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<document ID-DOI="10.3853/j.2201-4349.71.2019.1699" ID-ISSN="2201-4349" ID-Zenodo-Dep="4653272" ID-ZooBank="BC469398-0572-492D-A29F-37F5BFF9199C" checkinTime="1617216840585" checkinUser="felipe" docAuthor="Shea, Michael &amp; Köhler, Frank" docDate="2019" docId="03978791FFAFFFDAFC5DE8B9FBF6F870" docLanguage="en" docName="RecAustMus.71.4.111-120.pdf" docOrigin="Records of the Australian Museum 71 (4)" docStyle="DocumentStyle:997BA24B490466A2F99594A367FA147F.2:RecAustMus.2019-.journal_article" docStyleId="997BA24B490466A2F99594A367FA147F" docStyleName="RecAustMus.2019-.journal_article" docStyleVersion="2" docTitle="Austrochloritis porteri" docType="treatment" docVersion="3" lastPageNumber="115" masterDocId="FFAEFFE9FFADFFDEFFDFE82FFFD5FFA9" masterDocTitle="Towards a Systematic Revision of the Eastern Australian Land Snail Austrochloritis Pilsbry, 1891 (Eupulmonata, Camaenidae): Re-description of its Type Species, A. porteri (Cox, 1866)" masterLastPageNumber="120" masterPageNumber="111" pageNumber="113" updateTime="1674569275333" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>Towards a Systematic Revision of the Eastern Australian Land Snail Austrochloritis Pilsbry, 1891 (Eupulmonata, Camaenidae): Re-description of its Type Species, A. porteri (Cox, 1866)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shea, Michael</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Köhler, Frank</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Records of the Australian Museum</mods:title>
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<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-07-24</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>71</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
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<mods:extent unit="page">
<mods:start>111</mods:start>
<mods:end>120</mods:end>
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<mods:identifier type="Zenodo-Dep">4653272</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:03978791FFAFFFDAFC5DE8B9FBF6F870" httpUri="http://treatment.plazi.org/id/03978791FFAFFFDAFC5DE8B9FBF6F870" lastPageId="4" lastPageNumber="115" pageId="2" pageNumber="113">
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<heading bold="true" box="[898,1408,150,182]" centered="true" fontSize="12" level="2" pageId="2" pageNumber="113" reason="2">
<taxonomicName ID-CoL="JW4F" authority="(Cox, 1866)" baseAuthorityName="Cox" baseAuthorityYear="1866" box="[898,1408,150,182]" class="Gastropoda" family="Camaenidae" genus="Austrochloritis" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="113" phylum="Mollusca" rank="species" species="porteri">
<emphasis bold="true" box="[898,1408,150,182]" pageId="2" pageNumber="113">
<emphasis bold="true" box="[898,1225,150,182]" italics="true" pageId="2" pageNumber="113">Austrochloritis porteri</emphasis>
(Cox, 1866)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection pageId="2" pageNumber="113" type="reference_group">
<paragraph blockId="2.[869,1437,200,382]" pageId="2" pageNumber="113">
<treatmentCitationGroup pageId="2" pageNumber="113">
<taxonomicName ID-CoL="6LLM5" authority="Cox, 1866: 373" authorityName="Cox" authorityPageNumber="373" authorityYear="1866" box="[869,1153,201,223]" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="113" phylum="Mollusca" rank="species" species="porteri">
<emphasis box="[869,995,201,223]" italics="true" pageId="2" pageNumber="113">Helix porteri</emphasis>
Cox, 1866: 373
</taxonomicName>
[publ. 5 Sep];
<treatmentCitation author="Hedley, C." page="249" pageId="2" pageNumber="113" year="1890">
<bibRefCitation author="Hedley, C." pageId="2" pageNumber="113" pagination="249 - 251" refId="ref6365" refString="Hedley, C. 1890. Anatomical notes on the Helicidae, part III. Proceedings of the Royal Society of Queensland 6: 249 - 251." type="journal article" year="1890">Hedley, 1890: 249250</bibRefCitation>
</treatmentCitation>
, pl. 15.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="2.[869,1437,200,382]" pageId="2" pageNumber="113">
<treatmentCitationGroup pageId="2" pageNumber="113">
<taxonomicName authority="Pilsbry, 1891" box="[869,1226,254,276]" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="113" phylum="Mollusca" rank="species" species="porteri" subGenus="Austrochloritis">
<emphasis box="[869,962,254,275]" italics="true" pageId="2" pageNumber="113">Chloritis</emphasis>
(
<emphasis box="[980,1135,254,275]" italics="true" pageId="2" pageNumber="113">Austrochloritis</emphasis>
)
<emphasis box="[1154,1226,255,276]" italics="true" pageId="2" pageNumber="113">porteri</emphasis>
</taxonomicName>
.—Pilsbry, 1891 (in
<date box="[900,1012,281,303]" pageId="2" pageNumber="113" value="1890" valueMax="1891">18901891</date>
): 263, pl. 58, figs 2324; Pilsbry, 1894 (in
<date box="[900,988,308,330]" pageId="2" pageNumber="113" value="1893" valueMax="1895">189395</date>
): 121, pl. 28, figs 14;
<treatmentCitation author="Zilch, A." box="[1208,1371,308,330]" page="299" pageId="2" pageNumber="113" year="1966">
<bibRefCitation author="Zilch, A." box="[1208,1371,308,330]" pageId="2" pageNumber="113" pagination="293 - 319" refId="ref7009" refString="Zilch, A. 1966. Die Typen und Typoide des Natur-Museums Senckenberg, 35: Mollusca, Camaenidae. Archiv fur Molluskenkunde 95: 293 - 319." type="journal article" year="1966">Zilch, 1966: 299</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="2.[869,1437,200,382]" pageId="2" pageNumber="113">
<treatmentCitationGroup pageId="2" pageNumber="113">
<taxonomicName ID-CoL="JW4F" baseAuthorityName="Cox" baseAuthorityYear="1866" box="[869,1088,334,356]" class="Gastropoda" family="Camaenidae" genus="Austrochloritis" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="113" phylum="Mollusca" rank="species" species="porteri">
<emphasis box="[869,1088,334,356]" italics="true" pageId="2" pageNumber="113">Austrochloritis porteri</emphasis>
</taxonomicName>
.—
<treatmentCitation author="Iredale, T." box="[1116,1289,334,356]" page="94" pageId="2" pageNumber="113" year="1938">
<bibRefCitation author="Iredale, T." box="[1116,1289,334,356]" pageId="2" pageNumber="113" pagination="83 - 124" refId="ref6517" refString="Iredale, T. 1938. A basic list of the land Mollusca of Australia. Part III. Australian Zoologist 9: 83 - 124." type="journal article" year="1938">Iredale, 1938: 94</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Stanisic, J. &amp; M. Shea &amp; D. Potter &amp; O. Griffiths" page="390" pageId="2" pageNumber="113" year="2010">
<bibRefCitation author="Stanisic, J. &amp; M. Shea &amp; D. Potter &amp; O. Griffiths" pageId="2" pageNumber="113" refId="ref6926" refString="Stanisic, J., M. Shea, D. Potter, and O. Griffiths. 2010. Australian land snails. Volume 1. A field guide to eastern Australian species. Mauritius: Bioculture Press." type="book" year="2010">
Stanisic
<emphasis box="[1386,1431,334,356]" italics="true" pageId="2" pageNumber="113">et al</emphasis>
. 2010: 390
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="115" pageId="2" pageNumber="113" type="materials_examined">
<paragraph blockId="2.[1032,1274,400,426]" box="[1032,1274,400,426]" pageId="2" pageNumber="113">
<emphasis bold="true" box="[1032,1274,400,426]" pageId="2" pageNumber="113">Material examined</emphasis>
</paragraph>
<paragraph blockId="2.[839,1466,448,515]" pageId="2" pageNumber="113">
<emphasis box="[839,965,448,467]" italics="true" pageId="2" pageNumber="113">Type material</emphasis>
.
