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<mods:title id="D22798B14C945F065512835D5DDA8055">Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution</mods:title>
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<paragraph id="8B81369EB86EFEDFC1E36F15AF9D9E3A" blockId="282.[151,486,1830,1856]" box="[151,486,1830,1856]" pageId="282" pageNumber="283">
<heading id="D0C981F2B86EFEDFC1E36F15AF9D9E3A" bold="true" box="[151,486,1830,1856]" fontSize="11" level="1" pageId="282" pageNumber="283" reason="1">
<emphasis id="B94AEA8CB86EFEDFC1E36F15AF9D9E3A" bold="true" box="[151,486,1830,1856]" pageId="282" pageNumber="283">
Tribe
<taxonomicName id="4C3E4D1DB86EFEDFC1966F15AF9D9E3A" ID-CoL="8VR85" authority="Casey, 1906" authorityName="Casey" authorityYear="1906" box="[226,486,1830,1856]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="282" pageNumber="283" phylum="Arthropoda" rank="tribe" tribe="Diochini">
Diochini
<bibRefCitation id="EFAF4B6FB86EFEDFC0276F15AF9D9E3A" author="Casey, T. L." box="[339,486,1830,1856]" pageId="282" pageNumber="283" pagination="125 - 434" refId="ref182872" refString="Casey, T. L. (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini chiefly of America. Transactions of the Academy of Science of St. Louis, 16 (6), 125 - 434." type="journal article" year="1906">Casey, 1906</bibRefCitation>
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</paragraph>
<paragraph id="8B81369EB86EFEDFC1E36F46ACD79EF4" blockId="282.[151,704,1909,1974]" box="[151,684,1909,1934]" pageId="282" pageNumber="283">
<emphasis id="B94AEA8CB86EFEDFC1E36F46AEAD9EF4" bold="true" box="[151,214,1909,1934]" pageId="282" pageNumber="283">0943.</emphasis>
<taxonomicName id="4C3E4D1DB86EFEDFC1A96F46ACD79EF4" authority="Coiffait &amp; Saiz, 1969: 279" authorityName="Coiffait &amp; Saiz" authorityPageNumber="279" authorityYear="1969" box="[221,684,1909,1934]" class="Insecta" family="Staphylinidae" genus="Antarctothius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="282" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB86EFEDFC1A96F46AF079EF4" bold="true" box="[221,380,1909,1934]" italics="true" pageId="282" pageNumber="283">Antarctothius</emphasis>
<bibRefCitation id="EFAF4B6FB86EFEDFC0F76F46ACD79EF4" author="Coiffait, H. &amp; Saiz, F." box="[387,684,1909,1934]" pageId="282" pageNumber="283" pagination="279 - 286" refId="ref187652" refString="Coiffait, H. &amp; Saiz, F. (1969) Antarctothius nouveau genre de la Terre de Feu (Col. Staphylinidae). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 105, 279 - 286." type="journal article" year="1969">Coiffait &amp; Sáiz, 1969: 279</bibRefCitation>
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<paragraph id="8B81369EB86EFEDFC1E36FADACBB9ECF" blockId="282.[151,704,1909,1974]" box="[151,704,1948,1974]" pageId="282" pageNumber="283">
<typeStatus id="5485883CB86EFEDFC1E36FADAEAA9ECC" box="[151,209,1950,1974]" pageId="282" pageNumber="283">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB86EFEDFC0436FAFACBB9ECF" authority="Fairmaire, 1885" authorityName="Fairmaire" authorityYear="1885" box="[311,704,1948,1973]" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="282" pageNumber="283" phylum="Arthropoda" rank="species" species="antarcticus">
<emphasis id="B94AEA8CB86EFEDFC0436FAFAC7C9ECF" box="[311,519,1948,1973]" italics="true" pageId="282" pageNumber="283">Othius antarcticus</emphasis>
Fairmaire, 1885
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB86FFEDEC1E368ACAD2899CD" blockId="283.[151,916,158,300]" box="[151,851,158,183]" pageId="283" pageNumber="284">2 species are known from and are restricted to South America.</paragraph>
<paragraph id="8B81369EB86FFEDEC1E368F5ADEF99A4" blockId="283.[151,916,158,300]" box="[151,916,197,222]" pageId="283" pageNumber="284">
Key:
<bibRefCitation id="EFAF4B6FB86FFEDEC1A068F6AF8C99A4" author="Coiffait, H. &amp; Saiz, F." box="[212,503,197,222]" pageId="283" pageNumber="284" pagination="279 - 286" refId="ref187652" refString="Coiffait, H. &amp; Saiz, F. (1969) Antarctothius nouveau genre de la Terre de Feu (Col. Staphylinidae). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 105, 279 - 286." type="journal article" year="1969">Coiffait &amp; Sáiz 1969: 281</bibRefCitation>
(species of Tierra del Fuego,
<collectingCountry id="F329760EB86FFEDEC23268F6ADF399A4" box="[838,904,197,222]" name="Chile" pageId="283" pageNumber="284">Chile</collectingCountry>
).
