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<document id="1D2E7476400FE8DC8AB20BC36FC5EF0A" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFFA5730932FAFF2DE71FF961" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Zoosphaerium isalo Wesener 2009, new species" docType="treatment" docUuid="6ACBBAA4-F623-47AF-8A06-EBED167E252F" docUuidSource="ZooBank" docVersion="4" lastPageNumber="90" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="87" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="1622DD47607E45D53EDA1CF8F526433A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D9ECBA69441D97152F146446AF179320">
<mods:title id="95D5A7FAE83070A1CA18600E8E90A737">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
</mods:titleInfo>
<mods:name id="7FFC3C5E696B830075170409979855D2" type="personal">
<mods:role id="095B6044AE4EC797AE984AF6EF8519BE">
<mods:roleTerm id="92204AC3C1A158D5540C42020373EA77">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="69DDED60EA9C1CBBA23613C6D438139D">Wesener, Thomas</mods:namePart>
</mods:name>
<mods:typeOfResource id="CC6D91F5E9A0E89FA3DDB30E22440D9C">text</mods:typeOfResource>
<mods:relatedItem id="1B77EAC9018D0D210568E7BD492BE5B6" type="host">
<mods:titleInfo id="CBFCDB4C9F2349C4AF51AA7217D697E4">
<mods:title id="5E62CE4380EEFFA91C9FEDDCD790D93F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3ECD84613BB31B411DC42ECA016E35E1">
<mods:date id="9AD4E48ED861950C9B97666CD6C51178">2009</mods:date>
<mods:detail id="10D87D6911EB99B5F94F445C2EB10DF1" type="pubDate">
<mods:number id="9C8A15B26AAC621EE79EE85A4FE197F4">2009-05-11</mods:number>
</mods:detail>
<mods:detail id="4B5FDFB9EA158428D66CADCC9B1FDD25" type="volume">
<mods:number id="1312A1F9D4E20D59B5C9FD8120583C58">2097</mods:number>
</mods:detail>
<mods:detail id="9B666D721B16089B62068435F5185428" type="issue">
<mods:number id="EBC14885885C28B6EA269ED748AE30AA">1</mods:number>
</mods:detail>
<mods:extent id="B4678B47FCB6315ADB0247A4BF82BD28" unit="page">
<mods:start id="5A5D8896148060B9DCDC517794438B1A">1</mods:start>
<mods:end id="043B8573B0318C163C3948FD5AFB33C9">134</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="A582EE61BD67D9CF6A3EE59A07301658">
<mods:url id="6D3130C97C2603B91BE88E3D039EF817">https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1</mods:url>
</mods:location>
<mods:classification id="0BB29CA13C52D4E572D81869BF361965">journal article</mods:classification>
<mods:identifier id="FB5D38ED38D3260C859392251062A57C" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
<mods:identifier id="562183B9C6EEF450B06899BA34CAFFF0" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="CA1DACB389AB4E12535D8047010B56BF" type="Zenodo-Dep">5316536</mods:identifier>
<mods:identifier id="EE2FF787BA75B9EC04D536B109290C33" type="ZooBank">0E18167A-78EB-4079-8BF4-972A4AC88C69</mods:identifier>
</mods:mods>
<treatment id="03FFBE1DFFA5730932FAFF2DE71FF961" LSID="urn:lsid:zoobank.org:act:6ACBBAA4-F623-47AF-8A06-EBED167E252F" httpUri="http://treatment.plazi.org/id/03FFBE1DFFA5730932FAFF2DE71FF961" lastPageId="89" lastPageNumber="90" pageId="86" pageNumber="87">
<subSubSection id="C34C5C80FFA5730632FAFF2DE122FF68" box="[151,549,151,177]" pageId="86" pageNumber="87" type="nomenclature">
<paragraph id="8BE90F0BFFA5730632FAFF2DE122FF68" blockId="86.[151,549,151,177]" box="[151,549,151,177]" pageId="86" pageNumber="87">
<heading id="D0A1B867FFA5730632FAFF2DE122FF68" bold="true" box="[151,549,151,177]" fontSize="11" level="1" pageId="86" pageNumber="87" reason="1">
<emphasis id="B922D319FFA5730632FAFF2DE122FF68" bold="true" box="[151,549,151,177]" pageId="86" pageNumber="87">
<emphasis id="B922D319FFA5730632FAFF2DE289FF68" bold="true" box="[151,398,151,177]" italics="true" pageId="86" pageNumber="87">
<taxonomicName id="4C567488FFA5730632FAFF2DE28FFF68" ID-CoL="5D7HC" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,392,151,177]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="isalo" status="sp. nov.">Zoosphaerium isalo</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A2116E62FFA5730633F8FF2DE122FF68" box="[405,549,151,177]" pageId="86" pageNumber="87" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFA5730632FAFF5AE2D0FEFB" pageId="86" pageNumber="87" type="etymology">
<paragraph id="8BE90F0BFFA5730632FAFF5AE7CAFF22" blockId="86.[151,1436,224,406]" box="[151,1229,224,251]" pageId="86" pageNumber="87">
<emphasis id="B922D319FFA5730632FAFF5AE287FF23" bold="true" box="[151,384,224,250]" pageId="86" pageNumber="87">Derivatio nominis:</emphasis>
<taxonomicName id="4C567488FFA5730633E5FF5BE2C6FF23" authorityName="Wesener" authorityYear="2009" box="[392,449,225,250]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="isalo">
<emphasis id="B922D319FFA5730633E5FF5BE2C6FF23" box="[392,449,225,250]" italics="true" pageId="86" pageNumber="87">isalo</emphasis>
</taxonomicName>
, noun in apposition, after the
<typeStatus id="54EDB1A9FFA57306314DFF5BE055FF22" box="[800,850,225,251]" pageId="86" pageNumber="87">type</typeStatus>
locality,
<taxonomicName id="4C567488FFA5730631D3FF5BE735FF22" authority=", SW" authorityName="SW" box="[958,1074,225,251]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="isalo">Isalo, SW</taxonomicName>
<collectingCountry id="F3414F9BFFA573063654FF5BE7CEFF22" box="[1081,1225,225,251]" name="Madagascar" pageId="86" pageNumber="87">Madagascar</collectingCountry>
.
