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<document id="E05FB745854F4093B63C028B961C8336" ID-DOI="10.11646/zootaxa.3920.1.1" ID-GBIF-Dataset="7289c86d-0ed1-4e93-b7b3-bc596042d854" ID-ISSN="1175-5326" ID-Zenodo-Dep="287886" ID-ZooBank="025F9918-F9F6-4049-B7CD-5AC632094F5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460994190417" checkinUser="plazi" docAuthor="Kataev, Boris M. &amp; Liang, Hongbin" docDate="2015" docId="039787812E4AFFC694FD4985E83EE879" docLanguage="en" docName="zt03920p039.pdf" docOrigin="Zootaxa 3920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpalus (Pseudoophonus) eous Tschitscherine 1901" docType="treatment" docVersion="7" lastPageNumber="8" masterDocId="FFAEFFF92E4DFFC1946A4A2BE927EF1A" masterDocTitle="Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)" masterLastPageNumber="39" masterPageNumber="1" pageNumber="8" updateTime="1698562432667" updateUser="plazi">
<mods:mods id="8B5361A83AD132466D2AB297E434B6B2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="485C0DD5D29E3BD5779A8AA8D9C8AD98">
<mods:title id="00F4D637CCEE930A8E12B4508968E0F7">Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="2829602B1880184FF27E53937D6EA61F" type="personal">
<mods:role id="02173205E66F97E595A526E1EEFC78DF">
<mods:roleTerm id="19C419D819408931F711C5016F7ADE18">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C174F70307CD9623CCB018714C0F51E2">Kataev, Boris M.</mods:namePart>
</mods:name>
<mods:name id="CE8A65E2DC29627E7830F6736C306528" type="personal">
<mods:role id="589C22664219F99818F5F7BEE594D5E1">
<mods:roleTerm id="15DEA466B9B512A645EB96B97E063FD3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E97FC27BDB06D01FDF76A9D0AF0827CF">Liang, Hongbin</mods:namePart>
</mods:name>
<mods:typeOfResource id="4C1549333C1CCE4F68DB59299CCB3CB6">text</mods:typeOfResource>
<mods:relatedItem id="A750EADC7814CABC268C45BAE58C7FE3" type="host">
<mods:titleInfo id="2F37F5D5D76EEC6F887FA1E328D8365D">
<mods:title id="CC446257A0F51664CFB05E38DF134DDC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A612525F1E139462112EFDEEC1D69D95">
<mods:date id="04B2B31730B1CED920775D7E60A8F2B8">2015</mods:date>
<mods:detail id="3A7246475E87973D038669CABB6894B1" type="volume">
<mods:number id="612419852D3C151CE3167FEEEF6382BF">3920</mods:number>
</mods:detail>
<mods:detail id="41043A1F85180FA83EBDBAE9BF498E2A" type="issue">
<mods:number id="FF63B1ACFF0B4CB11B62EB93C853B2F1">1</mods:number>
</mods:detail>
<mods:extent id="6395C9D6C1402E1E5A9810EB0770ADF3" unit="page">
<mods:start id="6E554CB74470D823A0DB5580656D440C">1</mods:start>
<mods:end id="8646ED15A03E5D17BC84549490B0F1CF">39</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="09FF7E35239DAF0C5ED020283CC69F44">journal article</mods:classification>
<mods:identifier id="DEFB360935A5480A85D1AB47E7A397B3" type="DOI">10.11646/zootaxa.3920.1.1</mods:identifier>
<mods:identifier id="5364446087119716BDEA53B548462CB5" type="GBIF-Dataset">7289c86d-0ed1-4e93-b7b3-bc596042d854</mods:identifier>
<mods:identifier id="E9FECB01A3D766C860F40C28AD140DDB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4175E0E41EC395CBDB40DAE918CB0E4A" type="Zenodo-Dep">287886</mods:identifier>
<mods:identifier id="51E0209C89BBA5D3C2B3BA3CDF31964F" type="ZooBank">025F9918-F9F6-4049-B7CD-5AC632094F5E</mods:identifier>
</mods:mods>
<treatment id="039787812E4AFFC694FD4985E83EE879" ID-DOI="http://doi.org/10.5281/zenodo.6114244" ID-GBIF-Taxon="127671653" ID-Zenodo-Dep="6114244" LSID="urn:lsid:plazi:treatment:039787812E4AFFC694FD4985E83EE879" httpUri="http://treatment.plazi.org/id/039787812E4AFFC694FD4985E83EE879" lastPageNumber="8" pageId="7" pageNumber="8">
