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<mods:title id="32C30FE264DAB2B084FB4B2E13A3AE3A">The Early Miocene Bats (Chiroptera, Mammalia) From The Karstic Sites Of Erkertshofen And Petersbuch 2 (Southern Germany)</mods:title>
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<mods:namePart id="6B1C6EB46CCC5BF48BD98F0875D46829">Rosina, Valentina V.</mods:namePart>
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<paragraph id="8BE93E41F323FFBAD3D96296FDEB1B7A" blockId="9.[218,632,503,528]" box="[218,632,503,528]" pageId="9" pageNumber="421">
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<taxonomicName id="4C5645C2F323FFBAD3D96296FDEB1B7A" authority="DEPERET, 1892" authorityName="DEPERET" authorityYear="1892" box="[218,632,503,528]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="421" phylum="Chordata" rank="species" species="grivensis">
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<emphasis id="B922E253F323FFBAD3D96296FE421B7A" bold="true" box="[218,465,503,527]" italics="true" pageId="9" pageNumber="421">Rhinolophus grivensis</emphasis>
DEPÉRET, 1892
</emphasis>
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<paragraph id="8BE93E41F323FFBAD2506176FE6D1B5A" blockId="9.[339,510,535,559]" box="[339,510,535,559]" pageId="9" pageNumber="421">
<figureCitation id="136D22C4F323FFBAD2506176FE6D1B5A" box="[339,510,535,559]" captionStart="Text-fig" captionStartId="5.[126,207,1558,1579]" captionTargetBox="[126,1439,151,1538]" captionTargetId="figure-137@5.[126,1439,144,1546]" captionTargetPageId="5" captionText="Text-fig. 2. ae Rhinolophus aff. lemanensis: a PCMRCh65, left mxl with P4M3, Petersbuch 2, ventral view; b PCMRCh79 right mnd with C inf., Petersbuch 2, lingual view; c PCMRCh66, right C sup., Petersbuch 2, lingual view; d BSP 1976 XXII 5515, left mnd with m23, Petersbuch 2, occlusal view; e BSP 1974 XIV 1139, left mnd with p4m1; Erkertshofen 2, occlusal view; f, i, km R. dehmi: f BSP 1974 XIV 1118, right C sup., Erkertshofen 2, lingual view; i BSP 1976 XXII 5507, left mnd with p4m1, Petersbuch 2, occlusal view; k PCMRCh32, right mxl with C sup., P4M1, Petersbuch 2, ventral view; l PCMRCh37, left mnd with m12, Petersbuch 2, occlusal view; m PCMRCh89, right mnd with C inf., Petersbuch 2, lateral view; gh R. cf. delphinensis: g BSP 1976 XXII 5522, right mxl with P4, Petersbuch 2, ventral view; h BSP 1962 XIX 4153, left mnd with m13, Erkertshofen 1, occlusal view; j, n, o R. grivensis: j PCMRCh43, left C sup., Petersbuch 2, lingual view; n BSP 1976 XXII 5543, left mxl, Petersbuch 2, ventral view; o BSP 1976 XXII 5548, left mnd with m23, Petersbuch 2, occlusal view." figureDoi="http://doi.org/10.5281/zenodo.5383145" httpUri="https://zenodo.org/record/5383145/files/figure.png" pageId="9" pageNumber="421">Text-fig. 2j, no</figureCitation>
</paragraph>
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<subSubSection id="C34C6DCAF323FFBAD3936126FE361C48" pageId="9" pageNumber="421" type="description">
<paragraph id="8BE93E41F323FFBAD3936126FF051B0A" blockId="9.[106,745,583,927]" pageId="9" pageNumber="421">M a t e r i a l. Erkertshofen 1: BSP 1962 XIX 4194, left M2.</paragraph>
<paragraph id="8BE93E41F323FFBAD39361E6FD011BEB" blockId="9.[106,745,583,927]" box="[144,658,647,671]" pageId="9" pageNumber="421">Erkertshofen 2: BSP 1974 XIV 1114, left C sup.</paragraph>
<paragraph id="8BE93E41F323FFBAD39361C6FEF91AEB" blockId="9.[106,745,583,927]" pageId="9" pageNumber="421">Petersbuch 2: BSP 1976 XXII 5543, left mxl with P4; PCMRCh55a, right m1; PCMRCh56a, left m2; PCMRCh56b, right m2; PCMRCh57c, right m3; PCMRCh48, right M3; PCMRCh49a, left M1 (damaged); PCMRCh49b, left M1; PCMRCh49c, left M2; PCMRCh49d, right M2; PCMRCh27, left P4; PCMRCh29, right C sup.; PCMRCh30, right C sup.; PCMRCh31, left C sup.; PCMRCh42, right C sup.; PCMRCh43, left C sup.</paragraph>