</paragraph>
<paragraph id="8BE90F0BFFA5730632ABFEBDE2D0FEFB" blockId="86.[151,1436,224,406]" box="[198,471,263,290]" pageId="86" pageNumber="87">
<emphasis id="B922D319FFA5730632ABFEBDE298FEF8" bold="true" box="[198,415,263,289]" pageId="86" pageNumber="87">Studied material:</emphasis>
3 M
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFA5730732AAFE94E257FB1D" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87" type="materials_examined">
<paragraph id="8BE90F0BFFA5730632AAFE94E2F1FE4F" blockId="86.[151,1436,224,406]" pageId="86" pageNumber="87">
<materialsCitation id="3B3E0556FFA5730632AAFE94E2F1FE4F" collectingDate="2003-01-27" collectingDateMax="2003-01-31" collectingDateMin="2003-01-27" collectionCode="CAS" collectorName="Fisher &amp; Griswold" country="Madagascar" county="Locus" elevation="725" latitude="-22.481667" location="Parc National d'Isalo" longLatPrecision="21" longitude="45.461666" municipality="Province" pageId="86" pageNumber="87" specimenCode="BLF 7301" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B922D319FFA5730632AAFE94E245FE91" bold="true" box="[199,322,302,328]" pageId="86" pageNumber="87">
<typeStatus id="54EDB1A9FFA5730632AAFE94E23AFE91" box="[199,317,302,328]" pageId="86" pageNumber="87" type="holotype">Holotype</typeStatus>
:
</emphasis>
1 M (
<quantity id="4CAEA2EEFFA5730633E1FE94E2E7FE91" box="[396,480,302,328]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="86" pageNumber="87" unit="mm" value="24.0">24 mm</quantity>
long),
<emphasis id="B922D319FFA57306305BFE94E1F2FE91" bold="true" box="[566,757,302,328]" pageId="86" pageNumber="87">
<collectionCode id="ED4797CEFFA57306305BFE94E177FE91" box="[566,624,302,328]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="86" pageNumber="87" type="Museum">CAS</collectionCode>
<specimenCode id="DBF0A770FFA573063014FE94E1F2FE91" box="[633,757,302,328]" collectionCode="BLF" pageId="86" pageNumber="87">BLF 7301</specimenCode>
</emphasis>
,
<emphasis id="B922D319FFA573063169FE95E0A1FE91" box="[772,934,303,328]" italics="true" pageId="86" pageNumber="87">
<collectingCounty id="62887787FFA573063169FE95E04DFE91" box="[772,842,303,328]" pageId="86" pageNumber="87">Locus</collectingCounty>
typicus
</emphasis>
:
<collectingCountry id="F3414F9BFFA5730631DBFE94E742FE91" box="[950,1093,302,328]" name="Madagascar" pageId="86" pageNumber="87">Madagascar</collectingCountry>
,
<collectingMunicipality id="6B8D9571FFA57306363FFE94E7BDFE91" box="[1106,1210,302,328]" pageId="86" pageNumber="87">Province</collectingMunicipality>
<collectingRegion id="4992C1E9FFA5730636AEFE94E65BFE91" box="[1219,1372,302,328]" country="Madagascar" name="Fianarantsoa" pageId="86" pageNumber="87">Fianarantsoa</collectingRegion>
,
<location id="8E8959D0FFA573063705FE94E25FFEB6" LSID="urn:lsid:plazi:treatment:03FFBE1DFFA5730932FAFF2DE71FF961:8E8959D0FFA573063705FE94E25FFEB6" country="Madagascar" county="Locus" latitude="-22.481667" longLatPrecision="21" longitude="45.461666" municipality="Province" name="Parc National d'Isalo" pageId="86" pageNumber="87" stateProvince="Fianarantsoa">Parc National d'Isalo</location>
,
<locationDeviation id="9A5B1629FFA573063306FEEFE19DFEB6" box="[363,666,341,367]" location="Ranohira" pageId="86" pageNumber="87">
<quantity id="4CAEA2EEFFA573063306FEEFE2C5FEB6" box="[363,450,341,367]" metricMagnitude="3" metricUnit="m" metricValue="9.1" pageId="86" pageNumber="87" unit="km" value="9.1">9.1 km</quantity>
354°N Ranohira
</locationDeviation>
,
<quantity id="4CAEA2EEFFA5730630C6FEEFE1F9FEB6" box="[683,766,341,367]" metricMagnitude="2" metricUnit="m" metricValue="7.25" pageId="86" pageNumber="87" unit="m" value="725.0">
<elevation id="007BE838FFA5730630C6FEEFE1F9FEB6" box="[683,766,341,367]" metricMagnitude="2" metricUnit="m" metricValue="7.25" pageId="86" pageNumber="87" unit="m" value="725.0">725 m</elevation>
</quantity>
, gallery forest,
<geoCoordinate id="EE6269CCFFA5730631AAFEEFE748FEB6" box="[967,1103,341,367]" degrees="22" direction="south" minutes="28" orientation="latitude" pageId="86" pageNumber="87" precision="15" seconds="54" value="-22.481667">22°28'54&quot;S</geoCoordinate>
,
<geoCoordinate id="EE6269CCFFA57306360CFEEFE7FDFEB6" box="[1121,1274,341,367]" degrees="045" direction="east" minutes="27" orientation="longitude" pageId="86" pageNumber="87" precision="15" seconds="42" value="45.461666">045°27'42&quot;E</geoCoordinate>
, coll.