<subSubSection id="C324651C2E4AFFC694FD4985E801ECF0" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8B8136972E4AFFC694FD4985EA00ECD2" blockId="7.[151,807,942,1002]" box="[151,807,942,968]" pageId="7" pageNumber="8">
<heading id="D0C981FB2E4AFFC694FD4985EA00ECD2" bold="true" box="[151,807,942,968]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<taxonomicName id="4C3E4D142E4AFFC694FD4985EA00ECD2" ID-CoL="3JSTN" authority="Tschitscherine, 1901" authorityName="Tschitscherine" authorityYear="1901" box="[151,807,942,968]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="eous" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E4AFFC694FD4985EA00ECD2" bold="true" box="[151,807,942,968]" pageId="7" pageNumber="8">
<emphasis id="B94AEA852E4AFFC694FD4985E82EECD2" bold="true" box="[151,265,942,968]" italics="true" pageId="7" pageNumber="8">Harpalus</emphasis>
(
<emphasis id="B94AEA852E4AFFC695704985E8FDECD2" bold="true" box="[282,474,942,968]" italics="true" pageId="7" pageNumber="8">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E4AFFC695814985EB04ECD2" bold="true" box="[491,547,942,968]" italics="true" pageId="7" pageNumber="8">eous</emphasis>
<bibRefCitation id="EFAF4B662E4AFFC696404985EA00ECD2" author="Tschitscherine" box="[554,807,942,968]" pageId="7" pageNumber="8" refString="Tschitscherine, T. (1901) Genera des Harpalini des region Palearctique et Paleanarctique. Horae Societatis Entomologicae Rossicae, 35, 217 - 251." type="journal article" year="1901">Tschitschérine, 1901</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8136972E4AFFC694FD49FAE801ECF0" blockId="7.[151,807,942,1002]" box="[151,294,977,1002]" pageId="7" pageNumber="8">
(
<figureCitation id="13052A122E4AFFC694F549FAE839ECF0" box="[159,286,977,1002]" captionStart="FIGURES 11 – 14" captionStartId="6.[151,263,1553,1576]" captionTargetBox="[220,1362,388,1520]" captionTargetId="figure@6.[212,1374,379,1532]" captionTargetPageId="6" captionText="FIGURES 11 – 14. Harpalus, median lobe of aedeagus. 11 – 12, H. capito (Heilongjiang); 13 – 14, H. eous (Heilongjiang). 11, 13, dorsal view; 12, 14, view from the left side. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287887/files/figure.png" pageId="7" pageNumber="8">Figs 13–14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E4AFFC694FD4E32E8CDEB54" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="8B8136972E4AFFC694FD4E32E8CDEB54" blockId="7.[151,1436,1049,1102]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="AB2E11B92E4AFFC694FD4E32E8CDEB54" pageId="7" pageNumber="8">
<taxonomicName id="4C3E4D142E4AFFC694FD4E32EBC2EB35" ID-CoL="3JSTN" authority="Tschitscherine, 1901: 236" authorityName="Tschitscherine" authorityPageNumber="236" authorityYear="1901" box="[151,741,1049,1071]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="eous" subGenus="Pseudophonus">
<emphasis id="B94AEA852E4AFFC694FD4E32E9D0EB34" box="[151,247,1049,1070]" italics="true" pageId="7" pageNumber="8">Harpalus</emphasis>
(
<emphasis id="B94AEA852E4AFFC6956F4E32E8BEEB34" box="[261,409,1049,1070]" italics="true" pageId="7" pageNumber="8">Pseudophonus</emphasis>
)
<emphasis id="B94AEA852E4AFFC695C24E31E8F1EB35" box="[424,470,1050,1071]" italics="true" pageId="7" pageNumber="8">eous</emphasis>
<treatmentCitation id="0A9F10862E4AFFC695B74E32EBC2EB35" author="Tschitscherine" box="[477,741,1049,1071]" page="236" pageId="7" pageNumber="8" year="1901">