<materialsCitation collectionCode="AM" county="Upper Richmond River" latitude="-28.75" location="Pearces Ck" longLatPrecision="20" longitude="153.44806" municipality="Booyong" pageId="2" pageNumber="113" specimenCode="C.24556" specimenCount="1" stateProvince="New South Wales" typeStatus="neotype">
<typeStatus box="[980,1058,449,468]" pageId="2" pageNumber="113">Neotype</typeStatus>
, herein designated,
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<specimenCode box="[1288,1365,448,467]" pageId="2" pageNumber="113">C.24556</specimenCode>
from
<collectingRegion country="Australia" name="New South Wales" pageId="2" pageNumber="113">New South Wales</collectingRegion>
,
<collectingCounty box="[958,1162,472,492]" pageId="2" pageNumber="113">Upper Richmond River</collectingCounty>
,
<collectingMunicipality box="[1171,1251,473,492]" pageId="2" pageNumber="113">Booyong</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03978791FFAFFFDAFC5DE8B9FBF6F870:8EE1605CFFAFFFDCFB32E9F6FA85FE42" box="[1261,1360,472,492]" county="Upper Richmond River" latitude="-28.75" longLatPrecision="20" longitude="153.44806" municipality="Booyong" name="Pearces Ck" pageId="2" pageNumber="113" stateProvince="New South Wales">Pearces Ck</location>
,
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<geoCoordinate box="[839,948,496,515]" degrees="153" direction="east" minutes="26" orientation="longitude" pageId="2" pageNumber="113" precision="15" seconds="53" value="153.44806">153°26'53&quot;E</geoCoordinate>
(leg. 1906) (
<figureCitation box="[1058,1126,496,515]" captionStart="Figure 1" captionStartId="3.[120,188,1570,1594]" captionTargetBox="[120,1972,533,1550]" captionTargetId="figure-10@3.[120,1972,533,1550]" captionTargetPageId="3" captionText="Figure 1. Shells (front, top, and umbilical view). (AC) Austrochloritis porteri: (A) neotype of Helix porteriAM C.24556 (NSW, Upper Richmond River); (B) AM C.575455 (NSW, Mt Warning); (C) AM C.512605 (Queensland, Lamington NP). (DE) Austrochloritis speculoris: (D) AM C.575464 (NSW, Point Lookout); (E) AM C.575456 (NSW, Cathedral Rock NP). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4653274" httpUri="https://zenodo.org/record/4653274/files/figure.png" pageId="2" pageNumber="113">Fig. 1A</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph blockId="2.[839,1468,539,1254]" pageId="2" pageNumber="113">
<emphasis box="[839,1005,539,558]" italics="true" pageId="2" pageNumber="113">Non-type material</emphasis>
.
<materialsCitation collectionCode="AM" county="Upper Richmond River" elevation="75" latitude="-28.866667" location="Booyong" longLatPrecision="1230" longitude="153.05" municipality="Upper Richmond River" pageId="2" pageNumber="113" specimenCode="C.557014" specimenCount="1" stateProvince="NEW SOUTH WALES">
<collectingRegion box="[1020,1236,539,558]" country="Australia" name="New South Wales" pageId="2" pageNumber="113">NEW SOUTH WALES</collectingRegion>
:
<collectingCounty box="[1247,1463,539,558]" pageId="2" pageNumber="113">Upper Richmond River</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:03978791FFAFFFDAFC5DE8B9FBF6F870:8EE1605CFFAFFFDCFC98EA1CFC42FDEF" box="[839,919,563,582]" county="Upper Richmond River" latitude="-28.866667" longLatPrecision="1230" longitude="153.05" municipality="Upper Richmond River" name="Booyong" pageId="2" pageNumber="113" stateProvince="NEW SOUTH WALES">Booyong</location>
, Pearces Ck,
<geoCoordinate box="[1040,1137,563,582]" degrees="28" direction="south" minutes="45" orientation="latitude" pageId="2" pageNumber="113" precision="15" seconds="00" value="-28.75">28°45'00&quot;S</geoCoordinate>
<geoCoordinate box="[1144,1253,563,582]" degrees="28" direction="south" minutes="45" orientation="latitude" pageId="2" pageNumber="113" precision="15" seconds="00" value="-28.75">153°26'53&quot;E</geoCoordinate>
(
<collectionCode box="[1266,1302,563,582]" country="Australia" name="Australian Museum" pageId="2" pageNumber="113" type="Museum">AM</collectionCode>
<specimenCode box="[1309,1396,563,582]" pageId="2" pageNumber="113">C.557014</specimenCode>
);
</materialsCitation>
<materialsCitation collectionCode="AM" county="Upper Richmond River" elevation="75" latitude="-28.866667" location="Upper Richmond River" longLatPrecision="1230" longitude="153.05" municipality="Upper Richmond River" pageId="2" pageNumber="113" specimenCode="C.334934-35" specimenCount="1" stateProvince="New South Wales">
<collectingMunicipality pageId="2" pageNumber="113">Upper Richmond River</collectingMunicipality>
,
<geoCoordinate box="[988,1084,587,606]" degrees="28" direction="south" minutes="30" orientation="latitude" pageId="2" pageNumber="113" precision="15" seconds="30" value="-28.508333">28°30'30&quot;S</geoCoordinate>
<geoCoordinate box="[1089,1168,587,606]" degrees="152" direction="east" minutes="59" orientation="longitude" pageId="2" pageNumber="113" precision="925" value="152.98334">152°59'E</geoCoordinate>
(
<collectionCode box="[1178,1214,587,606]" country="Australia" name="Australian Museum" pageId="2" pageNumber="113" type="Museum">AM</collectionCode>
<specimenCode box="[1217,1337,587,606]" pageId="2" pageNumber="113">C.33493435</specimenCode>
);
</materialsCitation>
<materialsCitation collectionCode="AM" county="Upper Richmond River" elevation="75" latitude="-28.866667" location="Wilson River" longLatPrecision="1230" longitude="153.05" municipality="Upper Richmond River" pageId="2" pageNumber="113" specimenCode="C.112291" specimenCount="1" stateProvince="New South Wales">
<location LSID="urn:lsid:plazi:treatment:03978791FFAFFFDAFC5DE8B9FBF6F870:8EE1605CFFAFFFDCFA99EA64FA6DFDF7" box="[1350,1464,587,606]" county="Upper Richmond River" latitude="-28.866667" longLatPrecision="1230" longitude="153.05" municipality="Upper Richmond River" name="Wilson River" pageId="2" pageNumber="113" stateProvince="New South Wales">Wilson River</location>
, tributary of
