822 lines
97 KiB
XML
822 lines
97 KiB
XML
<document id="735AE17FE08F6CC6A70F92D50D6BF025" ID-DOI="10.5281/zenodo.274877" ID-GBIF-Dataset="cf4ac737-234c-4213-b723-04e7157bfa63" ID-ISSN="1175-5326" ID-Zenodo-Dep="274877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461077221482" checkinUser="plazi" docAuthor="Branch, William R. & Bayliss, Julian" docDate="2009" docId="03ED5B5FFFE7965BE7D427EDFA90779D" docLanguage="en" docName="zt02113p054.pdf" docOrigin="Zootaxa 2113" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Atheris mabuensis Branch & Bayliss, 2009, n. sp." docType="treatment" docVersion="7" lastPageNumber="49" masterDocId="FFD42327FFE49653E743277AFFB67214" masterDocTitle="A new species of Atheris (Serpentes: Viperidae) from northern Mozambique" masterLastPageNumber="54" masterPageNumber="41" pageNumber="44" updateTime="1698586653946" updateUser="plazi">
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<mods:mods id="0D98BACD0EABFD33D4206B5FC6205A7C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="769DF792A93C881323D38238761A277E">
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<mods:title id="415EF561DA333C0FDA299905560B1D04">A new species of Atheris (Serpentes: Viperidae) from northern Mozambique</mods:title>
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</mods:titleInfo>
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<mods:name id="52568CF409558D8FF29C8F3440C689BA" type="personal">
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<mods:role id="2DED46845F666E0A55E13733E9D63377">
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<mods:roleTerm id="6065315940044D877A7A6844B25C15C6">Author</mods:roleTerm>
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<mods:namePart id="2BD81EE4AFCEA2667D81716973A9FDF4">Branch, William R.</mods:namePart>
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</mods:name>
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<mods:name id="D442EC0DD9E747F9601F60316815AA3A" type="personal">
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<mods:roleTerm id="C460CECB958E8BE13C6EF7B4B222A3CE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D44B53914F65AF3D1672575A2F36BBDD">Bayliss, Julian</mods:namePart>
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<mods:typeOfResource id="9C093A01A78F408B4EB84AB6332D4212">text</mods:typeOfResource>
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<mods:title id="BFEF7CFA9E1937951451450387742FE7">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="45B3704DCCA678CE4A1A2BD6C816A958">
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<mods:date id="BDFB8ACF9BD00D398F47A6B96DE92CCD">2009</mods:date>
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<mods:detail id="7192F37EF9D658855E1AD80320C575E9" type="volume">
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<mods:number id="3A4619C8364FB85A5B25CF1FC140B728">2113</mods:number>
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<mods:classification id="E0059F6185A8B5D6532FBEBA2B2CD734">journal article</mods:classification>
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<mods:identifier id="B087ED9B3A29E0A15C96D1FE9A8914AC" type="DOI">10.5281/zenodo.274877</mods:identifier>
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<treatment id="03ED5B5FFFE7965BE7D427EDFA90779D" ID-DOI="http://doi.org/10.5281/zenodo.5691535" ID-GBIF-Taxon="119617789" ID-Zenodo-Dep="5691535" LSID="urn:lsid:plazi:treatment:03ED5B5FFFE7965BE7D427EDFA90779D" httpUri="http://treatment.plazi.org/id/03ED5B5FFFE7965BE7D427EDFA90779D" lastPageId="8" lastPageNumber="49" pageId="3" pageNumber="44">
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<subSubSection id="C35EB9C2FFE79650E7D427EDFEA172C0" pageId="3" pageNumber="44" type="nomenclature">
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<paragraph id="8BFBEA49FFE79650E7D427EDFE7672A5" blockId="3.[151,448,151,212]" box="[151,448,151,177]" pageId="3" pageNumber="44">
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<heading id="D0B35D25FFE79650E7D427EDFE7672A5" bold="true" box="[151,448,151,177]" fontSize="11" level="1" pageId="3" pageNumber="44" reason="1">
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<emphasis id="B930365BFFE79650E7D427EDFE7672A5" bold="true" box="[151,448,151,177]" pageId="3" pageNumber="44">
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<taxonomicName id="4C4491CAFFE79650E7D427EDFEC172A5" box="[151,375,151,177]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="mabuensis" status="sp. nov.">
|
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<emphasis id="B930365BFFE79650E7D427EDFEC172A5" bold="true" box="[151,375,151,177]" italics="true" pageId="3" pageNumber="44">Atheris mabuensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2038B20FFE79650E63D27EDFE7672A5" box="[382,448,151,177]" pageId="3" pageNumber="44" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
<paragraph id="8BFBEA49FFE79650E7D427C0FEA172C0" blockId="3.[151,448,151,212]" box="[151,279,186,212]" pageId="3" pageNumber="44">
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(
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||
<figureCitation id="137FF6CCFFE79650E7DC27C0FEB972C0" box="[159,271,186,212]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="4.[151,255,1976,2000]" captionStartId-1="5.[151,257,1630,1654]" captionStartId-2="6.[151,259,1959,1983]" captionStartId-3="8.[151,255,982,1006]" captionStartId-4="9.[151,258,1869,1893]" captionStartId-5="10.[151,257,1239,1263]" captionTargetBox-0="[151,1434,451,1949]" captionTargetBox-1="[155,1435,666,1617]" captionTargetBox-2="[151,1435,1078,1932]" captionTargetBox-3="[151,1435,194,957]" captionTargetBox-4="[152,1433,194,1846]" captionTargetBox-5="[151,1435,377,1215]" captionTargetId-0="figure@4.[151,1436,1095,1949]" captionTargetId-1="figure@5.[151,1436,654,1618]" captionTargetId-2="figure@6.[151,1436,1078,1932]" captionTargetId-3="figure@8.[151,1436,194,958]" captionTargetId-4="figure@9.[152,1435,194,1846]" captionTargetId-5="figure@10.[151,1436,377,1215]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionText-0="FIGURE 2. Right (top) and left (bottom) lateral views of the head of Atheris mabuensis n. sp. (Holotype PEM R 17901, Mount Mabu) before fixation. Note swollen gold keel tips on the head and body scales." captionText-1="FIGURE 3. Dorsal (left) and ventral (right) views of the head of Atheris mabuensis n. sp. (Holotype PEM R 17901, Mount Mabu) before fixation. Note the poorly-defined dark V-patch on crown of head" captionText-2="FIGURE 4. Atheris mabuensis n. sp. (Holotype PEM R 17901, Mount Mabu) in life showing habitus and general colouraion. Note: Retention of yellow tail tip in adult." captionText-3="FIGURE 5. Midbody, right lateral view of Atheris mabuensis n. sp. (Holotype PEM R 17901 Mount Mabu): note oblique orientation of lateral scale rows 1 – 5, and the progressive asymmetry in the shape and lack of serrated keels on these scales." captionText-4="FIGURE 6. Atheris mabuensis n. sp. (PEM R 17910): Top - small juvenile (<150 mm TL) photographed in situ on capture (25 January 2006, Julian Bayliss); note drab colouration imparting camouflage in habitat among leaf litter on forest floor. Bottom – same specimen (photographed 19 Nov 2008, after 34 months in captivity); note retention of juvenile colouration, particularly yellow tail tip, and development of gold tips to the keels on the head and forebody scales." captionText-5="FIGURE 7. Right lateral view of head and forebody Atheris mabuensis n. sp. (PEM R 17904, Mount Namuli): note reduction of gold tipped scales on head and body." httpUri-0="https://zenodo.org/record/274879/files/figure.png" httpUri-1="https://zenodo.org/record/274880/files/figure.png" httpUri-2="https://zenodo.org/record/274881/files/figure.png" httpUri-3="https://zenodo.org/record/274882/files/figure.png" httpUri-4="https://zenodo.org/record/274883/files/figure.png" httpUri-5="https://zenodo.org/record/274884/files/figure.png" pageId="3" pageNumber="44">Figs. 2–7</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C35EB9C2FFE79650E7D4267DFC79710C" pageId="3" pageNumber="44" type="materials_examined">
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<paragraph id="8BFBEA49FFE79650E7D4267DFF4670DF" blockId="3.[151,1437,263,2035]" pageId="3" pageNumber="44">
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<emphasis id="B930365BFFE79650E7D4267DFED77335" bold="true" box="[151,353,263,289]" pageId="3" pageNumber="44">
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||
<typeStatus id="54FF54EBFFE79650E7D4267DFF617335" box="[151,215,263,289]" pageId="3" pageNumber="44">Type</typeStatus>
|
||
Material:
|
||
</emphasis>
|
||
<emphasis id="B930365BFFE79650E62F2672FE5A7335" box="[364,492,264,289]" italics="true" pageId="3" pageNumber="44">
|
||
<typeStatus id="54FF54EBFFE79650E62F2672FE537335" box="[364,485,264,289]" pageId="3" pageNumber="44" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
adult male,
|
||
<collectionCode id="ED55728CFFE79650E5CC2672FD797336" box="[655,719,264,290]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="3" pageNumber="44">PEM</collectionCode>
|
||
R17901.
|
||
<emphasis id="B930365BFFE79650E40E2672FC447335" box="[845,1010,264,289]" italics="true" pageId="3" pageNumber="44">
|
||
<typeStatus id="54FF54EBFFE79650E40E2672FC3E7335" box="[845,904,264,289]" pageId="3" pageNumber="44">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
: main forest camp, Mount Mabu (
|
||
<geoCoordinate id="EE708C8EFFE79650E7E22654FE81735C" box="[161,311,302,328]" direction="south" orientation="latitude" pageId="3" pageNumber="44" precision="15" value="-16.286667">16°17’12”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE708C8EFFE79650E6062654FE6A735C" box="[325,476,302,328]" direction="east" orientation="longitude" pageId="3" pageNumber="44" precision="15" value="36.40389">36°24’14”E</geoCoordinate>
|
||
,
|
||
<quantity id="4CBC47ACFFE79650E6A92654FDF6735C" box="[490,576,302,328]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="44" unit="m" value="1000.0">1000m</quantity>
|
||
a.s.l.); collected by local hunter,
|
||
<date id="FFFACC89FFE79650E4A52654FB0A735C" box="[998,1212,302,328]" pageId="3" pageNumber="44" value="2008-10-20">20 October 2008</date>
|
||
; fixed in buffered formalin and stored in 70% EtOH.