<bibRefCitation id="EFAF4B662E4AFFC695B74E32EB95EB35" author="Tschitscherine" box="[477,690,1049,1071]" pageId="7" pageNumber="8" refString="Tschitscherine, T. (1901) Genera des Harpalini des region Palearctique et Paleanarctique. Horae Societatis Entomologicae Rossicae, 35, 217 - 251." type="journal article" year="1901">Tschitschérine, 1901</bibRefCitation>
: 236
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="548588352E4AFFC696854E31EA06EB35" box="[751,801,1050,1071]" pageId="7" pageNumber="8">Type</typeStatus>
locality: &quot;Mantschurie: G. Boyan SuSu&quot; (restricted here based on designation of
<typeStatus id="548588352E4AFFC695174E13E8F8EB54" box="[381,479,1080,1102]" pageId="7" pageNumber="8" type="lectotype">lectotype</typeStatus>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E4AFFC694FD4E51EB68EAA9" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8B8136972E4AFFC694FD4E51EB42EA39" blockId="7.[151,1436,1146,1891]" pageId="7" pageNumber="8">
<emphasis id="B94AEA852E4AFFC694FD4E51E865EB89" bold="true" box="[151,322,1146,1171]" pageId="7" pageNumber="8">
<typeStatus id="548588352E4AFFC694FD4E51E9F5EB89" box="[151,210,1146,1171]" pageId="7" pageNumber="8">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="548588352E4AFFC6953E4E50E8E3EB89" box="[340,452,1147,1171]" pageId="7" pageNumber="8" type="lectotype">Lectotype</typeStatus>
(present designation): ♂, labeled &quot;G. Boyan SuSu (Mn)
<date id="FF8010572E4AFFC690024E50EDF2EB89" box="[1128,1237,1147,1171]" pageId="7" pageNumber="8" value="1997-07-23">23/VII.97</date>
Grombchev&quot; [in Cyrillic], &quot;
<taxonomicName id="4C3E4D142E4AFFC6957B4EB4E86EEBAC" box="[273,329,1183,1206]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="eous">
<emphasis id="B94AEA852E4AFFC6957B4EB4E86EEBAC" box="[273,329,1183,1206]" italics="true" pageId="7" pageNumber="8">Eous</emphasis>
</taxonomicName>
m. Typ. Tschitscherin det.&quot; (
<collectionCode id="ED2FAE522E4AFFC696FE4EB4EBEFEBAD" box="[660,712,1183,1207]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="7" pageNumber="8">ZIN</collectionCode>
).
<typeStatus id="548588352E4AFFC696B74EB5EAA2EBAD" box="[733,901,1182,1207]" pageId="7" pageNumber="8" type="paralectotype">Paralectotypes</typeStatus>
:
<specimenCount id="9D38FD1E2E4AFFC697FB4EB4EA99EBAC" box="[913,958,1183,1207]" pageId="7" pageNumber="8" type="generic">4 ♂</specimenCount>
, 1 ♀, same data as preceding (
<collectionCode id="ED2FAE522E4AFFC6914A4EB4EC73EBAD" box="[1312,1364,1183,1207]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="7" pageNumber="8">ZIN</collectionCode>
); 1 ♀, &quot;Salidugan
<date id="FF8010572E4AFFC695714EE8E895EBC1" box="[283,434,1219,1243]" pageId="7" pageNumber="8" value="1990-07-10" valueMax="1990-07-20" valueMin="1990-07-10">10–20.VII.90</date>
Komarov&quot; [in Cyrillic], &quot;
<taxonomicName id="4C3E4D142E4AFFC696B64EE8EA33EBC0" box="[732,788,1219,1242]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="eous">
<emphasis id="B94AEA852E4AFFC696B64EE8EA33EBC0" box="[732,788,1219,1242]" italics="true" pageId="7" pageNumber="8">Eous</emphasis>
</taxonomicName>
m. Typ. Tschitscherin det.&quot; (
<collectionCode id="ED2FAE522E4AFFC6900B4EE8EDB2EBC1" box="[1121,1173,1219,1243]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="7" pageNumber="8">ZIN</collectionCode>
),
<specimenCount id="9D38FD1E2E4AFFC690C14EE8EDFEEBC0" box="[1195,1241,1219,1243]" pageId="7" pageNumber="8" type="generic">1 ♂</specimenCount>
