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<document id="2DE8821F665BCE53EEC9998EE71221D7" ID-DOI="10.11646/zootaxa.3920.1.1" ID-GBIF-Dataset="7289c86d-0ed1-4e93-b7b3-bc596042d854" ID-ISSN="1175-5326" ID-Zenodo-Dep="287886" ID-ZooBank="025F9918-F9F6-4049-B7CD-5AC632094F5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460994190417" checkinUser="plazi" docAuthor="Kataev, Boris M. &amp; Liang, Hongbin" docDate="2015" docId="039787812E57FFDD94FD4CB6EC36EE92" docLanguage="en" docName="zt03920p039.pdf" docOrigin="Zootaxa 3920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpalus (Pseudoophonus) indicus Bates 1891" docType="treatment" docVersion="7" lastPageNumber="29" masterDocId="FFAEFFF92E4DFFC1946A4A2BE927EF1A" masterDocTitle="Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)" masterLastPageNumber="39" masterPageNumber="1" pageNumber="27" updateTime="1698562432667" updateUser="plazi">
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<mods:title id="4C7C370B33493C62458D8254AF4D40F0">Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="AA7A690AFC7077FFBEF44478E96B0C9D">Kataev, Boris M.</mods:namePart>
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<mods:namePart id="9181ABB8B689C2F5C6CADBA403FC931F">Liang, Hongbin</mods:namePart>
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<mods:title id="BDCA41C8C0A725F9134ACC04E242A8A4">Zootaxa</mods:title>
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<mods:date id="9B6F99D39B2D967D725F32DE87F202BE">2015</mods:date>
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<mods:number id="5063F2FC9EEB7396BC105B52BBD1A978">3920</mods:number>
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<mods:identifier id="5F7A18F61D08475B34BD17580F377F96" type="DOI">10.11646/zootaxa.3920.1.1</mods:identifier>
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<treatment id="039787812E57FFDD94FD4CB6EC36EE92" ID-DOI="http://doi.org/10.5281/zenodo.6114290" ID-GBIF-Taxon="127671648" ID-Zenodo-Dep="6114290" LSID="urn:lsid:plazi:treatment:039787812E57FFDD94FD4CB6EC36EE92" httpUri="http://treatment.plazi.org/id/039787812E57FFDD94FD4CB6EC36EE92" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27">
<subSubSection id="C324651C2E57FFDB94FD4CB6E878E9C2" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8B8136972E57FFDB94FD4CB6EBFFE9AD" blockId="26.[151,728,1693,1752]" box="[151,728,1693,1719]" pageId="26" pageNumber="27">
<heading id="D0C981FB2E57FFDB94FD4CB6EBFFE9AD" bold="true" box="[151,728,1693,1719]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<taxonomicName id="4C3E4D142E57FFDB94FD4CB6EBFFE9AD" ID-CoL="3JSTW" authority="Bates, 1891" authorityName="Bates" authorityYear="1891" box="[151,728,1693,1719]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="indicus" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E57FFDB94FD4CB6EBFFE9AD" bold="true" box="[151,728,1693,1719]" pageId="26" pageNumber="27">
<emphasis id="B94AEA852E57FFDB94FD4CB6E82EE9AD" bold="true" box="[151,265,1693,1719]" italics="true" pageId="26" pageNumber="27">Harpalus</emphasis>
(
<emphasis id="B94AEA852E57FFDB95704CB6E8FDE9AD" bold="true" box="[282,474,1693,1719]" italics="true" pageId="26" pageNumber="27">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E57FFDB95814CB6EB64E9AD" bold="true" box="[491,579,1693,1719]" italics="true" pageId="26" pageNumber="27">indicus</emphasis>
Bates, 1891
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8136972E57FFDB94FD4C94E878E9C2" blockId="26.[151,728,1693,1752]" box="[151,351,1727,1752]" pageId="26" pageNumber="27">(Figs 2–3, 60–67)</paragraph>
</subSubSection>
<subSubSection id="C324651C2E57FFDD94FD4D2CEB82EFED" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="8B8136972E57FFDD94FD4D2CEB82EFED" blockId="26.[151,1436,1799,2005]" lastBlockId="28.[151,1436,151,392]" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27">
<emphasis id="B94AEA852E57FFDB94FD4D2CE82DE83A" bold="true" box="[151,266,1799,1824]" pageId="26" pageNumber="27">Diagnosis</emphasis>
. This species is somewhat similar to
<taxonomicName id="4C3E4D142E57FFDB96AD4D22EA14E83A" box="[711,819,1800,1824]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="B94AEA852E57FFDB96AD4D22EA14E83A" box="[711,819,1800,1824]" italics="true" pageId="26" pageNumber="27">H. davidi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3E4D142E57FFDB97184D22ED57E83A" box="[882,1136,1800,1824]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="pseudohauserianus">
<emphasis id="B94AEA852E57FFDB97184D22ED57E83A" box="[882,1136,1800,1824]" italics="true" pageId="26" pageNumber="27">H. pseudohauserianus</emphasis>
</taxonomicName>
in having pronotum with rounded basal angles, glabrous elytra with only two lateral intervals punctate and apical spur of protibia not dentate at margins (Figs 2–3), but is easily distinguished from both species by pale legs and very fine microsculpture consisting of narrow transverse meshes on elytral disc in baso-medial portion.
