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<document id="57B1F3880310DB851DE28A9EEC9344A4" ID-DOI="10.15553/c2016v711a7" ID-GBIF-Dataset="508bebc5-39aa-4f9c-b544-171e992dbbea" ID-ISSN="2235-3658" ID-Zenodo-Dep="161516" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476779283805" checkinUser="plazi" docAuthor="Wendy L. Applequist" docDate="2016" docId="03978783FF98FFE5FF2DFE40AFCC9BFE" docLanguage="en" docName="Candollea.71.1.33-60.pdf" docOrigin="Candollea 71 (1)" docStyle="DocumentStyle:1E5CC43F38D0DBC531513809D1267F7B.3:Candollea.2015-.journal_article" docStyleId="1E5CC43F38D0DBC531513809D1267F7B" docStyleName="Candollea.2015-.journal_article" docStyleVersion="3" docTitle="Homalium randrianasoloi Appleq., spec. nova" docType="treatment" docVersion="9" lastPageNumber="53" masterDocId="FFAEFFFBFF8CFFF1FFA6FF9DAA7C9D57" masterDocTitle="Revision of the Malagasy species of Homalium sect. Eumyriantheia Warb. (Salicaceae)" masterLastPageNumber="60" masterPageNumber="33" pageNumber="53" updateTime="1698390116944" updateUser="plazi">
<mods:mods id="077A41B9868A143D3AD4FA53590C4B98" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="388F4A234488A72245F337A090A9E830">
<mods:title id="E5D306A4B5205DBE733AE3A4BC4590BC">Revision of the Malagasy species of Homalium sect. Eumyriantheia Warb. (Salicaceae)</mods:title>
</mods:titleInfo>
<mods:name id="463A8C6E393A293F7996820B6ADEF847" type="personal">
<mods:role id="F56713FA712CC4236840727BCF775DC7">
<mods:roleTerm id="431A6F96A1F062621D9035AA73D03FC7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="752ECAB4FC36E2ABECBD095DDDD04D40">Wendy L. Applequist</mods:namePart>
</mods:name>
<mods:typeOfResource id="C515E06AF7D00167910B3EFEB8D01E70">text</mods:typeOfResource>
<mods:relatedItem id="471773DFFD5CA48E0652D07C4F39FAD9" type="host">
<mods:titleInfo id="B6C3B993471A758C3FF19FA1840FB3CC">
<mods:title id="595E8CAF3949A4B8008B7C1F0763651D">Candollea</mods:title>
</mods:titleInfo>
<mods:part id="C42FBFD77196E022462084F033C9BFEA">
<mods:date id="59FF9528745B3DB1D763B9D0DE465D87">2016</mods:date>
<mods:detail id="C02F6D5B2DBE16E5806A99F5655A7B0A" type="pubDate">
<mods:number id="61320A3D74049AA476087D44771167FA">2016-03-14</mods:number>
</mods:detail>
<mods:detail id="4F5154EB578F27F52872ED9C594BE525" type="volume">
<mods:number id="80067482545E5A898707A612FA96AA9E">71</mods:number>
</mods:detail>
<mods:detail id="65592644681A107458570622B5584E5F" type="issue">
<mods:number id="6006B44FA6EE967AD2BEB72B7A22CF22">1</mods:number>
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<mods:start id="60FDF47C0FA05C2069279910FFCDF661">33</mods:start>
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<mods:classification id="E1C1EC9054F5ED5313FA9FD291411F6A">journal article</mods:classification>
<mods:identifier id="62711F4E2F0F63F4C00B794A460A26A4" type="DOI">10.15553/c2016v711a7</mods:identifier>
<mods:identifier id="A11D5B7A186555CFCE1C4DD7A9E9A2EE" type="GBIF-Dataset">508bebc5-39aa-4f9c-b544-171e992dbbea</mods:identifier>
<mods:identifier id="C6ED7B5F8E345936D94CFA1BDB1D5E08" type="ISSN">2235-3658</mods:identifier>
<mods:identifier id="63CBE3D6ABF728183D668C5FF1A5A411" type="Zenodo-Dep">161516</mods:identifier>
</mods:mods>
<treatment id="03978783FF98FFE5FF2DFE40AFCC9BFE" ID-DOI="http://doi.org/10.5281/zenodo.4484271" ID-GBIF-Taxon="177200346" ID-Zenodo-Dep="4484271" LSID="urn:lsid:plazi:treatment:03978783FF98FFE5FF2DFE40AFCC9BFE" httpUri="http://treatment.plazi.org/id/03978783FF98FFE5FF2DFE40AFCC9BFE" lastPageId="20" lastPageNumber="53" pageId="20" pageNumber="53">
<subSubSection id="C324651EFF98FFE5FF2DFE40A8029CA1" box="[139,638,477,502]" pageId="20" pageNumber="53" type="nomenclature">
<paragraph id="8B813695FF98FFE5FF2DFE40A8029CA1" blockId="20.[139,790,477,747]" box="[139,638,477,502]" pageId="20" pageNumber="53">
<emphasis id="B94AEA87FF98FFE5FF2DFE40AAD79CA2" bold="true" box="[139,171,477,501]" pageId="20" pageNumber="53">12.</emphasis>
<taxonomicName id="4C3E4D16FF98FFE5FF09FE40A8769CA2" ID-CoL="3MCR2" authority="Appleq." authorityName="Appleq." box="[175,522,477,501]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="randrianasoloi" status="sp. nov.">
<emphasis id="B94AEA87FF98FFE5FF09FE40ABCE9CA2" bold="true" box="[175,434,477,501]" italics="true" pageId="20" pageNumber="53">Homalium randrianasoloi</emphasis>
Appleq.