|
||
<emphasis id="B930365BFFE79650E503262FFD72737A" box="[576,708,341,366]" italics="true" pageId="3" pageNumber="44">
|
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<typeStatus id="54FF54EBFFE79650E503262FFD0B737A" box="[576,701,341,366]" pageId="3" pageNumber="44" type="paratype">Paratypes</typeStatus>
|
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:
|
||
</emphasis>
|
||
<specimenCount id="9D4221C0FFE79650E58F262FFCD3737B" box="[716,869,341,367]" pageId="3" pageNumber="44" type="generic">3 specimens</specimenCount>
|
||
: adult female,
|
||
<collectionCode id="ED55728CFFE79650E354262FFBE2737B" box="[1047,1108,341,367]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="3" pageNumber="44">PEM</collectionCode>
|
||
R17902, collected by Tom Timberlake, same locality and date as
|
||
<typeStatus id="54FF54EBFFE79650E52A2606FD607382" box="[617,726,380,406]" pageId="3" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
; juvenile,
|
||
<collectionCode id="ED55728CFFE79650E4162606FC257382" box="[853,915,380,406]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="3" pageNumber="44">PEM</collectionCode>
|
||
R17903, collected by Julian Bayliss, same locality and date as
|
||
<typeStatus id="54FF54EBFFE79650E6C026D8FE5873A8" box="[387,494,418,444]" pageId="3" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
; subadult male,
|
||
<collectionCode id="ED55728CFFE79650E5EC26D8FD5A73A8" box="[687,748,418,444]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="3" pageNumber="44">PEM</collectionCode>
|
||
R19704, collected on path at 13h00 in leaf litter in Khara Forest (lower end on Manho Forest, Mount Namuli, Zambezia Province,
|
||
<collectingCountry id="F353AAD9FFE79650E37026B3FB6073F7" box="[1075,1238,457,483]" name="Mozambique" pageId="3" pageNumber="44">Mozambique</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE708C8EFFE79650E3AA26B3FA2173F7" box="[1257,1431,457,483]" direction="south" orientation="latitude" pageId="3" pageNumber="44" precision="1" value="-15.411083">15°24’39.9”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE708C8EFFE79650E7D4268AFEF7701E" box="[151,321,496,522]" direction="east" orientation="longitude" pageId="3" pageNumber="44" precision="1" value="37.037914">37°02’16.5”E</geoCoordinate>
|
||
,
|
||
<quantity id="4CBC47ACFFE79650E60C268AFE14701E" box="[335,418,496,522]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="3" pageNumber="44" unit="m" value="1550.0">1550m</quantity>
|
||
a.s.l.), Colin Congdon,
|
||
<date id="FFFACC89FFE79650E589268AFC0F701E" box="[714,953,496,522]" pageId="3" pageNumber="44" value="2008-11-26">26 November 2008</date>
|
||
. The two Mount Mabu specimens were retained for one month in captivity before dying; on death they were placed in 90% EtOH, after small incisions were made in the ventrum; both were subsequently transferred to 70% EtOH for storage. The Mount Namuli specimen was kept alive for 2 months, and then euthanased, fixed in buffered formalin and subsequently stored in 70% EtOH; a small ventral incision was made to collected fresh liver tissue for genetic studies.
|
||
</paragraph>
|
||
<paragraph id="8BFBEA49FFE79650E78525ADFC79710C" blockId="3.[151,1437,263,2035]" pageId="3" pageNumber="44">
|
||
<emphasis id="B930365BFFE79650E78525ADFE7A70E5" bold="true" box="[198,460,727,753]" pageId="3" pageNumber="44">Additional Material:</emphasis>
|
||
<collectionCode id="ED55728CFFE79650E69525A2FDA470E6" box="[470,530,728,754]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="3" pageNumber="44">PEM</collectionCode>
|
||
R17910 (still maintained in captivity; shed skins and digital images), adult male, same locality as
|
||
<typeStatus id="54FF54EBFFE79650E6E22584FDBC710C" box="[417,522,766,792]" pageId="3" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
, collected
|
||
<date id="FFFACC89FFE79650E5C42584FCFD710C" box="[647,843,766,792]" pageId="3" pageNumber="44" value="2006-01-25">25 January 2006</date>
|
||
, J. Bayliss.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35EB9C2FFE79650E785245FFEEC71A7" pageId="3" pageNumber="44" type="description">
|
||
<paragraph id="8BFBEA49FFE79650E785245FFCFD712B" blockId="3.[151,1437,263,2035]" box="[198,843,805,831]" pageId="3" pageNumber="44">
|
||
<emphasis id="B930365BFFE79650E785245FFEE2712B" bold="true" box="[198,340,805,831]" pageId="3" pageNumber="44">Synonymy:</emphasis>
|
||
<taxonomicName id="4C4491CAFFE79650E618245FFE06712A" box="[347,432,805,830]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="B930365BFFE79650E618245FFE06712A" box="[347,432,805,830]" italics="true" pageId="3" pageNumber="44">Atheris</emphasis>
|
||
</taxonomicName>
|
||
sp.
|
||
<bibRefCitation id="EFD597B8FFE79650E697245FFD4E712B" author="Dobiey" box="[468,760,805,831]" pageId="3" pageNumber="53" refString="Dobiey, M. & Vigel, G. (2007). Venomous Snakes of Africa / Giftschlangen Afrika. Terralog, vol. 15, Edition Chimaira, Frankfurt am Main, 150 pp." type="book" year="2007">Dobiey and Vogel, 2007</bibRefCitation>
|
||
, p 110.
|
||
</paragraph>
|
||
<paragraph id="8BFBEA49FFE79650E7852436FEEC71A7" blockId="3.[151,1437,263,2035]" pageId="3" pageNumber="44">
|
||
Numerous other images of the species under the same name have appeared in the world media following a press release from Kew highlighting the biodiversity discoveries during the Mount Mabu expedition (e.g.
|
||
<bibRefCitation id="EFD597B8FFE79650E7D424E3FEFB71A7" author="Marshall" box="[151,333,921,947]" pageId="3" pageNumber="54" refString="Marshall, L. 2009. Unspoilt Eden a rare find. Sunday Independent (newspaper, United Kingdom), 1 February 2009, p 7." type="journal article" year="2009">
|
||
<collectingCountry id="F353AAD9FFE79650E7D424E3FEB471A7" box="[151,258,921,947]" name="Marshall Islands" pageId="3" pageNumber="44">Marshall</collectingCountry>
|
||
, 2009
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35EB9C2FFE79650E78524C5FAE974D0" pageId="3" pageNumber="44" type="diagnosis">
|
||
<paragraph id="8BFBEA49FFE79650E78524C5FAE974D0" blockId="3.[151,1437,263,2035]" pageId="3" pageNumber="44">
|
||
<emphasis id="B930365BFFE79650E78524C5FEFE71CD" bold="true" box="[198,328,959,985]" pageId="3" pageNumber="44">Diagnosis:</emphasis>
|
||
<taxonomicName id="4C4491CAFFE79650E61224BAFD9C71CD" box="[337,554,960,985]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="mabuensis">
|
||
<emphasis id="B930365BFFE79650E61224BAFD9C71CD" box="[337,554,960,985]" italics="true" pageId="3" pageNumber="44">Atheris mabuensis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from all other members of the genus by a combination of characters: (1) its small size, maximum length
|
||
<quantity id="4CBC47ACFFE79650E5AE249FFCFA71EB" box="[749,844,997,1023]" metricMagnitude="-1" metricUnit="m" metricValue="3.84" pageId="3" pageNumber="44" unit="mm" value="384.0">384mm</quantity>
|
||
(all other
|
||
<taxonomicName id="4C4491CAFFE79650E490249FFB9871EA" box="[979,1070,997,1022]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="B930365BFFE79650E490249FFB9871EA" box="[979,1070,997,1022]" italics="true" pageId="3" pageNumber="44">Atheris</emphasis>
|
||
</taxonomicName>
|
||
exceed
|
||
<quantity id="4CBC47ACFFE79650E3D4249FFB4071EB" box="[1175,1270,997,1023]" metricMagnitude="-1" metricUnit="m" metricValue="5.8" pageId="3" pageNumber="44" unit="mm" value="580.0">580mm</quantity>
|
||
TL, with the exception of -
|
||
<emphasis id="B930365BFFE79650E60A2370FDB47637" box="[329,514,1034,1059]" italics="true" pageId="3" pageNumber="44">
|
||
<taxonomicName id="4C4491CAFFE79650E60A2370FE4B7637" box="[329,509,1034,1059]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="katangensis">A. katangensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
TL
|
||
<quantity id="4CBC47ACFFE79650E5742370FD207630" box="[567,662,1034,1060]" metricMagnitude="-1" metricUnit="m" metricValue="3.97" pageId="3" pageNumber="44" unit="mm" value="397.0">397mm</quantity>
|
||
;
|
||
<taxonomicName id="4C4491CAFFE79650E5E02370FC877637" box="[675,817,1034,1059]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="barbouri">
|
||
<emphasis id="B930365BFFE79650E5E02370FC877637" box="[675,817,1034,1059]" italics="true" pageId="3" pageNumber="44">A. barbouri</emphasis>
|
||
</taxonomicName>
|
||
, TL
|
||
<quantity id="4CBC47ACFFE79650E42D2370FC647630" box="[878,978,1034,1060]" metricMagnitude="-1" metricUnit="m" metricValue="3.69" pageId="3" pageNumber="44" unit="mm" value="369.0">369 mm</quantity>
|
||
(Barbour and
|
||
<bibRefCitation id="EFD597B8FFE79650E3C72370FA9B7630" author="Howell" box="[1156,1325,1034,1060]" pageId="3" pageNumber="54" refString="Rasmussen, J. B. & Howell, K. M .. (1998). A review of Barbour's Short-headed Viper, Adenorhinus barbouri (Serpentes: Viperidae). African Journal of Herpetology, 47 (2), 69 - 75." type="journal article" year="1998">Howell, 1998</bibRefCitation>
|
||
); and the unique
|
||
<typeStatus id="54FF54EBFFE79650E7AD234AFE97765E" box="[238,289,1072,1098]" pageId="3" pageNumber="44">type</typeStatus>
|
||
of
|
||
<emphasis id="B930365BFFE79650E605234AFE5F765D" box="[326,489,1072,1097]" italics="true" pageId="3" pageNumber="44">
|
||
