[192,1440,1910,1959]" pageId="4" pageNumber="3">
<emphasis id="B94AEAC58602FFAFBFC4F8CC9F58F864" bold="true" box="[192,337,1910,1934]" pageId="4" pageNumber="3">Figures 1–4:</emphasis>
<taxonomicName id="4C3E4D548602FFAFBE5FF8C39FD6F867" authorityName="Lowe" authorityYear="2010" box="[347,479,1912,1934]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="pellucidus" status="sp. nov.">
<emphasis id="B94AEAC58602FFAFBE5FF8C39F65F864" box="[347,364,1913,1934]" italics="true" pageId="4" pageNumber="3">H</emphasis>
.
<emphasis id="B94AEAC58602FFAFBE78F8C29FD6F867" box="[380,479,1912,1933]" italics="true" pageId="4" pageNumber="3">pellucidus</emphasis>
</taxonomicName>
,
<emphasis id="B94AEAC58602FFAFBEEAF8C39C36F864" bold="true" box="[494,575,1913,1934]" pageId="4" pageNumber="3">
<taxonomicNameLabel id="A27957BE8602FFAFBEEAF8C39C36F864" box="[494,575,1913,1934]" pageId="4" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, habitus,
<emphasis id="B94AEAC58602FFAFBDA0F8C29CCEF867" bold="true" box="[676,711,1912,1933]" pageId="4" pageNumber="3">1–2</emphasis>
, holotype male;
<emphasis id="B94AEAC58602FFAFBC70F8C29D91F867" bold="true" box="[884,920,1912,1933]" pageId="4" pageNumber="3">3–4</emphasis>
, paratype female.
<emphasis id="B94AEAC58602FFAFBB51F8C29A8BF867" bold="true" box="[1109,1154,1912,1934]" pageId="4" pageNumber="3">1, 3.</emphasis>
Dorsal aspect.
<emphasis id="B94AEAC58602FFAFBA25F8C29B47F864" bold="true" box="[1313,1358,1912,1934]" pageId="4" pageNumber="3">2, 4.</emphasis>
Ventral aspect. Scale bar: 10 mm.
</paragraph>
</caption>
<paragraph id="8B8136D78603FFAEBFC4FF7D9F1CFC75" blockId="5.[192,802,199,927]" pageId="5" pageNumber="4">
<emphasis id="B94AEAC58603FFAEBFC4FF7D9F27FF34" bold="true" box="[192,302,199,222]" pageId="5" pageNumber="4">Carapace</emphasis>
(
<figureCitation id="13052A528603FFAEBE3AFF7D9F83FF34" box="[318,394,199,222]" captionStart="Figures 1–4" captionStartId="4.[192,278,1910,1934]" captionTargetBox="[233,1392,156,1881]" captionTargetId="figure-12@4.[221,1404,156,1881]" captionTargetPageId="4" captionText="Figures 1–4: H. pellucidus, sp. nov., habitus, 1–2, holotype male; 3–4, paratype female. 1, 3. Dorsal aspect. 2, 4. Ventral aspect. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.12782244" httpUri="https://zenodo.org/record/12782244/files/figure.png" pageId="5" pageNumber="4">Figs. 1</figureCitation>
,
<figureCitation id="13052A528603FFAEBE9DFF7D9FA1FF34" box="[409,424,199,222]" captionStart="Figures 5–11" captionStartId="6.[192,278,1662,1685]" captionTargetBox="[228,1382,170,1606]" captionTargetId="figure-10@6.[192,1441,147,1632]" captionTargetPageId="6" captionText="Figures 5–11: H. pellucidus, sp. nov. 5, 8–11, holotype male; 6–7, paratype male (Jabal Bani Jabir). 5. Carapace and tergites I– II, dorsal aspect. 6–7, right chelicera. 6. Dorsal aspect. 7. Ventral aspect. 8–9, right basitarsus and telotarsus III. 8. Ventral aspect. 9. Retrolateral aspect. 10–11, metasoma and telson. 10. Right lateral aspect. 11. Ventral aspect. Scale bar: 5: 3.19 mm; 6–7: 2.125 mm; 8–9: 1.5 mm; 10–11: 6.22 mm." figureDoi="http://doi.org/10.5281/zenodo.12782246" httpUri="https://zenodo.org/record/12782246/files/figure.png" pageId="5" pageNumber="4">5</figureCitation>
). Subquadrate, only slightly wider than long, dorsoventrally compressed, lateral flanks moderately sloped; median eyes slightly raised; lateral eyes with 5 ocelli (3 large, 2 small); anterior margin concave, smooth except for weak median denticulations, bordered by coarse granules;
<emphasis id="B94AEAC58603FFAEBD44FEDA9CA6FE9D" box="[576,687,352,375]" italics="true" pageId="5" pageNumber="4">carination</emphasis>
: anterior median, central median and posterior median carinae strongly developed, coarsely granular; central lateral and lateral ocular carinae less distinct, granular; central median and posterior median carinae not collinear, but connected by short transverse series of 4–6 granules; supraocular part of superciliary carinae smooth, postocular part with enlarged granule; other carinae indistinct;
<emphasis id="B94AEAC58603FFAEBE3BFDEC9FBAFD87" box="[319,435,598,621]" italics="true" pageId="5" pageNumber="4">chaetotaxy</emphasis>
: short macrosetae present on anterior part of carapace, 12 along anterior margin, 7 among anterior bordering granules, 1–2 behind lateral eyes, 2 on anterior median carinae;
<emphasis id="B94AEAC58603FFAEBD50FD089CDBFD23" box="[596,722,690,713]" italics="true" pageId="5" pageNumber="4">granulation</emphasis>
: group of coarse granules on anterior interocular triangle behind lateral eyes; lateral flanks with coarse granulation; posterior median intercarinal area mostly smooth, with scattered fine granules; area between anterior median carinae smooth; posterior margin of carapace bordered by row of fine granules between posterior median carinae.
</paragraph>
<paragraph id="8B8136D78603FFAEBFC4FC7C9C7EF98B" blockId="5.[192,802,966,1633]" pageId="5" pageNumber="4">
<emphasis id="B94AEAC58603FFAEBFC4FC7C9F27FC37" bold="true" box="[192,302,966,989]" pageId="5" pageNumber="4">Chelicera</emphasis>
(
<typeStatus id="548588758603FFAEBE38FC7C9F93FC37" box="[316,410,966,989]" pageId="5" pageNumber="4" type="paratype">paratype</typeStatus>
male) (
<figureCitation id="13052A528603FFAEBEE9FC7C9C5FFC37" box="[493,598,966,989]" captionStart="Figures 5–11" captionStartId="6.[192,278,1662,1685]" captionTargetBox="[228,1382,170,1606]" captionTargetId="figure-10@6.[192,1441,147,1632]" captionTargetPageId="6" captionText="Figures 5–11: H. pellucidus, sp. nov. 5, 8–11, holotype male; 6–7, paratype male (Jabal Bani Jabir). 5. Carapace and tergites I– II, dorsal aspect. 6–7, right chelicera. 6. Dorsal aspect. 7. Ventral aspect. 8–9, right basitarsus and telotarsus III. 8. Ventral aspect. 9. Retrolateral aspect. 10–11, metasoma and telson. 10. Right lateral aspect. 11. Ventral aspect. Scale bar: 5: 3.19 mm; 6–7: 2.125 mm; 8–9: 1.5 mm; 10–11: 6.22 mm." figureDoi="http://doi.org/10.5281/zenodo.12782246" httpUri="https://zenodo.org/record/12782246/files/figure.png" pageId="5" pageNumber="4">Figs. 6–7</figureCitation>
). Dorsal surface of manus smooth, glabrous, with transverse row of coarse, subdistal granules; base of fixed finger with prominent dorsointernal carina bearing coarse granules;
<emphasis id="B94AEAC58603FFAEBDA3FB989D12FBD3" box="[679,795,1058,1081]" italics="true" pageId="5" pageNumber="4">chaetotaxy</emphasis>
: manus with pair of short pale microsetae on apical border; long reddish macroseta on dorsointernal carina, shorter reddish macroseta medially located on subdistal granule row; dorsal surface of movable finger with 5 short pale microsetae in distal half; manus ventrally smooth, bearing numerous long, fine microsetae, sparser medially, denser on medio-internal aspect, merging with dense setal brush on ventral aspect of fixed finger; dense fine setae extend to medial and dorsomedial surfaces at base of fixed finger; ventral surface of movable finger densely setose;
<emphasis id="B94AEAC58603FFAEBE73FAC99FDFFA60" box="[375,470,1395,1418]" italics="true" pageId="5" pageNumber="4">dentition</emphasis>
: fingers with normal buthid dentition (Vachon, 1963; Sissom, 1990); fixed finger with large distal tine, subdistal denticle and large proximal bicusp, two prominent denticles on ventral surface; movable finger with large dorsal and ventral distal tines; dorsal margin of movable finger with two large triangulate denticles and small proximal bicusp, ventral margin with two robust denticles.
</paragraph>
<paragraph id="8B8136D78603FFAEBFC4F93D9D9AFEF1" blockId="5.[192,802,1671,1909]" lastBlockId="5.[830,1441,199,283]" pageId="5" pageNumber="4">
<emphasis id="B94AEAC58603FFAEBFC4F93D9F82F974" bold="true" box="[192,395,1671,1694]" pageId="5" pageNumber="4">Coxosternal area</emphasis>
(
<figureCitation id="13052A528603FFAEBEA7F93D9FE4F974" box="[419,493,1671,1694]" captionStart="Figures 1–4" captionStartId="4.[192,278,1910,1934]" captionTargetBox="[233,1392,156,1881]" captionTargetId="figure-12@4.[221,1404,156,1881]" captionTargetPageId="4" captionText="Figures 1–4: H. pellucidus, sp. nov., habitus, 1–2, holotype male; 3–4, paratype female. 1, 3. Dorsal aspect. 2, 4. Ventral aspect. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.12782244" httpUri="https://zenodo.org/record/12782244/files/figure.png" pageId="5" pageNumber="4">Fig. 2</figureCitation>
). Coxae smooth; posterior margins of coxae III–IV finely granulated proximally; coxa I with 8–9 macrosetae, endite I of coxa I with 9–12 macrosetae, endite II with 3–6 macrosetae; coxa II with 8–10 macrosetae on anterior and distal margins; coxa III with 5–6 macrosetae on anterior margin; coxa IV with single basal macroseta; sternum subtriangular, smooth anteriorly, finely granulated on posterior margin, with deep posteromedian excavation; genital opercula smooth, each with 20 short macrosetae; genital papillae present.