<paragraph id="8BE93E41F323FFBAD39360D7FE7D1ABB" blockId="9.[144,494,950,974]" box="[144,494,950,974]" pageId="9" pageNumber="421">
M e a s u r e m e n t s. See
<tableCitation id="C6D40BFAF323FFBAD2A660D7FE7D1ABB" box="[421,494,950,974]" captionStart="Table 4" captionStartId="7.[125,181,152,173]" captionText="Table 4. Small Rhinolophus species from Petersbuch 2 (P 2), Erkertshofen 1 (E 1) and Erkertshofen 2 (E 2), sample statistics of the teeth and jaws." pageId="9" pageNumber="421">Tab. 4.</tableCitation>
</paragraph>
<paragraph id="8BE93E41F323FFBAD3936084FE361C48" blockId="9.[106,746,997,1341]" pageId="9" pageNumber="421">
D e s c r i p t i o n a n d c o m p a r i s o n.The preserved fossils share all the morphological traits of
<taxonomicName id="4C5645C2F323FFBAD1616764FF281D48" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="421" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B922E253F323FFBAD1616764FD7A1D68" box="[610,745,1029,1053]" italics="true" pageId="9" pageNumber="421">Rhinolophus</emphasis>
species
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: the upper canine is semilunar in occlusal view with a well-developed cingulum (
<figureCitation id="136D22C4F323FFBAD29D6724FD881D28" box="[414,539,1093,1117]" captionStart="Text-fig" captionStartId="5.[126,207,1558,1579]" captionTargetBox="[126,1439,151,1538]" captionTargetId="figure-137@5.[126,1439,144,1546]" captionTargetPageId="5" captionText="Text-fig. 2. ae Rhinolophus aff. lemanensis: a PCMRCh65, left mxl with P4M3, Petersbuch 2, ventral view; b PCMRCh79 right mnd with C inf., Petersbuch 2, lingual view; c PCMRCh66, right C sup., Petersbuch 2, lingual view; d BSP 1976 XXII 5515, left mnd with m23, Petersbuch 2, occlusal view; e BSP 1974 XIV 1139, left mnd with p4m1; Erkertshofen 2, occlusal view; f, i, km R. dehmi: f BSP 1974 XIV 1118, right C sup., Erkertshofen 2, lingual view; i BSP 1976 XXII 5507, left mnd with p4m1, Petersbuch 2, occlusal view; k PCMRCh32, right mxl with C sup., P4M1, Petersbuch 2, ventral view; l PCMRCh37, left mnd with m12, Petersbuch 2, occlusal view; m PCMRCh89, right mnd with C inf., Petersbuch 2, lateral view; gh R. cf. delphinensis: g BSP 1976 XXII 5522, right mxl with P4, Petersbuch 2, ventral view; h BSP 1962 XIX 4153, left mnd with m13, Erkertshofen 1, occlusal view; j, n, o R. grivensis: j PCMRCh43, left C sup., Petersbuch 2, lingual view; n BSP 1976 XXII 5543, left mxl, Petersbuch 2, ventral view; o BSP 1976 XXII 5548, left mnd with m23, Petersbuch 2, occlusal view." figureDoi="http://doi.org/10.5281/zenodo.5383145" httpUri="https://zenodo.org/record/5383145/files/figure.png" pageId="9" pageNumber="421">Text-fig. 2j</figureCitation>
), p3 was displaced buccally from the midline of the tooth-row, and the m3 talonid is only slightly smaller than the trigonid (
<figureCitation id="136D22C4F323FFBAD18F67E4FF1B1DC8" captionStart="Text-fig" captionStartId="5.[126,207,1558,1579]" captionTargetBox="[126,1439,151,1538]" captionTargetId="figure-137@5.[126,1439,144,1546]" captionTargetPageId="5" captionText="Text-fig. 2. ae Rhinolophus aff. lemanensis: a PCMRCh65, left mxl with P4M3, Petersbuch 2, ventral view; b PCMRCh79 right mnd with C inf., Petersbuch 2, lingual view; c PCMRCh66, right C sup., Petersbuch 2, lingual view; d BSP 1976 XXII 5515, left mnd with m23, Petersbuch 2, occlusal view; e BSP 1974 XIV 1139, left mnd with p4m1; Erkertshofen 2, occlusal view; f, i, km R. dehmi: f BSP 1974 XIV 1118, right C sup., Erkertshofen 2, lingual view; i BSP 1976 XXII 5507, left mnd with p4m1, Petersbuch 2, occlusal view; k PCMRCh32, right mxl with C sup., P4M1, Petersbuch 2, ventral view; l PCMRCh37, left mnd with m12, Petersbuch 2, occlusal view; m PCMRCh89, right mnd with C inf., Petersbuch 2, lateral view; gh R. cf. delphinensis: g BSP 1976 XXII 5522, right mxl with P4, Petersbuch 2, ventral view; h BSP 1962 XIX 4153, left mnd with m13, Erkertshofen 1, occlusal view; j, n, o R. grivensis: j PCMRCh43, left C sup., Petersbuch 2, lingual view; n BSP 1976 XXII 5543, left mxl, Petersbuch 2, ventral view; o BSP 1976 XXII 5548, left mnd with m23, Petersbuch 2, occlusal view." figureDoi="http://doi.org/10.5281/zenodo.5383145" httpUri="https://zenodo.org/record/5383145/files/figure.png" pageId="9" pageNumber="421">Text-fig. 2o</figureCitation>