<taxonomicName id="4C4491CAFFE79650E605234AFE53765D" box="[326,485,1072,1097]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="acuminata">A. acuminata</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
TL
|
||
<quantity id="4CBC47ACFFE79650E558234AFDCD765E" box="[539,635,1072,1098]" metricMagnitude="-1" metricUnit="m" metricValue="4.4" pageId="3" pageNumber="44" unit="mm" value="440.0">440mm</quantity>
|
||
); (2) the lack of ‘horns’ (enlarged supraocular scales are present in
|
||
<taxonomicName id="4C4491CAFFE79650E2C9234AFE9F767A" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="ceratophora">
|
||
<emphasis id="B930365BFFE79650E2C9234AFE9F767A" italics="true" pageId="3" pageNumber="44">A ceratophora</emphasis>
|
||
</taxonomicName>
|
||
); (3) the lack of lanceolate or acuminate scales on top of the head (present in
|
||
<taxonomicName id="4C4491CAFFE79650E38D232FFAF1767A" box="[1230,1351,1109,1134]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="hispida">
|
||
<emphasis id="B930365BFFE79650E38D232FFAF1767A" box="[1230,1351,1109,1134]" italics="true" pageId="3" pageNumber="44">A. hispida</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4491CAFFE79650E2C0232FFEA27680" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="acuminata">
|
||
<emphasis id="B930365BFFE79650E2C0232FFEA27680" italics="true" pageId="3" pageNumber="44">A. acuminata</emphasis>
|
||
</taxonomicName>
|
||
); (4) having weakly keeled gular scales (smooth only
|
||
<emphasis id="B930365BFFE79650E4E12301FB897680" box="[930,1087,1147,1172]" italics="true" pageId="3" pageNumber="44">
|
||
in
|
||
<taxonomicName id="4C4491CAFFE79650E4822301FB897680" box="[961,1087,1147,1172]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="nitschei">A. nitschei</taxonomicName>
|
||
</emphasis>
|
||
; gulars moderately keeled in the eastern species,
|
||
<emphasis id="B930365BFFE79650E6C223DAFCCE76AD" box="[385,888,1184,1209]" italics="true" pageId="3" pageNumber="44">
|
||
<taxonomicName id="4C4491CAFFE79650E6C223DAFD8176AD" box="[385,567,1184,1209]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="rungweensis">A. rungweensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4491CAFFE79650E50023DAFD0C76AD" box="[579,698,1184,1209]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="desaixi">A. desaixi</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4491CAFFE79650E58523DAFCCE76AD" box="[710,888,1184,1209]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="ceratophora">A. ceratophora</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4491CAFFE79650E4F223DAFBEA76AD" box="[945,1116,1184,1209]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="katangensis">
|
||
<emphasis id="B930365BFFE79650E4F223DAFBEA76AD" box="[945,1116,1184,1209]" italics="true" pageId="3" pageNumber="44">A. katangensis</emphasis>
|
||
</taxonomicName>
|
||
, and strongly keeled in the remaining central and western species); (5) lacking interoculabials (
|
||
<emphasis id="B930365BFFE79650E4F223BFFC4476CA" box="[945,1010,1221,1246]" italics="true" pageId="3" pageNumber="44">sensu</emphasis>
|
||
<bibRefCitation id="EFD597B8FFE79650E4BA23BFFB1876CB" author="Broadley" box="[1017,1198,1221,1247]" pageId="3" pageNumber="53" refString="Broadley, D. G. (1998). A review of the genus Atheris Cope (Serpentes: Viperidae), with the description of a new species from Uganda. Herpetological Journal, 8,117 - 135." type="book" year="1998">Broadley, 1998</bibRefCitation>
|
||
, i.e. the supralabials are in contact with circumorbitals; 1 or
|
||
<quantity id="4CBC47ACFFE79650E5242390FD227710" box="[615,660,1258,1284]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="44" unit="in" value="2.0">2 in</quantity>
|
||
<taxonomicName id="4C4491CAFFE79650E5D82391FCB97710" box="[667,783,1259,1284]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="desaixi">
|
||
<emphasis id="B930365BFFE79650E5D82391FCB97710" box="[667,783,1259,1284]" italics="true" pageId="3" pageNumber="44">A. desaixi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4491CAFFE79650E40B2391FC4A7710" box="[840,1020,1259,1284]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="rungweensis">
|
||
<emphasis id="B930365BFFE79650E40B2391FC4A7710" box="[840,1020,1259,1284]" italics="true" pageId="3" pageNumber="44">A. rungweensis</emphasis>
|
||
</taxonomicName>
|
||
); (6) having 19–21 transverse head scales (
|
||
<emphasis id="B930365BFFE79650E7B0226AFE81773D" box="[243,311,1296,1321]" italics="true" pageId="3" pageNumber="44">sensu</emphasis>
|
||
<bibRefCitation id="EFD597B8FFE79650E67C226AFE4B773E" author="Broadley" box="[319,509,1296,1322]" pageId="3" pageNumber="53" refString="Broadley, D. G. (1998). A review of the genus Atheris Cope (Serpentes: Viperidae), with the description of a new species from Uganda. Herpetological Journal, 8,117 - 135." type="book" year="1998">Broadley, 1998</bibRefCitation>
|
||
, i.e. number of scales across head between posterior supralabials; these are reduced in highly arboreal species, e.g.
|
||
<emphasis id="B930365BFFE79650E53C224FFC8F775A" box="[639,825,1333,1358]" italics="true" pageId="3" pageNumber="44">
|
||
<taxonomicName id="4C4491CAFFE79650E53C224FFC85775A" box="[639,819,1333,1358]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="squamigera">A. squamigera</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
15–22,
|
||
<emphasis id="B930365BFFE79650E4DF224FFB97775A" box="[924,1057,1333,1358]" italics="true" pageId="3" pageNumber="44">
|
||
<taxonomicName id="4C4491CAFFE79650E4DF224FFBAA775A" box="[924,1052,1333,1358]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="hispida">A. hispida</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
12; and
|
||
<emphasis id="B930365BFFE79650E3CF224FFA8E775A" box="[1164,1336,1333,1358]" italics="true" pageId="3" pageNumber="44">
|
||
<taxonomicName id="4C4491CAFFE79650E3CF224FFA85775A" box="[1164,1331,1333,1358]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="acuminata">A. acuminata</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
10); (7) having 21–23
|
||
<collectingCountry id="F353AAD9FFE79650E60F2220FE3B7760" box="[332,397,1370,1396]" name="Egypt" pageId="3" pageNumber="44">MSR</collectingCountry>
|
||
(most species have 27+
|
||
<collectingCountry id="F353AAD9FFE79650E58B2220FCBF7760" box="[712,777,1370,1396]" name="Egypt" pageId="3" pageNumber="44">MSR</collectingCountry>
|
||
rows; reduced in highly arboreal species such as
|
||
<emphasis id="B930365BFFE79650E2C12221FE9C778D" italics="true" pageId="3" pageNumber="44">
|
||
<taxonomicName id="4C4491CAFFE79650E2C12221FE90778D" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="squamigera">A. squamigera</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
15–25,
|
||
<emphasis id="B930365BFFE79650E6CE22FAFDB9778D" box="[397,527,1408,1433]" italics="true" pageId="3" pageNumber="44">
|
||
<taxonomicName id="4C4491CAFFE79650E6CE22FAFDBC778D" box="[397,522,1408,1433]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="hispida">A. hispida</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
15–19, and
|
||
<emphasis id="B930365BFFE79650E5E022FAFCFC778D" box="[675,842,1408,1433]" italics="true" pageId="3" pageNumber="44">
|
||
<taxonomicName id="4C4491CAFFE79650E5E022FAFCF0778D" box="[675,838,1408,1433]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="acuminata">A. acuminata</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
14); (8) lateral body scales not serrated (strongly serrated in
|
||
<emphasis id="B930365BFFE79650E65B22DFFD6C77AA" box="[280,730,1445,1470]" italics="true" pageId="3" pageNumber="44">
|
||
<taxonomicName id="4C4491CAFFE79650E65B22DFFE7A77AA" box="[280,460,1445,1470]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="ceratophora">A. ceratophora</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4491CAFFE79650E69422DFFDFB77AA" box="[471,589,1445,1470]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="desaixi">A. desaixi</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4491CAFFE79650E51B22DFFD6077AA" box="[600,726,1445,1470]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="nitschei">A. nitschei</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4491CAFFE79650E45122DFFC7277AA" box="[786,964,1445,1470]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="rungweensis">
|
||
<emphasis id="B930365BFFE79650E45122DFFC7277AA" box="[786,964,1445,1470]" italics="true" pageId="3" pageNumber="44">A. rungweensis</emphasis>
|
||
</taxonomicName>
|
||
, and weakly serrated in
|
||
<taxonomicName id="4C4491CAFFE79650E3A322DFFA3C77AA" box="[1248,1418,1445,1470]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="katangensis">
|
||
<emphasis id="B930365BFFE79650E3A322DFFA3C77AA" box="[1248,1418,1445,1470]" italics="true" pageId="3" pageNumber="44">A. katangensis</emphasis>
|
||
</taxonomicName>
|
||
); (9) having 8–9 supralabials (six in
|
||
<taxonomicName id="4C4491CAFFE79650E51222B1FD4E77F0" box="[593,760,1483,1508]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="acuminata">
|
||