</taxonomicName>
,
<emphasis id="B94AEA87FF98FFE5FDB5FE43A8029CA1" bold="true" box="[531,638,478,502]" pageId="20" pageNumber="53">
<taxonomicNameLabel id="A27957FCFF98FFE5FDB5FE43A8029CA1" box="[531,638,478,502]" pageId="20" pageNumber="53">spec. nova</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C324651EFF98FFE5FD23FE40A8839CA1" box="[645,767,477,502]" pageId="20" pageNumber="53" type="description">
<paragraph id="8B813695FF98FFE5FD23FE40A8839CA1" blockId="20.[139,790,477,747]" box="[645,767,477,502]" pageId="20" pageNumber="53">
(
<figureCitation id="13052A10FF98FFE5FD2AFE40A8B99CA1" box="[652,709,477,502]" captionStart="Fig. 2" captionStartId="10.[139,166,1866,1885]" captionTargetBox="[139,1469,285,1846]" captionTargetId="figure@10.[137,811,744,1299]" captionTargetPageId="10" captionText="Fig. 2. – Flowers of newly described species of Homalium sect. Eumyriantheia Warb. A. Homalium dorrii Appleq.: flowers and nearly mature fruit with single seed; B. Homalium pseudoboinense Appleq.: abaxial surface of perianth; C. Homalium pseudoboinense: open flower; D. Homalium randrianasoloi Appleq.; E. Homalium ranomafanicum Appleq.; F. Homalium schatzii Appleq.: open flower; G. Homalium schatzii: abaxial surface of perianth." httpUri="https://zenodo.org/record/161518/files/figure.png" pageId="20" pageNumber="53">Fig. 2</figureCitation>
D, 4).
</paragraph>
</subSubSection>
<subSubSection id="C324651EFF98FFE5FF17FD95AB289FD7" pageId="20" pageNumber="53" type="materials_examined">
<paragraph id="8B813695FF98FFE5FF17FD95AB289FD7" blockId="20.[139,790,477,747]" pageId="20" pageNumber="53">
<materialsCitation id="3B563CC8FF98FFE5FF17FD95A8B99F37" collectingDate="1990-10-29" collectionCode="MO" collectorName="Randrianasolo et al." country="Madagascar" latitude="-24.1" location="foret de Marovony" longLatPrecision="1252" longitude="47.333332" municipality="Manantenenina" pageId="20" pageNumber="53" specimenCode="MO-4028861" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
<emphasis id="B94AEA87FF98FFE5FF17FD95A8249F77" bold="true" box="[177,600,520,544]" pageId="20" pageNumber="53">
<typeStatus id="54858837FF98FFE5FF17FD95AA849F77" box="[177,248,520,544]" pageId="20" pageNumber="53">Typus</typeStatus>
:
<collectingCountry id="F3297605FF98FFE5FEAFFD95ABD69F77" box="[265,426,520,544]" name="Madagascar" pageId="20" pageNumber="53">Madagascar</collectingCountry>
. Prov.