<taxonomicName id="4C3E4D142E57FFDB901A4D5FEC1CE896" box="[1136,1339,1908,1932]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94AEA852E57FFDB901A4D5FEC1CE896" box="[1136,1339,1908,1932]" italics="true" pageId="26" pageNumber="27">Harpalus indicus</emphasis>
</taxonomicName>
is also characterized by shallow lateral depressions of pronotum, fused with somewhat wide and shallow basal foveae, very indistinct humeral denticle, poorly visible only from behind, and metafemur with two to three setigerous pores along posterior margin. Internal sac of medial lobe of aedeagus is with long field &quot;
<emphasis id="B94AEA852E51FFDD90584AB2ED67EFAA" box="[1074,1088,153,176]" italics="true" pageId="28" pageNumber="29">a</emphasis>
&quot; in basal half and usually with peculiar folded formation &quot;
<emphasis id="B94AEA852E51FFDD95A04A97E8FFEFCE" box="[458,472,188,212]" italics="true" pageId="28" pageNumber="29">b</emphasis>
&quot; in apical half; either without any sclerotized elements or armed with a medial spiny patch (
<figureCitation id="13052A122E51FFDD94884AF4E841EFE2" box="[226,358,223,248]" captionStart="FIGURES 60 – 67" captionStartId="27.[151,263,1952,1975]" captionTargetBox="[214,1376,206,1918]" captionTargetId="figure@27.[196,1390,193,1932]" captionTargetPageId="27" captionText="FIGURES 60 – 67. Harpalus indicus, median lobe of aedeagus. 60 – 63, H. indicus indicus (Nepal); 64 – 67, H. indicus orientalis (64, Vietnam; 65 – 67, Taiwan). 60, 66, view from the left side; 61, 67, dorsal view; 62 – 65, view from the right side). a, b, folded formations of internal sac. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287898/files/figure.png" pageId="28" pageNumber="29">Figs 60–67</figureCitation>
). Body length
<quantity id="4CC69B722E51FFDD96634AF4EB86EFED" box="[521,673,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.125" metricValueMax="1.3599999999999999" metricValueMin="0.89" pageId="28" pageNumber="29" unit="mm" value="11.25" valueMax="13.6" valueMin="8.9">8.9–13.6 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF41661F2E56FFDA94FD4D8BEB6AE8EE" httpUri="https://zenodo.org/record/287898/files/figure.png" pageId="27" pageNumber="28" targetBox="[214,1376,206,1918]" targetPageId="27">
<paragraph id="8B8136972E56FFDA94FD4D8BEB6AE8EE" blockId="27.[151,1436,1952,2036]" pageId="27" pageNumber="28">
<emphasis id="B94AEA852E56FFDA94FD4D8BE874E8AC" bold="true" box="[151,339,1952,1975]" pageId="27" pageNumber="28">FIGURES 60–67.</emphasis>
<taxonomicName id="4C3E4D142E56FFDA95334D8AEB2FE8AC" box="[345,520,1953,1974]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94AEA852E56FFDA95334D8AEB2FE8AC" box="[345,520,1953,1974]" italics="true" pageId="27" pageNumber="28">Harpalus indicus</emphasis>
</taxonomicName>
, median lobe of aedeagus. 60–63,
<taxonomicName id="4C3E4D142E56FFDA97024D89ED51E8AD" authority="(Nepal)" baseAuthorityName="Nepal" box="[872,1142,1953,1975]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="indicus" subSpecies="indicus">
<emphasis id="B94AEA852E56FFDA97024D89ED38E8AC" box="[872,1055,1953,1975]" italics="true" pageId="27" pageNumber="28">H. indicus indicus</emphasis>
(Nepal)
</taxonomicName>
; 64–67,
<taxonomicName id="4C3E4D142E56FFDA90A64D89ECBBE8AC" box="[1228,1436,1953,1975]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="indicus" subSpecies="orientalis">
<emphasis id="B94AEA852E56FFDA90A64D89ECBBE8AC" box="[1228,1436,1953,1975]" italics="true" pageId="27" pageNumber="28">H. indicus orientalis</emphasis>
</taxonomicName>
(64, Vietnam; 65–67, Taiwan). 60, 66, view from the left side; 61, 67, dorsal view; 62–65, view from the right side).
<emphasis id="B94AEA852E56FFDA91404DEBEC10E8CF" box="[1322,1335,1984,2005]" italics="true" pageId="27" pageNumber="28">a</emphasis>
,
<emphasis id="B94AEA852E56FFDA91284DEBEC68E8CF" box="[1346,1359,1984,2005]" italics="true" pageId="27" pageNumber="28">b</emphasis>
, folded formations of internal sac. Scale = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C324651C2E51FFDD94AD4B2FEC36EE92" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8B8136972E51FFDD94AD4B2FEC36EE92" blockId="28.[151,1436,151,392]" pageId="28" pageNumber="29">
<emphasis id="B94AEA852E51FFDD94AD4B2FE813EE07" bold="true" box="[199,308,260,285]" pageId="28" pageNumber="29">Remarks</emphasis>
.