</paragraph>
<paragraph id="8B8136D78603FFAEBC3AFEF89B41FD61" blockId="5.[830,1441,322,651]" pageId="5" pageNumber="4">
<emphasis id="B94AEAC58603FFAEBC3AFEF89D92FEB3" bold="true" box="[830,923,322,345]" pageId="5" pageNumber="4">Pectines</emphasis>
(
<figureCitation id="13052A528603FFAEBCB9FEF89A15FEB3" box="[957,1052,322,345]" captionStart="Figures 1–4" captionStartId="4.[192,278,1910,1934]" captionTargetBox="[233,1392,156,1881]" captionTargetId="figure-12@4.[221,1404,156,1881]" captionTargetPageId="4" captionText="Figures 1–4: H. pellucidus, sp. nov., habitus, 1–2, holotype male; 3–4, paratype female. 1, 3. Dorsal aspect. 2, 4. Ventral aspect. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.12782244" httpUri="https://zenodo.org/record/12782244/files/figure.png" pageId="5" pageNumber="4">Figs. 2</figureCitation>
,
<figureCitation id="13052A528603FFAEBB3EFEF89A51FEB3" box="[1082,1112,322,345]" captionStart="Figures 12–20" captionStartId="7.[192,278,1626,1649]" captionTargetBox="[237,1379,217,1541]" captionTargetId="figure-6@7.[192,1442,168,1594]" captionTargetPageId="7" captionText="Figures 12–20: H. pellucidus, sp.nov., 12–16, 19, holotype male; 17–18, paratype male; 20, paratype female. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17–18. Right hemispermatophore, lobes at base of flagellum. 17. Convex aspect. 18. Inner aspect. 19. Left male pectine, ventral aspect. 20. Left female pectine ventral aspect. Scale bar: 12–16, 19–20: 4.1 mm; 17–18: 1.06 mm." figureDoi="http://doi.org/10.5281/zenodo.12782248" httpUri="https://zenodo.org/record/12782248/files/figure.png" pageId="5" pageNumber="4">19</figureCitation>
). Basal piece with deep anteromedian incision, surface smooth; 2 short macrosetae on left half; distal tips of pectines extending slightly past middle of trochanter IV; pectines with 3 marginal lamellae, 9 or 10 middle lamellae, plus small intermediate lamella at distal end of basal marginal lamella, 31–33 teeth; marginal lamellae, middle lamellae and fulcra with numerous short reddish macrosetae, 2–5 on fulcra; when anterior margins of both pectines are aligned with posterior margins of coxae IV, basal 3 teeth overlap with gap between basal middle lamellae.
</paragraph>
<paragraph id="8B8136D78603FFAEBC3AFD089ABCFA43" blockId="5.[830,1441,690,1449]" pageId="5" pageNumber="4">
<emphasis id="B94AEAC58603FFAEBC3AFD089DBFFD23" bold="true" box="[830,950,690,713]" pageId="5" pageNumber="4">Mesosoma</emphasis>
(
<figureCitation id="13052A528603FFAEBCCCFD089A38FD23" box="[968,1073,690,713]" captionStart="Figures 1–4" captionStartId="4.[192,278,1910,1934]" captionTargetBox="[233,1392,156,1881]" captionTargetId="figure-12@4.[221,1404,156,1881]" captionTargetPageId="4" captionText="Figures 1–4: H. pellucidus, sp. nov., habitus, 1–2, holotype male; 3–4, paratype female. 1, 3. Dorsal aspect. 2, 4. Ventral aspect. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.12782244" httpUri="https://zenodo.org/record/12782244/files/figure.png" pageId="5" pageNumber="4">Figs. 1–2</figureCitation>
,
<figureCitation id="13052A528603FFAEBB44FD089A46FD23" box="[1088,1103,690,713]" captionStart="Figures 5–11" captionStartId="6.[192,278,1662,1685]" captionTargetBox="[228,1382,170,1606]" captionTargetId="figure-10@6.[192,1441,147,1632]" captionTargetPageId="6" captionText="Figures 5–11: H. pellucidus, sp. nov. 5, 8–11, holotype male; 6–7, paratype male (Jabal Bani Jabir). 5. Carapace and tergites I– II, dorsal aspect. 6–7, right chelicera. 6. Dorsal aspect. 7. Ventral aspect. 8–9, right basitarsus and telotarsus III. 8. Ventral aspect. 9. Retrolateral aspect. 10–11, metasoma and telson. 10. Right lateral aspect. 11. Ventral aspect. Scale bar: 5: 3.19 mm; 6–7: 2.125 mm; 8–9: 1.5 mm; 10–11: 6.22 mm." figureDoi="http://doi.org/10.5281/zenodo.12782246" httpUri="https://zenodo.org/record/12782246/files/figure.png" pageId="5" pageNumber="4">5</figureCitation>
).
<emphasis id="B94AEAC58603FFAEBB63FD089AB7FD23" box="[1127,1214,690,713]" italics="true" pageId="5" pageNumber="4">Tergites</emphasis>
: pretergites smooth; tergites I–VI with 3 granulose, moderately strong carinae; median carinae of III–VI and lateral carinae of I–VI with terminal spinoid granule projecting slightly beyond posterior margins of tergites; lateral carinae of tergite I short, reduced to terminal projection; tergite VII with two pairs of coarsely granular lateral carinae, and granulated median hump; margins of all tergites with well-spaced small granules; lateral flanks of all tergites moderately sloped, with numerous coarse granules; median intercarinal surfaces mostly smooth, with sparse fine and coarse granules; tergites mostly bare with less than 4 macrosetae; tergites I–VI with paired marginal macrosetae on medial side of lateral carinae; additional small marginal macrosetae on lateral sides of lateral carinae on tergites III and VI; tergite VII devoid of macrosetae.
<emphasis id="B94AEAC58603FFAEBCCEFB269A2EFB59" box="[970,1063,1180,1203]" italics="true" pageId="5" pageNumber="4">Sternites</emphasis>
: all sternites smooth, sternite III without carinae, sternites IV–VI with smooth, weak lateral carinae; sternite VII with 4 carinae, medial pair weak, smooth, lateral pair moderate, weakly granular; macrosetal counts on medial and lateral surfaces + posterior margin: III 29+15, IV 11+17, V 16+15, VI 19+7, VII 21+3; lateral margins shagreened on sternite III, denticulate on IV–VII, posterior margins microdenticulate on III–VI, granular on VIII.
</paragraph>
<paragraph id="8B8136D78603FFAEBC3AFA759B02F8F3" blockId="5.[830,1441,1487,1817]" pageId="5" pageNumber="4">
<emphasis id="B94AEAC58603FFAEBC3AFA759AFDFA0C" bold="true" box="[830,1268,1487,1510]" pageId="5" pageNumber="4">
Hemispermatophore (
<typeStatus id="548588758603FFAEBB3BFA759AAEFA0C" box="[1087,1191,1487,1510]" pageId="5" pageNumber="4" type="paratype">paratype</typeStatus>
male)
</emphasis>
(
<figureCitation id="13052A528603FFAEBA0EFA759B9DFA0C" box="[1290,1428,1487,1510]" captionStart="Figures 12–20" captionStartId="7.[192,278,1626,1649]" captionTargetBox="[237,1379,217,1541]" captionTargetId="figure-6@7.[192,1442,168,1594]" captionTargetPageId="7" captionText="Figures 12–20: H. pellucidus, sp.nov., 12–16, 19, holotype male; 17–18, paratype male; 20, paratype female. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17–18. Right hemispermatophore, lobes at base of flagellum. 17. Convex aspect. 18. Inner aspect. 19. Left male pectine, ventral aspect. 20. Left female pectine ventral aspect. Scale bar: 12–16, 19–20: 4.1 mm; 17–18: 1.06 mm." figureDoi="http://doi.org/10.5281/zenodo.12782248" httpUri="https://zenodo.org/record/12782248/files/figure.png" pageId="5" pageNumber="4">Figs. 17–18</figureCitation>
). Flagelliform, trunk elongate, slender; flagellum filiform, relatively short, pars recta 0.2 times length of trunk, pars reflecta 0.4 times length of trunk; inner lobe a broad untapered lamina with truncate rounded apex; median lobe wide, triangulate with curved inner edge, outer lobe tapered, apically bent; basal lobe a stout blunt hook. Measurements (
<typeStatus id="548588758603FFAEBCEDF91C9A45F957" box="[1001,1100,1702,1725]" pageId="5" pageNumber="4" type="paratype">paratype</typeStatus>
): trunk L (to base of flagellum)
<quantity id="4CC69B328603FFAEBC3AF97E9DA8F931" box="[830,929,1732,1755]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="5" pageNumber="4" unit="mm" value="11.5">11.5 mm</quantity>
, pars recta
<quantity id="4CC69B328603FFAEBB18F97E9A78F931" box="[1052,1137,1732,1755]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="4" unit="mm" value="2.1">2.1 mm</quantity>
, pars reflecta
<quantity id="4CC69B328603FFAEBA0DF97E9B57F931" box="[1289,1374,1732,1755]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="4" unit="mm" value="4.6">4.6 mm</quantity>
, inner lobe (from base of flagellum) 860 µm, median lobe 530 µm, outer lobe 400 µm, basal lobe 330 µm.