<collectingRegion id="49FAF877FF98FFE5FE5FFD95A8319F77" box="[505,589,520,544]" country="Madagascar" name="Toliara" pageId="20" pageNumber="53">Toliara</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BE5ACEFFF98FFE5FDC6FD95A96D9F77" box="[608,785,520,544]" pageId="20" pageNumber="53">Manantenenina</collectingMunicipality>
,
<location id="8EE1604EFF98FFE5FF17FDB5ABF69F17" LSID="urn:lsid:plazi:treatment:03978783FF98FFE5FF2DFE40AFCC9BFE:8EE1604EFF98FFE5FF17FDB5ABF69F17" box="[177,394,552,576]" country="Madagascar" latitude="-24.1" longLatPrecision="1252" longitude="47.333332" municipality="Manantenenina" name="foret de Marovony" pageId="20" pageNumber="53" stateProvince="Toliara">forêt de Marovony</location>
[
<geoCoordinate id="EE0A5052FF98FFE5FE3FFDB5AB889F17" box="[409,500,552,576]" direction="south" orientation="latitude" pageId="20" pageNumber="53" precision="925" value="-24.1">24°06’S</geoCoordinate>
<geoCoordinate id="EE0A5052FF98FFE5FE5BFDB5A8209F17" box="[509,604,552,576]" direction="east" orientation="longitude" pageId="20" pageNumber="53" precision="925" value="47.333332">47°20’E</geoCoordinate>
],
<date id="FF801055FF98FFE5FDD7FDB5A8979F17" box="[625,747,552,576]" pageId="20" pageNumber="53" value="1990-10-29">
<collectingDate id="EFC4E9BDFF98FFE5FDD7FDB5A8979F17" box="[625,747,552,576]" pageId="20" pageNumber="53" value="1990-10-29">29.X.1990</collectingDate>
</date>
, fl.,
<emphasis id="B94AEA87FF98FFE5FF17FDD5ABCA9F37" box="[177,438,584,608]" italics="true" pageId="20" pageNumber="53">
<collectorName id="26CB5343FF98FFE5FF17FDD5ABFB9F37" box="[177,391,584,608]" pageId="20" pageNumber="53">Randrianasolo et al.</collectorName>
191
</emphasis>
(holo-:
<specimenCode id="DB989EEEFF98FFE5FDB1FDD5A8C49F37" box="[535,696,584,608]" pageId="20" pageNumber="53">MO-4028861</specimenCode>
!;
</materialsCitation>
<materialsCitation id="3B563CC8FF98FFE5FD6BFDD5AB289FD7" ID-GBIF-Occurrence="3027618303" collectingDate="1990-10-29" collectionCode="P" country="Madagascar" latitude="-24.1" location="Manantenenina, forêt de Marovony" longLatPrecision="1252" longitude="47.333332" pageId="20" pageNumber="53" specimenCode="P04734076" specimenCount="1" stateProvince="Toliara" typeStatus="isotype">
<typeStatus id="54858837FF98FFE5FD6BFDD5A8859F37" box="[717,761,584,608]" pageId="20" pageNumber="53" type="isotype">iso-</typeStatus>
,
<collectionCode id="ED2FAE50FF98FFE5FCA3FDD4A9699F37" box="[773,789,585,608]" pageId="20" pageNumber="53">P</collectionCode>
[
<specimenCode id="DB989EEEFF98FFE5FF11FDF5AB429FD7" box="[183,318,616,640]" pageId="20" pageNumber="53">P04734076</specimenCode>
]!).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C324651EFF98FFE5FF17FD0EA8429FBD" pageId="20" pageNumber="53" type="diagnosis">
<paragraph id="8B813695FF98FFE5FF17FD0EA8429FBD" blockId="20.[139,790,477,747]" pageId="20" pageNumber="53">
<emphasis id="B94AEA87FF98FFE5FF17FD0EA8429FBD" italics="true" pageId="20" pageNumber="53">
<taxonomicName id="4C3E4D16FF98FFE5FF17FD0EA86C9FFC" authority="Appleq." authorityName="Appleq." box="[177,528,659,683]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="randrianasoloi">Homalium randrianasoloi Appleq.</taxonomicName>
differs from
<taxonomicName id="4C3E4D16FF98FFE5FD35FD09AAA69F9C" authority="Sleumer" authorityName="Sleumer" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="capuronii">H. capuronii Sleumer</taxonomicName>
in having larger leaves, some panicles with well-developed side branches, sepals and petals sometimes 5.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C324651EFF98FFE5FF17FC89A8E79BBD" pageId="20" pageNumber="53" type="description">
<paragraph id="8B813695FF98FFE5FF17FC89A8E79BBD" blockId="20.[139,790,787,1771]" pageId="20" pageNumber="53">
<emphasis id="B94AEA87FF98FFE5FF17FC89AAA29E7C" box="[177,222,788,811]" italics="true" pageId="20" pageNumber="53">Tree</emphasis>
to
<quantity id="4CC69B70FF98FFE5FEA7FC8EAB449E7D" box="[257,312,787,811]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="53" unit="m" value="10.0">10 m</quantity>
tall; large twigs grayish; young twigs brown, glabrous; stipules linear to narrowly deltoid, &lt;
<quantity id="4CC69B70FF98FFE5FD3DFCAEA8A19E1D" box="[667,733,819,843]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="53" unit="mm" value="1.0">1 mm</quantity>
(few seen), glabrate.