<taxonomicName id="4C3E4D142E51FFDD952B4B2FEB25EE06" box="[321,514,260,284]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B94AEA852E51FFDD952B4B2FEB25EE06" box="[321,514,260,284]" italics="true" pageId="28" pageNumber="29">Harpalus indicus</emphasis>
</taxonomicName>
is widely distributed from North-Western
<collectingCountry id="F32976072E51FFDD97B64B2FED31EE07" box="[988,1046,260,285]" name="India" pageId="28" pageNumber="29">India</collectingCountry>
and North
<collectingCountry id="F32976072E51FFDD90F94B2FEDD5EE07" box="[1171,1266,260,285]" name="Pakistan" pageId="28" pageNumber="29">Pakistan</collectingCountry>
throughout the Himalayan region to southern
<collectingCountry id="F32976072E51FFDD95904B0CEB67EE5A" box="[506,576,295,320]" name="China" pageId="28" pageNumber="29">China</collectingCountry>
,
<collectingCountry id="F32976072E51FFDD96254B0CEB89EE5A" box="[591,686,295,320]" name="Vietnam" pageId="28" pageNumber="29">Vietnam</collectingCountry>
and
<collectingCountry id="F32976072E51FFDD96814B03EA02EE25" box="[747,805,296,319]" name="Laos" pageId="28" pageNumber="29">Laos</collectingCountry>
. This species forms two subspecies (
<bibRefCitation id="EFAF4B662E51FFDD90BC4B03EC48EE5A" author="Kataev" box="[1238,1391,296,320]" pageId="28" pageNumber="29" refString="Kataev, B. M. (2014) Systematic and nomenclatorial notes on some taxa of Zabrini and Harpalini from the Palaearctic, Oriental and Australian regions (Coleoptera: Carabidae). Proceedings of the Zoological Institute of the Russian Academy of Sciences, 318 (3), 252 - 267." type="journal article" year="2014">Kataev 2014</bibRefCitation>
). In
<collectingCountry id="F32976072E51FFDD94FD4B67E9FAEE7F" box="[151,221,332,357]" name="China" pageId="28" pageNumber="29">China</collectingCountry>
, only the subspecies
<taxonomicName id="4C3E4D142E51FFDD95A44B66EA74EE7E" authority="Kataev, 2014" authorityName="Kataev" authorityYear="2014" box="[462,851,332,356]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="indicus" subSpecies="orientalis">
<emphasis id="B94AEA852E51FFDD95A44B66EB91EE7E" box="[462,694,332,356]" italics="true" pageId="28" pageNumber="29">H. indicus orientalis</emphasis>
<bibRefCitation id="EFAF4B662E51FFDD96D44B66EA74EE7E" author="Kataev" box="[702,851,332,356]" pageId="28" pageNumber="29" refString="Kataev, B. M. (2014) Systematic and nomenclatorial notes on some taxa of Zabrini and Harpalini from the Palaearctic, Oriental and Australian regions (Coleoptera: Carabidae). Proceedings of the Zoological Institute of the Russian Academy of Sciences, 318 (3), 252 - 267." type="journal article" year="2014">Kataev, 2014</bibRefCitation>
</taxonomicName>
is present. The nominotypical subspecies occurs in
<collectingCountry id="F32976072E51FFDD94FD4B44E9F4EE92" box="[151,211,367,392]" name="India" pageId="28" pageNumber="29">India</collectingCountry>
,
<collectingCountry id="F32976072E51FFDD94B44B44E867EE92" box="[222,320,367,392]" name="Pakistan" pageId="28" pageNumber="29">Pakistan</collectingCountry>
,
<collectingCountry id="F32976072E51FFDD95214B44E8A9EE92" box="[331,398,367,392]" name="Nepal" pageId="28" pageNumber="29">Nepal</collectingCountry>
and Buthan (
<bibRefCitation id="EFAF4B662E51FFDD964E4B5BEB90EE92" author="Kataev" box="[548,695,368,392]" pageId="28" pageNumber="29" refString="Kataev, B. M. (2001) To the knowledge of species of the subgenus Pseudoophonus (genus Harpalus) from the Himalayan region and Southern China (Coleoptera: Carabidae). Zoosystematica Rossica, 9 (2), 2000, 391 - 407." type="journal article" year="2001">Kataev 2001</bibRefCitation>
,
<bibRefCitation id="EFAF4B662E51FFDD96A84B5BEBDBEE92" author="Kataev" box="[706,764,368,392]" pageId="28" pageNumber="29" refString="Kataev, B. M. (2014) Systematic and nomenclatorial notes on some taxa of Zabrini and Harpalini from the Palaearctic, Oriental and Australian regions (Coleoptera: Carabidae). Proceedings of the Zoological Institute of the Russian Academy of Sciences, 318 (3), 252 - 267." type="journal article" year="2014">2014</bibRefCitation>
) and will perhaps be found in southern Xizang.
</paragraph>
</subSubSection>
</treatment>
</document>
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