, &quot;M. Salidugan i Erdugan
<date id="FF8010572E4AFFC6956B4ECDE8B0EBE5" box="[257,407,1254,1279]" pageId="7" pageNumber="8" value="1996-07-10" valueMax="1996-07-20" valueMin="1996-07-10">10–20.VII.96</date>
Komarov&quot; [in Cyrillic], &quot;
<taxonomicName id="4C3E4D142E4AFFC696A04ECCEA25EBE4" box="[714,770,1255,1278]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="eous">
<emphasis id="B94AEA852E4AFFC696A04ECCEA25EBE4" box="[714,770,1255,1278]" italics="true" pageId="7" pageNumber="8">Eous</emphasis>
</taxonomicName>
m. Typ. Tschitscherin det.&quot; (
<collectionCode id="ED2FAE522E4AFFC690304ECCEDA9EBE5" box="[1114,1166,1255,1279]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="7" pageNumber="8">ZIN</collectionCode>
), 1 ♀, &quot;Vlsdsk., 7.90&quot;, &quot;
<taxonomicName id="4C3E4D142E4AFFC694C84F20E9FDEA38" box="[162,218,1291,1314]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="eous">
<emphasis id="B94AEA852E4AFFC694C84F20E9FDEA38" box="[162,218,1291,1314]" italics="true" pageId="7" pageNumber="8">Eous</emphasis>
</taxonomicName>
m., Typ. Tschitscherin det.&quot; (
<collectionCode id="ED2FAE522E4AFFC6964F4F20EB7EEA39" box="[549,601,1291,1315]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="7" pageNumber="8">ZIN</collectionCode>
).
</paragraph>
<paragraph id="8B8136972E4AFFC694AD4F05EB68EAA9" blockId="7.[151,1436,1146,1891]" pageId="7" pageNumber="8">
<emphasis id="B94AEA852E4AFFC694AD4F05EB1AEA5D" bold="true" box="[199,573,1326,1351]" pageId="7" pageNumber="8">Additional material examined</emphasis>
. More than
<specimenCount id="9D38FD1E2E4AFFC696B44F04EAABEA5D" box="[734,908,1326,1351]" pageId="7" pageNumber="8" type="generic">420 specimens</specimenCount>
(140 from
<collectingCountry id="F32976072E4AFFC690484F05ED4EEA5D" box="[1058,1129,1326,1351]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
), including the following specimens from Hubei and Hunan.
<collectingCountry id="F32976072E4AFFC6964A4F79EB4EEA71" box="[544,617,1362,1387]" name="China" pageId="7" pageNumber="8">
<emphasis id="B94AEA852E4AFFC6964A4F79EB4EEA71" bold="true" box="[544,617,1362,1387]" pageId="7" pageNumber="8">China</emphasis>
</collectingCountry>
.
<collectionCode id="ED2FAE522E4AFFC6961D4F78EBEBEA70" box="[631,716,1363,1386]" pageId="7" pageNumber="8">HUBEI</collectionCode>
:
<specimenCount id="9D38FD1E2E4AFFC696B14F78EA21EA70" box="[731,774,1363,1387]" pageId="7" pageNumber="8" type="generic">1 ♂</specimenCount>
, Shennongjia,
<quantity id="4CC69B722E4AFFC697C14F79ED25EA70" box="[939,1026,1362,1387]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="7" pageNumber="8" unit="m" value="1650.0">1650 m</quantity>
,
<date id="FF8010572E4AFFC690674F78EDBFEA71" box="[1037,1176,1363,1387]" pageId="7" pageNumber="8" value="1980-07-12">12.VII.1980</date>
, Yu Peiyu leg. (
<collectionCode id="ED2FAE522E4AFFC6912D4F78EC5CEA71" box="[1351,1403,1363,1387]" pageId="7" pageNumber="8">IOZ</collectionCode>
); 2 ♀, same data as preceding, but
<quantity id="4CC69B722E4AFFC695934F5CEB76EA94" box="[505,593,1399,1423]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="8" unit="m" value="1800.0">1800 m</quantity>
,
<date id="FF8010572E4AFFC696374F5CEBC4EA95" box="[605,739,1399,1423]" pageId="7" pageNumber="8" value="1981-08-02">2.VIII.1981</date>
, Han Yinheng leg. (
<collectionCode id="ED2FAE522E4AFFC697AF4F5CEADEEA95" box="[965,1017,1399,1423]" pageId="7" pageNumber="8">IOZ</collectionCode>
).