</paragraph>
<paragraph id="8B8136D78603FFA3BC3AF8859C97F981" blockId="5.[830,1440,1855,1909]" lastBlockId="8.[192,803,1221,1643]" lastPageId="8" lastPageNumber="7" pageId="5" pageNumber="4">
<emphasis id="B94AEAC58603FFAEBC3AF8859DBDF8BC" bold="true" box="[830,948,1855,1878]" pageId="5" pageNumber="4">Metasoma</emphasis>
(
<figureCitation id="13052A528603FFAEBCC9F8859A34F8BC" box="[973,1085,1855,1878]" captionStart="Figures 1–4" captionStartId="4.[192,278,1910,1934]" captionTargetBox="[233,1392,156,1881]" captionTargetId="figure-12@4.[221,1404,156,1881]" captionTargetPageId="4" captionText="Figures 1–4: H. pellucidus, sp. nov., habitus, 1–2, holotype male; 3–4, paratype female. 1, 3. Dorsal aspect. 2, 4. Ventral aspect. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.12782244" httpUri="https://zenodo.org/record/12782244/files/figure.png" pageId="5" pageNumber="4">Figs. 1–2</figureCitation>
,
<figureCitation id="13052A528603FFAEBB56F8859A91F8BC" box="[1106,1176,1855,1878]" captionStart="Figures 5–11" captionStartId="6.[192,278,1662,1685]" captionTargetBox="[228,1382,170,1606]" captionTargetId="figure-10@6.[192,1441,147,1632]" captionTargetPageId="6" captionText="Figures 5–11: H. pellucidus, sp. nov. 5, 8–11, holotype male; 6–7, paratype male (Jabal Bani Jabir). 5. Carapace and tergites I– II, dorsal aspect. 6–7, right chelicera. 6. Dorsal aspect. 7. Ventral aspect. 8–9, right basitarsus and telotarsus III. 8. Ventral aspect. 9. Retrolateral aspect. 10–11, metasoma and telson. 10. Right lateral aspect. 11. Ventral aspect. Scale bar: 5: 3.19 mm; 6–7: 2.125 mm; 8–9: 1.5 mm; 10–11: 6.22 mm." figureDoi="http://doi.org/10.5281/zenodo.12782246" httpUri="https://zenodo.org/record/12782246/files/figure.png" pageId="5" pageNumber="4">10–11</figureCitation>
). Moderately elongate, total length plus telson length 5.4 times carapace length; segments I–III with 10 carinae, segment IV with 8 carinae; segment V with 5 complete carinae; dorsosubmedian carinae strong, coarsely granular on I–VI; dorsolateral carinae strong, on I–IV, weak on V, coarsely granular on all segments; median lateral carinae strong, complete on segment I–II, weak, indistinct on anterior 1/4 of segment III, coarsely granulate on I–III; ventrolateral carinae strong, granulose on I–III, granulocrenulate on IV–V; ventrosubmedian carinae moderate, smooth on I–II, moderate to strong, granulose on III–IV; ventromedian carina strong, broadly granulated on V; ventrosubmedian carinae on V weak, nearly obsolete, only discernible on anterior 1/3 of segment, marked by linear series of coarse dentate granules; dorsal intercarinal surfaces smooth on segments I–IV, smooth medially and weakly granular laterally on V; dorsolateral intercarinal surfaces weakly granular on I, strongly, coarsely granular on II–IV; segment I granulated on lateral surface above median lateral carina, ventrolateral and ventral intercarinal surfaces smooth; segments II–V with coarse granulation on lateral and ventral intercarinal surfaces; lateral anal lobes not dissected; ventral anal arc with 26 crenulations;
<emphasis id="B94AEAC5860EFFA3BD11FAC79C80FA7E" box="[533,649,1405,1428]" italics="true" pageId="8" pageNumber="7">chaetotaxy</emphasis>
: ventrolateral and ventral surfaces on all segments with numerous long macrosetae on carinae and intercarinal surfaces; dorsolateral intercarinal surfaces with macrosetae absent on I– III, present on IV; dorsosubmedian carinae with macrosetae absent on I, present on II–IV; dorsal intercarinal surfaces of all segments lacking macrosetae; segment V with macrosetae around dorsolateral carinae.
</paragraph>
<caption id="DF41665F8600FFADBFC4F9C49C17F935" ID-DOI="http://doi.org/10.5281/zenodo.12782246" ID-Zenodo-Dep="12782246" httpUri="https://zenodo.org/record/12782246/files/figure.png" pageId="6" pageNumber="5" startId="6.[192,278,1662,1685]" targetBox="[228,1382,170,1606]" targetPageId="6" targetType="figure">
<paragraph id="8B8136D78600FFADBFC4F9C49C17F935" blockId="6.[192,1441,1662,1759]" pageId="6" pageNumber="5">
<emphasis id="B94AEAC58600FFADBFC4F9C49F52F97F" bold="true" box="[192,347,1662,1685]" pageId="6" pageNumber="5">Figures 5–11:</emphasis>
<taxonomicName id="4C3E4D548600FFADBE65F9C59FEBF97E" authorityName="Lowe" authorityYear="2010" box="[353,482,1663,1684]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="pellucidus" status="sp. nov.">
<emphasis id="B94AEAC58600FFADBE65F9C59F7BF97E" box="[353,370,1663,1684]" italics="true" pageId="6" pageNumber="5">H</emphasis>
.
<emphasis id="B94AEAC58600FFADBE7BF9C59FEBF97E" box="[383,482,1663,1684]" italics="true" pageId="6" pageNumber="5">pellucidus</emphasis>
</taxonomicName>
,
<emphasis id="B94AEAC58600FFADBEEAF9C59C83F97E" bold="true" box="[494,650,1663,1684]" pageId="6" pageNumber="5">
<taxonomicNameLabel id="A27957BE8600FFADBEEAF9C59C32F97E" box="[494,571,1663,1684]" pageId="6" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
5, 8–11
</emphasis>
, holotype male;
<emphasis id="B94AEAC58600FFADBC28F9C59D59F97E" bold="true" box="[812,848,1663,1684]" pageId="6" pageNumber="5">6–7</emphasis>
, paratype male (Jabal Bani Jabir).
<emphasis id="B94AEAC58600FFADBB9BF9C59AB8F97E" bold="true" box="[1183,1201,1663,1684]" pageId="6" pageNumber="5">5.</emphasis>
Carapace and tergites I
<emphasis id="B94AEAC58600FFADBA90F9C59BA9F97E" bold="true" box="[1428,1440,1663,1684]" pageId="6" pageNumber="5">–</emphasis>
II, dorsal aspect.
<emphasis id="B94AEAC58600FFADBE61F9229F86F947" bold="true" box="[357,399,1688,1709]" pageId="6" pageNumber="5">6–7,</emphasis>
right chelicera.
<emphasis id="B94AEAC58600FFADBD28F9229C37F947" bold="true" box="[556,574,1688,1709]" pageId="6" pageNumber="5">6.</emphasis>
Dorsal aspect.
<emphasis id="B94AEAC58600FFADBDD6F9229CEDF947" bold="true" box="[722,740,1688,1709]" pageId="6" pageNumber="5">7.</emphasis>
Ventral aspect.
<emphasis id="B94AEAC58600FFADBC84F9229DA3F947" bold="true" box="[896,938,1688,1709]" pageId="6" pageNumber="5">8–9,</emphasis>
right basitarsus and telotarsus III.
<emphasis id="B94AEAC58600FFADBBFCF9229B03F947" bold="true" box="[1272,1290,1688,1709]" pageId="6" pageNumber="5">8.</emphasis>
Ventral aspect.
<emphasis id="B94AEAC58600FFADBFC4F90B9EDBF92C" bold="true" box="[192,210,1713,1734]" pageId="6" pageNumber="5">9.</emphasis>
Retrolateral aspect.
<emphasis id="B94AEAC58600FFADBE9CF90B9FD3F92C" bold="true" box="[408,474,1713,1734]" pageId="6" pageNumber="5">10–11,</emphasis>
metasoma and telson.
<emphasis id="B94AEAC58600FFADBDB1F90B9CDDF92C" bold="true" box="[693,724,1713,1734]" pageId="6" pageNumber="5">10.</emphasis>
Right lateral aspect.
<emphasis id="B94AEAC58600FFADBC9BF90B9DB4F92C" bold="true" box="[927,957,1713,1734]" pageId="6" pageNumber="5">11.</emphasis>
Ventral aspect. Scale bar: 5: 3.19 mm; 6–7: 2.125 mm; 8–9: 1.5 mm; 10–11: 6.22 mm.
</paragraph>
</caption>
<caption id="DF41665F8601FFACBFC4F9E09C7EF935" ID-DOI="http://doi.org/10.5281/zenodo.12782248" ID-Zenodo-Dep="12782248" httpUri="https://zenodo.org/record/12782248/files/figure.png" pageId="7" pageNumber="6" startId="7.[192,278,1626,1649]" targetBox="[237,1379,217,1541]" targetPageId="7" targetType="figure">
<paragraph id="8B8136D78601FFACBFC4F9E09C7EF935" blockId="7.[192,1440,1626,1759]" pageId="7" pageNumber="6">
<emphasis id="B94AEAC58601FFACBFC4F9E09F64F99B" bold="true" box="[192,365,1626,1649]" pageId="7" pageNumber="6">Figures 12–20:</emphasis>
<taxonomicName id="4C3E4D548601FFACBE7CF9E69FF4F99B" authorityName="Lowe" authorityYear="2010" box="[376,509,1628,1649]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="pellucidus" status="sp.nov.">
<emphasis id="B94AEAC58601FFACBE7CF9E69F80F99B" box="[376,393,1628,1649]" italics="true" pageId="7" pageNumber="6">H</emphasis>
.