</paragraph>
<paragraph id="8B81369EB86FFEDEC1E368DFAF5E987E" blockId="283.[151,916,158,300]" box="[151,293,236,261]" pageId="283" pageNumber="284">Distribution.</paragraph>
<paragraph id="8B81369EB86FFEDEC1DE6920AC5F9856" blockId="283.[151,916,158,300]" box="[170,548,275,300]" pageId="283" pageNumber="284">
South America:
<collectingCountry id="F329760EB86FFEDEC0156920AFDB9856" box="[353,416,275,300]" name="Chile" pageId="283" pageNumber="284">Chile</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC0D86920AC5B9856" box="[428,544,275,300]" name="Argentina" pageId="283" pageNumber="284">Argentina</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB86FFEDEC1E36952ACB59803" blockId="283.[151,1258,353,495]" box="[151,718,353,378]" pageId="283" pageNumber="284">
<emphasis id="B94AEA8CB86FFEDEC1E36952AEAD9800" bold="true" box="[151,214,353,378]" pageId="283" pageNumber="284">0944.</emphasis>
<taxonomicName id="4C3E4D1DB86FFEDEC1A96952AC489803" authority="Erichson, 1839 a: 300" authorityName="Erichson" authorityPageNumber="300" authorityYear="1839" box="[221,563,353,378]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="283" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB86FFEDEC1A96952AF469800" bold="true" box="[221,317,353,378]" italics="true" pageId="283" pageNumber="284">Diochus</emphasis>
<bibRefCitation id="EFAF4B6FB86FFEDEC0306952AC489803" author="Erichson, G. F." box="[324,563,353,378]" pageId="283" pageNumber="284" refId="ref190452" refString="Erichson, G. F. (1839 a) Genera et species staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin. viii + 400 pp." type="book" year="1839">Erichson, 1839a: 300</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13052A1BB86FFEDEC3136952ACBD9803" box="[615,710,353,378]" captionStart="FIGURES 184195" captionStartId="336.[151,264,1860,1883]" captionTargetBox="[155,1435,193,1832]" captionTargetId="figure@336.[150,1436,193,1840]" captionTargetPageId="336" captionText="FIGURES 184195. Habitus; Staphylininae (184195); 184, Peru: department of Lima (MUSM); 185, Brazil: state of Paraná (MUSM); 186, after Chatzimanolis 2014a; 187, after Santiago-Jiménez 2008; 188, after Smetana 1982; 189, after Chatzimanolis 2012; 190, Peru: department of Madre de Dios (CEMT); 191, after Navarrete-Heredia &amp; Márquez 1998; 192, after Klimaszewski et al. 2013; 193, after Márquez 2001; 194, after Erichson 1840; 195, after Márquez 2003; slightly modified. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3257260/files/figure.png" pageId="283" pageNumber="284">Fig. 188</figureCitation>
)
</paragraph>
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<typeStatus id="5485883CB86FFEDEC1E369BAAEAA98DB" box="[151,209,393,417]" pageId="283" pageNumber="284">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB86FFEDEC04369BBACF498DA" authority="Erichson, 1839" authorityName="Erichson" authorityYear="1839" box="[311,655,392,417]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="283" pageNumber="284" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B94AEA8CB86FFEDEC04369BBAFA598DA" box="[311,478,392,417]" italics="true" pageId="283" pageNumber="284">Diochus nanus</emphasis>
Erichson, 1839
</taxonomicName>
</paragraph>
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<taxonomicName id="4C3E4D1DB86FFEDEC1BB699CACDA98B2" ID-CoL="8XPMP" authority="Motschulsky, 1858 d: 657" authorityName="Motschulsky" authorityPageNumber="657" authorityYear="1858" box="[207,673,431,456]" class="Insecta" family="Staphylinidae" genus="Rhegmatocerus" kingdom="Animalia" order="Coleoptera" pageId="283" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB86FFEDEC1BB699CAF0698B2" box="[207,381,431,456]" italics="true" pageId="283" pageNumber="284">Rhegmatocerus</emphasis>