<collectorName id="26A36ADDFFA573063720FEEFE69FFEB6" box="[1357,1432,341,367]" pageId="86" pageNumber="87">Fisher</collectorName>
,
<collectorName id="26A36ADDFFA5730632FAFEC6E203FE4F" box="[151,260,380,406]" pageId="86" pageNumber="87">Griswold</collectorName>
<emphasis id="B922D319FFA573063366FEC6E241FE4C" box="[267,326,380,405]" italics="true" pageId="86" pageNumber="87">et al.</emphasis>
,
<date id="FFE829CBFFA573063338FEC6E2F4FE4F" box="[341,499,380,406]" pageId="86" pageNumber="87" value="2003-01-27" valueMax="2003-01-31" valueMin="2003-01-27">
<collectingDate id="EFACD023FFA573063338FEC6E2F4FE4F" box="[341,499,380,406]" pageId="86" pageNumber="87" value="2003-01-27" valueMax="2003-01-31" valueMin="2003-01-27">27–31.i.2003</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE90F0BFFA5730632FAF8BAE11CF87B" blockId="86.[151,1436,1792,1954]" pageId="86" pageNumber="87">
<emphasis id="B922D319FFA5730632FAF8BAE228F8C1" bold="true" box="[151,303,1792,1816]" pageId="86" pageNumber="87">FIGURE 058</emphasis>
.
<taxonomicName id="4C567488FFA573063353F8BAE112F8CE" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[318,533,1792,1815]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="isalo" status="sp. nov.">
<emphasis id="B922D319FFA573063353F8BAE112F8CE" box="[318,533,1792,1815]" italics="true" pageId="86" pageNumber="87">Zoosphaerium isalo</emphasis>
</taxonomicName>
<emphasis id="B922D319FFA573063070F8BAE15CF8C1" bold="true" box="[541,603,1792,1816]" pageId="86" pageNumber="87">
<taxonomicNameLabel id="A2116E62FFA573063070F8BAE15CF8C1" box="[541,603,1792,1816]" pageId="86" pageNumber="87" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, A–D
<typeStatus id="54EDB1A9FFA5730630CBF8BAE001F8CE" box="[678,774,1792,1815]" pageId="86" pageNumber="87" type="holotype">holotype</typeStatus>
, E, F specimen from Zombitse (
<collectionCode id="ED4797CEFFA573063605F8BAE79AF8CE" box="[1128,1181,1792,1815]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="86" pageNumber="87" type="Museum">CAS</collectionCode>
Mei-99 Ma-14)
<emphasis id="B922D319FFA573063738F8BAE66FF8C1" bold="true" box="[1365,1384,1792,1816]" pageId="86" pageNumber="87">A</emphasis>
: left anterior telopods, anterior view;
<emphasis id="B922D319FFA573063396F898E10AF8E3" bold="true" box="[507,525,1826,1850]" pageId="86" pageNumber="87">B</emphasis>
: last two podomeres of anterior telopod, lateral view;
<emphasis id="B922D319FFA573063632F898E775F8E3" bold="true" box="[1119,1138,1826,1850]" pageId="86" pageNumber="87">C</emphasis>
: posterior telopod, anterior view;
<emphasis id="B922D319FFA5730632B6F8FFE3E9F884" bold="true" box="[219,238,1861,1885]" pageId="86" pageNumber="87">D</emphasis>
: chela of posterior telopod, posterior view;
<emphasis id="B922D319FFA5730630BEF8FFE1E2F884" bold="true" box="[723,741,1861,1885]" pageId="86" pageNumber="87">E</emphasis>
: posterior telopod, anterior view;
<emphasis id="B922D319FFA573063631F8FFE76BF884" bold="true" box="[1116,1132,1861,1885]" pageId="86" pageNumber="87">F</emphasis>
: chela of posterior telopod, posterior view. Abbreviations:
<emphasis id="B922D319FFA57306338FF8D2E10EF859" bold="true" box="[482,521,1896,1920]" pageId="86" pageNumber="87">cav</emphasis>
= cavity;
<emphasis id="B922D319FFA573063018F8D2E195F859" bold="true" box="[629,658,1896,1920]" pageId="86" pageNumber="87">ml</emphasis>
= membranous lobe;
<emphasis id="B922D319FFA573063111F8D2E099F859" bold="true" box="[892,926,1896,1920]" pageId="86" pageNumber="87">s-p</emphasis>
= sclerotized spot;
<emphasis id="B922D319FFA57306361CF8D2E7A6F859" bold="true" box="[1137,1185,1896,1920]" pageId="86" pageNumber="87">scl-t</emphasis>
= sclerotized teeth;
<emphasis id="B922D319FFA573063717F8D2E69BF859" bold="true" box="[1402,1436,1896,1920]" pageId="86" pageNumber="87">SR</emphasis>
= stridulation rib. Scale bar =
<quantity id="4CAEA2EEFFA5730633BAF831E110F87B" box="[471,535,1931,1954]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="86" pageNumber="87" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="8BE90F0BFFA4730732FAFC74E1C5FBD1" blockId="87.[151,1435,974,1032]" pageId="87" pageNumber="88">
<emphasis id="B922D319FFA4730732FAFC74E237FC3F" bold="true" box="[151,304,974,998]" pageId="87" pageNumber="88">FIGURE 059</emphasis>
.