<location LSID="urn:lsid:plazi:treatment:03978791FFAFFFDAFC5DE8B9FBF6F870:8EE1605CFFAFFFDCFC6BEA4CFB9DFDDF" box="[948,1096,611,630]" county="Upper Richmond River" latitude="-28.866667" longLatPrecision="1230" longitude="153.05" municipality="Upper Richmond River" name="Richmond River" pageId="2" pageNumber="113" stateProvince="New South Wales">Richmond River</location>
,
<location LSID="urn:lsid:plazi:treatment:03978791FFAFFFDAFC5DE8B9FBF6F870:8EE1605CFFAFFFDCFB8CEA4CFB71FDDF" box="[1107,1188,611,630]" county="Upper Richmond River" latitude="-28.866667" longLatPrecision="1230" longitude="153.05" municipality="Upper Richmond River" name="Booyong" pageId="2" pageNumber="113" stateProvince="New South Wales">Booyong</location>
,
<geoCoordinate box="[1200,1298,611,630]" degrees="28" direction="south" minutes="44" orientation="latitude" pageId="2" pageNumber="113" precision="15" seconds="53" value="-28.748056">28°44'53&quot;S</geoCoordinate>
<geoCoordinate box="[1306,1416,611,630]" degrees="153" direction="east" minutes="26" orientation="longitude" pageId="2" pageNumber="113" precision="15" seconds="53" value="153.44806">153°26'53&quot;E</geoCoordinate>
(
<collectionCode box="[1430,1466,611,630]" country="Australia" name="Australian Museum" pageId="2" pageNumber="113" type="Museum">AM</collectionCode>
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<materialsCitation collectionCode="AM" county="Upper Richmond River" elevation="75" latitude="-28.866667" location="Richmond River" longLatPrecision="1230" longitude="153.05" municipality="Upper Richmond River" pageId="2" pageNumber="113" specimenCode="C.33390, C.339397, C.334941, C.334943, 45, C.339394, C.339744" specimenCount="1" stateProvince="New South Wales">
<location LSID="urn:lsid:plazi:treatment:03978791FFAFFFDAFC5DE8B9FBF6F870:8EE1605CFFAFFFDCFC74EA54FBE2FD27" box="[939,1079,635,654]" county="Upper Richmond River" latitude="-28.866667" longLatPrecision="1230" longitude="153.05" municipality="Upper Richmond River" name="Richmond River" pageId="2" pageNumber="113" stateProvince="New South Wales">Richmond River</location>
(
<collectionCode box="[1089,1125,635,654]" country="Australia" name="Australian Museum" pageId="2" pageNumber="113" type="Museum">AM</collectionCode>
<specimenCode box="[1128,1204,635,654]" pageId="2" pageNumber="113">C.33390</specimenCode>
,
<collectionCode box="[1210,1244,635,654]" country="Australia" name="Australian Museum" pageId="2" pageNumber="113" type="Museum">AM</collectionCode>
<specimenCode box="[1248,1334,635,654]" pageId="2" pageNumber="113">C.339397</specimenCode>
,
<collectionCode box="[1340,1374,635,654]" country="Australia" name="Australian Museum" pageId="2" pageNumber="113" type="Museum">AM</collectionCode>
<specimenCode box="[1378,1463,635,654]" pageId="2" pageNumber="113">C.334941</specimenCode>
,
<collectionCode box="[839,873,659,678]" country="Australia" name="Australian Museum" pageId="2" pageNumber="113" type="Museum">AM</collectionCode>
<specimenCode box="[881,966,659,678]" pageId="2" pageNumber="113">C.334943</specimenCode>
<specimenCode box="[978,1000,659,678]" pageId="2" pageNumber="113">45</specimenCode>
);
</materialsCitation>
<materialsCitation collectionCode="AM" county="Upper Richmond River" elevation="75" latitude="-28.866667" location="Richmond River" longLatPrecision="1230" longitude="153.05" municipality="Upper Richmond River" pageId="2" pageNumber="113" specimenCode="C.33390, C.339397, C.334941, C.334943, 45, C.339394, C.339744" specimenCount="1" stateProvince="New South Wales">
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<quantity box="[1016,1072,659,678]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="113" unit="km" value="25.0">25 km</quantity>
NE of Kyogle
</locationDeviation>
,
<location LSID="urn:lsid:plazi:treatment:03978791FFAFFFDAFC5DE8B9FBF6F870:8EE1605CFFAFFFDCFB60EABCFAC2FD0F" box="[1215,1303,659,678]" county="Upper Richmond River" latitude="-28.866667" longLatPrecision="1230" longitude="153.05" municipality="Upper Richmond River" name="Wiangarie" pageId="2" pageNumber="113" stateProvince="New South Wales">Wiangarie</location>
SF, off
<collectorName box="[1376,1463,659,678]" pageId="2" pageNumber="113">Forest Rd</collectorName>
,
<geoCoordinate box="[839,936,683,702]" degrees="28" direction="south" minutes="23" orientation="latitude" pageId="2" pageNumber="113" precision="15" seconds="23" value="-28.389723">28°23'23&quot;S</geoCoordinate>
<geoCoordinate box="[940,1038,683,702]" degrees="153" direction="east" minutes="6" orientation="longitude" pageId="2" pageNumber="113" precision="15" seconds="24" value="153.10667">153°6'24&quot;E</geoCoordinate>
(
<collectionCode box="[1048,1084,683,702]" country="Australia" name="Australian Museum" pageId="2" pageNumber="113" type="Museum">AM</collectionCode>
<specimenCode box="[1088,1173,683,702]" pageId="2" pageNumber="113">C.339394</specimenCode>
,
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<materialsCitation collectionCode="AM" county="Upper Richmond River" elevation="75" latitude="-28.866667" location="Red Scrub Flora Reserve" longLatPrecision="1230" longitude="153.05" municipality="Upper Richmond River" pageId="2" pageNumber="113" specimenCode="C.339391, C.339745" specimenCount="1" stateProvince="New South Wales">
<collectorName box="[1318,1463,683,702]" pageId="2" pageNumber="113">Whian Whian SF</collectorName>
,
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,
<geoCoordinate box="[1070,1167,707,726]" degrees="28" direction="south" minutes="38" orientation="latitude" pageId="2" pageNumber="113" precision="15" seconds="12" value="-28.636667">28°38'12&quot;S</geoCoordinate>
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(
<collectionCode box="[1295,1331,707,726]" country="Australia" name="Australian Museum" pageId="2" pageNumber="113" type="Museum">AM</collectionCode>
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,