<treatmentCitation id="0A9F108FB86FFEDEC0F0699CACDA98B2" author="Motschulsky, V." box="[388,673,431,456]" page="657" pageId="283" pageNumber="284" year="1858">
<bibRefCitation id="EFAF4B6FB86FFEDEC0F0699CACDA98B2" author="Motschulsky, V." box="[388,673,431,456]" pageId="283" pageNumber="284" pagination="634 - 670" refId="ref217715" refString="Motschulsky, V. (1858 d) Enumeration des nouvelles especes de Coleopteres rapportes de ses voyages 2 - d. Article (continuation). Bulletin de la Societe Imperiale des Naturalistes de Moscou, 31 (2), 634 - 670." type="journal article" year="1858">Motschulsky, 1858d: 657</bibRefCitation>
</treatmentCitation>
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[synonymized by Gemminger &amp; Harold 1 868: 608]
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</paragraph>
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<paragraph id="8B81369EB86FFEDEC07369E4ADB29895" blockId="283.[151,1258,353,495]" box="[263,969,470,495]" pageId="283" pageNumber="284">
<typeStatus id="5485883CB86FFEDEC07369E4AF3A9895" box="[263,321,471,495]" pageId="283" pageNumber="284">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB86FFEDEC0D369E5ADB29895" authority="Motschulsky, 1858" authorityName="Motschulsky" authorityYear="1858" box="[423,969,470,495]" class="Insecta" family="Staphylinidae" genus="Rhegmatocerus" kingdom="Animalia" order="Coleoptera" pageId="283" pageNumber="284" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B94AEA8CB86FFEDEC0D369E5AC909894" box="[423,747,470,495]" italics="true" pageId="283" pageNumber="284">Rhegmatocerus punctipennis</emphasis>
Motschulsky, 1858
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB86FFEDEC1E36A2FAD649B4E" blockId="283.[151,1436,539,1026]" box="[151,799,539,564]" pageId="283" pageNumber="284">42 species are included in this genus from Latin America.</paragraph>
<paragraph id="8B81369EB86FFEDEC1E36A71AC029BD3" blockId="283.[151,1436,539,1026]" pageId="283" pageNumber="284">
Characters/Description:
<bibRefCitation id="EFAF4B6FB86FFEDEC0D36A71AC889B21" author="Lynch-Arribalzaga, F." box="[423,755,578,603]" pageId="283" pageNumber="284" pagination="5 - 392" refId="ref214921" refString="Lynch-Arribalzaga, F. (1884) Estafilinos de Buenos Aires. Boletin de la Academia Nacional de Ciencias en Cordoba, 7 (1 - 3), 5 - 392." type="journal article" year="1884">Lynch-Arribálzaga 1884: 171</bibRefCitation>
(characters);
<bibRefCitation id="EFAF4B6FB86FFEDEC2FF6A71AAF49B21" author="Blackwelder, R. E." box="[907,1167,578,603]" pageId="283" pageNumber="284" refId="ref175651" refString="Blackwelder, R. E. (1943) Monograph of the West-Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, viii + 658 pp. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book" year="1943">Blackwelder 1943: 394</bibRefCitation>
(characters in key), 455 (characters);
<bibRefCitation id="EFAF4B6FB86FFEDEC0106A59AC4B9BF8" author="Smetana, A." box="[356,560,618,642]" pageId="283" pageNumber="284" pagination="1 - 389" refId="ref239533" refString="Smetana, A. (1982) Revision of the subfamily Xantholininae of America north of Mexico (Coleoptera: Staphylinidae). Memoirs of the Entomological Society of Canada, 120, iv + 1 - 389. https: // doi. org / 10.4039 / entm 114120 fv" type="journal article" year="1982">Smetana 1982: 27</bibRefCitation>
(characters);