<taxonomicName id="4C567488FFA47307332DFC74E11FFC3C" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[320,536,974,997]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="isalo" status="sp. nov.">
<emphasis id="B922D319FFA47307332DFC74E11FFC3C" box="[320,536,974,997]" italics="true" pageId="87" pageNumber="88">Zoosphaerium isalo</emphasis>
</taxonomicName>
<emphasis id="B922D319FFA47307304CFC74E167FC3F" bold="true" box="[545,608,974,998]" pageId="87" pageNumber="88">
<taxonomicNameLabel id="A2116E62FFA47307304CFC74E167FC3F" box="[545,608,974,998]" pageId="87" pageNumber="88" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FFA47307301DFC74E1C8FC3C" box="[624,719,974,997]" pageId="87" pageNumber="88" type="paratype">paratype</typeStatus>
,
<collectionCode id="ED4797CEFFA4730730B1FC74E012FC3C" box="[732,789,974,997]" pageId="87" pageNumber="88">SEM</collectionCode>
, antenna
<emphasis id="B922D319FFA473073112FC74E095FC3F" bold="true" box="[895,914,974,998]" pageId="87" pageNumber="88">A</emphasis>
: first antennomere with
<emphasis id="B922D319FFA4730736CEFC74E676FC3C" box="[1187,1393,974,997]" italics="true" pageId="87" pageNumber="88">sensilla basiconica</emphasis>
;
<emphasis id="B922D319FFA4730737EEFC74E692FC3F" bold="true" box="[1411,1429,974,998]" pageId="87" pageNumber="88">B</emphasis>
: lateral view. Abbreviation:
<emphasis id="B922D319FFA4730733D7FC4AE2D4FBD1" bold="true" box="[442,467,1008,1032]" pageId="87" pageNumber="88">sb</emphasis>
=
<emphasis id="B922D319FFA47307339DFC4BE1BCFBD1" box="[496,699,1009,1032]" italics="true" pageId="87" pageNumber="88">sensilla basiconica</emphasis>
.
</paragraph>
<paragraph id="8BE90F0BFFA4730732AAFB8CE05AFB89" blockId="87.[151,1436,1078,2016]" box="[199,861,1078,1104]" pageId="87" pageNumber="88">
<materialsCitation id="3B3E0556FFA4730732AAFB8CE05EFB89" box="[199,857,1078,1104]" collectingDate="2003-01-27" collectingDateMax="2003-01-31" collectingDateMin="2003-01-27" collectionCode="CAS" collectorName="Fisher &amp; Griswold" country="Madagascar" county="Locus" elevation="725" latitude="-22.481667" location="Parc National d'Isalo" longLatPrecision="21" longitude="45.461666" municipality="Province" pageId="87" pageNumber="88" specimenCode="BLF 7301" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<emphasis id="B922D319FFA4730732AAFB8CE245FB89" bold="true" box="[199,322,1078,1104]" pageId="87" pageNumber="88">
<typeStatus id="54EDB1A9FFA4730732AAFB8CE23AFB89" box="[199,317,1078,1104]" pageId="87" pageNumber="88" type="paratype">Paratype</typeStatus>
:
</emphasis>
1 M,
<emphasis id="B922D319FFA4730733E5FB8CE141FB89" bold="true" box="[392,582,1078,1104]" pageId="87" pageNumber="88">
<collectionCode id="ED4797CEFFA4730733E5FB8CE2C4FB89" box="[392,451,1078,1104]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="87" pageNumber="88" type="Museum">CAS</collectionCode>
<specimenCode id="DBF0A770FFA4730733A7FB8CE141FB89" box="[458,582,1078,1104]" collectionCode="BLF" pageId="87" pageNumber="88">BLF 7301</specimenCode>
</emphasis>
, same data as holotype
</materialsCitation>
;
</paragraph>
<paragraph id="8BE90F0BFFA4730732AAFBE6E257FB1D" blockId="87.[151,1436,1078,2016]" pageId="87" pageNumber="88">
<emphasis id="B922D319FFA4730732AAFBE6E289FBAF" bold="true" box="[199,398,1116,1142]" pageId="87" pageNumber="88">Other material:</emphasis>
1 M,
<materialsCitation id="3B3E0556FFA4730733B4FBE6E24BFB1D" collectingDate="1999-12-15" collectingDateMax="1999-12-18" collectingDateMin="1999-12-15" collectionCode="CAS, BLF" collectorName="E. L. Schlinger &amp; Schlinger, M. E" elevation="825" latitude="-22.88231" location="Zombitse Nature Reserve" longLatPrecision="1" longitude="44.70062" municipality="Province" pageId="87" pageNumber="88" specimenCount="1" stateProvince="Toliara">
<emphasis id="B922D319FFA4730733B4FBE6E016FBAF" bold="true" box="[473,785,1116,1142]" pageId="87" pageNumber="88">
<collectionCode id="ED4797CEFFA4730733B4FBE6E113FBAF" box="[473,532,1116,1142]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="87" pageNumber="88" type="Museum">CAS</collectionCode>
<collectionCode id="ED4797CEFFA473073070FBE6E151FBAF" box="[541,598,1116,1142]" pageId="87" pageNumber="88">BLF</collectionCode>
Mei-99 Ma-14
</emphasis>
,
<collectingMunicipality id="6B8D9571FFA47307314FFBE7E08DFBAE" box="[802,906,1117,1143]" pageId="87" pageNumber="88">Province</collectingMunicipality>
<collectingRegion id="4992C1E9FFA4730731FEFBE7E0EFFBAE" box="[915,1000,1117,1143]" country="Madagascar" name="Toliara" pageId="87" pageNumber="88">Toliara</collectingRegion>
,
<location id="8E8959D0FFA47307319BFBE7E620FBAE" LSID="urn:lsid:plazi:treatment:03FFBE1DFFA5730932FAFF2DE71FF961:8E8959D0FFA47307319BFBE7E620FBAE" box="[1014,1319,1117,1143]" latitude="-22.88231" longLatPrecision="1" longitude="44.70062" municipality="Province" name="Zombitse Nature Reserve" pageId="87" pageNumber="88" stateProvince="Toliara">Zombitse Nature Reserve</location>
,
<locationDeviation id="9A5B1629FFA473073758FBE7E202FB44" location="Sakaraha" pageId="87" pageNumber="88">
<quantity id="4CAEA2EEFFA473073758FBE7E687FBAE" box="[1333,1408,1117,1143]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="87" pageNumber="88" unit="km" value="16.0">16 km</quantity>
E Sakaraha
</locationDeviation>
,
<quantity id="4CAEA2EEFFA47307337AFB39E26DFB44" box="[279,362,1155,1181]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="87" pageNumber="88" unit="m" value="825.0">
<elevation id="007BE838FFA47307337AFB39E26DFB44" box="[279,362,1155,1181]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="87" pageNumber="88" unit="m" value="825.0">825 m</elevation>
</quantity>
, tropical forest on sand,
<geoCoordinate id="EE6269CCFFA4730730C6FB39E03FFB44" box="[683,824,1155,1181]" degrees="22.88231" direction="south" orientation="latitude" pageId="87" pageNumber="88" precision="1" value="-22.88231">22.88231°S</geoCoordinate>
,
<geoCoordinate id="EE6269CCFFA473073127FB39E0DFFB44" box="[842,984,1155,1181]" degrees="44.70062" direction="east" orientation="longitude" pageId="87" pageNumber="88" precision="1" value="44.70062">44.70062°E</geoCoordinate>
, coll.