<collectionCode id="ED2FAE522E4AFFC690674F5CED50EA95" box="[1037,1143,1399,1423]" pageId="7" pageNumber="8">HUNAN</collectionCode>
:
<specimenCount id="9D38FD1E2E4AFFC690EB4F5CED89EA94" box="[1153,1198,1399,1423]" pageId="7" pageNumber="8" type="generic">1 ♂</specimenCount>
, Quyuan Nongchang (farm),
<date id="FF8010572E4AFFC694864FB0E849EAA9" box="[236,366,1435,1459]" pageId="7" pageNumber="8" value="1987-06-18">18.VI.1987</date>
, no collector (
<collectionCode id="ED2FAE522E4AFFC696644FB0EB65EAA9" box="[526,578,1435,1459]" pageId="7" pageNumber="8">IOZ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E4AFFC694AD4F95EADBE958" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8B8136972E4AFFC694AD4F95EADBE958" blockId="7.[151,1436,1146,1891]" pageId="7" pageNumber="8">
<emphasis id="B94AEA852E4AFFC694AD4F95E81DEACD" bold="true" box="[199,314,1470,1495]" pageId="7" pageNumber="8">Diagnosis</emphasis>
. This species is recognizable by combination of the following characters: head large, tempora setaceous, pronotum with several lateral setae along sides, and elytra densely punctate and pubescent on all intervals, without row of discal pores on intervals 1, 3, 5 and 7. Median lobe of aedeagus (
<figureCitation id="13052A122E4AFFC690FB4C2DEC36E905" box="[1169,1297,1542,1567]" captionStart="FIGURES 11 – 14" captionStartId="6.[151,263,1553,1576]" captionTargetBox="[220,1362,388,1520]" captionTargetId="figure@6.[212,1374,379,1532]" captionTargetPageId="6" captionText="FIGURES 11 – 14. Harpalus, median lobe of aedeagus. 11 – 12, H. capito (Heilongjiang); 13 – 14, H. eous (Heilongjiang). 11, 13, dorsal view; 12, 14, view from the left side. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287887/files/figure.png" pageId="7" pageNumber="8">Figs 13–14</figureCitation>
) has several somewhat large spiny patches in the internal sac. Body length
<quantity id="4CC69B722E4AFFC697384C01EADFE958" box="[850,1016,1578,1603]" metricMagnitude="-2" metricUnit="m" metricValue="1.365" metricValueMax="1.48" metricValueMin="1.25" pageId="7" pageNumber="8" unit="mm" value="13.65" valueMax="14.8" valueMin="12.5">12.5–14.8 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E4AFFC694AD4C65E83EE879" pageId="7" pageNumber="8" type="distribution">
<paragraph id="8B8136972E4AFFC694AD4C65E9C6E9C9" blockId="7.[151,1436,1146,1891]" pageId="7" pageNumber="8">
<emphasis id="B94AEA852E4AFFC694AD4C65E87EE97D" bold="true" box="[199,345,1614,1639]" pageId="7" pageNumber="8">Distribution</emphasis>
.
<collectingCountry id="F32976072E4AFFC695044C65E892E97D" box="[366,437,1614,1639]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
: mainly in the north-eastern and central parts. It was recorded from Nei Mongol, Heilongjiang, Jilin, Liaoning, Beijing, Hebei, Shaanxi, Sichuan, Anhui, Jiangsu, Shanghai, Zhejiang, Hunan (
<bibRefCitation id="EFAF4B662E4AFFC691334C59E83DE9B5" author="Liang" pageId="7" pageNumber="8" refString="Liang, H. - B. &amp; Yu, P. - Y. (2000) Species of ground beetles (Coleoptera: Carabidae) predating oriental armyworm (Lepidoptera: Noctuidae) in China. Natural Enemies of Insects, 22 (4), 160 - 167. [in Chinese with English summary]" type="journal article" year="2000">Liang &amp; Yu 2000</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E4AFFC695434CBCEB24E9B5" author="Kataev" box="[297,515,1686,1711]" pageId="7" pageNumber="8" refString="Kataev, B. M., Wrase, D. W. &amp; Ito, N. (2003) Harpalina. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
Kataev
<emphasis id="B94AEA852E4AFFC695E84CBCE892E9B4" box="[386,437,1686,1710]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
. 2003
</bibRefCitation>
), and Hubei (
<bibRefCitation id="EFAF4B662E4AFFC696CB4CBCEA3FE9B5" author="Hua" box="[673,792,1687,1711]" pageId="7" pageNumber="8" refString="Hua, L. (2002) List of Chinese Insects. Vol. II. Zhongshan (Sun Yat-sen) University Press, Guangzhou, 612 pp." type="book" year="2002">Hua 2002</bibRefCitation>
). Additional localities from Hubei and Hunan are listed above.