<emphasis id="B930365BFFE79650E51222B1FD4E77F0" box="[593,760,1483,1508]" italics="true" pageId="3" pageNumber="44">A. acuminata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="4CBC47ACFFE79650E44A22B0FCC177F0" box="[777,887,1482,1508]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="3" pageNumber="44" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">10–12 in</quantity>
|
||
<taxonomicName id="4C4491CAFFE79650E4C222B1FC4877F0" box="[897,1022,1483,1508]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="desaixi">
|
||
<emphasis id="B930365BFFE79650E4C222B1FC4877F0" box="[897,1022,1483,1508]" italics="true" pageId="3" pageNumber="44">A. desaixi</emphasis>
|
||
</taxonomicName>
|
||
); (10) having low ventral counts 128–137 (this is the lowest in the genus; usually over
|
||
<quantity id="4CBC47ACFFE79650E468228AFCCE741E" box="[811,888,1520,1546]" metricMagnitude="0" metricUnit="m" metricValue="3.556" pageId="3" pageNumber="44" unit="in" value="140.0">140 in</quantity>
|
||
both sexes in
|
||
<emphasis id="B930365BFFE79650E36B228AFDA8743A" italics="true" pageId="3" pageNumber="44">
|
||
<taxonomicName id="4C4491CAFFE79650E36B228AFB1D741D" box="[1064,1195,1520,1545]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="nitschei">A. nitschei</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4491CAFFE79650E3FA228AFAC3741D" box="[1209,1397,1520,1545]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="rungweensis">A. rungweensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4491CAFFE79650E2C1228AFF5B743A" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="desaixi">A. desaixi</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4491CAFFE79650E7BB216FFE23743A" box="[248,405,1557,1582]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="chlorechis">A. chlorechis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4491CAFFE79650E6E3216FFDAC743A" box="[416,538,1557,1582]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="hispida">A. hispida</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4491CAFFE79650E514216FFCB7743A" box="[599,769,1557,1582]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="subocularis">
|
||
<emphasis id="B930365BFFE79650E514216FFCB7743A" box="[599,769,1557,1582]" italics="true" pageId="3" pageNumber="44">A. subocularis</emphasis>
|
||
</taxonomicName>
|
||
, and in the only known males of
|
||
<taxonomicName id="4C4491CAFFE79650E3C4216FFA95743A" box="[1159,1315,1557,1582]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="acuminata">
|
||
<emphasis id="B930365BFFE79650E3C4216FFA95743A" box="[1159,1315,1557,1582]" italics="true" pageId="3" pageNumber="44">A. acuminata</emphasis>
|
||
</taxonomicName>
|
||
(160) and
|
||
<taxonomicName id="4C4491CAFFE79650E7D42141FEA67440" box="[151,272,1595,1620]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="hirsuta">
|
||
<emphasis id="B930365BFFE79650E7D42141FEA67440" box="[151,272,1595,1620]" italics="true" pageId="3" pageNumber="44">A. hirsuta</emphasis>
|
||
</taxonomicName>
|
||
(160); (11) having low subcaudal counts - 39–47 (always higher than
|
||
<quantity id="4CBC47ACFFE79650E3322140FB197440" box="[1137,1199,1594,1620]" metricMagnitude="0" metricUnit="m" metricValue="1.143" pageId="3" pageNumber="44" unit="in" value="45.0">45 in</quantity>
|
||
<emphasis id="B930365BFFE79650E3FB2141FD12746D" italics="true" pageId="3" pageNumber="44">
|
||
<taxonomicName id="4C4491CAFFE79650E3FB2141FAC37440" box="[1208,1397,1595,1620]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="rungweensis">A. rungweensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4491CAFFE79650E2C12141FE87746D" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="ceratophora">A. ceratophora</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4491CAFFE79650E67D211AFE52746D" box="[318,484,1632,1657]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="chlorechis">A. chlorechis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4491CAFFE79650E6B2211AFD12746D" box="[497,676,1632,1657]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="squamigera">A. squamigera</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C4491CAFFE79650E5A2211AFCE8746D" box="[737,862,1632,1657]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="hispida">
|
||
<emphasis id="B930365BFFE79650E5A2211AFCE8746D" box="[737,862,1632,1657]" italics="true" pageId="3" pageNumber="44">A. hispida</emphasis>
|
||
</taxonomicName>
|
||
; and 54 and
|
||
<quantity id="4CBC47ACFFE79650E343211AFB89746E" box="[1024,1087,1632,1658]" metricMagnitude="0" metricUnit="m" metricValue="1.4731999999999998" pageId="3" pageNumber="44" unit="in" value="58.0">58 in</quantity>
|
||
the only known males of
|
||
<taxonomicName id="4C4491CAFFE79650E2C1211AFEA5748A" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="acuminata">
|
||
<emphasis id="B930365BFFE79650E2C1211AFEA5748A" italics="true" pageId="3" pageNumber="44">A. acuminata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4491CAFFE79650E60821FFFE09748A" box="[331,447,1669,1694]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="hirsuta">
|
||
<emphasis id="B930365BFFE79650E60821FFFE09748A" box="[331,447,1669,1694]" italics="true" pageId="3" pageNumber="44">A. hirsuta</emphasis>
|
||
</taxonomicName>
|
||
, respectively); and (12) having a prehensile tail (non-prehensile in
|
||
<taxonomicName id="4C4491CAFFE79650E39221FFFAEF748A" box="[1233,1369,1669,1694]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="barbouri">
|
||
<emphasis id="B930365BFFE79650E39221FFFAEF748A" box="[1233,1369,1669,1694]" italics="true" pageId="3" pageNumber="44">A. barbouri</emphasis>
|
||
</taxonomicName>
|
||
), and higher subcaudal (
|
||
<taxonomicName id="4C4491CAFFE79650E62C21D1FE4174D0" box="[367,503,1707,1732]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="barbouri">
|
||
<emphasis id="B930365BFFE79650E62C21D1FE4174D0" box="[367,503,1707,1732]" italics="true" pageId="3" pageNumber="44">A. barbouri</emphasis>
|
||
</taxonomicName>
|
||
15–21) and labial (
|
||
<emphasis id="B930365BFFE79650E59E21D1FCDB74D0" box="[733,877,1707,1732]" italics="true" pageId="3" pageNumber="44">
|
||
<taxonomicName id="4C4491CAFFE79650E59E21D1FCDF74D0" box="[733,873,1707,1732]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="barbouri">A. barbouri</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
supralabials 5–6, infralabials 4–5) counts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35EB9C2FFE79650E78521B5FF55754E" pageId="3" pageNumber="44" type="discussion">
|
||
<paragraph id="8BFBEA49FFE79650E78521B5FF55754E" blockId="3.[151,1437,263,2035]" pageId="3" pageNumber="44">
|
||
<emphasis id="B930365BFFE79650E78521B5FEF474FD" bold="true" box="[198,322,1743,1769]" pageId="3" pageNumber="44">Remarks:</emphasis>
|
||
Within the Atherini the Mount Mabu snake is referred to the genus
|
||
<taxonomicName id="4C4491CAFFE79650E32F21AAFAB174FE" authority="Cope" authorityName="Cope" box="[1132,1287,1744,1770]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="B930365BFFE79650E32F21AAFB7774FD" box="[1132,1217,1744,1769]" italics="true" pageId="3" pageNumber="44">Atheris</emphasis>
|
||
Cope
|
||
</taxonomicName>
|
||
on the basis of the following diagnostic features: no enlarged supraocular shield (present in
|
||
<taxonomicName id="4C4491CAFFE79650E30D218FFA38751B" authority="Broadley, 1966" authorityName="Broadley" authorityYear="1966" box="[1102,1422,1781,1807]" class="Reptilia" family="Viperidae" genus="Proatheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="B930365BFFE79650E30D218FFB7A751A" box="[1102,1228,1781,1806]" italics="true" pageId="3" pageNumber="44">Proatheris</emphasis>
|
||
Broadley, 1966
|
||
</taxonomicName>
|
||
), tail prehensile and subcaudals single (tail non-prehensile and subcaudals paired in
|
||
<taxonomicName id="4C4491CAFFE79650E3C82061FF60754E" authority="Broadley, 1996" authorityName="Broadley" authorityYear="1996" class="Reptilia" family="Viperidae" genus="Montatheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="B930365BFFE79650E3C82061FAA97520" box="[1163,1311,1819,1844]" italics="true" pageId="3" pageNumber="44">Montatheris</emphasis>
|
||
<bibRefCitation id="EFD597B8FFE79650E2642060FF60754E" author="Broadley" pageId="3" pageNumber="53" refString="Broadley, D. G. (1996). A review of the tribe Atherini (Serpentes: Viperidae), with the description of two new genera. African Journal of Herpetology, 45, 40 - 48." type="journal article" year="1996">Broadley, 1996</bibRefCitation>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35EB9C2FFE79655E785201FFBAD718B" lastPageId="6" lastPageNumber="47" pageId="3" pageNumber="44" type="description">
|
||
<paragraph id="8BFBEA49FFE79650E785201FFC4175B2" blockId="3.[151,1437,263,2035]" pageId="3" pageNumber="44">
|
||
<emphasis id="B930365BFFE79650E785201FFED7756B" bold="true" box="[198,353,1893,1919]" pageId="3" pageNumber="44">Description:</emphasis>
|
||
<typeStatus id="54FF54EBFFE79650E628201FFE6E756B" box="[363,472,1893,1919]" pageId="3" pageNumber="44" type="holotype">Holotype</typeStatus>
|
||
PEM R17901; adult male preserved in formalin and transferred to 70% EtOH; body bent into ‘U’-shape, and with a small incision in the belly (
|
||
<date id="FFFACC89FFE79650E4C820F6FC7475B2" box="[907,962,1932,1958]" pageId="3" pageNumber="44" value="1948-05">V48</date>
|
||
-52).