<collectorName id="26A36ADDFFA473073641FB39E7EAFB44" box="[1068,1261,1155,1181]" pageId="87" pageNumber="88">E. L. Schlinger</collectorName>
, M. E. Irwin,
<date id="FFE829CBFFA4730732FAFB10E24BFB1D" box="[151,332,1194,1220]" pageId="87" pageNumber="88" value="1999-12-15" valueMax="1999-12-18" valueMin="1999-12-15">
<collectingDate id="EFACD023FFA4730732FAFB10E24BFB1D" box="[151,332,1194,1220]" pageId="87" pageNumber="88" value="1999-12-15" valueMax="1999-12-18" valueMin="1999-12-15">15–18.xii.1999</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFA4730732ABFB6AE65DFB32" box="[198,1370,1232,1259]" pageId="87" pageNumber="88" type="distribution">
<paragraph id="8BE90F0BFFA4730732ABFB6AE65DFB32" blockId="87.[151,1436,1078,2016]" box="[198,1370,1232,1259]" pageId="87" pageNumber="88">
<emphasis id="B922D319FFA4730732ABFB6AE26EFB33" bold="true" box="[198,361,1232,1258]" pageId="87" pageNumber="88">Distribution:</emphasis>
Recorded from the forest Isalo and Zombitse in South-Western
<collectingCountry id="F3414F9BFFA473073635FB6BE7E1FB32" box="[1112,1254,1233,1259]" name="Madagascar" pageId="87" pageNumber="88">Madagascar</collectingCountry>
(Fig. 50).
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFA4730732ABFB4DE7BAF84A" pageId="87" pageNumber="88" type="diagnosis">
<paragraph id="8BE90F0BFFA4730732ABFB4DE0F3F975" blockId="87.[151,1436,1078,2016]" pageId="87" pageNumber="88">
<emphasis id="B922D319FFA4730732ABFB4DE24FFAC8" bold="true" box="[198,328,1271,1297]" pageId="87" pageNumber="88">Diagnosis:</emphasis>
Small species, up to
<quantity id="4CAEA2EEFFA473073021FB42E1A7FACB" box="[588,672,1272,1298]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="87" pageNumber="88" unit="mm" value="25.0">25 mm</quantity>
long. Head, tergites and anal shield shiny blackish, appendages dark olive-green. Surface of tergites shiny, at high magnification leather-like texture becomes visible. Anal shield excavated at hindmost point, in contrast to glabrous tergites covered with numerous minute hairs. First antennomere with groove and
<emphasis id="B922D319FFA473073397FAD2E1DDFA58" box="[506,730,1384,1409]" italics="true" pageId="87" pageNumber="88">sensilla basiconica</emphasis>
(Fig. 59A), disc with four apical cones (Fig. 59B). Movable finger of posterior telopods broad, 2.3 times longer than wide (Fig. 58D), with two triangular membranous lobes and
<emphasis id="B922D319FFA473073360FA08E24FFA12" box="[269,328,1458,1483]" italics="true" pageId="87" pageNumber="88">circa</emphasis>
4 or 5 sclerotized spines (Fig. 58C), at posterior side with
<emphasis id="B922D319FFA473073199FA08E728FA12" box="[1012,1071,1458,1483]" italics="true" pageId="87" pageNumber="88">circa</emphasis>
9 or 10 small, sclerotized teeth (Fig. 58D). Fixed finger long and broad, basally with one membranous lobe and one small spine, distally at inner margin with numerous sclerotized spots (Fig. 58C). Fixed finger reaching 4/5 of movable fingers length. One stridulation rib on each male harp (Fig. 58A). Cavity of anterior telopods third podomere mesally with plateau covered with numerous sclerotized spots (Fig. 58B). Anal shield with two locking carinae, first small, second three times longer than first. Endotergum with one row of large elliptical cuticular impressions and two rows of marginal bristles, reaching up to posterior margin (Fig. 63E).
</paragraph>
<paragraph id="8BE90F0BFFA4730732ABF90DE7BAF84A" blockId="87.[151,1436,1078,2016]" pageId="87" pageNumber="88">
<emphasis id="B922D319FFA4730732ABF90DE296F908" bold="true" box="[198,401,1719,1745]" pageId="87" pageNumber="88">Similar species:</emphasis>
<emphasis id="B922D319FFA4730733F2F902E19FF908" box="[415,664,1720,1745]" italics="true" pageId="87" pageNumber="88">
<taxonomicName id="4C567488FFA4730733F2F902E194F908" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[415,659,1720,1745]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="isalo" status="sp. nov.">Zoosphaerium isalo</taxonomicName>
,
</emphasis>
<emphasis id="B922D319FFA4730730C8F90DE1E8F908" bold="true" box="[677,751,1719,1745]" pageId="87" pageNumber="88">
<taxonomicNameLabel id="A2116E62FFA4730730C8F90DE1E8F908" box="[677,751,1719,1745]" pageId="87" pageNumber="88" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
features a unique combination of characters in the
<taxonomicName id="4C567488FFA4730737E9F902E241F921" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="coquerelianum">
<emphasis id="B922D319FFA4730737E9F902E241F921" italics="true" pageId="87" pageNumber="88">Z. coquerelianum</emphasis>
</taxonomicName>
species-group, like one stridulation rib on the male harp, four apical apical cones on the antenna, a triangular membranous lobe on the basis of posterior telopods second podomere, and the presence of two triangular, membranous lobes on the third podomere of posterior telopods (Figs 58A–F).