<bibRefCitation id="EFAF4B6FB86FFEDEC3B96A5AAA239BF8" author="Navarrete-Heredia, J. L. &amp; Newton, A. F. &amp; Thayer, M. K. &amp; Ashe, J. S. &amp; Chandler, D. S." box="[717,1112,617,642]" pageId="283" pageNumber="284" refId="ref220178" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Universidad de Guadalajara y CONABIO, Mexico. xii + 401 pp." type="book" year="2002">
Navarrete-Heredia
<emphasis id="B94AEA8CB86FFEDEC2D16A58ADA59BF8" box="[933,990,617,642]" italics="true" pageId="283" pageNumber="284">et al.</emphasis>
2002: 295
</bibRefCitation>
(characters in key);
<bibRefCitation id="EFAF4B6FB86FFEDEC4376A59AFE19BD3" author="Newton, A. F. &amp; Peck, S. B." pageId="283" pageNumber="284" pagination="131 - 158" refId="ref220547" refString="Newton, A. F. &amp; Peck, S. B. (2006) Family Staphylinidae, the rove beetles. In: Peck, S. B. (Ed.), The beetles of the Galapagos Islands, Ecuador: evolution, ecology, and diversity (Insecta: Coleoptera). NRC Research Press, Ottawa, pp. 131 - 158." type="book chapter" year="2006">Newton &amp; Peck 2006: 152</bibRefCitation>
(characters in key).
</paragraph>
<paragraph id="8B81369EB86FFEDEC1E36A8BAC7F9B8D" blockId="283.[151,1436,539,1026]" pageId="283" pageNumber="284">
Key:
<bibRefCitation id="EFAF4B6FB86FFEDEC1A06A84AFF99BAA" author="Irmler, U." box="[212,386,695,720]" pageId="283" pageNumber="284" pagination="1 - 62" refId="ref203402" refString="Irmler, U. (2017 a) A review of the Neotropical genus Diochus Erichson, 1840 (Coleoptera: Staphylinidae: Staphylininae). Beitrage zur Entomologie, 67 (1), 1 - 62. https: // doi. org / 10.21248 / contrib. entomol. 67.1.001 - 062" type="journal article" year="2017">Irmler 2017a: 4</bibRefCitation>
(Neotropical species groups of the genus), 57 (males of the Neotropical species), 59 (females of the Neotropical species).
</paragraph>
<paragraph id="8B81369EB86FFEDEC1E36B36AF5E9A67" blockId="283.[151,1436,539,1026]" box="[151,293,773,798]" pageId="283" pageNumber="284">Distribution.</paragraph>
<paragraph id="8B81369EB86FFEDEC1DE6B1EAA179A16" blockId="283.[151,1436,539,1026]" pageId="283" pageNumber="284">
West Indies:
<collectingCountry id="F329760EB86FFEDEC0316B1FAFF89A3F" box="[325,387,812,837]" name="Cuba" pageId="283" pageNumber="284">Cuba</collectingCountry>
, Cayman Is.,
<collectingCountry id="F329760EB86FFEDEC35C6B1FACFE9A3F" box="[552,645,812,837]" name="Jamaica" pageId="283" pageNumber="284">Jamaica</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC3E06B1FADFF9A3F" box="[660,900,812,837]" name="Dominican Republic" pageId="283" pageNumber="284">Dominican Republic</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC2E76B1EAA649A3F" box="[915,1055,812,837]" name="Puerto Rico" pageId="283" pageNumber="284">Puerto Rico</collectingCountry>
, British Virgin Isl.,
<collectingCountry id="F329760EB86FFEDEC47A6B1FABE39A3F" box="[1294,1432,812,837]" name="Guadeloupe" pageId="283" pageNumber="284">Guadeloupe</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC1BB6B60AF3B9A16" box="[207,320,851,876]" name="Dominica" pageId="283" pageNumber="284">Dominica</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC0386B60AF9D9A16" box="[332,486,851,876]" name="Saint Vincent and the Grenadines" pageId="283" pageNumber="284">Saint Vincent</collectingCountry>
&amp;
<collectingCountry id="F329760EB86FFEDEC37D6B60ACCE9A16" box="[521,693,851,876]" name="Saint Vincent and the Grenadines" pageId="283" pageNumber="284">the Grenadines</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC3B46B60AD569A16" box="[704,813,851,876]" name="Barbados" pageId="283" pageNumber="284">Barbados</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC24C6B60ADE19A16" box="[824,922,851,876]" name="Grenada" pageId="283" pageNumber="284">Grenada</collectingCountry>
, Trinidad-Tobago.