<emphasis id="B94AEAC58601FFACBE9FF9E69FF4F99B" box="[411,509,1628,1649]" italics="true" pageId="7" pageNumber="6">pellucidus</emphasis>
</taxonomicName>
,
<emphasis id="B94AEAC58601FFACBD0AF9E69C5FF99B" bold="true" box="[526,598,1628,1649]" pageId="7" pageNumber="6">
<taxonomicNameLabel id="A27957BE8601FFACBD0AF9E69C5FF99B" box="[526,598,1628,1649]" pageId="7" pageNumber="6" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B94AEAC58601FFACBD6CF9E69CAAF99B" bold="true" box="[616,675,1628,1649]" pageId="7" pageNumber="6">12–16</emphasis>
,
<emphasis id="B94AEAC58601FFACBDB1F9E69CC4F99B" bold="true" box="[693,717,1628,1649]" pageId="7" pageNumber="6">19</emphasis>
, holotype male;
<emphasis id="B94AEAC58601FFACBC79F9E69DB6F99B" bold="true" box="[893,959,1628,1649]" pageId="7" pageNumber="6">17–18,</emphasis>
paratype male;
<emphasis id="B94AEAC58601FFACBB6CF9E69A89F99B" bold="true" box="[1128,1152,1628,1649]" pageId="7" pageNumber="6">20</emphasis>
, paratype female.
<emphasis id="B94AEAC58601FFACBA46F9E69B69F99B" bold="true" box="[1346,1376,1628,1649]" pageId="7" pageNumber="6">12.</emphasis>
Right pedipalp femur, dorsal aspect.
<emphasis id="B94AEAC58601FFACBEF7F9CD9C18F966" bold="true" box="[499,529,1655,1676]" pageId="7" pageNumber="6">13.</emphasis>
Right pedipalp patella, dorsal aspect.
<emphasis id="B94AEAC58601FFACBC96F9CD9DB9F966" bold="true" box="[914,944,1655,1676]" pageId="7" pageNumber="6">14.</emphasis>
Right pedipalp patella, external aspect.
<emphasis id="B94AEAC58601FFACBA40F9CD9B6BF966" bold="true" box="[1348,1378,1655,1676]" pageId="7" pageNumber="6">15.</emphasis>
Right pedipalp chela, external aspect.
<emphasis id="B94AEAC58601FFACBD07F9299C2BF942" bold="true" box="[515,546,1683,1704]" pageId="7" pageNumber="6">16.</emphasis>
Right pedipalp chela, ventral aspect.
<emphasis id="B94AEAC58601FFACBCA0F9299DEFF942" bold="true" box="[932,998,1683,1704]" pageId="7" pageNumber="6">17–18.</emphasis>
Right hemispermatophore, lobes at base of flagellum.
<emphasis id="B94AEAC58601FFACBE2CF9159F4EF92E" bold="true" box="[296,327,1711,1732]" pageId="7" pageNumber="6">17.</emphasis>
Convex aspect.
<emphasis id="B94AEAC58601FFACBEE2F9159C0DF92E" bold="true" box="[486,516,1711,1732]" pageId="7" pageNumber="6">18.</emphasis>
Inner aspect.
<emphasis id="B94AEAC58601FFACBD8FF9159CA0F92E" bold="true" box="[651,681,1711,1732]" pageId="7" pageNumber="6">19.</emphasis>
Left male pectine, ventral aspect.
<emphasis id="B94AEAC58601FFACBCF2F9159A1AF92E" bold="true" box="[1014,1043,1711,1732]" pageId="7" pageNumber="6">20.</emphasis>
Left female pectine ventral aspect. Scale bar: 12–16, 19–20: 4.1 mm; 17–18: 1.06 mm.
</paragraph>
</caption>
<caption id="DF41665F860EFFA3BFC4FC659F80FB76" ID-DOI="http://doi.org/10.5281/zenodo.12782250" ID-Zenodo-Dep="12782250" httpUri="https://zenodo.org/record/12782250/files/figure.png" pageId="8" pageNumber="7" startId="8.[192,278,991,1014]" targetBox="[199,774,194,958]" targetPageId="8" targetType="figure">
<paragraph id="8B8136D7860EFFA3BFC4FC659F80FB76" blockId="8.[192,802,991,1180]" pageId="8" pageNumber="7">
<emphasis id="B94AEAC5860EFFA3BFC4FC659F62FC1C" bold="true" box="[192,363,991,1014]" pageId="8" pageNumber="7">Figures 21–26:</emphasis>
<taxonomicName id="4C3E4D54860EFFA3BE70FC5B9FFEFC1C" authorityName="Lowe" authorityYear="2010" box="[372,503,993,1014]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="pellucidus" status="sp. nov.">
<emphasis id="B94AEAC5860EFFA3BE70FC5B9F8CFC1C" box="[372,389,993,1014]" italics="true" pageId="8" pageNumber="7">H</emphasis>
.
<emphasis id="B94AEAC5860EFFA3BE91FC5B9FFEFC1C" box="[405,503,993,1014]" italics="true" pageId="8" pageNumber="7">pellucidus</emphasis>
</taxonomicName>
,
<emphasis id="B94AEAC5860EFFA3BD02FC5B9C5EFC1C" bold="true" box="[518,599,993,1014]" pageId="8" pageNumber="7">
<taxonomicNameLabel id="A27957BE860EFFA3BD02FC5B9C5EFC1C" box="[518,599,993,1014]" pageId="8" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, trichobothrial map, holotype male.
<emphasis id="B94AEAC5860EFFA3BE5BFC479F74FBF8" bold="true" box="[351,381,1021,1042]" pageId="8" pageNumber="7">21.</emphasis>
Right pedipalp femur, dorsal aspect.
<emphasis id="B94AEAC5860EFFA3BC00FC479D2BFBF8" bold="true" box="[772,802,1021,1042]" pageId="8" pageNumber="7">22.</emphasis>
Right pedipalp femur, proximal internal aspect.
<emphasis id="B94AEAC5860EFFA3BDC4FBA29CD7FBC7" bold="true" box="[704,734,1048,1069]" pageId="8" pageNumber="7">23.</emphasis>
Right pedipalp patella, dorsal aspect.
<emphasis id="B94AEAC5860EFFA3BD08FB8E9C23FBA3" bold="true" box="[524,554,1076,1097]" pageId="8" pageNumber="7">24.</emphasis>
Right pedipalp patella, external aspect.
<emphasis id="B94AEAC5860EFFA3BE65FBF59F76FB8E" bold="true" box="[353,383,1103,1124]" pageId="8" pageNumber="7">25.</emphasis>
Right pedipalp chela, external aspect.
<emphasis id="B94AEAC5860EFFA3BC00FBF59D2BFB8E" bold="true" box="[772,802,1103,1124]" pageId="8" pageNumber="7">26.</emphasis>
Right pedipalp chela, ventral aspect. Trichobothrial notation after Vachon (1974).
</paragraph>
</caption>
<paragraph id="8B8136D7860EFFA3BFC4F92B9C9AF8C9" blockId="8.[192,802,1681,1827]" pageId="8" pageNumber="7">
<emphasis id="B94AEAC5860EFFA3BFC4F92B9F05F942" bold="true" box="[192,268,1681,1704]" pageId="8" pageNumber="7">Telson</emphasis>
(
<figureCitation id="13052A52860EFFA3BE24F92B9FACF942" box="[288,421,1681,1704]" captionStart="Figures 5–11" captionStartId="6.[192,278,1662,1685]" captionTargetBox="[228,1382,170,1606]" captionTargetId="figure-10@6.[192,1441,147,1632]" captionTargetPageId="6" captionText="Figures 5–11: H. pellucidus, sp. nov. 5, 8–11, holotype male; 6–7, paratype male (Jabal Bani Jabir). 5. Carapace and tergites I– II, dorsal aspect. 6–7, right chelicera. 6. Dorsal aspect. 7. Ventral aspect. 8–9, right basitarsus and telotarsus III. 8. Ventral aspect. 9. Retrolateral aspect. 10–11, metasoma and telson. 10. Right lateral aspect. 11. Ventral aspect. Scale bar: 5: 3.19 mm; 6–7: 2.125 mm; 8–9: 1.5 mm; 10–11: 6.22 mm." figureDoi="http://doi.org/10.5281/zenodo.12782246" httpUri="https://zenodo.org/record/12782246/files/figure.png" pageId="8" pageNumber="7">Figs. 10–11</figureCitation>
). Vesicle bulbous, steeply sloped posteriorly; dorsal surface of vesicle smooth, without setae, lateral and ventral surfaces studded with coarse granules, bearing numerous long macrosetae; aculeus robust, curved, slightly shorter than vesicle.