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Rd,
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(
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<locationDeviation box="[998,1204,1139,1158]" location="Murwillumbah" pageId="2" pageNumber="113">NW of Murwillumbah</locationDeviation>
, nr Springbrook,
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(
<collectionCode box="[971,1009,1163,1182]" country="Australia" name="Australian Museum" pageId="2" pageNumber="113" type="Museum">AM</collectionCode>
<specimenCode box="[1017,1112,1163,1182]" pageId="2" pageNumber="113">C.444266</specimenCode>
);
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,
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(
<collectionCode box="[845,881,1187,1206]" country="Australia" name="Australian Museum" pageId="2" pageNumber="113" type="Museum">AM</collectionCode>
<specimenCode box="[889,978,1187,1206]" pageId="2" pageNumber="113">C.444267</specimenCode>
);
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,
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<quantity box="[1104,1170,1187,1206]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="2" pageNumber="113" unit="mi" value="4.0">4 miles</quantity>
W of coast
</locationDeviation>
,
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<geoCoordinate box="[1356,1467,1187,1206]" degrees="153" direction="east" minutes="25" orientation="longitude" pageId="2" pageNumber="113" precision="15" seconds="19" value="153.42195">153°25'19&quot;E</geoCoordinate>
(
<collectionCode box="[845,881,1211,1230]" country="Australia" name="Australian Museum" pageId="2" pageNumber="113" type="Museum">AM</collectionCode>
<specimenCode box="[887,975,1211,1230]" pageId="2" pageNumber="113">C.363858</specimenCode>
);
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<materialsCitation collectionCode="AM" county="Numinbah Valley" latitude="-28.216667" location="Collins Gap" longLatPrecision="1233" longitude="153.23334" municipality="Valley Rd" pageId="2" pageNumber="113" specimenCode="C.444259" specimenCount="1" stateProvince="Queensland">
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,
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,
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<geoCoordinate box="[1340,1418,1211,1230]" degrees="152" direction="east" minutes="42" orientation="longitude" pageId="2" pageNumber="113" precision="925" value="152.7">152°42'E</geoCoordinate>
(
<collectionCode box="[1431,1467,1211,1230]" country="Australia" name="Australian Museum" pageId="2" pageNumber="113" type="Museum">AM</collectionCode>
<specimenCode box="[839,926,1235,1254]" pageId="2" pageNumber="113">C.444259</specimenCode>
);
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<materialsCitation box="[941,1405,1235,1254]" collectionCode="AM" county="Numinbah Valley" latitude="-28.216667" location="Natural Bridge NP" longLatPrecision="1233" longitude="153.23334" municipality="Valley Rd" pageId="2" pageNumber="113" specimenCode="C.129279" specimenCount="1" stateProvince="Natural Bridge NP">
<collectingRegion box="[941,1101,1235,1254]" pageId="2" pageNumber="113">Natural Bridge NP</collectingRegion>
,
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<geoCoordinate box="[1181,1261,1235,1254]" degrees="153" direction="east" minutes="14" orientation="longitude" pageId="2" pageNumber="113" precision="925" value="153.23334">153°14'E</geoCoordinate>
(
<collectionCode box="[1271,1307,1235,1254]" country="Australia" name="Australian Museum" pageId="2" pageNumber="113" type="Museum">AM</collectionCode>
<specimenCode box="[1312,1400,1235,1254]" pageId="2" pageNumber="113">C.129279</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="2.[902,1404,1282,1309]" box="[902,1404,1282,1309]" pageId="2" pageNumber="113">
<emphasis bold="true" box="[902,1404,1282,1309]" pageId="2" pageNumber="113">Nomenclatural and taxonomic remarks</emphasis>
</paragraph>
<paragraph blockId="2.[839,1467,1327,2026]" pageId="2" pageNumber="113">
The original
<typeStatus box="[972,1017,1328,1352]" pageId="2" pageNumber="113">type</typeStatus>
material could not be traced in the
<collectionCode box="[1377,1422,1327,1351]" country="Australia" name="Australian Museum" pageId="2" pageNumber="113" type="Museum">AM</collectionCode>
, nor in the
<collectionCode box="[905,951,1357,1381]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="2" pageNumber="113" type="Museum">QM</collectionCode>
,
<collectionCode box="[961,1023,1357,1380]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="2" pageNumber="113" type="Museum">NMV</collectionCode>
,
<collectionCode box="[1033,1093,1357,1380]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="2" pageNumber="113" type="Museum">ZMB</collectionCode>
,
<collectionCode box="[1104,1158,1357,1381]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="113">SMF</collectionCode>
,
<collectionCode box="[1168,1272,1357,1380]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="2" pageNumber="113" type="Museum">NHMUK</collectionCode>
, and is considered lost. Coxs
<typeStatus box="[973,1032,1386,1410]" pageId="2" pageNumber="113">types</typeStatus>
have generally been deposited in the collection of the Australian Museum. Indeed, other historic shell material collected by both Porter and Macgillivray in northern NSW in or before 1866 is still extant in this collection. Repeated efforts by various persons (Alan Solem, John Stanisic, ourselves) to find
<typeStatus box="[1201,1247,1533,1557]" pageId="2" pageNumber="113">type</typeStatus>
material at the
<collectionCode box="[1423,1466,1533,1557]" country="Australia" name="Australian Museum" pageId="2" pageNumber="113" type="Museum">AM</collectionCode>
or other institutions have been unsuccessful. No
<typeStatus box="[1369,1425,1562,1586]" pageId="2" pageNumber="113">types</typeStatus>
are known or suspected to be kept in other museums.