<emphasis id="B94AEA87FF98FFE5FE94FCC9AB049E3C" box="[306,376,852,875]" italics="true" pageId="20" pageNumber="53">Leaves</emphasis>
alternate; petiole
<quantity id="4CC69B70FF98FFE5FD99FCCEA8D99E3D" box="[575,677,851,875]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="20" pageNumber="53" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5-10 mm</quantity>
, glabrous; blade elliptical to obovate or broadly (narrowly) elliptical, 6-12.5 ×
<quantity id="4CC69B70FF98FFE5FF53FC0EAB0E9EFD" box="[245,370,915,939]" metricMagnitude="-2" metricUnit="m" metricValue="4.699999999999999" metricValueMax="6.6" metricValueMin="2.8" pageId="20" pageNumber="53" unit="cm" value="4.699999999999999" valueMax="6.6" valueMin="2.8">2.8-6.6 cm</quantity>
; base convex to rounded (cuneate in young leaves), at extreme base short-attenuate; apex roundedacute to cuspidate, rounded, or shallowly emarginate; margins minutely revolute with 0 to 2 basal glands just inside margin; abaxial surface glabrous, drying greenish; adaxial surface glabrous, drying greenish.
<emphasis id="B94AEA87FF98FFE5FE1FFBAEA83A991C" box="[441,582,1075,1099]" italics="true" pageId="20" pageNumber="53">Inflorescences</emphasis>
paniculate, shortbranched, ridged, with most flowers borne in small clusters, sometimes with short but well-developed proximal branches, lateral mostly near twig apices,
<quantity id="4CC69B70FF98FFE5FE76FB0EA84199FD" box="[464,573,1171,1195]" metricMagnitude="-2" metricUnit="m" metricValue="8.75" metricValueMax="10.5" metricValueMin="7.0" pageId="20" pageNumber="53" unit="cm" value="8.75" valueMax="10.5" valueMin="7.0">7-10.5 cm</quantity>
, glabrate or glabrous proximally, the distal rachis and branches sparsely (moderately) minutely pubescent; peduncle
<quantity id="4CC69B70FF98FFE5FD8EFB4EA8F299BD" box="[552,654,1235,1259]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="4.8" metricValueMin="1.0" pageId="20" pageNumber="53" unit="cm" value="2.9" valueMax="4.8" valueMin="1.0">1-4.8 cm</quantity>
, moderately sturdy; pedicel
<quantity id="4CC69B70FF98FFE5FE93FB6EABC5985D" box="[309,441,1267,1291]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="4.3" metricValueMin="1.5" pageId="20" pageNumber="53" unit="mm" value="2.9" valueMax="4.3" valueMin="1.5">1.5-4.3 mm</quantity>
, sparsely (moderately) minutely pubescent; bracts ovate,
<quantity id="4CC69B70FF98FFE5FE39FA8EA859987D" box="[415,549,1299,1323]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="2.5" metricValueMin="0.3" pageId="20" pageNumber="53" unit="mm" value="1.4" valueMax="2.5" valueMin="0.3">0.3-2.5 mm</quantity>
, minutely pubescent; bracteoles ovate,
<quantity id="4CC69B70FF98FFE5FEEBFAAEABC3981D" box="[333,447,1331,1355]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="20" pageNumber="53" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5-1 mm</quantity>
, minutely pubescent.