</paragraph>
<paragraph id="8B8136D7860EFFA3BFC4F8F39B42FADA" blockId="8.[192,802,1865,1919]" lastBlockId="8.[830,1441,199,1328]" pageId="8" pageNumber="7">
<emphasis id="B94AEAC5860EFFA3BFC4F8F39F2CF88A" bold="true" box="[192,293,1865,1888]" pageId="8" pageNumber="7">Pedipalp</emphasis>
(
<figureCitation id="13052A52860EFFA3BE31F8F39FBCF88A" box="[309,437,1865,1888]" captionStart="Figures 12–20" captionStartId="7.[192,278,1626,1649]" captionTargetBox="[237,1379,217,1541]" captionTargetId="figure-6@7.[192,1442,168,1594]" captionTargetPageId="7" captionText="Figures 12–20: H. pellucidus, sp.nov., 12–16, 19, holotype male; 17–18, paratype male; 20, paratype female. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17–18. Right hemispermatophore, lobes at base of flagellum. 17. Convex aspect. 18. Inner aspect. 19. Left male pectine, ventral aspect. 20. Left female pectine ventral aspect. Scale bar: 12–16, 19–20: 4.1 mm; 17–18: 1.06 mm." figureDoi="http://doi.org/10.5281/zenodo.12782248" httpUri="https://zenodo.org/record/12782248/files/figure.png" pageId="8" pageNumber="7">Figs. 12–16</figureCitation>
,
<figureCitation id="13052A52860EFFA3BEC7F8F39C01F88A" box="[451,520,1865,1888]" captionStart="Figures 21–26" captionStartId="8.[192,278,991,1014]" captionTargetBox="[199,774,194,958]" captionTargetId="figure-10@8.[180,804,154,984]" captionTargetPageId="8" captionText="Figures 21–26: H. pellucidus, sp. nov., trichobothrial map, holotype male. 21. Right pedipalp femur, dorsal aspect. 22. Right pedipalp femur, proximal internal aspect. 23. Right pedipalp patella, dorsal aspect. 24. Right pedipalp patella, external aspect. 25. Right pedipalp chela, external aspect. 26. Right pedipalp chela, ventral aspect. Trichobothrial notation after Vachon (1974)." figureDoi="http://doi.org/10.5281/zenodo.12782250" httpUri="https://zenodo.org/record/12782250/files/figure.png" pageId="8" pageNumber="7">21–26</figureCitation>
,
<figureCitation id="13052A52860EFFA3BD10F8F39C51F88A" box="[532,600,1865,1888]" captionStart="Figures 53–56" captionStartId="17.[192,278,1091,1114]" captionTargetBox="[206,1424,171,1059]" captionTargetId="figure-6@17.[192,1439,168,1059]" captionTargetPageId="17" captionText="Figures 53–56: Pedipalp finger dentition of new Hottentotta species. 53–54, H. pellucidus, sp. nov., holotype male. 55–56, H. saxinatans, sp. nov., holotype male. 53, 55: right fixed finger, ventral aspect. 54, 56: right movable finger, dorsal aspect. Scale bar: 1 mm. Images acquired under UV fluorescence." figureDoi="http://doi.org/10.5281/zenodo.12782262" httpUri="https://zenodo.org/record/12782262/files/figure.png" pageId="8" pageNumber="7">53–54</figureCitation>
).
<emphasis id="B94AEAC5860EFFA3BD6AF8F39CBCF88A" box="[622,693,1865,1888]" italics="true" pageId="8" pageNumber="7">Femur</emphasis>
(
<figureCitation id="13052A52860EFFA3BDC7F8F39D1BF88A" box="[707,786,1865,1888]" captionStart="Figures 12–20" captionStartId="7.[192,278,1626,1649]" captionTargetBox="[237,1379,217,1541]" captionTargetId="figure-6@7.[192,1442,168,1594]" captionTargetPageId="7" captionText="Figures 12–20: H. pellucidus, sp.nov., 12–16, 19, holotype male; 17–18, paratype male; 20, paratype female. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17–18. Right hemispermatophore, lobes at base of flagellum. 17. Convex aspect. 18. Inner aspect. 19. Left male pectine, ventral aspect. 20. Left female pectine ventral aspect. Scale bar: 12–16, 19–20: 4.1 mm; 17–18: 1.06 mm." figureDoi="http://doi.org/10.5281/zenodo.12782248" httpUri="https://zenodo.org/record/12782248/files/figure.png" pageId="8" pageNumber="7">Fig. 12</figureCitation>
): slender, 4.03 times longer than wide; dorsoexternal, dorsointernal and ventrointernal carinae moderate, with regular small dentate granules; other carinae obsolete; dorsal surface flat, external surface convex, ventral surface concave, all smooth; internal surface convex, with scattered enlarged dentate granules; distal external macrosetae arranged in two longitudinal rows on distal half of femur (16–20 shorter ventral setae, 9–11 longer dorsal setae); numerous long and short macrosetae on dorsoexternal, dorsointernal and ventrointernal carinae, and dorsal and internal surfaces; ventral surface without setae.
<emphasis id="B94AEAC5860EFFA3BC87FE409DD9FDFB" box="[899,976,506,529]" italics="true" pageId="8" pageNumber="7">Patella</emphasis>
(
<figureCitation id="13052A52860EFFA3BCE4FE409A6CFDFB" box="[992,1125,506,529]" captionStart="Figures 12–20" captionStartId="7.[192,278,1626,1649]" captionTargetBox="[237,1379,217,1541]" captionTargetId="figure-6@7.[192,1442,168,1594]" captionTargetPageId="7" captionText="Figures 12–20: H. pellucidus, sp.nov., 12–16, 19, holotype male; 17–18, paratype male; 20, paratype female. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17–18. Right hemispermatophore, lobes at base of flagellum. 17. Convex aspect. 18. Inner aspect. 19. Left male pectine, ventral aspect. 20. Left female pectine ventral aspect. Scale bar: 12–16, 19–20: 4.1 mm; 17–18: 1.06 mm." figureDoi="http://doi.org/10.5281/zenodo.12782248" httpUri="https://zenodo.org/record/12782248/files/figure.png" pageId="8" pageNumber="7">Figs. 13–14</figureCitation>
): elongate, 3.14 times longer than wide; dorsointernal carina moderate, weakly granulose; dorsomedian and ventromedian carinae weak, smooth to weakly granular; internal carina weak, marked by widely spaced granules; other carinae obsolete; all intercarinal surfaces smooth; external, dorsal and internal surfaces with numerous long and short macrosetae, ventral surface without setae.
<emphasis id="B94AEAC5860EFFA3BBCEFD6A9B01FD0D" box="[1226,1288,720,743]" italics="true" pageId="8" pageNumber="7">Chela</emphasis>
(
<figureCitation id="13052A52860EFFA3BA1DFD6A9B93FD0D" box="[1305,1434,720,743]" captionStart="Figures 12–20" captionStartId="7.[192,278,1626,1649]" captionTargetBox="[237,1379,217,1541]" captionTargetId="figure-6@7.[192,1442,168,1594]" captionTargetPageId="7" captionText="Figures 12–20: H. pellucidus, sp.nov., 12–16, 19, holotype male; 17–18, paratype male; 20, paratype female. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17–18. Right hemispermatophore, lobes at base of flagellum. 17. Convex aspect. 18. Inner aspect. 19. Left male pectine, ventral aspect. 20. Left female pectine ventral aspect. Scale bar: 12–16, 19–20: 4.1 mm; 17–18: 1.06 mm." figureDoi="http://doi.org/10.5281/zenodo.12782248" httpUri="https://zenodo.org/record/12782248/files/figure.png" pageId="8" pageNumber="7">Figs. 15–16</figureCitation>
,
<figureCitation id="13052A52860EFFA3BC44FD559D88FCEC" box="[832,897,751,774]" captionStart="Figures 53–56" captionStartId="17.[192,278,1091,1114]" captionTargetBox="[206,1424,171,1059]" captionTargetId="figure-6@17.[192,1439,168,1059]" captionTargetPageId="17" captionText="Figures 53–56: Pedipalp finger dentition of new Hottentotta species. 53–54, H. pellucidus, sp. nov., holotype male. 55–56, H. saxinatans, sp. nov., holotype male. 53, 55: right fixed finger, ventral aspect. 54, 56: right movable finger, dorsal aspect. Scale bar: 1 mm. Images acquired under UV fluorescence." figureDoi="http://doi.org/10.5281/zenodo.12782262" httpUri="https://zenodo.org/record/12782262/files/figure.png" pageId="8" pageNumber="7">53–54</figureCitation>
): slender with tenuous fingers, 6.12 times longer than wide; movable finger 2.45 times manus ventral length; manus smooth, carinae obsolete; fixed and movable fingers smooth, lacking scalloping of proximal dentate margins; short macrosetae dense on dorsal, external and ventral surfaces of manus and fingers, sparse or absent on internal surfaces; 14 primary denticle subrows on both fixed and movable fingers (
<figureCitation id="13052A52860EFFA3BA3CFC7C9D50FC11" captionStart="Figures 53–56" captionStartId="17.[192,278,1091,1114]" captionTargetBox="[206,1424,171,1059]" captionTargetId="figure-6@17.[192,1439,168,1059]" captionTargetPageId="17" captionText="Figures 53–56: Pedipalp finger dentition of new Hottentotta species. 53–54, H. pellucidus, sp. nov., holotype male. 55–56, H. saxinatans, sp. nov., holotype male. 53, 55: right fixed finger, ventral aspect. 54, 56: right movable finger, dorsal aspect. Scale bar: 1 mm. Images acquired under UV fluorescence." figureDoi="http://doi.org/10.5281/zenodo.12782262" httpUri="https://zenodo.org/record/12782262/files/figure.png" pageId="8" pageNumber="7">Figs. 53– 54</figureCitation>
); all denticle subrows flanked by internal and external accessory denticles, except for most proximal subrow of fixed finger; movable finger with 4 enlarged subdistal denticles.