<bibRefCitation author="Zilch, A." pageId="2" pageNumber="113" pagination="293 - 319" refId="ref7009" refString="Zilch, A. 1966. Die Typen und Typoide des Natur-Museums Senckenberg, 35: Mollusca, Camaenidae. Archiv fur Molluskenkunde 95: 293 - 319." type="journal article" year="1966">Zilch (1966)</bibRefCitation>
did not report any
<typeStatus box="[1114,1170,1621,1645]" pageId="2" pageNumber="113">types</typeStatus>
to be held by the
<collectionCode box="[1361,1416,1621,1645]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="113">SMF</collectionCode>
.
</paragraph>
<paragraph blockId="2.[839,1467,1327,2026]" lastBlockId="4.[166,794,151,469]" lastPageId="4" lastPageNumber="115" pageId="2" pageNumber="113">
The original description was based on material collected by two different collectors at two separate sites: “Upper Clarence River at Guy Faux Station (Porter)”and “Upper Richmond River, at Cowlong […] (Macgillivray)”. Using contemporary georeferenced materials, we have located these two sites with reasonable precision. The first site is Guy Fawkes Station, N of Ebor and W of Point Lookout within the uppermost Clarence River catchment (c.
<geoCoordinate box="[1178,1301,1855,1879]" degrees="30" direction="south" minutes="21" orientation="latitude" pageId="2" pageNumber="113" precision="15" seconds="00" value="-30.35">30°21'00&quot;S</geoCoordinate>
<geoCoordinate box="[1312,1455,1855,1879]" degrees="152" direction="east" minutes="22" orientation="longitude" pageId="2" pageNumber="113" precision="15" seconds="12" value="152.37">152°22'12&quot;E</geoCoordinate>
). The second is NE of Lismore, S of Booyong (c.
<geoCoordinate box="[1347,1467,1884,1908]" degrees="28" direction="south" minutes="46" orientation="latitude" pageId="2" pageNumber="113" precision="15" seconds="12" value="-28.77">28°46'12&quot;S</geoCoordinate>
<geoCoordinate box="[839,973,1914,1938]" degrees="153" direction="east" minutes="25" orientation="longitude" pageId="2" pageNumber="113" precision="15" seconds="12" value="153.42">153°25'12&quot;E</geoCoordinate>
) (
<figureCitation box="[994,1057,1914,1938]" captionStart="Figure 2" captionStartId="4.[839,906,1357,1379]" captionTargetBox="[839,1466,151,1343]" captionTargetId="figure-558@4.[839,1466,151,1343]" captionTargetPageId="4" captionText="Figure 2. Occurrence records of Austrochloritis porteri (blue dots) and A. speculoris (red dots) from the malacological collection of theAM.Legend:blue triangle—type locality of A. porteri based on neotypeAM C.24556, red triangle—type location of A. speculoris, original type locality of Helix porteri: 1—Guy Fawkes Station near Ebor; 2—Cowlong. Source of base map: Google Maps." figureDoi="http://doi.org/10.5281/zenodo.4653276" httpUri="https://zenodo.org/record/4653276/files/figure.png" pageId="2" pageNumber="113">Fig. 2</figureCitation>
). Both locations are about
<quantity box="[1331,1409,1914,1938]" metricMagnitude="5" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="113" unit="km" value="230.0">230 km</quantity>
apart from each other. Since the type material was collected in 1866, the original forest cover that existed at this time has largely been destroyed through clearing for agriculture. However, we have examined material from as close to these two locations as possible and found that the
<taxonomicName authorityName="Pilsbry" authorityYear="1891" box="[487,643,181,205]" class="Gastropoda" family="Camaenidae" genus="Austrochloritis" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="115" phylum="Mollusca" rank="genus">
<emphasis box="[487,643,181,205]" italics="true" pageId="4" pageNumber="115">Austrochloritis</emphasis>
</taxonomicName>
species found near one (Guy Fawkes) is not identical with that found near the other site (Cowlong) even though they are both similar in external appearance. It is not impossible to identify
<taxonomicName baseAuthorityName="Cox" baseAuthorityYear="1866" box="[694,793,269,293]" class="Gastropoda" family="Camaenidae" genus="Austrochloritis" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="115" phylum="Mollusca" rank="species" species="porteri">
<emphasis box="[694,793,269,293]" italics="true" pageId="4" pageNumber="115">A. porteri</emphasis>
</taxonomicName>
among these two species based on the diagnosis given in the original description. In order to remove the uncertain identity of
<taxonomicName authority="Cox, 1866" authorityName="Cox" authorityYear="1866" box="[194,447,357,381]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="115" phylum="Mollusca" rank="species" species="porteri">
<emphasis box="[194,330,357,381]" italics="true" pageId="4" pageNumber="115">Helix porteri</emphasis>
Cox, 1866
</taxonomicName>
, to stabilize this taxonomic name and to fix the type locality, we here designate a
<typeStatus box="[682,765,387,411]" pageId="4" pageNumber="115">neotype</typeStatus>
in accordance with Art. 75 of The Code (
<bibRefCitation author="ICZN" box="[601,737,416,440]" pageId="4" pageNumber="115" refId="ref6450" refString="ICZN. 1999. International Code of Zoological Nomenclature, 4 th Edition. 1 - 306, London: International Trust for Zoological Nomenclature." type="book" year="1999">ICZN, 1999</bibRefCitation>
) and provide an updated diagnosis.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4653274" ID-Zenodo-Dep="4653274" httpUri="https://zenodo.org/record/4653274/files/figure.png" pageId="3" pageNumber="114" startId="3.[120,188,1570,1594]" targetBox="[120,1972,533,1550]" targetPageId="3">
<paragraph blockId="3.[120,1974,1570,1619]" pageId="3" pageNumber="114">
<emphasis bold="true" box="[120,205,1570,1594]" pageId="3" pageNumber="114">Figure 1</emphasis>
. Shells (front, top, and umbilical view).
<emphasis box="[593,871,1570,1593]" italics="true" pageId="3" pageNumber="114">
(AC)
<taxonomicName baseAuthorityName="Cox" baseAuthorityYear="1866" box="[656,871,1570,1593]" class="Gastropoda" family="Camaenidae" genus="Austrochloritis" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="114" phylum="Mollusca" rank="species" species="porteri">Austrochloritis porteri</taxonomicName>
</emphasis>
:
<emphasis box="[883,913,1570,1593]" italics="true" pageId="3" pageNumber="114">(A)</emphasis>
neotype of
<taxonomicName authority="AM C." authorityName="AM C." box="[1024,1215,1570,1593]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="114" phylum="Mollusca" rank="species" species="porteri">
<emphasis box="[1024,1148,1570,1593]" italics="true" pageId="3" pageNumber="114">Helix porteri</emphasis>
AM C.