</paragraph>
<paragraph id="8B81369EB86FFEDEC1DE6B49AAEC9AE8" blockId="283.[151,1436,539,1026]" box="[170,1175,890,915]" pageId="283" pageNumber="284">
Central America:
<collectingCountry id="F329760EB86FFEDEC0066B49AFB19AE9" box="[370,458,890,915]" name="Mexico" pageId="283" pageNumber="284">Mexico</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC0A26B49AC299AE9" box="[470,594,890,915]" name="Guatemala" pageId="283" pageNumber="284">Guatemala</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC3296B49ACDC9AE9" box="[605,679,890,915]" name="Belize" pageId="283" pageNumber="284">Belize</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC3C76B49AD589AE9" box="[691,803,890,915]" name="Honduras" pageId="283" pageNumber="284">Honduras</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC25A6B49ADDF9AE9" box="[814,932,890,915]" name="Nicaragua" pageId="283" pageNumber="284">Nicaragua</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC2DB6B48AA579AE9" box="[943,1068,890,915]" name="Costa Rica" pageId="283" pageNumber="284">Costa Rica</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC5436B48AAE89AE8" box="[1079,1171,891,914]" name="Panama" pageId="283" pageNumber="284">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB86FFEDEC1D06BA8AC7E9AA2" blockId="283.[151,1436,539,1026]" pageId="283" pageNumber="284">
South America:
<collectingCountry id="F329760EB86FFEDEC0286BA8AFB69ACE" box="[348,461,923,948]" name="Colombia" pageId="283" pageNumber="284">Colombia</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC0AC6BA8AC359ACE" box="[472,590,923,948]" name="Venezuela" pageId="283" pageNumber="284">Venezuela</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC32D6BAFACC89ACE" box="[601,691,924,948]" name="Guyana" pageId="283" pageNumber="284">Guyana</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC3CA6BA8AD1D9ACE" box="[702,870,923,948]" name="French Guiana" pageId="283" pageNumber="284">French Guiana</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC2056BA8ADA49ACE" box="[881,991,923,948]" name="Suriname" pageId="283" pageNumber="284">Suriname</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC29E6BA8AA329ACE" box="[1002,1097,923,948]" name="Ecuador" pageId="283" pageNumber="284">Ecuador</collectingCountry>
,
<collectingRegion id="49FAF87CB86FFEDEC5206BA8AAB19ACE" box="[1108,1226,923,948]" country="Ecuador" name="Galapagos" pageId="283" pageNumber="284">Galapagos</collectingRegion>
Isl.,
<collectingCountry id="F329760EB86FFEDEC4766BAFAB439AC9" box="[1282,1336,924,947]" name="Peru" pageId="283" pageNumber="284">Peru</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC4376BA8ABE39ACE" box="[1347,1432,923,948]" name="Bolivia" pageId="283" pageNumber="284">Bolivia</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC1B36B8CAF759AA2" box="[199,270,959,984]" name="Brazil" pageId="283" pageNumber="284">Brazil</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC06D6BF3AFF99AA2" box="[281,386,960,984]" name="Paraguay" pageId="283" pageNumber="284">Paraguay</collectingCountry>
,
<collectingCountry id="F329760EB86FFEDEC0F96B8CAC7B9AA2" box="[397,512,959,984]" name="Argentina" pageId="283" pageNumber="284">Argentina</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB86FFEDEC1DE6BDAAAD79D78" blockId="283.[151,1436,539,1026]" box="[170,1196,1001,1026]" pageId="283" pageNumber="284">
Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan,
<collectingRegion id="49FAF87CB86FFEDEC2286BDAADC19D78" box="[860,954,1001,1026]" country="Republic of the Congo" name="Orientale" pageId="283" pageNumber="284">Oriental</collectingRegion>
, Australian, Oceanic.
</paragraph>
</subSubSection>
</treatment>
</document>