<emphasis id="B94AEAC5860EFFA3BCB5FBFA9AA1FBBD" box="[945,1192,1088,1111]" italics="true" pageId="8" pageNumber="7">Trichobothrial pattern</emphasis>
(
<figureCitation id="13052A52860EFFA3BBB9FBFA9B4FFBBD" box="[1213,1350,1088,1111]" captionStart="Figures 21–26" captionStartId="8.[192,278,991,1014]" captionTargetBox="[199,774,194,958]" captionTargetId="figure-10@8.[180,804,154,984]" captionTargetPageId="8" captionText="Figures 21–26: H. pellucidus, sp. nov., trichobothrial map, holotype male. 21. Right pedipalp femur, dorsal aspect. 22. Right pedipalp femur, proximal internal aspect. 23. Right pedipalp patella, dorsal aspect. 24. Right pedipalp patella, external aspect. 25. Right pedipalp chela, external aspect. 26. Right pedipalp chela, ventral aspect. Trichobothrial notation after Vachon (1974)." figureDoi="http://doi.org/10.5281/zenodo.12782250" httpUri="https://zenodo.org/record/12782250/files/figure.png" pageId="8" pageNumber="7">Figs. 21–26</figureCitation>
): orthobothriotaxic,
<typeStatus id="54858875860EFFA3BCD4FBDB9DF7FB92" box="[976,1022,1121,1144]" pageId="8" pageNumber="7">type</typeStatus>
Aβ (Vachon, 1974; 1975); femur
<emphasis id="B94AEAC5860EFFA3BA87FBDB9B99FB92" box="[1411,1424,1121,1144]" italics="true" pageId="8" pageNumber="7">d</emphasis>
<subScript id="17BA3492860EFFA3BA95FBD19B93FB90" attach="right" box="[1425,1434,1131,1146]" fontSize="6" pageId="8" pageNumber="7">2</subScript>
, patella
<emphasis id="B94AEAC5860EFFA3BC8AFB3A9D92FB7D" box="[910,923,1152,1175]" italics="true" pageId="8" pageNumber="7">d</emphasis>
<subScript id="17BA3492860EFFA3BC98FB309DACFB73" attach="right" box="[924,933,1162,1177]" fontSize="6" pageId="8" pageNumber="7">2</subScript>
, chela
<emphasis id="B94AEAC5860EFFA3BCF3FB3A9A1AFB7D" box="[1015,1043,1152,1175]" italics="true" pageId="8" pageNumber="7">Eb</emphasis>
<subScript id="17BA3492860EFFA3BB16FB309A12FB73" attach="left" box="[1042,1051,1162,1177]" fontSize="6" pageId="8" pageNumber="7">3</subScript>
,
<emphasis id="B94AEAC5860EFFA3BB2EFB3A9A5BFB7D" box="[1066,1106,1152,1175]" italics="true" pageId="8" pageNumber="7">Esb</emphasis>
and
<emphasis id="B94AEAC5860EFFA3BB89FB3A9AB8FB7D" box="[1165,1201,1152,1175]" italics="true" pageId="8" pageNumber="7">esb</emphasis>
petite; patella with
<emphasis id="B94AEAC5860EFFA3BA88FB3A9B9EFB7D" box="[1420,1431,1152,1175]" italics="true" pageId="8" pageNumber="7">d</emphasis>
<subScript id="17BA3492860EFFA3BA9CFB309BA8FB73" attach="right" box="[1432,1441,1162,1177]" fontSize="6" pageId="8" pageNumber="7">3</subScript>
internal to dorsomedian carina (Fet, Soleglad &amp; Lowe, 2005);
<emphasis id="B94AEAC5860EFFA3BC94FB079DA2FB3E" box="[912,939,1213,1236]" italics="true" pageId="8" pageNumber="7">db</emphasis>
positioned on fixed finger midway between
<emphasis id="B94AEAC5860EFFA3BC3AFB669D55FB19" box="[830,860,1244,1267]" italics="true" pageId="8" pageNumber="7">est</emphasis>
and
<emphasis id="B94AEAC5860EFFA3BC9EFB669DA4FB19" box="[922,941,1244,1267]" italics="true" pageId="8" pageNumber="7">et</emphasis>
(Kovařík, 2007); line joining
<emphasis id="B94AEAC5860EFFA3BA0DFB669B10FB19" box="[1289,1305,1244,1267]" italics="true" pageId="8" pageNumber="7">V</emphasis>
<subScript id="17BA3492860EFFA3BA1DFB5C9B2BFB1F" attach="left" box="[1305,1314,1254,1269]" fontSize="6" pageId="8" pageNumber="7">1</subScript>
and
<emphasis id="B94AEAC5860EFFA3BA64FB669B79FB19" box="[1376,1392,1244,1267]" italics="true" pageId="8" pageNumber="7">V</emphasis>
<subScript id="17BA3492860EFFA3BA75FB5C9B73FB1F" attach="left" box="[1393,1402,1254,1269]" fontSize="6" pageId="8" pageNumber="7">2</subScript>
on manus oblique, not perpendicular to axis of movable finger articulation (Sun, Zhu &amp; Lourenço, 2010).
</paragraph>
<paragraph id="8B8136D7860EFFA3BC3AFAE39DC5F949" blockId="8.[830,1441,1369,1699]" pageId="8" pageNumber="7">
<emphasis id="B94AEAC5860EFFA3BC3AFAE39D7AFA9A" bold="true" box="[830,883,1369,1392]" pageId="8" pageNumber="7">Legs</emphasis>
(
<figureCitation id="13052A52860EFFA3BC91FAE39A07FA9A" box="[917,1038,1369,1392]" captionStart="Figures 1–4" captionStartId="4.[192,278,1910,1934]" captionTargetBox="[233,1392,156,1881]" captionTargetId="figure-12@4.[221,1404,156,1881]" captionTargetPageId="4" captionText="Figures 1–4: H. pellucidus, sp. nov., habitus, 1–2, holotype male; 3–4, paratype female. 1, 3. Dorsal aspect. 2, 4. Ventral aspect. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.12782244" httpUri="https://zenodo.org/record/12782244/files/figure.png" pageId="8" pageNumber="7">Figs. 1–2</figureCitation>
,
<figureCitation id="13052A52860EFFA3BB28FAE39A5DFA9A" box="[1068,1108,1369,1392]" captionStart="Figures 5–11" captionStartId="6.[192,278,1662,1685]" captionTargetBox="[228,1382,170,1606]" captionTargetId="figure-10@6.[192,1441,147,1632]" captionTargetPageId="6" captionText="Figures 5–11: H. pellucidus, sp. nov. 5, 8–11, holotype male; 6–7, paratype male (Jabal Bani Jabir). 5. Carapace and tergites I– II, dorsal aspect. 6–7, right chelicera. 6. Dorsal aspect. 7. Ventral aspect. 8–9, right basitarsus and telotarsus III. 8. Ventral aspect. 9. Retrolateral aspect. 10–11, metasoma and telson. 10. Right lateral aspect. 11. Ventral aspect. Scale bar: 5: 3.19 mm; 6–7: 2.125 mm; 8–9: 1.5 mm; 10–11: 6.22 mm." figureDoi="http://doi.org/10.5281/zenodo.12782246" httpUri="https://zenodo.org/record/12782246/files/figure.png" pageId="8" pageNumber="7">8–9</figureCitation>
). Slender; ventral carinae prominently denticulate on femora, weakly crenulate on tibiae; legs III–IV with tibial spurs; retrolateral pedal spurs simple, non-setose; prolateral pedal spurs basally bifurcate, bearing 0 (I) or 1 (II–IV) macrosetae; prolateral and retrolateral surfaces of basitarsi and telotarsi with numerous macrosetae, not arranged into wide bristle-combs; ventral aspect of basitarsi I–III with single row of short, stout macrosetae; ventral aspect of telotarsi I–IV paired rows of stout spiniform macrosetae; ungues stout.
</paragraph>
<paragraph id="8B8136D7860EFFA2BC3AF9769F60FEB3" blockId="8.[830,1441,1740,1917]" lastBlockId="9.[192,802,199,345]" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">
<emphasis id="B94AEAC5860EFFA3BC3AF9769AA6F909" bold="true" box="[830,1199,1740,1763]" pageId="8" pageNumber="7">
Measurements of
<typeStatus id="54858875860EFFA3BB14F9769A79F909" box="[1040,1136,1740,1763]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
male
</emphasis>
(mm). Total L 60.00; metasoma and telson L 37.50; carapace L 6.97, anterior
<geoCoordinate id="EE0A5010860EFFA3BC3AF8B09D86F8CB" box="[830,911,1802,1825]" degrees="3.57" direction="west" orientation="longitude" pageId="8" pageNumber="7" precision="555" value="-3.57">W 3.57</geoCoordinate>
, posterior
<geoCoordinate id="EE0A5010860EFFA3BB02F8B09A51F8CB" box="[1030,1112,1802,1825]" degrees="7.37" direction="west" orientation="longitude" pageId="8" pageNumber="7" precision="555" value="-7.37">W 7.37</geoCoordinate>
, preocular L 2.79; metasomal segments (L/D/W) I 4.68/3.51/4.37, II 5.42/3.43/4.30, III 5.89/3.52/4.23, IV 6.84/3.61/4.05, V 8.00/3.41/3.72; telson L 7.47; vesicle L 4.44, D 3.12,
<geoCoordinate id="EE0A5010860EFFA3BBE1F8DC9B31F897" box="[1253,1336,1894,1917]" degrees="3.11" direction="west" orientation="longitude" pageId="8" pageNumber="7" precision="555" value="-3.11">W 3.11</geoCoordinate>
; pedipalp chela L 14.38, manus ventral L 4.26, chela
<geoCoordinate id="EE0A5010860FFFA2BDAFFF7D9D09FF34" box="[683,768,199,222]" degrees="2.35" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-2.35">W 2.35</geoCoordinate>
, D 2.42, fixed finger L 9.05, movable finger L 10.42; pedipalp femur L 7.62,
<geoCoordinate id="EE0A5010860FFFA2BEC0FEBE9C1FFEF1" box="[452,534,260,283]" degrees="1.89" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-1.89">W 1.89</geoCoordinate>
, patella L 8.03,
<geoCoordinate id="EE0A5010860FFFA2BDCEFEBE9D17FEF1" box="[714,798,260,283]" degrees="2.56" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-2.56">W 2.56</geoCoordinate>
; pectine L 8.44; leg III patella L 6.33, D 1.58, sternite VII L 4.43,
<geoCoordinate id="EE0A5010860FFFA2BE17FEF89F6CFEB3" box="[275,357,322,345]" degrees="7.27" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-7.27">W 7.27</geoCoordinate>
.