<emphasis id="B94AEA87FF98FFE5FD10FAAEA96A981C" box="[694,790,1331,1355]" italics="true" pageId="20" pageNumber="53">Flowers:</emphasis>
sepals 4(-5), oblong with rounded apex,
<quantity id="4CC69B70FF98FFE5FD98FACEA8C3983D" box="[574,703,1363,1387]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="4.2" metricValueMin="2.4" pageId="20" pageNumber="53" unit="mm" value="3.3" valueMax="4.2" valueMin="2.4">2.4-4.2 mm</quantity>
, abaxial surface sparsely minutely pubescent; calyx cup funnelform, moderately (sparsely) minutely pubescent; sepal glands irregularly oblong, 0.6-1 ×
<quantity id="4CC69B70FF98FFE5FED4FA2EAB8A989D" box="[370,502,1459,1483]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="20" pageNumber="53" unit="mm" value="0.5" valueMax="0.6" valueMin="0.4">0.4-0.6 mm</quantity>
, upper surface glabrous or sparsely short-pubescent to moderately short-pubescent in the depressed center; petals white or pink (possibly only after anthesis), obovate to narrowly obovate with a narrow base and rounded apex, 3.7-6 ×
<quantity id="4CC69B70FF98FFE5FE26F9AEA87E9B1D" box="[384,514,1587,1611]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.6" metricValueMin="1.6" pageId="20" pageNumber="53" unit="mm" value="2.1" valueMax="2.6" valueMin="1.6">1.6-2.6 mm</quantity>
, significantly longer than sepals, abaxial surface minutely pubescent especially at base and in center, adaxial surface glabrous; filaments
<quantity id="4CC69B70FF98FFE5FD37F9EEA96D9BDD" box="[657,785,1651,1675]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.4" metricValueMin="1.7" pageId="20" pageNumber="53" unit="mm" value="2.05" valueMax="2.4" valueMin="1.7">1.7-2.4 mm</quantity>
, glabrous; anthers
<quantity id="4CC69B70FF98FFE5FEECF90EABE19BFD" box="[330,413,1683,1707]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="53" unit="mm" value="0.3">0.3 mm</quantity>
; ovary flat-conic, short-pubescent; styles 4(-5),
<quantity id="4CC69B70FF98FFE5FEABF92EABF19B9D" box="[269,397,1715,1739]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="20" pageNumber="53" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5-2.5 mm</quantity>
, glabrous or sparsely pubescent especially on short fused basal portion.
<emphasis id="B94AEA87FF98FFE5FE59F94EA84F9BBC" box="[511,563,1747,1771]" italics="true" pageId="20" pageNumber="53">Seeds</emphasis>
not seen.
</paragraph>
</subSubSection>
<subSubSection id="C324651EFF98FFE5FF17F88EAE759FE2" pageId="20" pageNumber="53" type="distribution">
<paragraph id="8B813695FF98FFE5FF17F88EAE759FE2" blockId="20.[139,790,1811,1963]" lastBlockId="20.[827,1478,285,693]" pageId="20" pageNumber="53">
<emphasis id="B94AEA87FF98FFE5FF17F88EAB589A1C" italics="true" pageId="20" pageNumber="53">
Distribution, ecology and conservation status. –
<taxonomicName id="4C3E4D16FF98FFE5FD03F88EAB589A1C" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="randrianasoloi">Homalium randrianasoloi</taxonomicName>
</emphasis>
is known from only two collection numbers made two days apart at the low-elevation humid forest of Marovony near the southeast coast north of Fort-Dauphin and southeast of Befotaka. This area has not been frequently collected, so the paucity of specimens is not clear evidence of local rarity. However, the
<typeStatus id="54858837FF98FFE5FBDCFEA3AED69C02" box="[1146,1194,318,341]" pageId="20" pageNumber="53">type</typeStatus>
locality is under extreme and immediate threat: only small fragments of the Marovony forest remained at the time of the most recent vegetation mapping (
<bibRefCitation id="EFAF4B64FF98FFE5FC16FE00A90F9C82" author="Madagascar Vegetation Mapping Project" pageId="20" pageNumber="53" publicationUrl="http://www.kew.org/gis/projects/mad_veg/default.html" refId="ref29039" refString="Madagascar Vegetation Mapping Project (2006). The CEPF Madagascar Vegetation Mapping Project [http: // www. kew. org / gis / projects / mad _ veg / default. html]." title="The CEPF Madagascar Vegetation Mapping Project" type="url" year="2006">
<collectingCountry id="F3297605FF98FFE5FC16FE00AE319CE2" box="[944,1101,413,437]" name="Madagascar" pageId="20" pageNumber="53">Madagascar</collectingCountry>
Vegetation Mapping Project, 2006
</bibRefCitation>
), and the larger low-elevation forests to its west and southwest were at that time already completely or almost completely gone. Thus, a preliminary conservation status assessment of “Critically Endangered” [CR B1ab(iii)+2ab(iii)] seems justified. An attempt to locate another population of the species in the larger remaining forested areas of Alan’i Tsidikamboro, to the north of Amparihy Atsinanana, would be highly desirable.