|
||
</paragraph>
|
||
<paragraph id="8BFBEA49FFE79650E78520C9FB1175E7" blockId="3.[151,1437,263,2035]" pageId="3" pageNumber="44">
|
||
<emphasis id="B930365BFFE79650E78520C9FE8575D8" box="[198,307,1971,1996]" italics="true" pageId="3" pageNumber="44">Meristics</emphasis>
|
||
: snout-vent length (SVL)
|
||
<quantity id="4CBC47ACFFE79650E53020C8FD6675D8" box="[627,720,1970,1996]" metricMagnitude="-1" metricUnit="m" metricValue="2.56" pageId="3" pageNumber="44" unit="mm" value="256.0">256mm</quantity>
|
||
, tail
|
||
<quantity id="4CBC47ACFFE79650E44D20C8FCEF75D8" box="[782,857,1970,1996]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="3" pageNumber="44" unit="mm" value="47.0">47mm</quantity>
|
||
(TL
|
||
<quantity id="4CBC47ACFFE79650E4D920C8FBB575D8" box="[922,1027,1970,1996]" metricMagnitude="-1" metricUnit="m" metricValue="3.03" pageId="3" pageNumber="44" unit="mm" value="303.0">303 mm</quantity>
|
||
); tail into SVL 5.45 times; rostral width
|
||
<quantity id="4CBC47ACFFE79650E7A220A3FE8075E7" box="[225,310,2009,2035]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="44" unit="mm" value="2.9">2.9mm</quantity>
|
||
, rostral height
|
||
<quantity id="4CBC47ACFFE79650E6A720A3FD8C75E7" box="[484,570,2009,2035]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="44" unit="mm" value="0.8">0.8mm</quantity>
|
||
; eye diameter (vertical)
|
||
<quantity id="4CBC47ACFFE79650E41520A3FC1B75E7" box="[854,941,2009,2035]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="44" unit="mm" value="3.2">3.2mm</quantity>
|
||
; snout to eye
|
||
<quantity id="4CBC47ACFFE79650E30E20A3FB1575E7" box="[1101,1187,2009,2035]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="44" unit="mm" value="3.3">3.3mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFBEA49FFE09657E78527E2FB297332" blockId="4.[151,1437,152,410]" pageId="4" pageNumber="45">
|
||
<emphasis id="B930365BFFE09657E78527E2FE9572A5" box="[198,291,152,177]" italics="true" pageId="4" pageNumber="45">Habitus</emphasis>
|
||
: Body relatively slender and subtriangular in cross-section, with a thin, prehensile tail (SVL/Tail approximately 5–6 times; 4.92–6.16, mean 5.56, n = 5); head triangular, with a very distinct neck, bluntly rounded snout and swollen supraorbital region that does not bear elongate scales; eyes relatively large, laterally placed, and with a horizontal diameter approximately equal to the snout length.
|
||
</paragraph>
|
||
<paragraph id="8BFBEA49FFE09656E7852648FD1A7065" blockId="4.[151,1437,152,410]" lastBlockId="5.[151,1437,152,625]" lastPageId="5" lastPageNumber="46" pageId="4" pageNumber="45">
|
||
<emphasis id="B930365BFFE09657E7852648FE83735F" box="[198,309,306,331]" italics="true" pageId="4" pageNumber="45">Scalation</emphasis>
|
||
: Crown of head covered in small scales, that are larger over the temporal region and somewhat irregular in outline between the eyes; they bear a prominent keel that is extended into a rounded, pale knob; the rostral is flattened, subtriangular, about 2.4–3.4 times broader than high, contacting first supralabials and three large, unkeeled suprarostrals, of which the middle is the smallest and about 1.25 times wider than high; right and left suprarostrals slightly higher than wide; nasal wider than high, with raised, embossed posterior edge, particularly on the left; nostril circular and approximately in the centre of the nasal; internasals 5, all strongly keeled; interrictals 21; interorbitals 6–7, keeled, central scales more irregular in shape; circumoculars 15–15, all keeled and terminating in blunt knobs; suboculars absent; circumoculars separated from nasals by a single row of feebly knobbed scales; a row of three irregular scales, bordering supralabials between nasal and lower circumoculars; supralabials 8–8, 3 largest, and 6–8 feebly-keeled; infralabials 9–9, posteriormost feebly keeled and first in contact at the midline behind the mental, and separating the latter from 3 pairs of chin shields, the first largest; mental triangular, approximately twice as wide as deep; gulars bordering chin shields feebly-keeled, but prominently-keeled towards the rictus; 2 preventrals, first largest; 132 ventrals; 47 undivided subcaudals (including spine); anal entire; dorsal scales about 1.5 times as long as wide, becoming shorter posteriorly, and 20 rows anteriorly, 25–26 rows at midbody, 17 rows posteriorly; keel on dorsal scales increasing in height from base, terminating in a rounded, pale knob that is more conspicuous on scales of the forebody; paraventrals larger than other dorsals, with a feeble, asymmetric keel; additional scales occur irregularly in rows 2–5 on the forebody, but do not form continuous duplicated scale rows.
|
||
</paragraph>
|
||
<caption id="DF3BBAC1FFE09657E7D420C2FB8F75E6" httpUri="https://zenodo.org/record/274879/files/figure.png" pageId="4" pageNumber="45" targetBox="[151,1434,451,1949]" targetPageId="4">
|
||
<paragraph id="8BFBEA49FFE09657E7D420C2FB8F75E6" blockId="4.[151,1436,1976,2034]" pageId="4" pageNumber="45">
|
||
<emphasis id="B930365BFFE09657E7D420C2FEAF75C4" bold="true" box="[151,281,1976,2000]" pageId="4" pageNumber="45">FIGURE 2.</emphasis>
|
||
Right (top) and left (bottom) lateral views of the head of
|
||
<taxonomicName id="4C4491CAFFE09657E4C120C3FBF175C4" box="[898,1095,1977,2000]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="45" phylum="Chordata" rank="species" species="mabuensis" status="sp. nov.">
|
||
<emphasis id="B930365BFFE09657E4C120C3FBF175C4" box="[898,1095,1977,2000]" italics="true" pageId="4" pageNumber="45">Atheris mabuensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B930365BFFE09657E30E20C2FB3F75C4" bold="true" box="[1101,1161,1976,2000]" pageId="4" pageNumber="45">
|
||
<taxonomicNameLabel id="A2038B20FFE09657E30E20C2FB3F75C4" box="[1101,1161,1976,2000]" pageId="4" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Holotype PEM R17901, Mount Mabu) before fixation. Note swollen gold keel tips on the head and body scales.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3BBAC1FFE19656E7D42124FB98748D" httpUri="https://zenodo.org/record/274880/files/figure.png" pageId="5" pageNumber="46" targetBox="[155,1435,666,1617]" targetPageId="5">
|
||
<paragraph id="8BFBEA49FFE19656E7D42124FB98748D" blockId="5.[151,1436,1630,1689]" pageId="5" pageNumber="46">
|
||
<emphasis id="B930365BFFE19656E7D42124FEAB7462" bold="true" box="[151,285,1630,1654]" pageId="5" pageNumber="46">FIGURE 3.</emphasis>
|
||
Dorsal (left) and ventral (right) views of the head of
|
||
<taxonomicName id="4C4491CAFFE19656E42C2125FB8C7462" box="[879,1082,1631,1654]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="46" phylum="Chordata" rank="species" species="mabuensis" status="sp. nov.">
|
||
<emphasis id="B930365BFFE19656E42C2125FB8C7462" box="[879,1082,1631,1654]" italics="true" pageId="5" pageNumber="46">Atheris mabuensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B930365BFFE19656E3012124FB367462" bold="true" box="[1090,1152,1630,1654]" pageId="5" pageNumber="46">
|
||
<taxonomicNameLabel id="A2038B20FFE19656E3012124FB367462" box="[1090,1152,1630,1654]" pageId="5" pageNumber="46" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Holotype PEM R17901, Mount Mabu) before fixation. Note the poorly-defined dark V-patch on crown of head
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFBEA49FFE19656E78521BDFCBD7545" blockId="5.[151,1436,1735,2022]" pageId="5" pageNumber="46">
|
||
<emphasis id="B930365BFFE19656E78521BDFEF574F4" box="[198,323,1735,1760]" italics="true" pageId="5" pageNumber="46">Hemipenis</emphasis>
|
||
: Both hemipenes have only partially everted, despite the rector muscles having been cut through a small lateral incision in subcaudals 13–17; the basal portion of the hemipenis reveals a typically divided organ with a number of basal spines on the lateral surfaces, and indications of a distal calyculate zone (this needs confirmation on fully everted hemipenes).
|
||
</paragraph>
|
||
<paragraph id="8BFBEA49FFE19655E7852026FC717380" blockId="5.[151,1436,1735,2022]" lastBlockId="6.[151,1437,152,1076]" lastPageId="6" lastPageNumber="47" pageId="5" pageNumber="46">
|
||
<emphasis id="B930365BFFE19656E7852026FED67561" box="[198,352,1884,1909]" italics="true" pageId="5" pageNumber="46">Colouration:</emphasis>
|
||
Eye dirty bronze, darker in centre and with bright yellow edge around iris; crown greyish brown with each scale tipped with yellow; head pattern vague, comprising a broken inverted ‘V’-shape represented by two small dark patches over the jaw muscles; side of head dark greyish in front of eye, becoming darker towards angle of jaw; dorsum pale brown with a pale, vague and irregular dorsal chain pattern, with faint paler edges and centre paler than body background colour; a dark edge to the chain pattern is represented only by a lateral series of subtriangular dark brown blotches that may lie adjacent to each other on either side of body, or become staggered; a series of small (sometimes only 1–2 scales), faded brown blotches occur along the flanks and align with the widest region of the dorsal ‘chain’ pattern; all scales on crown, body and upper surface of tail tipped in pale yellow to light orange, that is more prominent on the head and forebody; throat, belly and basal 2/3rd of underside of tail blue-grey, uniformly and heavily stippled with fine dark spots; underneath of tail becoming yellowish on distal third.