</paragraph>
<paragraph id="8B8136D7860FFFA2BFC4FEC59F30FD40" blockId="9.[192,802,383,682]" pageId="9" pageNumber="8">
<emphasis id="B94AEAC5860FFFA2BFC4FEC59F5DFE7C" bold="true" box="[192,340,383,406]" pageId="9" pageNumber="8">Adult female</emphasis>
(
<typeStatus id="54858875860FFFA2BE66FEC59FCBFE7C" box="[354,450,383,406]" pageId="9" pageNumber="8" type="paratype">paratype</typeStatus>
, Jabal Bani Jabir) (
<figureCitation id="13052A52860FFFA2BD8BFEC59CFCFE7C" box="[655,757,383,406]" captionStart="Figures 1–4" captionStartId="4.[192,278,1910,1934]" captionTargetBox="[233,1392,156,1881]" captionTargetId="figure-12@4.[221,1404,156,1881]" captionTargetPageId="4" captionText="Figures 1–4: H. pellucidus, sp. nov., habitus, 1–2, holotype male; 3–4, paratype female. 1, 3. Dorsal aspect. 2, 4. Ventral aspect. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.12782244" httpUri="https://zenodo.org/record/12782244/files/figure.png" pageId="9" pageNumber="8">Figs. 3–4</figureCitation>
,
<figureCitation id="13052A52860FFFA2BC04FEC59D14FE7C" box="[768,797,383,406]" captionStart="Figures 12–20" captionStartId="7.[192,278,1626,1649]" captionTargetBox="[237,1379,217,1541]" captionTargetId="figure-6@7.[192,1442,168,1594]" captionTargetPageId="7" captionText="Figures 12–20: H. pellucidus, sp.nov., 12–16, 19, holotype male; 17–18, paratype male; 20, paratype female. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17–18. Right hemispermatophore, lobes at base of flagellum. 17. Convex aspect. 18. Inner aspect. 19. Left male pectine, ventral aspect. 20. Left female pectine ventral aspect. Scale bar: 12–16, 19–20: 4.1 mm; 17–18: 1.06 mm." figureDoi="http://doi.org/10.5281/zenodo.12782248" httpUri="https://zenodo.org/record/12782248/files/figure.png" pageId="9" pageNumber="8">20</figureCitation>
,
<figureCitation id="13052A52860FFFA2BFC4FE249ED4FE5F" box="[192,221,414,437]" captionStart="Figures 5–11" captionStartId="6.[192,278,1662,1685]" captionTargetBox="[228,1382,170,1606]" captionTargetId="figure-10@6.[192,1441,147,1632]" captionTargetPageId="6" captionText="Figures 5–11: H. pellucidus, sp. nov. 5, 8–11, holotype male; 6–7, paratype male (Jabal Bani Jabir). 5. Carapace and tergites I– II, dorsal aspect. 6–7, right chelicera. 6. Dorsal aspect. 7. Ventral aspect. 8–9, right basitarsus and telotarsus III. 8. Ventral aspect. 9. Retrolateral aspect. 10–11, metasoma and telson. 10. Right lateral aspect. 11. Ventral aspect. Scale bar: 5: 3.19 mm; 6–7: 2.125 mm; 8–9: 1.5 mm; 10–11: 6.22 mm." figureDoi="http://doi.org/10.5281/zenodo.12782246" httpUri="https://zenodo.org/record/12782246/files/figure.png" pageId="9" pageNumber="8">5 9</figureCitation>
). Similar to male, but differs as follows: a larger scorpion, coarser granules on carapace and tergites; pectines shorter, distal tips extending to proximal 1/4 of trochanter IV, with 7–8 middle lamellae; pectine teeth smaller, shorter; when anterior margins of both pectines are aligned with posterior margins of coxae IV, basal teeth do not overlap; basal piece with more shallow anteromedian incision; lateral carinae obsolete on sternite IV.
</paragraph>
<paragraph id="8B8136D7860FFFA2BFC4FD6A9C2FFBF0" blockId="9.[192,802,720,1050]" pageId="9" pageNumber="8">
<emphasis id="B94AEAC5860FFFA2BFC4FD6A9C6DFD0D" bold="true" box="[192,612,720,743]" pageId="9" pageNumber="8">
Measurements of
<typeStatus id="54858875860FFFA2BEA4FD6A9C0FFD0D" box="[416,518,720,743]" pageId="9" pageNumber="8" type="paratype">paratype</typeStatus>
female
</emphasis>
(mm). Total L 6 4.00; metasoma and telson L 36.50; carapace L 7.40, anterior
<geoCoordinate id="EE0A5010860FFFA2BE24FCB49F7EFCCF" box="[288,375,782,805]" degrees="3.52" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-3.52">W 3.52</geoCoordinate>
, posterior
<geoCoordinate id="EE0A5010860FFFA2BEF0FCB49C45FCCF" box="[500,588,782,805]" degrees="7.66" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-7.66">W 7.66</geoCoordinate>
, preocular L 2.91; metasomal segments (L/D/W) I 4.48/3.58/4.51, II 5.22/3.55/4.33, III 5.54/3.58/4.16, IV 6.51/3.56/3.96, V 8.00/3.39/3.73; telson L 7.37; vesicle L 4.50, D 2.91,
<geoCoordinate id="EE0A5010860FFFA2BC0CFCD09EFBFC75" degrees="3.12" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-3.12">W 3.12</geoCoordinate>
; pedipalp chela L 14.16, manus ventral L 4.00, chela
<geoCoordinate id="EE0A5010860FFFA2BE02FC1D9F57FC54" box="[262,350,935,958]" degrees="2.43" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-2.43">W 2.43</geoCoordinate>
, D 2.43, fixed finger L 9.05, movable finger L 10.18; pedipalp femur L 6.90,
<geoCoordinate id="EE0A5010860FFFA2BD62FC7C9CB1FC37" box="[614,696,966,989]" degrees="1.77" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-1.77">W 1.77</geoCoordinate>
, patella L 8.22,
<geoCoordinate id="EE0A5010860FFFA2BFFBFC5E9F5CFC11" box="[255,341,996,1019]" degrees="2.69" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-2.69">W 2.69</geoCoordinate>
; pectine L 7.31; leg III patella L 6.19, D 1.65; sternite VII L 4.12,
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.
</paragraph>
<paragraph id="8B8136D7860FFFA2BFC4FBF89C0EFAA5" blockId="9.[192,803,1090,1880]" pageId="9" pageNumber="8">
<emphasis id="B94AEAC5860FFFA2BFC4FBF89F27FBB3" bold="true" box="[192,302,1090,1113]" pageId="9" pageNumber="8">Variation</emphasis>
.
<emphasis id="B94AEAC5860FFFA2BE45FBF89FADFBB3" box="[321,420,1090,1113]" italics="true" pageId="9" pageNumber="8">Juveniles</emphasis>
: differed from adults as follows: carapace and tergites with weaker carination, intercarinal surfaces smooth; metasoma I–IV with weaker, finely granulated carinae, intercarinal surfaces smooth; metasoma V smooth except for granules on ventral surface; metasomal setation more sparse, most setae located on carinae; setation on pedipalps, basitarsi and telotarsi more sparse; metasoma V with faint fuscous markings on lateral and ventral surfaces.
</paragraph>
<paragraph id="8B8136D7860FFFA2BFF4FAEC9A2EFF17" blockId="9.[192,803,1090,1880]" lastBlockId="9.[830,1440,199,253]" pageId="9" pageNumber="8">
<emphasis id="B94AEAC5860FFFA2BFF4FAEC9FB7FA87" box="[240,446,1366,1389]" italics="true" pageId="9" pageNumber="8">Sexual dimorphism</emphasis>
: Morphometric data for adults of both sexes are summarized in
<tableCitation id="C6BC036C860FFFA2BD1AFACF9C7CFA66" box="[542,629,1397,1420]" captionStart="Table 1" captionStartId="16.[192,257,1818,1841]" captionText="Table 1: Variation in meristics and selected morphometric ratios for Hottentotta pellucidus, sp.nov. and Hottentotta saxinatans, sp.nov. Indicated are ranges, mean ± standard deviation with sample sizes in parentheses. All samples are from adults except pedipalp denticle subrow counts and pectinal tooth counts, which also include measurements from immatures and from both left and right appendages. Values of variables significantly different between the two sexes (P &lt;0.01 for H. pellucidus, P &lt;0.05 for H. saxinatans; Mann Whitney test) are cited separately for males and females, and those not significantly different are pooled for both sexes." pageId="9" pageNumber="8">Table 1</tableCitation>
. Males had on average significantly longer (relative to carapace) and more slender metasomal segments, pedipalps, legs, pectines and sternite VII than females. Pectine teeth: males 29–35 (mode 33, 62/68 = 91.2% of combs with 31–34 teeth), females 25–33 (mode 29, 60/78 = 76.9% of combs with 27–30 teeth). The number of denticle subrows on the pedipalp fingers of males was not significantly different from that of females. The mean number of subrows for movable fingers was significantly higher than that for fixed fingers (P &lt;0.001, paired t-test,
<emphasis id="B94AEAC5860FFFA2BE5AF97C9F62F937" box="[350,363,1734,1757]" italics="true" pageId="9" pageNumber="8">n</emphasis>
= 142). In 143 fixed fingers (sexes pooled), there were 39/143 (27.3 %) with 13 subrows, 101/143 (70.6 %) with 14 subrows, and 3/143 (2.1 %) with 10 or 12 subrows; in 143 movable fingers (sexes pooled), there were 107/143 (74.8 %) with 14 subrows, 33/143 (23.1 %) with 15 subrows, and 3/143 (2.1 %) with 6 or 13 subrows.
</paragraph>
<paragraph id="8B8136D7860FFFA2BC3AFE999B11FDC5" blockId="9.[830,1440,291,559]" pageId="9" pageNumber="8">
<emphasis id="B94AEAC5860FFFA2BC3AFE999DEAFED0" bold="true" box="[830,995,291,314]" pageId="9" pageNumber="8">Reproduction.</emphasis>
An adult female from Jabal Bani Jabir, collected on
<date id="FF801017860FFFA2BCD5FEF89AA7FEB3" box="[977,1198,322,345]" pageId="9" pageNumber="8" value="1995-09-14">14 September 1995</date>
, was held in isolated captivity, and on
<date id="FF801017860FFFA2BB0DFEDA9AC9FE9D" box="[1033,1216,352,375]" pageId="9" pageNumber="8" value="1998-08-21">21 August 1998</date>
was observed after parturition with a live second instar juvenile and 4 dead embryos. This suggests that this species is capable of storing viable spermatozoa for up to 3 years (Kovoor, Lourenço &amp; Muñoz-Cuevas, 1987). The abundance of males implies that this is not a parthenogenetic population (Matthiesen, 1962; Lourenço, 2002).