</taxonomicName>
24556 (NSW, Upper Richmond River);
<emphasis box="[1595,1625,1570,1593]" italics="true" pageId="3" pageNumber="114">(B)</emphasis>
AM C.575455 (NSW, Mt Warning);
<emphasis box="[120,152,1596,1619]" italics="true" pageId="3" pageNumber="114">(C)</emphasis>
AM C.512605 (Queensland, Lamington NP).
<emphasis box="[596,909,1596,1619]" italics="true" pageId="3" pageNumber="114">
(DE)
<taxonomicName authorityName="Shea &amp; Griffiths" authorityYear="2010" box="[662,909,1596,1619]" class="Gastropoda" family="Camaenidae" genus="Austrochloritis" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="114" phylum="Mollusca" rank="species" species="speculoris">Austrochloritis speculoris</taxonomicName>
</emphasis>
:
<emphasis box="[921,954,1596,1619]" italics="true" pageId="3" pageNumber="114">(D)</emphasis>
AM C.575464 (NSW, Point Lookout);
<emphasis box="[1332,1362,1596,1619]" italics="true" pageId="3" pageNumber="114">(E)</emphasis>
AM C.575456 (NSW, Cathedral Rock NP). Scale bar = 10 mm.
</paragraph>
</caption>
<paragraph blockId="4.[166,794,503,939]" pageId="4" pageNumber="115">
<bibRefCitation author="Hedley, C." box="[197,350,503,528]" pageId="4" pageNumber="115" pagination="249 - 251" refId="ref6365" refString="Hedley, C. 1890. Anatomical notes on the Helicidae, part III. Proceedings of the Royal Society of Queensland 6: 249 - 251." type="journal article" year="1890">Hedley (1890)</bibRefCitation>
and Pilsbry (1894) published descriptions and figures of the digestive and reproductive anatomy and attributed these to
<taxonomicName box="[382,493,563,586]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="115" phylum="Mollusca" rank="species" species="porteri">
<emphasis box="[382,493,563,586]" italics="true" pageId="4" pageNumber="115">A. porteri</emphasis>
</taxonomicName>
.
<bibRefCitation author="Hedley, C." box="[511,693,562,586]" pageId="4" pageNumber="115" pagination="249 - 251" refId="ref6365" refString="Hedley, C. 1890. Anatomical notes on the Helicidae, part III. Proceedings of the Royal Society of Queensland 6: 249 - 251." type="journal article" year="1890">Hedleys (1890)</bibRefCitation>
material originated from near Brisbane, which is not very close to any of the two original locations. The whereabouts of Pilsbrys (1894) material have remained uncertain.
<bibRefCitation author="Stanisic, J. &amp; M. Shea &amp; D. Potter &amp; O. Griffiths" pageId="4" pageNumber="115" refId="ref6926" refString="Stanisic, J., M. Shea, D. Potter, and O. Griffiths. 2010. Australian land snails. Volume 1. A field guide to eastern Australian species. Mauritius: Bioculture Press." type="book" year="2010">
Stanisic
<emphasis box="[166,217,679,703]" italics="true" pageId="4" pageNumber="115">et al</emphasis>
. (2010)
</bibRefCitation>
reported the species to be distributed from SE of Lismore (Victoria Park, NSW) to W of
<collectingCountry box="[671,794,709,733]" name="Ghana" pageId="4" pageNumber="115">Gold Coast</collectingCountry>
(Springbrook, Qld). This distribution incorporates the second site mentioned in the original description (Cowlong), but not the first. By contrast, the species occurring at the first site, Guy Fawkes Station, has subsequently been described as
<taxonomicName authority="Shea &amp; Griffiths, 2010" authorityName="Shea &amp; Griffiths" authorityYear="2010" class="Gastropoda" family="Camaenidae" genus="Austrochloritis" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="115" phylum="Mollusca" rank="species" species="speculoris">
<emphasis italics="true" pageId="4" pageNumber="115">A. speculoris</emphasis>
Shea &amp; Griffiths, 2010
</taxonomicName>
(see below). To conserve the subsequent use of these two species names, we chose the
<typeStatus box="[166,250,915,939]" pageId="4" pageNumber="115">neotype</typeStatus>
to come from close to site 2 (Cowlong).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="115" type="description">
<paragraph blockId="4.[406,554,971,997]" box="[406,554,971,997]" pageId="4" pageNumber="115">
<emphasis bold="true" box="[406,554,971,997]" pageId="4" pageNumber="115">Description</emphasis>
</paragraph>
<paragraph blockId="4.[166,794,1018,2010]" pageId="4" pageNumber="115">
<emphasis box="[166,219,1018,1042]" italics="true" pageId="4" pageNumber="115">Shell</emphasis>
(
<figureCitation box="[235,294,1018,1042]" captionStart="Figure 1" captionStartId="3.[120,188,1570,1594]" captionTargetBox="[120,1972,533,1550]" captionTargetId="figure-10@3.[120,1972,533,1550]" captionTargetPageId="3" captionText="Figure 1. Shells (front, top, and umbilical view). (AC) Austrochloritis porteri: (A) neotype of Helix porteriAM C.24556 (NSW, Upper Richmond River); (B) AM C.575455 (NSW, Mt Warning); (C) AM C.512605 (Queensland, Lamington NP). (DE) Austrochloritis speculoris: (D) AM C.575464 (NSW, Point Lookout); (E) AM C.575456 (NSW, Cathedral Rock NP). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4653274" httpUri="https://zenodo.org/record/4653274/files/figure.png" pageId="4" pageNumber="115">Fig. 1</figureCitation>
AC,
<figureCitation box="[362,427,1018,1042]" captionStart="Figure 3" captionStartId="5.[121,189,1712,1734]" captionTargetBox="[121,1421,300,1697]" captionTargetId="figure-10@5.[121,1421,300,1697]" captionTargetPageId="5" captionText="Figure 3. Scanning electron micrographs of shells of Austrochloritis porteri showing protoconch (left) and detail of teleoconch sculpture (right). (A, B) AM C.557014 (topotype, Upper Richmond River); (C, D) AM C.575262 (NSW, Mt Warning); (E, F) AM C.444258 (Queensland, Lamington NP). Scale bars = 0.2 or 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4653280" httpUri="https://zenodo.org/record/4653280/files/figure.png" pageId="4" pageNumber="115">Fig. 3</figureCitation>