|
||
</paragraph>
|
||
<paragraph id="8BFBEA49FFE29655E78526DAFD577140" blockId="6.[151,1437,152,1076]" pageId="6" pageNumber="47">
|
||
<emphasis id="B930365BFFE29655E78526DAFE8C73AD" box="[198,314,416,441]" italics="true" pageId="6" pageNumber="47">Va r ia ti o n:</emphasis>
|
||
Details and meristics for the
|
||
<typeStatus id="54FF54EBFFE29655E5F726DAFD5173AE" box="[692,743,416,442]" pageId="6" pageNumber="47">type</typeStatus>
|
||
series are summarized in
|
||
<tableCitation id="C6C6DFF2FFE29655E37226DAFB3873AE" box="[1073,1166,416,442]" captionStart="TABLE 1" captionStartId="7.[151,239,151,175]" captionTargetBox="[151,1427,202,842]" captionTargetPageId="7" captionText="TABLE 1. Measurements and scalation for Atheris mabuensis. * includes terminal spine; ** (right / left)." httpUri="http://table.plazi.org/id/DF3BBAC1FFE39654E7D427EDFB4272BB" pageId="6" pageNumber="47" tableUuid="DF3BBAC1FFE39654E7D427EDFB4272BB">Table 1</tableCitation>
|
||
. The most significant differences in colouration include: body colouration in the solitary subadult from Mount Namuli (PEM R17904) is slightly duller (bluish) with the scales on the head and forebody lacking the conspicuous yellow tips of specimens from Mount Mabu. Whether this is typical of this isolated population awaits the collection of additional material. Pale, narrow eye stripes radiating from the orbit to the lip (see
|
||
<figureCitation id="137FF6CCFFE29655E3CF254FFB67705B" box="[1164,1233,565,591]" captionStart="FIGURE 6" captionStartId="9.[151,258,1869,1893]" captionTargetBox="[152,1433,194,1846]" captionTargetId="figure@9.[152,1435,194,1846]" captionTargetPageId="9" captionText="FIGURE 6. Atheris mabuensis n. sp. (PEM R 17910): Top - small juvenile (<150 mm TL) photographed in situ on capture (25 January 2006, Julian Bayliss); note drab colouration imparting camouflage in habitat among leaf litter on forest floor. Bottom – same specimen (photographed 19 Nov 2008, after 34 months in captivity); note retention of juvenile colouration, particularly yellow tail tip, and development of gold tips to the keels on the head and forebody scales." httpUri="https://zenodo.org/record/274883/files/figure.png" pageId="6" pageNumber="47">Fig. 6</figureCitation>
|
||
[top] and
|
||
<figureCitation id="137FF6CCFFE29655E20E254FFA24705B" box="[1357,1426,565,591]" captionStart="FIGURE 7" captionStartId="10.[151,257,1239,1263]" captionTargetBox="[151,1435,377,1215]" captionTargetId="figure@10.[151,1436,377,1215]" captionTargetPageId="10" captionText="FIGURE 7. Right lateral view of head and forebody Atheris mabuensis n. sp. (PEM R 17904, Mount Namuli): note reduction of gold tipped scales on head and body." httpUri="https://zenodo.org/record/274884/files/figure.png" pageId="6" pageNumber="47">Fig. 7</figureCitation>
|
||
) are found in all snakes, except the
|
||
<typeStatus id="54FF54EBFFE29655E5642520FD267060" box="[551,656,602,628]" pageId="6" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
, although in some (e.g. PEM R17903-4) only the rear stripe is well developed. The head pattern forms a well-developed inverted ‘V’-shape (see
|
||
<figureCitation id="137FF6CCFFE29655E37525FAFBC8708E" box="[1078,1150,640,666]" captionStart="FIGURE 7" captionStartId="10.[151,257,1239,1263]" captionTargetBox="[151,1435,377,1215]" captionTargetId="figure@10.[151,1436,377,1215]" captionTargetPageId="10" captionText="FIGURE 7. Right lateral view of head and forebody Atheris mabuensis n. sp. (PEM R 17904, Mount Namuli): note reduction of gold tipped scales on head and body." httpUri="https://zenodo.org/record/274884/files/figure.png" pageId="6" pageNumber="47">Fig. 7</figureCitation>
|
||
, PEM R17910) in three snakes, but is reduced to two isolated dark blotches at the back of the head in the
|
||
<typeStatus id="54FF54EBFFE29655E3C225DFFB5870AB" box="[1153,1262,677,703]" pageId="6" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
, and is almost completely absent in the subadult from Mount Namuli. There is little ontogenetic colour change (based on PEM R17910); when first captured (
|
||
<figureCitation id="137FF6CCFFE29655E512258AFD2D711E" box="[593,667,752,778]" captionStart="FIGURE 6" captionStartId="9.[151,258,1869,1893]" captionTargetBox="[152,1433,194,1846]" captionTargetId="figure@9.[152,1435,194,1846]" captionTargetPageId="9" captionText="FIGURE 6. Atheris mabuensis n. sp. (PEM R 17910): Top - small juvenile (<150 mm TL) photographed in situ on capture (25 January 2006, Julian Bayliss); note drab colouration imparting camouflage in habitat among leaf litter on forest floor. Bottom – same specimen (photographed 19 Nov 2008, after 34 months in captivity); note retention of juvenile colouration, particularly yellow tail tip, and development of gold tips to the keels on the head and forebody scales." httpUri="https://zenodo.org/record/274883/files/figure.png" pageId="6" pageNumber="47">Fig. 6</figureCitation>
|
||
, top) the juvenile snake (<
|
||
<quantity id="4CBC47ACFFE29655E49C258AFBF5711E" box="[991,1091,752,778]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="47" unit="mm" value="150.0">150 mm</quantity>
|
||
TL) had darker brown body colouration that after three years in captivity has lightened slightly (
|
||
<figureCitation id="137FF6CCFFE29655E482246FFBBC713B" box="[961,1034,789,815]" captionStart="FIGURE 6" captionStartId="9.[151,258,1869,1893]" captionTargetBox="[152,1433,194,1846]" captionTargetId="figure@9.[152,1435,194,1846]" captionTargetPageId="9" captionText="FIGURE 6. Atheris mabuensis n. sp. (PEM R 17910): Top - small juvenile (<150 mm TL) photographed in situ on capture (25 January 2006, Julian Bayliss); note drab colouration imparting camouflage in habitat among leaf litter on forest floor. Bottom – same specimen (photographed 19 Nov 2008, after 34 months in captivity); note retention of juvenile colouration, particularly yellow tail tip, and development of gold tips to the keels on the head and forebody scales." httpUri="https://zenodo.org/record/274883/files/figure.png" pageId="6" pageNumber="47">Fig. 6</figureCitation>
|
||
, bottom), although the extent and intensity of yellow on the tail remains unchanged.
|
||
</paragraph>
|
||
<paragraph id="8BFBEA49FFE29655E785241AFBAD718B" blockId="6.[151,1437,152,1076]" pageId="6" pageNumber="47">
|
||
<emphasis id="B930365BFFE29655E785241AFEB4716E" bold="true" box="[198,258,864,890]" pageId="6" pageNumber="47">Size:</emphasis>
|
||
Largest female (PEM R17902) 328 + 56 =
|
||
<quantity id="4CBC47ACFFE29655E5BD241AFCD4716E" box="[766,866,864,890]" metricMagnitude="-1" metricUnit="m" metricValue="3.84" pageId="6" pageNumber="47" unit="mm" value="384.0">384 mm</quantity>
|
||
; largest male (PEM R7910) 305 + 62 =
|
||
<quantity id="4CBC47ACFFE29655E277241AFA2E716E" box="[1332,1432,864,890]" metricMagnitude="-1" metricUnit="m" metricValue="3.67" pageId="6" pageNumber="47" unit="mm" value="367.0">367 mm</quantity>
|
||
. The series is too small to determine whether females grow larger then males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35EB9C2FFE29655E78524D0FCF371FE" pageId="6" pageNumber="47" type="etymology">
|
||
<paragraph id="8BFBEA49FFE29655E78524D0FAD771D0" blockId="6.[151,1437,152,1076]" box="[198,1377,938,964]" pageId="6" pageNumber="47">
|
||
<emphasis id="B930365BFFE29655E78524D0FEE171D0" bold="true" box="[198,343,938,964]" pageId="6" pageNumber="47">Etymology:</emphasis>
|
||
Named after the
|
||
<typeStatus id="54FF54EBFFE29655E55824D0FDFB71D0" box="[539,589,938,964]" pageId="6" pageNumber="47">type</typeStatus>
|
||
locality, Mount Mabu, Zambezia Province, northern
|
||
<collectingCountry id="F353AAD9FFE29655E3FD24D0FAEB71D0" box="[1214,1373,938,964]" name="Mozambique" pageId="6" pageNumber="47">Mozambique</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFBEA49FFE29655E78524AAFCF371FE" blockId="6.[151,1437,152,1076]" box="[198,837,976,1002]" pageId="6" pageNumber="47">
|
||
<emphasis id="B930365BFFE29655E78524AAFDBD71FE" bold="true" box="[198,523,976,1002]" pageId="6" pageNumber="47">Suggested common name:</emphasis>
|
||
Mount Mabu Forest Viper
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35EB9C2FFE29654E785248FFCA6748B" lastPageId="7" lastPageNumber="48" pageId="6" pageNumber="47" type="distribution">
|
||
<paragraph id="8BFBEA49FFE29655E785248FFD3B7620" blockId="6.[151,1437,152,1076]" pageId="6" pageNumber="47">
|
||
<emphasis id="B930365BFFE29655E785248FFEDF761B" bold="true" box="[198,361,1013,1039]" pageId="6" pageNumber="47">Distribution:</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54FF54EBFFE29655E5D5248FFD7E761B" box="[662,712,1013,1039]" pageId="6" pageNumber="47">type</typeStatus>
|
||
locality, Mount Mabu, and adjacent Mount Namuli, both montane isolates in northern
|
||
<collectingCountry id="F353AAD9FFE29655E6A82360FD3C7620" box="[491,650,1050,1076]" name="Mozambique" pageId="6" pageNumber="47">Mozambique</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF3BBAC1FFE29655E7D420DDFD6075F6" httpUri="https://zenodo.org/record/274881/files/figure.png" pageId="6" pageNumber="47" targetBox="[151,1435,1078,1932]" targetPageId="6">