</paragraph>
</subSubSection>
<subSubSection id="C324651EFF98FFE5FCC7FD43AFE69B02" pageId="20" pageNumber="53" type="discussion">
<paragraph id="8B813695FF98FFE5FCC7FD43A9B89862" blockId="20.[827,1478,733,1621]" pageId="20" pageNumber="53">
<emphasis id="B94AEA87FF98FFE5FCC7FD43AEC49FA2" box="[865,1208,733,757]" italics="true" pageId="20" pageNumber="53">
Notes. –
<taxonomicName id="4C3E4D16FF98FFE5FC10FD40AEC49FA2" box="[950,1208,733,757]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="randrianasoloi">Homalium randrianasoloi</taxonomicName>
</emphasis>
is part of the
<taxonomicName id="4C3E4D16FF98FFE5FAEEFD43AFB99FA2" box="[1352,1477,734,757]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="capuronii">
<emphasis id="B94AEA87FF98FFE5FAEEFD43AFB99FA2" box="[1352,1477,734,757]" italics="true" pageId="20" pageNumber="53">H. capuronii</emphasis>
</taxonomicName>
group (see
<tableCitation id="C6BC032EFF98FFE5FC15FD60AE789E41" box="[947,1028,765,790]" captionStart="Table 1" captionStartId="7.[124,170,238,257]" captionTargetBox="[124,1479,364,1130]" captionTargetPageId="7" captionText="Table 1. – Comparison of key features of species in the Homalium capuronii Sleumer group. Apparent differences in stamen and style size are excluded because of limited material." httpUri="http://table.plazi.org/id/DF41661DFF8BFFF6FFDAFF73AB0F9C4D" pageId="20" pageNumber="53" tableUuid="DF41661DFF8BFFF6FFDAFF73AB0F9C4D">Table 1</tableCitation>
). It is most similar to
<taxonomicName id="4C3E4D16FF98FFE5FB58FD63AFFE9E42" box="[1278,1410,766,789]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="capuronii">
<emphasis id="B94AEA87FF98FFE5FB58FD63AFFE9E42" box="[1278,1410,766,789]" italics="true" pageId="20" pageNumber="53">H. capuronii</emphasis>
</taxonomicName>
, f rom which it is visually distinguished by its typically larger, broader leaves with sometimes rounded bases and sometimes betterdeveloped panicles. The leaves of the
<typeStatus id="54858837FF98FFE5FB68FCC3AE819E22" box="[1230,1277,862,885]" pageId="20" pageNumber="53">type</typeStatus>
are comparable in size to the aberrant extreme of
<taxonomicName id="4C3E4D16FF98FFE5FB22FCE3AF789EC2" box="[1156,1284,894,917]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="capuronii">
<emphasis id="B94AEA87FF98FFE5FB22FCE3AF789EC2" box="[1156,1284,894,917]" italics="true" pageId="20" pageNumber="53">H. capuronii</emphasis>
</taxonomicName>
’s size range, while the
<typeStatus id="54858837FF98FFE5FCC2FC03A9BC9EE2" box="[868,960,926,949]" pageId="20" pageNumber="53" type="paratype">paratype</typeStatus>
has even larger leaves, elliptical with broad bases and rounded-acute apices, but only short-branched panicles. Flowers of
<taxonomicName id="4C3E4D16FF98FFE5FC1FFC43AE419EA2" box="[953,1085,990,1013]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="capuronii">
<emphasis id="B94AEA87FF98FFE5FC1FFC43AE419EA2" box="[953,1085,990,1013]" italics="true" pageId="20" pageNumber="53">H. capuronii</emphasis>
</taxonomicName>
are consistently 4-merous, whereas some flowers of
<taxonomicName id="4C3E4D16FF98FFE5FC43FC63AEEA9942" box="[997,1174,1021,1045]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="randrianasoloi">
<emphasis id="B94AEA87FF98FFE5FC43FC63AEEA9942" box="[997,1174,1021,1045]" italics="true" pageId="20" pageNumber="53">H. randrianasoloi</emphasis>
</taxonomicName>
are 5-merous. Known specimens of
<taxonomicName id="4C3E4D16FF98FFE5FC3FFB83AE329962" box="[921,1102,1053,1077]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="randrianasoloi">