). Large (D = 1424 (average 17.8) mm, H = 1015 (average 12) mm; for n = 34 lots), subglobose to turbinate in shape with a low to moderately raised spire, with on average 4.25 rounded to sub-angulate whorl that increase in diameter, sutures moderately impressed; protoconch sculpture of scattered pustules and rugose pustulose radial ridges; teleoconch sculpture of low irregular growth lines with microsculpture of small crowded pustules, periostracal sculpture of crowded short curved setae, microsculpture of wavy periostracal ridgelets; end of last whorl descending below whorl plane; aperture moderately tilted from axis of coiling, with moderately thickened and reflected white outer lip with sulcus behind, columella slope about 45°; umbilicus narrow and partially covered by reflected columella, V-shaped in profile; shell colour from pale reddish brown to dark reddish brown, with or without a narrow reddish brown spiral colour band at whorl periphery.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4653276" ID-Zenodo-Dep="4653276" httpUri="https://zenodo.org/record/4653276/files/figure.png" pageId="4" pageNumber="115" startId="4.[839,906,1357,1379]" targetBox="[839,1466,151,1343]" targetPageId="4">
<paragraph blockId="4.[839,1467,1357,1513]" pageId="4" pageNumber="115">
<emphasis bold="true" box="[839,922,1357,1379]" pageId="4" pageNumber="115">Figure 2</emphasis>
. Occurrence records of
<taxonomicName baseAuthorityName="Cox" baseAuthorityYear="1866" box="[1149,1361,1358,1379]" class="Gastropoda" family="Camaenidae" genus="Austrochloritis" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="115" phylum="Mollusca" rank="species" species="porteri">
<emphasis box="[1149,1361,1358,1379]" italics="true" pageId="4" pageNumber="115">Austrochloritis porteri</emphasis>
</taxonomicName>
(blue dots) and
<taxonomicName authorityName="Shea &amp; Griffiths" authorityYear="2010" box="[881,1007,1384,1406]" class="Gastropoda" family="Camaenidae" genus="Austrochloritis" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="115" phylum="Mollusca" rank="species" species="speculoris">
<emphasis box="[881,1007,1384,1406]" italics="true" pageId="4" pageNumber="115">A. speculoris</emphasis>
</taxonomicName>
(red dots) from the malacological collection of the AM. Legend: blue triangle—type locality of
<taxonomicName baseAuthorityName="Cox" baseAuthorityYear="1866" box="[1289,1380,1411,1433]" class="Gastropoda" family="Camaenidae" genus="Austrochloritis" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="115" phylum="Mollusca" rank="species" species="porteri">
<emphasis box="[1289,1380,1411,1433]" italics="true" pageId="4" pageNumber="115">A. porteri</emphasis>
</taxonomicName>
based on neotype AM C.24556, red triangle—type location of
<taxonomicName authorityName="Shea &amp; Griffiths" authorityYear="2010" box="[1337,1460,1438,1459]" class="Gastropoda" family="Camaenidae" genus="Austrochloritis" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="115" phylum="Mollusca" rank="species" species="speculoris">
<emphasis box="[1337,1460,1438,1459]" italics="true" pageId="4" pageNumber="115">A. speculoris</emphasis>
</taxonomicName>
, original type locality of
<taxonomicName box="[1062,1184,1464,1486]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="115" phylum="Mollusca" rank="species" species="porteri">
<emphasis box="[1062,1184,1464,1486]" italics="true" pageId="4" pageNumber="115">Helix porteri</emphasis>
</taxonomicName>
: 1—Guy Fawkes Station near Ebor; 2—Cowlong. Source of base map: Google Maps.
</paragraph>
</caption>
<paragraph blockId="4.[166,794,1018,2010]" pageId="4" pageNumber="115">
<emphasis box="[197,378,1517,1541]" italics="true" pageId="4" pageNumber="115">General anatomy</emphasis>
. A retractable head wart is present at the inner bases of the ocular tentacles.Animal head-foot dark grey.
</paragraph>
<paragraph blockId="4.[166,794,1018,2010]" lastBlockId="4.[839,1467,1588,1788]" pageId="4" pageNumber="115">
<emphasis box="[197,447,1575,1599]" italics="true" pageId="4" pageNumber="115">Reproductive anatomy</emphasis>
(
<figureCitation box="[466,536,1575,1600]" captionStart="Figure 4" captionStartId="6.[166,234,1710,1732]" captionTargetBox="[169,1441,198,1678]" captionTargetId="figure-9@6.[166,1466,176,1706]" captionTargetPageId="6" captionText="Figure 4. Reproductive anatomy of Austrochloritis porteri. (AC) Drawings showing the entire reproductive system: (A) AM C.363835 (NSW, Upper Richmond River); (B) AM C.575455 (NSW, Mt Warning); (C) AM C.512605 (Queensland, Lamington NP). Scale bars = 5 mm. (DF) Details of penial anatomy: (D) AM C.363835 (NSW, Upper Richmond River); (E) AM C.575455 (NSW, Mt Warning); (F) AM C.512605 (Queensland, Lamington NP). Scale bars = 3 mm.Abbreviations: ag—albumen gland; at—atrium; bc—bursa copulatirx; ep—epiphallus; fl—epiphallic flagellum; hd—hermaphroditic duct; ipw—inner penial wall; p—penis; pm—penial retractor; pv—penial verge; va—vagina; vd—vas deferens." figureDoi="http://doi.org/10.5281/zenodo.4653282" httpUri="https://zenodo.org/record/4653282/files/figure.png" pageId="4" pageNumber="115">Fig. 4</figureCitation>
). Penis cylindrical, no penial sheath, inner penial wall with corrugated interlocking longitudinal pilasters, distally giving rise to longitudinal rows of interlocking pilasters; epiphallus cylindrical, about 1.5 to 2 times as long as penis, distal end with well-developed epiphallic flagellum, broad at base, epiphallus tapering to very narrow toward apex, opening into penial lumen through a conical longitudinally grooved (
<collectingRegion box="[531,619,1781,1805]" country="Australia" name="Victoria" pageId="4" pageNumber="115">Victoria</collectingRegion>
Park and Binna Burra;
<figureCitation box="[241,348,1810,1834]" captionStart="Figure 4" captionStartId="6.[166,234,1710,1732]" captionTargetBox="[169,1441,198,1678]" captionTargetId="figure-9@6.[166,1466,176,1706]" captionTargetPageId="6" captionText="Figure 4. Reproductive anatomy of Austrochloritis porteri. (AC) Drawings showing the entire reproductive system: (A) AM C.363835 (NSW, Upper Richmond River); (B) AM C.575455 (NSW, Mt Warning); (C) AM C.512605 (Queensland, Lamington NP). Scale bars = 5 mm. (DF) Details of penial anatomy: (D) AM C.363835 (NSW, Upper Richmond River); (E) AM C.575455 (NSW, Mt Warning); (F) AM C.512605 (Queensland, Lamington NP). Scale bars = 3 mm.Abbreviations: ag—albumen gland; at—atrium; bc—bursa copulatirx; ep—epiphallus; fl—epiphallic flagellum; hd—hermaphroditic duct; ipw—inner penial wall; p—penis; pm—penial retractor; pv—penial verge; va—vagina; vd—vas deferens." figureDoi="http://doi.org/10.5281/zenodo.4653282" httpUri="https://zenodo.org/record/4653282/files/figure.png" pageId="4" pageNumber="115">Figs4 D,F</figureCitation>