|
||
<paragraph id="8BFBEA49FFE29655E7D420DDFD6075F6" blockId="6.[151,1437,1959,2018]" pageId="6" pageNumber="47">
|
||
<emphasis id="B930365BFFE29655E7D420DDFE9675AB" bold="true" box="[151,288,1959,1983]" pageId="6" pageNumber="47">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C4491CAFFE29655E66420DDFE4175AA" box="[295,503,1959,1982]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="mabuensis" status="sp. nov.">
|
||
<emphasis id="B930365BFFE29655E66420DDFE4175AA" box="[295,503,1959,1982]" italics="true" pageId="6" pageNumber="47">Atheris mabuensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B930365BFFE29655E6BC20DDFD8975AB" bold="true" box="[511,575,1959,1983]" pageId="6" pageNumber="47">
|
||
<taxonomicNameLabel id="A2038B20FFE29655E6BC20DDFD8975AB" box="[511,575,1959,1983]" pageId="6" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Holotype PEM R17901, Mount Mabu) in life showing habitus and general colouraion. Note: Retention of yellow tail tip in adult.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3BBAC1FFE39654E7D427EDFB4272BB" ID-Table-UUID="DF3BBAC1FFE39654E7D427EDFB4272BB" box="[151,1268,151,175]" httpUri="http://table.plazi.org/id/DF3BBAC1FFE39654E7D427EDFB4272BB" pageId="7" pageNumber="48" targetBox="[151,1427,202,842]" targetIsTable="true" targetPageId="7">
|
||
<paragraph id="8BFBEA49FFE39654E7D427EDFB4272BB" blockId="7.[151,1268,151,175]" box="[151,1268,151,175]" pageId="7" pageNumber="48">
|
||
<emphasis id="B930365BFFE39654E7D427EDFEBE72BB" bold="true" box="[151,264,151,175]" pageId="7" pageNumber="48">TABLE 1.</emphasis>
|
||
Measurements and scalation for
|
||
<emphasis id="B930365BFFE39654E52827E2FC8172BB" box="[619,823,152,175]" italics="true" pageId="7" pageNumber="48">
|
||
<taxonomicName id="4C4491CAFFE39654E52827E2FC8472BB" box="[619,818,152,175]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="mabuensis">Atheris mabuensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
* includes terminal spine; ** (right/left).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFBEA49FFE39654E63727B0FB5B715E" pageId="7" pageNumber="48">
|
||
<table id="F94418E9FFE369ACE7D427B0FA25715E" box="[151,1427,202,842]" gridcols="6" gridrows="15" pageId="7" pageNumber="48">
|
||
<tr id="3574E80BFFE369ACE7D427B0FA2572E8" box="[151,1427,202,252]" gridrow="0" pageId="7" pageNumber="48" rowspan-0="1">
|
||
<th id="76A58177FFE369ACE63727B0FD8472E8" box="[372,562,202,252]" gridcol="1" gridrow="0" pageId="7" pageNumber="48">Holotype PEM R17901</th>
|
||
<th id="76A58177FFE369ACE50F27B0FCBC72E8" box="[588,778,202,252]" gridcol="2" gridrow="0" pageId="7" pageNumber="48">Paratype PEM R17904</th>
|
||
<th id="76A58177FFE369ACE46727B0FC5472E8" box="[804,994,202,252]" gridcol="3" gridrow="0" pageId="7" pageNumber="48">Paratype PEM R17903</th>
|
||
<th id="76A58177FFE369ACE4BE27B0FB0C72E8" box="[1021,1210,202,252]" gridcol="4" gridrow="0" pageId="7" pageNumber="48">Captive PEM R17910</th>
|
||
<th id="76A58177FFE369ACE39627B0FA2572E8" box="[1237,1427,202,252]" gridcol="5" gridrow="0" pageId="7" pageNumber="48">Paratype PEM R17904</th>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D42675FA257330" box="[151,1427,271,292]" gridrow="1" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D42675FEF47330" box="[151,322,271,292]" gridcol="0" gridrow="1" pageId="7" pageNumber="48">Sex</th>
|
||
<td id="76A58177FFE369ACE6372675FD847330" box="[372,562,271,292]" gridcol="1" gridrow="1" pageId="7" pageNumber="48">Male</td>
|
||
<td id="76A58177FFE369ACE50F2675FCBC7330" box="[588,778,271,292]" gridcol="2" gridrow="1" pageId="7" pageNumber="48">Female</td>
|
||
<td id="76A58177FFE369ACE4672675FC547330" box="[804,994,271,292]" gridcol="3" gridrow="1" pageId="7" pageNumber="48">Juvenile</td>
|
||
<td id="76A58177FFE369ACE4BE2675FB0C7330" box="[1021,1210,271,292]" gridcol="4" gridrow="1" pageId="7" pageNumber="48">Male</td>
|
||
<td id="76A58177FFE369ACE3962675FA257330" box="[1237,1427,271,292]" gridcol="5" gridrow="1" pageId="7" pageNumber="48">Male</td>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D4264DFA257358" box="[151,1427,311,332]" gridrow="2" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D4264DFEF47358" box="[151,322,311,332]" gridcol="0" gridrow="2" pageId="7" pageNumber="48">Locality</th>
|
||
<td id="76A58177FFE369ACE637264DFD847358" box="[372,562,311,332]" gridcol="1" gridrow="2" pageId="7" pageNumber="48">Mount Mabu</td>
|
||
<td id="76A58177FFE369ACE50F264DFCBC7358" box="[588,778,311,332]" gridcol="2" gridrow="2" pageId="7" pageNumber="48">Mount Mabu</td>
|
||
<td id="76A58177FFE369ACE467264DFC547358" box="[804,994,311,332]" gridcol="3" gridrow="2" pageId="7" pageNumber="48">Mount Mabu</td>
|
||
<td id="76A58177FFE369ACE4BE264DFB0C7358" box="[1021,1210,311,332]" gridcol="4" gridrow="2" pageId="7" pageNumber="48">Mount Mabu</td>
|
||
<td id="76A58177FFE369ACE396264DFA257358" box="[1237,1427,311,332]" gridcol="5" gridrow="2" pageId="7" pageNumber="48">Mount Namuli</td>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D42625FA257360" box="[151,1427,351,372]" gridrow="3" pageId="7" pageNumber="48" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
|
||
<th id="76A58177FFE369ACE7D42625FEF47360" box="[151,322,351,372]" gridcol="0" gridrow="3" pageId="7" pageNumber="48">Measurements</th>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D426FDFA257388" box="[151,1427,391,412]" gridrow="4" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D426FDFEF47388" box="[151,322,391,412]" gridcol="0" gridrow="4" pageId="7" pageNumber="48">SVL + Tail = TL</th>
|
||
<td id="76A58177FFE369ACE63726FDFD847388" box="[372,562,391,412]" gridcol="1" gridrow="4" pageId="7" pageNumber="48">256 + 47 = 303 mm</td>
|
||
<td id="76A58177FFE369ACE50F26FDFCBC7388" box="[588,778,391,412]" gridcol="2" gridrow="4" pageId="7" pageNumber="48">328 + 56 = 384 mm</td>
|
||
<td id="76A58177FFE369ACE46726FDFC547388" box="[804,994,391,412]" gridcol="3" gridrow="4" pageId="7" pageNumber="48">154 + 25 = 179 mm</td>
|
||
<td id="76A58177FFE369ACE4BE26FDFB0C7388" box="[1021,1210,391,412]" gridcol="4" gridrow="4" pageId="7" pageNumber="48">305 + 62 = 367 mm</td>
|
||
<td id="76A58177FFE369ACE39626FDFA257388" box="[1237,1427,391,412]" gridcol="5" gridrow="4" pageId="7" pageNumber="48">194 + 36 = 233 mm</td>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D426D5FA2573D0" box="[151,1427,431,452]" gridrow="5" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D426D5FEF473D0" box="[151,322,431,452]" gridcol="0" gridrow="5" pageId="7" pageNumber="48">SVL/Tail</th>
|
||
<td id="76A58177FFE369ACE63726D5FD8473D0" box="[372,562,431,452]" gridcol="1" gridrow="5" pageId="7" pageNumber="48">5.45</td>
|
||
<td id="76A58177FFE369ACE50F26D5FCBC73D0" box="[588,778,431,452]" gridcol="2" gridrow="5" pageId="7" pageNumber="48">5.86</td>
|
||
<td id="76A58177FFE369ACE46726D5FC5473D0" box="[804,994,431,452]" gridcol="3" gridrow="5" pageId="7" pageNumber="48">6.16</td>
|
||
<td id="76A58177FFE369ACE4BE26D5FB0C73D0" box="[1021,1210,431,452]" gridcol="4" gridrow="5" pageId="7" pageNumber="48">4.92</td>
|
||
<td id="76A58177FFE369ACE39626D5FA2573D0" box="[1237,1427,431,452]" gridcol="5" gridrow="5" pageId="7" pageNumber="48">5.39</td>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D426ADFA2573F8" box="[151,1427,471,492]" gridrow="6" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D426ADFEF473F8" box="[151,322,471,492]" gridcol="0" gridrow="6" pageId="7" pageNumber="48">Eye Diameter</th>
|
||
<td id="76A58177FFE369ACE63726ADFD8473F8" box="[372,562,471,492]" gridcol="1" gridrow="6" pageId="7" pageNumber="48">3.2</td>
|
||
<td id="76A58177FFE369ACE50F26ADFCBC73F8" box="[588,778,471,492]" gridcol="2" gridrow="6" pageId="7" pageNumber="48">3.6</td>
|
||
<td id="76A58177FFE369ACE46726ADFC5473F8" box="[804,994,471,492]" gridcol="3" gridrow="6" pageId="7" pageNumber="48">2.3</td>
|
||
<td id="76A58177FFE369ACE4BE26ADFB0C73F8" box="[1021,1210,471,492]" gridcol="4" gridrow="6" pageId="7" pageNumber="48">na</td>
|
||
<td id="76A58177FFE369ACE39626ADFA2573F8" box="[1237,1427,471,492]" gridcol="5" gridrow="6" pageId="7" pageNumber="48">2.6</td>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D42685FA257000" box="[151,1427,511,532]" gridrow="7" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D42685FEF47000" box="[151,322,511,532]" gridcol="0" gridrow="7" pageId="7" pageNumber="48">Snout</th>
|
||
<td id="76A58177FFE369ACE6372685FD847000" box="[372,562,511,532]" gridcol="1" gridrow="7" pageId="7" pageNumber="48">3.3</td>
|
||
<td id="76A58177FFE369ACE50F2685FCBC7000" box="[588,778,511,532]" gridcol="2" gridrow="7" pageId="7" pageNumber="48">4.8</td>
|
||
<td id="76A58177FFE369ACE4672685FC547000" box="[804,994,511,532]" gridcol="3" gridrow="7" pageId="7" pageNumber="48">2.5</td>
|
||
<td id="76A58177FFE369ACE4BE2685FB0C7000" box="[1021,1210,511,532]" gridcol="4" gridrow="7" pageId="7" pageNumber="48">na</td>