<emphasis id="B94AEA87FF98FFE5FC3FFB83AE329962" box="[921,1102,1053,1077]" italics="true" pageId="20" pageNumber="53">H. randrianasoloi</emphasis>
</taxonomicName>
dry greenish, which is uncommon in
<taxonomicName id="4C3E4D16FF98FFE5FCF1FBA3A9A49902" box="[855,984,1086,1109]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="capuronii">
<emphasis id="B94AEA87FF98FFE5FCF1FBA3A9A49902" box="[855,984,1086,1109]" italics="true" pageId="20" pageNumber="53">H. capuronii</emphasis>
</taxonomicName>
(though this character is not consistent in the latter species, and might not be in the former). In addition,
<taxonomicName id="4C3E4D16FF98FFE5FC9DFBE3A9C699C2" box="[827,954,1150,1173]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="capuronii">
<emphasis id="B94AEA87FF98FFE5FC9DFBE3A9C699C2" box="[827,954,1150,1173]" italics="true" pageId="20" pageNumber="53">H. capuronii</emphasis>
</taxonomicName>
is native to the northern province of
<collectingRegion id="49FAF877FF98FFE5FAE5FBE0AFB999C2" box="[1347,1477,1149,1173]" country="Madagascar" name="Antsiranana" pageId="20" pageNumber="53">Antsiranana</collectingRegion>
and occurs only at medium to high elevations. This provides further reason to presume that these taxa are genetically distinct, though their similarity and the variability of some features within
<taxonomicName id="4C3E4D16FF98FFE5FC54FB63AE019842" box="[1010,1149,1278,1301]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="capuronii">
<emphasis id="B94AEA87FF98FFE5FC54FB63AE019842" box="[1010,1149,1278,1301]" italics="true" pageId="20" pageNumber="53">H. capuronii</emphasis>
</taxonomicName>
may make them difficult to differentiate.
</paragraph>
<paragraph id="8B813695FF98FFE5FCC7FAA3AFE69B02" blockId="20.[827,1478,733,1621]" pageId="20" pageNumber="53">
The other low-elevation southeastern species in this group,
<taxonomicName id="4C3E4D16FF98FFE5FC9DFAC3A9C79822" box="[827,955,1373,1397]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="pulchrum">
<emphasis id="B94AEA87FF98FFE5FC9DFAC3A9C79822" box="[827,955,1373,1397]" italics="true" pageId="20" pageNumber="53">H. pulchrum</emphasis>
</taxonomicName>
, has narrower leaves, oblanceolate to elliptical or narrowly elliptical with usually cuneate to convex bases and subentire margins having 2 to 4 glands per side, and its inflorescences are racemose and usually glabrous. It does not seem plausible that the observed morphology of
<taxonomicName id="4C3E4D16FF98FFE5FAA9FA43AFBA98A2" box="[1295,1478,1501,1525]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="randrianasoloi">
<emphasis id="B94AEA87FF98FFE5FAA9FA43AFBA98A2" box="[1295,1478,1501,1525]" italics="true" pageId="20" pageNumber="53">H. randrianasoloi</emphasis>
</taxonomicName>
could have resulted from hybridization of
<taxonomicName id="4C3E4D16FF98FFE5FAACFA63AFF09B42" box="[1290,1420,1533,1557]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="pulchrum">
<emphasis id="B94AEA87FF98FFE5FAACFA63AFF09B42" box="[1290,1420,1533,1557]" italics="true" pageId="20" pageNumber="53">H. pulchrum</emphasis>
</taxonomicName>
with any of the other species known from the region, which include
<taxonomicName id="4C3E4D16FF98FFE5FC9DF9A3AE629B02" box="[827,1054,1597,1621]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="brevipedunculatum">