<emphasis id="B922D319FFA473073698F896E653F89C" box="[1269,1364,1836,1861]" italics="true" pageId="87" pageNumber="88">
<taxonomicName id="4C567488FFA473073698F896E657F89C" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1269,1360,1836,1861]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="isalo" status="sp. nov.">Z. isalo</taxonomicName>
,
</emphasis>
<emphasis id="B922D319FFA473073736F891E69BF89C" bold="true" box="[1371,1436,1835,1861]" pageId="87" pageNumber="88">
<taxonomicNameLabel id="A2116E62FFA473073736F891E69BF89C" box="[1371,1436,1835,1861]" pageId="87" pageNumber="88" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is very similar to
<emphasis id="B922D319FFA473073305F8E8E2E9F8B2" box="[360,494,1874,1899]" italics="true" pageId="87" pageNumber="88">
<taxonomicName id="4C567488FFA473073305F8E8E2ECF8B2" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[360,491,1874,1899]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="bilobum" status="sp. nov.">Z. bilobum</taxonomicName>
,
</emphasis>
<emphasis id="B922D319FFA47307339AF8E8E13DF8B5" bold="true" box="[503,570,1874,1900]" pageId="87" pageNumber="88">
<taxonomicNameLabel id="A2116E62FFA47307339AF8E8E13DF8B5" box="[503,570,1874,1900]" pageId="87" pageNumber="88" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, however in the latter species overreach the endotergum bristles strongly the tergite margin and the anterior telopod second podomere is not visible in anterior view.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FFA4730932ABF825E71FF961" lastPageId="89" lastPageNumber="90" pageId="87" pageNumber="88" type="description">
<paragraph id="8BE90F0BFFA4730732ABF825E2ACF839" blockId="87.[151,1436,1078,2016]" pageId="87" pageNumber="88">
<emphasis id="B922D319FFA4730732ABF825E267F860" bold="true" box="[198,352,1951,1977]" pageId="87" pageNumber="88">Description:</emphasis>
<emphasis id="B922D319FFA473073306F81AE2FCF860" box="[363,507,1952,1977]" italics="true" pageId="87" pageNumber="88">Body length</emphasis>
: Male
<typeStatus id="54EDB1A9FFA473073038F81AE1C7F863" box="[597,704,1952,1978]" pageId="87" pageNumber="88" type="holotype">holotype</typeStatus>
: length
<quantity id="4CAEA2EEFFA47307314CF81AE07EF863" box="[801,889,1952,1978]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="87" pageNumber="88" unit="mm" value="24.0">24 mm</quantity>
, width of thoracic shield
<quantity id="4CAEA2EEFFA4730736DEF81AE627F863" box="[1203,1312,1952,1978]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="87" pageNumber="88" unit="mm" value="12.7">12.7 mm</quantity>
, height of thoracic shield
<quantity id="4CAEA2EEFFA473073327F87CE2A0F839" box="[330,423,1990,2016]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="87" pageNumber="88" unit="mm" value="7.2">7.2 mm</quantity>
.
</paragraph>
<paragraph id="8BE90F0BFFAB730832AAFF22E133FF01" blockId="88.[151,1435,152,255]" pageId="88" pageNumber="89">
<emphasis id="B922D319FFAB730832AAFF22E256FF68" box="[199,337,152,177]" italics="true" pageId="88" pageNumber="89">Coloration:</emphasis>
Faded in alcohol. Head, collum, thoracic shield, tergites and anal shield uniformly blackish, antennae and legs dark olive green.
</paragraph>
<paragraph id="8BE90F0BFFAB730832AAFF5FE116FF26" blockId="88.[151,1435,152,255]" box="[199,529,229,255]" pageId="88" pageNumber="89">
<emphasis id="B922D319FFAB730832AAFF5FE217FF27" box="[199,272,229,254]" italics="true" pageId="88" pageNumber="89">Head:</emphasis>
Eyes with&gt;75 ocelli.
</paragraph>
<paragraph id="8BE90F0BFFAB730832FAF82CE624F82B" blockId="88.[151,1436,1942,2035]" pageId="88" pageNumber="89">
<emphasis id="B922D319FFAB730832FAF82CE237F877" bold="true" box="[151,304,1942,1966]" pageId="88" pageNumber="89">FIGURE 060</emphasis>
.