). However, there is a significant space between the canine and P
<quantity id="4CAE93A4F323FFBAD3A767A4FF581DA8" box="[164,203,1221,1245]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="9" pageNumber="421" unit="in" value="4.0">4 in</quantity>
the maxillary (
<figureCitation id="136D22C4F323FFBAD26467A4FE761DA8" box="[359,485,1221,1245]" captionStart="Text-fig" captionStartId="5.[126,207,1558,1579]" captionTargetBox="[126,1439,151,1538]" captionTargetId="figure-137@5.[126,1439,144,1546]" captionTargetPageId="5" captionText="Text-fig. 2. ae Rhinolophus aff. lemanensis: a PCMRCh65, left mxl with P4M3, Petersbuch 2, ventral view; b PCMRCh79 right mnd with C inf., Petersbuch 2, lingual view; c PCMRCh66, right C sup., Petersbuch 2, lingual view; d BSP 1976 XXII 5515, left mnd with m23, Petersbuch 2, occlusal view; e BSP 1974 XIV 1139, left mnd with p4m1; Erkertshofen 2, occlusal view; f, i, km R. dehmi: f BSP 1974 XIV 1118, right C sup., Erkertshofen 2, lingual view; i BSP 1976 XXII 5507, left mnd with p4m1, Petersbuch 2, occlusal view; k PCMRCh32, right mxl with C sup., P4M1, Petersbuch 2, ventral view; l PCMRCh37, left mnd with m12, Petersbuch 2, occlusal view; m PCMRCh89, right mnd with C inf., Petersbuch 2, lateral view; gh R. cf. delphinensis: g BSP 1976 XXII 5522, right mxl with P4, Petersbuch 2, ventral view; h BSP 1962 XIX 4153, left mnd with m13, Erkertshofen 1, occlusal view; j, n, o R. grivensis: j PCMRCh43, left C sup., Petersbuch 2, lingual view; n BSP 1976 XXII 5543, left mxl, Petersbuch 2, ventral view; o BSP 1976 XXII 5548, left mnd with m23, Petersbuch 2, occlusal view." figureDoi="http://doi.org/10.5281/zenodo.5383145" httpUri="https://zenodo.org/record/5383145/files/figure.png" pageId="9" pageNumber="421">Text-fig. 2n</figureCitation>
). This is the smallest and most gracile
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<emphasis id="B922E253F323FFBAD3F06784FEE91D88" box="[243,378,1253,1277]" italics="true" pageId="9" pageNumber="421">Rhinolophus</emphasis>
species
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from the studied localities (
<tableCitation id="C6D40BFAF323FFBAD3726664FF291C68" box="[113,186,1285,1309]" captionStart="Table 4" captionStartId="7.[125,181,152,173]" captionText="Table 4. Small Rhinolophus species from Petersbuch 2 (P 2), Erkertshofen 1 (E 1) and Erkertshofen 2 (E 2), sample statistics of the teeth and jaws." pageId="9" pageNumber="421">Tab. 4</tableCitation>
) and morphologically corresponds to
<taxonomicName id="4C5645C2F323FFBAD1656667FD7B1C68" authorityName="DEPERET" authorityYear="1892" box="[614,744,1286,1309]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="421" phylum="Chordata" rank="species" species="grivensis">
<emphasis id="B922E253F323FFBAD1656667FD7B1C68" box="[614,744,1286,1309]" italics="true" pageId="9" pageNumber="421">R. grivensis</emphasis>
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(e.g.
<bibRefCitation id="EFC743B0F323FFBAD3A36644FE0A1C48" author="Ziegler, R." box="[160,409,1317,1341]" pageId="9" pageNumber="421" pagination="447 - 490" refId="ref24261" refString="Ziegler, R. (2003): Bats (Chiroptera, Mammalia) from Mid- dle Miocene karstic fissure fillings of Petersbuch near Eichstatt, Southern Franconian Alb (Bavaria). - Geo- bios, 36: 447 - 490. https: // doi. org / 10.1016 / S 0016 - 6995 (03) 00043 - 3" type="journal article" year="2003">Ziegler 2003: 456459</bibRefCitation>
).
</paragraph>
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