</paragraph>
<paragraph id="8B8136972E4AFFC694AD4CF5E83EE879" blockId="7.[151,1436,1146,1891]" pageId="7" pageNumber="8">
The species is also known from Eastern
<collectingCountry id="F32976072E4AFFC696DD4CF5EA25E9ED" box="[695,770,1758,1783]" name="Russia" pageId="7" pageNumber="8">Russia</collectingCountry>
(Buryatia, Chita and Amur Provinces, Khabarovsk and Maritime Territories), North and
<collectingCountry id="F32976072E4AFFC696484D29EB93E800" box="[546,692,1794,1819]" name="South Korea" pageId="7" pageNumber="8">South Korea</collectingCountry>
including Jejudo, and
<collectingCountry id="F32976072E4AFFC697A64D28ED2BE800" box="[972,1036,1795,1818]" name="Japan" pageId="7" pageNumber="8">Japan</collectingCountry>
(Hokkaido, Honshu, Sikoku, and Kyushu Islands) (
<bibRefCitation id="EFAF4B662E4AFFC6950A4D0DE8C5E825" author="Habu" box="[352,482,1830,1855]" pageId="7" pageNumber="8" refString="Habu, A. (1973) Carabidae: Harpalini (Insecta, Coleoptera). Fauna Japonica. Keigaku Publishing Co., Tokio, xiii + 430 pp." type="book" year="1973">Habu 1973</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E4AFFC695854D0DEB56E825" author="Lafer" box="[495,625,1830,1855]" pageId="7" pageNumber="8" refString="Lafer, G. Sh. (1989) Fam. Carabidae-The ground-beetles. In: Lehr, P. A. (Ed.), Opredelitel' nasekomykh Dal'nego Vostoka SSSR v shesti tomakh. Tom III. Zhestkokrylye, ili zhuki. Chast' 1. Nauka, Leningrad, pp. 70 - 222. [in Russian]" type="journal article" year="1989">Lafer 1989</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E4AFFC696144D0DEA8EE825" author="Kryzhanovskij" box="[638,937,1830,1855]" pageId="7" pageNumber="8" refString="Kryzhanovskij, O. L., Belousov, I. A., Kabak, I. I., Kataev, B. M., Makarov, K. V., Shilenkov, V. G. (1995) A checklist of the groundbeetles of Russia and adjacent lands (Insecta, Coleoptera, Carabidae). Pensoft Publishers, Sofia - Moscow, 272 pp." type="book" year="1995">
Kryzhanovskij
<emphasis id="B94AEA852E4AFFC697414D0CEA7AE824" box="[811,861,1830,1854]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
. 1995
</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E4AFFC697DC4D0CED6EE825" author="Kataev" box="[950,1097,1831,1855]" pageId="7" pageNumber="8" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev 1997</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E4AFFC6903D4D0CEC0AE825" author="Kataev" box="[1111,1325,1830,1855]" pageId="7" pageNumber="8" refString="Kataev, B. M., Wrase, D. W. &amp; Ito, N. (2003) Harpalina. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
Kataev
<emphasis id="B94AEA852E4AFFC690C54D0CEDC5E824" box="[1199,1250,1830,1854]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
. 2003
</bibRefCitation>
; Moon &amp;
<bibRefCitation id="EFAF4B662E4AFFC694FD4D61E82AE879" author="Paik" box="[151,269,1866,1891]" pageId="7" pageNumber="8" refString="Moon, Ch. - S. &amp; Paik, J. - Ch. (2006) Notes on Harpalini (Coleoptera, Carabidae) from Korea (1 - 3). Korean Journal of Soil Zoology, 11 (1 - 2), 30 - 84." type="journal article" year="2006">Paik 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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