<taxonomicName id="4C567488FFAB7308332DF82CE110F874" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[320,535,1942,1965]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="isalo" status="sp. nov.">
<emphasis id="B922D319FFAB7308332DF82CE110F874" box="[320,535,1942,1965]" italics="true" pageId="88" pageNumber="89">Zoosphaerium isalo</emphasis>
</taxonomicName>
<emphasis id="B922D319FFAB7308304DF82CE159F877" bold="true" box="[544,606,1942,1966]" pageId="88" pageNumber="89">
<taxonomicNameLabel id="A2116E62FFAB7308304DF82CE159F877" box="[544,606,1942,1966]" pageId="88" pageNumber="89" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FFAB73083003F82CE1CAF874" box="[622,717,1942,1965]" pageId="88" pageNumber="89" type="paratype">paratype</typeStatus>
, SEM,
<emphasis id="B922D319FFAB7308314CF82CE033F877" bold="true" box="[801,820,1942,1966]" pageId="88" pageNumber="89">A</emphasis>
: gnathochilarium, ventral view;
<emphasis id="B922D319FFAB730836F6F82CE7AAF877" bold="true" box="[1179,1197,1942,1966]" pageId="88" pageNumber="89">B</emphasis>
: left mandible, mesal view. Abbreviations:
<emphasis id="B922D319FFAB73083312F802E2A1F809" bold="true" box="[383,422,1976,2000]" pageId="88" pageNumber="89">3iT</emphasis>
= 3 inner teeth;
<emphasis id="B922D319FFAB73083033F802E179F809" bold="true" box="[606,638,1976,2000]" pageId="88" pageNumber="89">Ca</emphasis>
= rudiments of cardines;
<emphasis id="B922D319FFAB730831F7F802E0BCF809" bold="true" box="[922,955,1976,2000]" pageId="88" pageNumber="89">Co</emphasis>
= condylus;
<emphasis id="B922D319FFAB73083626F802E76EF809" bold="true" box="[1099,1129,1976,2000]" pageId="88" pageNumber="89">eT</emphasis>
= external tooth;
<emphasis id="B922D319FFAB73083740F802E642F809" bold="true" box="[1325,1349,1976,2000]" pageId="88" pageNumber="89">iP</emphasis>
= inner palpus;
<emphasis id="B922D319FFAB73083287F861E20AF82A" bold="true" box="[234,269,2011,2035]" pageId="88" pageNumber="89">LL</emphasis>
= fused lamella linguales;
<emphasis id="B922D319FFAB73083043F861E154F82A" bold="true" box="[558,595,2011,2035]" pageId="88" pageNumber="89">Me</emphasis>
= mentum;
<emphasis id="B922D319FFAB730830B8F861E1FDF82A" bold="true" box="[725,762,2011,2035]" pageId="88" pageNumber="89">mp</emphasis>
= molar plate;
<emphasis id="B922D319FFAB730831F0F861E0B4F82A" bold="true" box="[925,947,2011,2035]" pageId="88" pageNumber="89">St</emphasis>
= stipites;
<emphasis id="B922D319FFAB73083647F861E74DF82A" bold="true" box="[1066,1098,2011,2035]" pageId="88" pageNumber="89">pL</emphasis>
= pectinate lamella.
</paragraph>
<paragraph id="8BE90F0BFFAA730932AAFF22E60EFF26" blockId="89.[151,1436,152,1720]" pageId="89" pageNumber="90">
<emphasis id="B922D319FFAA730932AAFF22E238FF68" box="[199,319,152,177]" italics="true" pageId="89" pageNumber="90">Antennae:</emphasis>
Length of antennomeres: 1&gt;2&gt;3&gt;4&lt;5&lt;6, sixth antennomere longest, cylindrical, disc with four sensory cones (Fig. 59B). First antennomere remarkably broader than others, short, with groove and 1 or 2 rows of
<emphasis id="B922D319FFAA7309329AFF5FE2D0FF27" box="[247,471,229,254]" italics="true" pageId="89" pageNumber="90">sensilla basiconica</emphasis>
(Fig. 59A). Sclerotized teeth at base of antennomeres 1–4 (Fig. 59B).
</paragraph>
<paragraph id="8BE90F0BFFAA730932AAFEB6E0CEFE95" blockId="89.[151,1436,152,1720]" pageId="89" pageNumber="90">
<emphasis id="B922D319FFAA730932AAFEB6E247FEFC" box="[199,320,268,293]" italics="true" pageId="89" pageNumber="90">Mandible:</emphasis>
Five rows of pectinate lamellae, number of teeth declining apically to proximally (Fig. 60B). Condylus with strongly developed step near apical margin (Fig. 60B).
</paragraph>
<paragraph id="8BE90F0BFFAA730932AAFEE3E745FE43" blockId="89.[151,1436,152,1720]" pageId="89" pageNumber="90">
<emphasis id="B922D319FFAA730932AAFEE3E296FEAB" box="[199,401,345,370]" italics="true" pageId="89" pageNumber="90">Gnathochilarium</emphasis>
: Ventral side with numerous, long setae (Fig. 60A). Lateral of palpi four sensory cones, located together. Depression on posterior side of palpi with some sensory cones.
</paragraph>
<paragraph id="8BE90F0BFFAA730932AAFE1DE09BFE19" blockId="89.[151,1436,152,1720]" box="[199,924,422,448]" pageId="89" pageNumber="90">
<emphasis id="B922D319FFAA730932AAFE1DE220FE19" box="[199,295,423,448]" italics="true" pageId="89" pageNumber="90">Collum:</emphasis>
Only at margins with isolated hair of medium length.
</paragraph>
<paragraph id="8BE90F0BFFAA730932AAFE77E03DFE3E" blockId="89.[151,1436,152,1720]" box="[199,826,461,487]" pageId="89" pageNumber="90">
<emphasis id="B922D319FFAA730932AAFE77E279FE3F" box="[199,382,461,486]" italics="true" pageId="89" pageNumber="90">Thoracic shield</emphasis>
shaped like in other species of genus.
</paragraph>
<paragraph id="8BE90F0BFFAA730932AAFE4EE12AFDED" blockId="89.[151,1436,152,1720]" pageId="89" pageNumber="90">
<emphasis id="B922D319FFAA730932AAFE4EE224FDD4" box="[199,291,500,525]" italics="true" pageId="89" pageNumber="90">Tergites</emphasis>
: Surface shiny, at high magnification (90x) leather-like texture becomes visible. Paratergites straight, not projecting posteriorly.
</paragraph>
<paragraph id="8BE90F0BFFAA730932AAFDFBE210FD71" blockId="89.[151,1436,152,1720]" pageId="89" pageNumber="90">
<emphasis id="B922D319FFAA730932AAFDFBE267FD83" box="[199,352,577,602]" italics="true" pageId="89" pageNumber="90">Endotergum:</emphasis>
Inner area with thin conical spines and long hair. Single row of few scattered, large, elliptical cuticular impressions (Fig. 63E). Two rows of marginal bristles, slightly protruding to tergite margin (Fig. 63E).
</paragraph>
<paragraph id="8BE90F0BFFAA730932FAFD0FE7C4FCC3" blockId="89.[151,1436,152,1720]" pageId="89" pageNumber="90">
<emphasis id="B922D319FFAA730932FAFD0FE220FD17" box="[151,295,693,718]" italics="true" pageId="89" pageNumber="90">Anal shield:</emphasis>
Tapered at hindmost point. Covered with numerous hairs, each located in minute pit. Underside carries two well-developed black locking carinae, anterior one small, but well developed. Posterior carina three times longer than anterior one, straight. Carinae located closely to anal shield margin.