</paragraph>
</subSubSection>
<subSubSection id="C324655C860FFFA2BC3AFDEC9B7DFC11" pageId="9" pageNumber="8" type="distribution">
<paragraph id="8B8136D7860FFFA2BC3AFDEC9B7DFC11" blockId="9.[830,1441,598,1019]" pageId="9" pageNumber="8">
<emphasis id="B94AEAC5860FFFA2BC3AFDEC9DC0FD87" bold="true" box="[830,969,598,621]" pageId="9" pageNumber="8">Distribution</emphasis>
(
<figureCitation id="13052A52860FFFA2BCDBFDEC9A60FD87" box="[991,1129,598,621]" captionStart="Figures 65–66" captionStartId="20.[192,278,726,749]" captionTargetBox="[200,1434,148,689]" captionTargetId="figure-10@20.[192,1440,144,695]" captionTargetPageId="20" captionText="Figures 65–66: Maps showing collection sites of new species of Hottentotta in northern Oman. 65. Records plotted on larger scale terrain and elevation map of southeast Arabia with geopolitical borders. 66. Records plotted on color topographic map of northern Oman showing locations in the Al Hajar mountain ranges. Magenta circles: H. pellucidus, sp. nov.; red circles: H. saxinatans, sp. nov. The isolated record of H. pellucidus at Muscat may be spurious." figureDoi="http://doi.org/10.5281/zenodo.12782266" httpUri="https://zenodo.org/record/12782266/files/figure.png" pageId="9" pageNumber="8">Figs. 65–66</figureCitation>
). This species is found on Jabal Bani Jabir in the eastern Al Hajar Mountains (Al Hajar Ash Sharqi) of
<collectingCountry id="F3297647860FFFA2BB35FD299A7CFD40" box="[1073,1141,659,682]" name="Oman" pageId="9" pageNumber="8">Oman</collectingCountry>
, where all specimens were collected at high elevation (&gt;
<quantity id="4CC69B32860FFFA2BB94FD089AE2FD23" box="[1168,1259,690,713]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="8" unit="m" value="1600.0">1,600 m</quantity>
a.s.l.). A single isolated specimen from Ras al Hamra, along the edge of the sea near
<collectingCountry id="F3297647860FFFA2BCD4FD559A28FCEC" box="[976,1057,751,774]" name="Oman" pageId="9" pageNumber="8">Muscat</collectingCountry>
, may be a spurious record due to human transport. There were no other specimens of this species in a large sample of scorpions (2,048) collected from lower elevations (&lt;
<quantity id="4CC69B32860FFFA2BB5AFCF19AB0FC88" box="[1118,1209,843,866]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="8" unit="m" value="1500.0">1,500 m</quantity>
a.s.l.), including the heavily populated coastal areas in the vicinity of
<collectingCountry id="F3297647860FFFA2BA4FFCD09B94FC6B" box="[1355,1437,874,897]" name="Oman" pageId="9" pageNumber="8">Muscat</collectingCountry>
. The lack of records from the western Al Hajar ranges suggests that this species does not cross the Sumail Gap dividing the eastern and western ranges, and may be confined to higher elevations in the eastern Al Hajar.
</paragraph>
</subSubSection>
<subSubSection id="C324655C860FFFA2BC3AFB989B65F995" pageId="9" pageNumber="8" type="biology_ecology">
<paragraph id="8B8136D7860FFFA2BC3AFB989B65F995" blockId="9.[830,1441,1058,1663]" pageId="9" pageNumber="8">
<emphasis id="B94AEAC5860FFFA2BC3AFB989D91FBD3" bold="true" box="[830,920,1058,1081]" pageId="9" pageNumber="8">Ecology</emphasis>
. Ultraviolet detection on Jabal Bani Jabir revealed a dense population of
<taxonomicName id="4C3E4D54860FFFA2BBB6FBFA9B41FBBD" authorityName="Lowe" authorityYear="2010" box="[1202,1352,1088,1111]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="pellucidus">
<emphasis id="B94AEAC5860FFFA2BBB6FBFA9ACCFBBD" box="[1202,1221,1088,1111]" italics="true" pageId="9" pageNumber="8">H</emphasis>
.
<emphasis id="B94AEAC5860FFFA2BBDEFBFA9B41FBBD" box="[1242,1352,1088,1111]" italics="true" pageId="9" pageNumber="8">pellucidus</emphasis>
</taxonomicName>
on the remote high plateau of Shir (
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<geoCoordinate id="EE0A5010860FFFA2BB83FBE59ACAFB9C" box="[1159,1219,1119,1142]" degrees="1,600" direction="north" orientation="latitude" pageId="9" pageNumber="8" precision="55" value="1.6">1,600</geoCoordinate>
<geoCoordinate id="EE0A5010860FFFA2BBC7FBE59B04FB9C" box="[1219,1293,1119,1142]" degrees="1,800" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="55" value="-1.8">–1,800</geoCoordinate>
m
</quantity>
a.s.l.). At the collection sites the terrain was open, very dry, sparsely vegetated and littered with abundant loose rock broken into blocks and slabs (
<figureCitation id="13052A52860FFFA2BBC1FB019B5FFB38" box="[1221,1366,1211,1234]" captionStart="Figures 57–64" captionStartId="19.[192,278,1836,1860]" captionTargetBox="[201,1429,158,1805]" captionTargetId="figure-6@19.[192,1439,154,1805]" captionTargetPageId="19" captionText="Figures 57–64: In vivo habitus and habitats of new Hottentotta species in Oman. 57–59. H. pellucidus, sp. nov. 57–58. Paratype male. 59. Paratype female. 60. H. saxinatans, sp. nov., paratype female. 61–63. Shir Plateau, Jabal Bani Jabir, type locality of H. pellucidus. 61–62. Terrain photos showing extensive fields of loose rock. 63. Prehistoric tower tomb constructed from broken rock.64. Jabal Shams, type locality of H. saxinatans." figureDoi="http://doi.org/10.5281/zenodo.12782264" httpUri="https://zenodo.org/record/12782264/files/figure.png" pageId="9" pageNumber="8">Figs. 61–63</figureCitation>
). The scorpions were found at night sitting stationary, some clinging to rocks, others on the ground under the edges of loose rocks. Many males were wandering out in the open. Other less common lithophilic or lapidicolous scorpions collected in the same area included
<taxonomicName id="4C3E4D54860FFFA2BA5CFAEE9AF1FA60" authority="(Kraepelin, 1900)" baseAuthorityName="Kraepelin" baseAuthorityYear="1900" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis id="B94AEAC5860FFFA2BA5CFAEE9A14FA60" italics="true" pageId="9" pageNumber="8">Compsobuthus maindroni</emphasis>
(Kraepelin, 1900)
</taxonomicName>
,
<taxonomicName id="4C3E4D54860FFFA2BA08FAC99A61FA43" authority="Lowe, 2010" authorityName="Lowe" authorityYear="2010" class="Arachnida" family="Hemiscorpiidae" genus="Hemiscorpius" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="flagelliraptor">
<emphasis id="B94AEAC5860FFFA2BA08FAC99DC7FA43" italics="true" pageId="9" pageNumber="8">Hemiscorpius flagelliraptor</emphasis>
Lowe, 2010
</taxonomicName>
,
<taxonomicName id="4C3E4D54860FFFA2BB78FA289D7AFA2D" authority="Francke, 1980" authorityName="Francke" authorityYear="1980" class="Arachnida" family="Diplocentridae" genus="Nebo" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="omanensis">
<emphasis id="B94AEAC5860FFFA2BB78FA289B3CFA43" box="[1148,1333,1426,1449]" italics="true" pageId="9" pageNumber="8">Nebo omanensis</emphasis>
Francke, 1980
</taxonomicName>
and
<taxonomicName id="4C3E4D54860FFFA2BCB4FA0A9B3BFA2D" authority="Lowe, 2010" authorityName="Lowe" authorityYear="2010" box="[944,1330,1456,1479]" class="Arachnida" family="Buthidae" genus="Microbuthus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="gardneri">
<emphasis id="B94AEAC5860FFFA2BCB4FA0A9AA9FA2D" box="[944,1184,1456,1479]" italics="true" pageId="9" pageNumber="8">Microbuthus gardneri</emphasis>
Lowe, 2010
</taxonomicName>
. The Shir Plateau is an important archaeological site where prehistoric residents of the 3
<superScript id="7C4B9B9F860FFFA2BB5EFA529A61FA1D" attach="left" box="[1114,1128,1512,1527]" fontSize="6" pageId="9" pageNumber="8">rd</superScript>
millennium BC utilized the same broken rock that provides so much scorpion microhabitat to construct tower tombs that still stand as memorials to respected members of their society (
<figureCitation id="13052A52860FFFA2BA71F9F09D53F995" captionStart="Figures 57–64" captionStartId="19.[192,278,1836,1860]" captionTargetBox="[201,1429,158,1805]" captionTargetId="figure-6@19.[192,1439,154,1805]" captionTargetPageId="19" captionText="Figures 57–64: In vivo habitus and habitats of new Hottentotta species in Oman. 57–59. H. pellucidus, sp. nov. 57–58. Paratype male. 59. Paratype female. 60. H. saxinatans, sp. nov., paratype female. 61–63. Shir Plateau, Jabal Bani Jabir, type locality of H. pellucidus. 61–62. Terrain photos showing extensive fields of loose rock. 63. Prehistoric tower tomb constructed from broken rock.64. Jabal Shams, type locality of H. saxinatans." figureDoi="http://doi.org/10.5281/zenodo.12782264" httpUri="https://zenodo.org/record/12782264/files/figure.png" pageId="9" pageNumber="8">Fig. 63</figureCitation>
) (Yule &amp; Weisgerber, 1998; Seibert et al., 2005).
</paragraph>
</subSubSection>
</treatment>
</document>
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