<emphasis id="B94AEA87FF98FFE5FC9DF9A3AE629B02" box="[827,1054,1597,1621]" italics="true" pageId="20" pageNumber="53">H. brevipedunculatum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3E4D16FF98FFE5FB8CF9A3AEA59B02" box="[1066,1241,1597,1621]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="oppositifolium">
<emphasis id="B94AEA87FF98FFE5FB8CF9A3AEA59B02" box="[1066,1241,1597,1621]" italics="true" pageId="20" pageNumber="53">H. oppositifolium</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C3E4D16FF98FFE5FAB4F9A3AFEF9B02" box="[1298,1427,1598,1621]" class="Magnoliopsida" family="Salicaceae" genus="Homalium" kingdom="Plantae" order="Malpighiales" pageId="20" pageNumber="53" phylum="Tracheophyta" rank="species" species="trigynum">
<emphasis id="B94AEA87FF98FFE5FAB4F9A3AFEF9B02" box="[1298,1427,1598,1621]" italics="true" pageId="20" pageNumber="53">H. trigynum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C324651EFF98FFE5FCC7F9E1AFCC9BFE" pageId="20" pageNumber="53" type="materials_examined">
<paragraph id="8B813695FF98FFE5FCC7F9E1AFCC9BFE" blockId="20.[827,1477,1659,1705]" pageId="20" pageNumber="53">
<emphasis id="B94AEA87FF98FFE5FCC7F9E1A9BA9BD9" box="[865,966,1660,1678]" italics="true" pageId="20" pageNumber="53">
<typeStatus id="54858837FF98FFE5FCC7F9E1A9C49BD9" box="[865,952,1660,1678]" pageId="20" pageNumber="53" type="paratype">Paratypus</typeStatus>
.–
</emphasis>
<materialsCitation id="3B563CC8FF98FFE5FC6AF9E6AFCC9BFE" ID-GBIF-Occurrence="3461735365" collectingDate="1990-10-27" collectionCode="P" collectorName="Dumetz" country="Madagascar" location="Marovony Foret" municipality="Fort-Dauphin" pageId="20" pageNumber="53" specimenCode="P04705534" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
<emphasis id="B94AEA87FF98FFE5FC6AF9E6AEA59BD9" bold="true" box="[972,1241,1659,1678]" pageId="20" pageNumber="53">
<collectingCountry id="F3297605FF98FFE5FC6AF9E6AE319BD9" box="[972,1101,1659,1678]" name="Madagascar" pageId="20" pageNumber="53">Madagascar</collectingCountry>
. Prov.
<collectingRegion id="49FAF877FF98FFE5FB2AF9E6AEB39BD9" box="[1164,1231,1659,1678]" country="Madagascar" name="Toliara" pageId="20" pageNumber="53">Toliara</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BE5ACEFFF98FFE5FB79F9E6AF1C9BD9" box="[1247,1376,1659,1678]" pageId="20" pageNumber="53">Fort-Dauphin</collectingMunicipality>
,
<location id="8EE1604EFF98FFE5FACCF9E6A9159BFE" LSID="urn:lsid:plazi:treatment:03978783FF98FFE5FF2DFE40AFCC9BFE:8EE1604EFF98FFE5FACCF9E6A9159BFE" country="Madagascar" municipality="Fort-Dauphin" name="Marovony Foret" pageId="20" pageNumber="53" stateProvince="Toliara">Marovony Forêt</location>
,
<date id="FF801055FF98FFE5FCD4F90BA9B09BFE" box="[882,972,1686,1705]" pageId="20" pageNumber="53" value="1990-10-27">
<collectingDate id="EFC4E9BDFF98FFE5FCD4F90BA9B09BFE" box="[882,972,1686,1705]" pageId="20" pageNumber="53" value="1990-10-27">27.X.1990</collectingDate>
</date>
, post-fl.,
<emphasis id="B94AEA87FF98FFE5FBBAF90BAEF39BFE" box="[1052,1167,1686,1705]" italics="true" pageId="20" pageNumber="53">
<collectorName id="26CB5343FF98FFE5FBBAF90BAE239BFF" box="[1052,1119,1686,1704]" pageId="20" pageNumber="53">Dumetz</collectorName>
1348
</emphasis>
(MO-3853942,
<collectionCode id="ED2FAE50FF98FFE5FAB8F90BAF579BFE" box="[1310,1323,1686,1705]" pageId="20" pageNumber="53">P</collectionCode>
[
<specimenCode id="DB989EEEFF98FFE5FA90F90BAFDC9BFE" box="[1334,1440,1686,1705]" pageId="20" pageNumber="53">P04705534</specimenCode>
]).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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