</paragraph>
<paragraph id="8BE90F0BFFAA730932AAFC9FE264FC6A" blockId="89.[151,1436,152,1720]" pageId="89" pageNumber="90">
<emphasis id="B922D319FFAA730932AAFC9FE20FFCE7" box="[199,264,805,830]" italics="true" pageId="89" pageNumber="90">Legs:</emphasis>
Tarsi of first pair of legs with three or four, second with four or five and third with six to eight ventral spines. First two leg pairs with only weakly curved claws and without apical spine. Tarsi of legs 3–21 with curved claws, 10–12 ventral spines and one apical spine. In 9th leg femur 1.9 times, tarsi 4.25 times longer than wide.
</paragraph>
<paragraph id="8BE90F0BFFAA730932AAFC7AE04DFC03" blockId="89.[151,1436,152,1720]" box="[199,842,960,986]" pageId="89" pageNumber="90">
<emphasis id="B922D319FFAA730932AAFC7AE28BFC00" box="[199,396,960,985]" italics="true" pageId="89" pageNumber="90">Stigmatic plates:</emphasis>
First lobe triangular, with pointed tip.
</paragraph>
<paragraph id="8BE90F0BFFAA730932AAFC5DE297FBD9" blockId="89.[151,1436,152,1720]" box="[199,400,999,1024]" pageId="89" pageNumber="90">
<emphasis id="B922D319FFAA730932AAFC5DE297FBD9" box="[199,400,999,1024]" italics="true" pageId="89" pageNumber="90">Female unknown</emphasis>
</paragraph>
<paragraph id="8BE90F0BFFAA730932AAFBB7E771FBFE" blockId="89.[151,1436,152,1720]" box="[199,1142,1037,1063]" pageId="89" pageNumber="90">
<emphasis id="B922D319FFAA730932AAFBB7E2DDFBFF" box="[199,474,1037,1062]" italics="true" pageId="89" pageNumber="90">Male sexual characters</emphasis>
: Male gonopore covered by sclerotized, undivided plate.
</paragraph>
<paragraph id="8BE90F0BFFAA730932AAFB8EE68AFB31" blockId="89.[151,1436,152,1720]" pageId="89" pageNumber="90">
<emphasis id="B922D319FFAA730932AAFB8EE29EFB94" box="[199,409,1076,1101]" italics="true" pageId="89" pageNumber="90">Anterior telopods</emphasis>
(Figs 58A, B): Width of first podomere slightly larger than height. Harp with one straight and well-developed stridulation rib, located basally at lateral part of first podomere. Second podomere process lobe-like and little curved with rounded edges, protruding to 2/3 of third podomere height, visible laterally in anterior view, apically with sclerotized spots and one spine. Third podomere elongated, cavity mesally with several sclerotized spots and two sclerotized spines. One sclerotized spot located closely to tip.
</paragraph>
<paragraph id="8BE90F0BFFAA730932AAFB4FE293F9C7" blockId="89.[151,1436,152,1720]" pageId="89" pageNumber="90">
<emphasis id="B922D319FFAA730932AAFB4FE2A3FAD7" box="[199,420,1269,1294]" italics="true" pageId="89" pageNumber="90">Posterior telopods</emphasis>
(Figs 58C, D): Movable finger of chela thick, 2.3 times longer than wide, weakly curved, apically slightly tapering. Hollowed-out inner margin with two triangular membranous lobes and 4 or 5 sclerotized spines, posterior aspect with
<emphasis id="B922D319FFAA730930E3FAF9E1CEFA85" box="[654,713,1347,1372]" italics="true" pageId="89" pageNumber="90">circa</emphasis>
9 or 10 small, sclerotized teeth. Fixed finger wide, but more slender and shorter than movable finger. Tip of fixed finger almost straight, only weakly curved; anterior side basally with one short sclerotized spine, on tip with numerous sclerotized spots. Fixed finger on posterior side covered with numerous, on anterior side with very few long hairs. Third podomere on both sides almost glabrous. Inner horns of syncoxite apically curved posteriorly. Tips pointed, whole horns covered with numerous short hairs.
</paragraph>
<paragraph id="8BE90F0BFFAA730932AAF990E71FF961" blockId="89.[151,1436,152,1720]" pageId="89" pageNumber="90">
<emphasis id="B922D319FFAA730932AAF990E2EBF99D" bold="true" box="[199,492,1578,1604]" pageId="89" pageNumber="90">Intraspecific variation:</emphasis>
The specimen from Zombitse is very similar to the
<typeStatus id="54EDB1A9FFAA73093614F991E7E3F99C" box="[1145,1252,1579,1605]" pageId="89" pageNumber="90" type="holotype">holotype</typeStatus>
. Especially the posterior telopods (Fig. 58E, F) are almost identical with those of the
<typeStatus id="54EDB1A9FFAA730931A6F9EBE736F9B2" box="[971,1073,1617,1643]" pageId="89" pageNumber="90" type="holotype">holotype</typeStatus>
(Fig. 58C, D). However, more specimens from both, the Isalo type locality and from Zombitse are necessary to evaluate if the specimens from Zombitse belong to
<emphasis id="B922D319FFAA730933AFF925E125F961" box="[450,546,1695,1720]" italics="true" pageId="89" pageNumber="90">
<taxonomicName id="4C567488FFAA730933AFF925E11BF961" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[450,540,1695,1720]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="isalo" status="sp. nov.">Z. isalo</taxonomicName>
,
</emphasis>
<emphasis id="B922D319FFAA73093044F924E16CF961" bold="true" box="[553,619,1694,1720]" pageId="89" pageNumber="90">
<taxonomicNameLabel id="A2116E62FFAA73093044F924E16CF961" box="[553,619,1694,1720]" pageId="89" pageNumber="90" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
or represent an undescribed species.
</paragraph>
</subSubSection>
</treatment>
</document>
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