) to pustulose (Mt Warning;
<figureCitation box="[646,733,1810,1834]" captionStart="Figure 4" captionStartId="6.[166,234,1710,1732]" captionTargetBox="[169,1441,198,1678]" captionTargetId="figure-9@6.[166,1466,176,1706]" captionTargetPageId="6" captionText="Figure 4. Reproductive anatomy of Austrochloritis porteri. (AC) Drawings showing the entire reproductive system: (A) AM C.363835 (NSW, Upper Richmond River); (B) AM C.575455 (NSW, Mt Warning); (C) AM C.512605 (Queensland, Lamington NP). Scale bars = 5 mm. (DF) Details of penial anatomy: (D) AM C.363835 (NSW, Upper Richmond River); (E) AM C.575455 (NSW, Mt Warning); (F) AM C.512605 (Queensland, Lamington NP). Scale bars = 3 mm.Abbreviations: ag—albumen gland; at—atrium; bc—bursa copulatirx; ep—epiphallus; fl—epiphallic flagellum; hd—hermaphroditic duct; ipw—inner penial wall; p—penis; pm—penial retractor; pv—penial verge; va—vagina; vd—vas deferens." figureDoi="http://doi.org/10.5281/zenodo.4653282" httpUri="https://zenodo.org/record/4653282/files/figure.png" pageId="4" pageNumber="115">Fig. 4 E</figureCitation>
), free verge, opening laterally; penial retractor attached to midepiphallus; vas deferens entering head of epiphallus through single pore just below base of epiphallic flagellum; vas deferens quite broad at its junction with apex of epiphallus but later tapering to a narrow tubule; vagina cylindrical, twice as long as penis, inner wall with prominent longitudinal anastomising pilasters usually thickened around vaginal entrance; free oviduct very short; bursa copulatrix long and broad, particularly at its base, folded or kinked several times, about as long as spermoviduct (
<collectingRegion box="[1195,1284,1676,1700]" country="Australia" name="Victoria" pageId="4" pageNumber="115">Victoria</collectingRegion>
Park and Binna Burra;
<figureCitation box="[911,1018,1705,1729]" captionStart="Figure 4" captionStartId="6.[166,234,1710,1732]" captionTargetBox="[169,1441,198,1678]" captionTargetId="figure-9@6.[166,1466,176,1706]" captionTargetPageId="6" captionText="Figure 4. Reproductive anatomy of Austrochloritis porteri. (AC) Drawings showing the entire reproductive system: (A) AM C.363835 (NSW, Upper Richmond River); (B) AM C.575455 (NSW, Mt Warning); (C) AM C.512605 (Queensland, Lamington NP). Scale bars = 5 mm. (DF) Details of penial anatomy: (D) AM C.363835 (NSW, Upper Richmond River); (E) AM C.575455 (NSW, Mt Warning); (F) AM C.512605 (Queensland, Lamington NP). Scale bars = 3 mm.Abbreviations: ag—albumen gland; at—atrium; bc—bursa copulatirx; ep—epiphallus; fl—epiphallic flagellum; hd—hermaphroditic duct; ipw—inner penial wall; p—penis; pm—penial retractor; pv—penial verge; va—vagina; vd—vas deferens." figureDoi="http://doi.org/10.5281/zenodo.4653282" httpUri="https://zenodo.org/record/4653282/files/figure.png" pageId="4" pageNumber="115">Figs 4A,C</figureCitation>
) to 1.5 times longer (Mt Warning;
<figureCitation box="[1374,1454,1705,1729]" captionStart="Figure 4" captionStartId="6.[166,234,1710,1732]" captionTargetBox="[169,1441,198,1678]" captionTargetId="figure-9@6.[166,1466,176,1706]" captionTargetPageId="6" captionText="Figure 4. Reproductive anatomy of Austrochloritis porteri. (AC) Drawings showing the entire reproductive system: (A) AM C.363835 (NSW, Upper Richmond River); (B) AM C.575455 (NSW, Mt Warning); (C) AM C.512605 (Queensland, Lamington NP). Scale bars = 5 mm. (DF) Details of penial anatomy: (D) AM C.363835 (NSW, Upper Richmond River); (E) AM C.575455 (NSW, Mt Warning); (F) AM C.512605 (Queensland, Lamington NP). Scale bars = 3 mm.Abbreviations: ag—albumen gland; at—atrium; bc—bursa copulatirx; ep—epiphallus; fl—epiphallic flagellum; hd—hermaphroditic duct; ipw—inner penial wall; p—penis; pm—penial retractor; pv—penial verge; va—vagina; vd—vas deferens." figureDoi="http://doi.org/10.5281/zenodo.4653282" httpUri="https://zenodo.org/record/4653282/files/figure.png" pageId="4" pageNumber="115">Fig. 4B</figureCitation>
), with inflated bulb-like head, aligning with base of albumen gland; hermaphroditic duct inserting into head of talon.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="115" type="distribution">
<paragraph blockId="4.[995,1311,1820,1847]" box="[995,1311,1820,1847]" pageId="4" pageNumber="115">
<emphasis bold="true" box="[995,1311,1820,1847]" pageId="4" pageNumber="115">Distribution and ecology</emphasis>
</paragraph>
<paragraph blockId="4.[839,1466,1868,2009]" pageId="4" pageNumber="115">
Lives in subtropical rainforests from the Richmond River valley of NE NSW north to the upper Nerang Creek valley of SE Qld. Mainly found at altitudes of up to
<quantity box="[1282,1348,1927,1951]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="115" unit="m" value="900.0">900 m</quantity>
on basaltic bedrock; found under logs, rocks and inside sheaths of fallen palm leaves (
<figureCitation box="[980,1046,1985,2009]" captionStart="Figure 2" captionStartId="4.[839,906,1357,1379]" captionTargetBox="[839,1466,151,1343]" captionTargetId="figure-558@4.[839,1466,151,1343]" captionTargetPageId="4" captionText="Figure 2. Occurrence records of Austrochloritis porteri (blue dots) and A. speculoris (red dots) from the malacological collection of theAM.Legend:blue triangle—type locality of A. porteri based on neotypeAM C.24556, red triangle—type location of A. speculoris, original type locality of Helix porteri: 1—Guy Fawkes Station near Ebor; 2—Cowlong. Source of base map: Google Maps." figureDoi="http://doi.org/10.5281/zenodo.4653276" httpUri="https://zenodo.org/record/4653276/files/figure.png" pageId="4" pageNumber="115">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>