|
||
<td id="76A58177FFE369ACE3962685FA257000" box="[1237,1427,511,532]" gridcol="5" gridrow="7" pageId="7" pageNumber="48">3.2</td>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D4255DFA25704E" box="[151,1427,551,602]" gridrow="8" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D4255DFEF4704E" box="[151,322,551,602]" gridcol="0" gridrow="8" pageId="7" pageNumber="48">Rostral (width & height)</th>
|
||
<td id="76A58177FFE369ACE637255DFD84704E" box="[372,562,551,602]" gridcol="1" gridrow="8" pageId="7" pageNumber="48">2.9 & 0.8</td>
|
||
<td id="76A58177FFE369ACE50F255DFCBC704E" box="[588,778,551,602]" gridcol="2" gridrow="8" pageId="7" pageNumber="48">2.7 & 1.2</td>
|
||
<td id="76A58177FFE369ACE467255DFC54704E" box="[804,994,551,602]" gridcol="3" gridrow="8" pageId="7" pageNumber="48">1.9 & 0.8</td>
|
||
<td id="76A58177FFE369ACE4BE255DFB0C704E" box="[1021,1210,551,602]" gridcol="4" gridrow="8" pageId="7" pageNumber="48">na</td>
|
||
<td id="76A58177FFE369ACE396255DFA25704E" box="[1237,1427,551,602]" gridcol="5" gridrow="8" pageId="7" pageNumber="48">2.6 & 1.0</td>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D42517FA257096" box="[151,1427,621,642]" gridrow="9" pageId="7" pageNumber="48" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
|
||
<th id="76A58177FFE369ACE7D42517FEF47096" box="[151,322,621,642]" gridcol="0" gridrow="9" pageId="7" pageNumber="48">Scalation</th>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D425EFFA2570BE" box="[151,1427,661,682]" gridrow="10" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D425EFFEF470BE" box="[151,322,661,682]" gridcol="0" gridrow="10" pageId="7" pageNumber="48">Ventrals</th>
|
||
<td id="76A58177FFE369ACE63725EFFD8470BE" box="[372,562,661,682]" gridcol="1" gridrow="10" pageId="7" pageNumber="48">132</td>
|
||
<td id="76A58177FFE369ACE50F25EFFCBC70BE" box="[588,778,661,682]" gridcol="2" gridrow="10" pageId="7" pageNumber="48">135</td>
|
||
<td id="76A58177FFE369ACE46725EFFC5470BE" box="[804,994,661,682]" gridcol="3" gridrow="10" pageId="7" pageNumber="48">137</td>
|
||
<td id="76A58177FFE369ACE4BE25EFFB0C70BE" box="[1021,1210,661,682]" gridcol="4" gridrow="10" pageId="7" pageNumber="48">135</td>
|
||
<td id="76A58177FFE369ACE39625EFFA2570BE" box="[1237,1427,661,682]" gridcol="5" gridrow="10" pageId="7" pageNumber="48">128</td>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D425C7FA2570C6" box="[151,1427,701,722]" gridrow="11" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D425C7FEF470C6" box="[151,322,701,722]" gridcol="0" gridrow="11" pageId="7" pageNumber="48">Subcaudals *</th>
|
||
<td id="76A58177FFE369ACE63725C7FD8470C6" box="[372,562,701,722]" gridcol="1" gridrow="11" pageId="7" pageNumber="48">47</td>
|
||
<td id="76A58177FFE369ACE50F25C7FCBC70C6" box="[588,778,701,722]" gridcol="2" gridrow="11" pageId="7" pageNumber="48">40</td>
|
||
<td id="76A58177FFE369ACE46725C7FC5470C6" box="[804,994,701,722]" gridcol="3" gridrow="11" pageId="7" pageNumber="48">39</td>
|
||
<td id="76A58177FFE369ACE4BE25C7FB0C70C6" box="[1021,1210,701,722]" gridcol="4" gridrow="11" pageId="7" pageNumber="48">43</td>
|
||
<td id="76A58177FFE369ACE39625C7FA2570C6" box="[1237,1427,701,722]" gridcol="5" gridrow="11" pageId="7" pageNumber="48">41</td>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D4259FFA2570EE" box="[151,1427,741,762]" gridrow="12" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D4259FFEF470EE" box="[151,322,741,762]" gridcol="0" gridrow="12" pageId="7" pageNumber="48">ASR</th>
|
||
<td id="76A58177FFE369ACE637259FFD8470EE" box="[372,562,741,762]" gridcol="1" gridrow="12" pageId="7" pageNumber="48">20</td>
|
||
<td id="76A58177FFE369ACE50F259FFCBC70EE" box="[588,778,741,762]" gridcol="2" gridrow="12" pageId="7" pageNumber="48">20</td>
|
||
<td id="76A58177FFE369ACE467259FFC5470EE" box="[804,994,741,762]" gridcol="3" gridrow="12" pageId="7" pageNumber="48">20</td>
|
||
<td id="76A58177FFE369ACE4BE259FFB0C70EE" box="[1021,1210,741,762]" gridcol="4" gridrow="12" pageId="7" pageNumber="48">21</td>
|
||
<td id="76A58177FFE369ACE396259FFA2570EE" box="[1237,1427,741,762]" gridcol="5" gridrow="12" pageId="7" pageNumber="48">21</td>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D42477FA257136" box="[151,1427,781,802]" gridrow="13" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D42477FEF47136" box="[151,322,781,802]" gridcol="0" gridrow="13" pageId="7" pageNumber="48">MSR</th>
|
||
<td id="76A58177FFE369ACE6372477FD847136" box="[372,562,781,802]" gridcol="1" gridrow="13" pageId="7" pageNumber="48">25/26</td>
|
||
<td id="76A58177FFE369ACE50F2477FCBC7136" box="[588,778,781,802]" gridcol="2" gridrow="13" pageId="7" pageNumber="48">23</td>
|
||
<td id="76A58177FFE369ACE4672477FC547136" box="[804,994,781,802]" gridcol="3" gridrow="13" pageId="7" pageNumber="48">23/24</td>
|
||
<td id="76A58177FFE369ACE4BE2477FB0C7136" box="[1021,1210,781,802]" gridcol="4" gridrow="13" pageId="7" pageNumber="48">23</td>
|
||
<td id="76A58177FFE369ACE3962477FA257136" box="[1237,1427,781,802]" gridcol="5" gridrow="13" pageId="7" pageNumber="48">22</td>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D4244FFA25715E" box="[151,1427,821,842]" gridrow="14" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D4244FFEF4715E" box="[151,322,821,842]" gridcol="0" gridrow="14" pageId="7" pageNumber="48">PSR</th>
|
||
<td id="76A58177FFE369ACE637244FFD84715E" box="[372,562,821,842]" gridcol="1" gridrow="14" pageId="7" pageNumber="48">17</td>
|
||
<td id="76A58177FFE369ACE50F244FFCBC715E" box="[588,778,821,842]" gridcol="2" gridrow="14" pageId="7" pageNumber="48">17</td>
|
||
<td id="76A58177FFE369ACE467244FFC54715E" box="[804,994,821,842]" gridcol="3" gridrow="14" pageId="7" pageNumber="48">18</td>
|
||
<td id="76A58177FFE369ACE4BE244FFB0C715E" box="[1021,1210,821,842]" gridcol="4" gridrow="14" pageId="7" pageNumber="48">18</td>
|
||
<td id="76A58177FFE369ACE396244FFA25715E" box="[1237,1427,821,842]" gridcol="5" gridrow="14" pageId="7" pageNumber="48">17</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BFBEA49FFE39654E7DC2427FB5B7660" pageId="7" pageNumber="48">
|
||
<table id="F94418E9FFE369ACE7D42427FA257686" box="[151,1427,861,1170]" gridcols="5" gridrows="6" pageId="7" pageNumber="48">
|
||
<tr id="3574E80BFFE369ACE7D42427FA25719B" box="[151,1427,861,911]" gridrow="0" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D42427FE1C719B" box="[151,426,861,911]" gridcol="0" gridrow="0" pageId="7" pageNumber="48">Circumoculars (CO) 15/15 **</th>
|
||
<th id="76A58177FFE369ACE50F2427FD35719B" box="[588,643,861,911]" gridcol="1" gridrow="0" pageId="7" pageNumber="48">15/16</th>
|
||
<th id="76A58177FFE369ACE4672427FCED719B" box="[804,859,861,911]" gridcol="2" gridrow="0" pageId="7" pageNumber="48">16/16</th>
|
||
<th id="76A58177FFE369ACE4BE2427FB85719B" box="[1021,1075,861,911]" gridcol="3" gridrow="0" pageId="7" pageNumber="48">16/15</th>
|
||
<th id="76A58177FFE369ACE3962427FA25719B" box="[1237,1427,861,911]" gridcol="4" gridrow="0" pageId="7" pageNumber="48">16/14</th>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D424D8FA2571A3" box="[151,1427,930,951]" gridrow="1" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D424D8FE1C71A3" box="[151,426,930,951]" gridcol="0" gridrow="1" pageId="7" pageNumber="48">Suprarostrals 3</th>
|
||
<td id="76A58177FFE369ACE50F24D8FD3571A3" box="[588,643,930,951]" gridcol="1" gridrow="1" pageId="7" pageNumber="48">3</td>
|
||
<td id="76A58177FFE369ACE46724D8FCED71A3" box="[804,859,930,951]" gridcol="2" gridrow="1" pageId="7" pageNumber="48">3</td>
|
||
<td id="76A58177FFE369ACE4BE24D8FB8571A3" box="[1021,1075,930,951]" gridcol="3" gridrow="1" pageId="7" pageNumber="48">4</td>
|
||
<td id="76A58177FFE369ACE39624D8FA2571A3" box="[1237,1427,930,951]" gridcol="4" gridrow="1" pageId="7" pageNumber="48">3</td>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D424B0FA2571CB" box="[151,1427,970,991]" gridrow="2" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D424B0FE1C71CB" box="[151,426,970,991]" gridcol="0" gridrow="2" pageId="7" pageNumber="48">Supralabials 8/8</th>
|
||
<td id="76A58177FFE369ACE50F24B0FD3571CB" box="[588,643,970,991]" gridcol="1" gridrow="2" pageId="7" pageNumber="48">8/8</td>
|
||
<td id="76A58177FFE369ACE46724B0FCED71CB" box="[804,859,970,991]" gridcol="2" gridrow="2" pageId="7" pageNumber="48">7/8</td>
|
||
<td id="76A58177FFE369ACE4BE24B0FB8571CB" box="[1021,1075,970,991]" gridcol="3" gridrow="2" pageId="7" pageNumber="48">9/8</td>
|
||
<td id="76A58177FFE369ACE39624B0FA2571CB" box="[1237,1427,970,991]" gridcol="4" gridrow="2" pageId="7" pageNumber="48">8/8</td>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D42488FA257613" box="[151,1427,1010,1031]" gridrow="3" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D42488FE1C7613" box="[151,426,1010,1031]" gridcol="0" gridrow="3" pageId="7" pageNumber="48">Infralabials 8/9</th>
|
||
<td id="76A58177FFE369ACE50F2488FD357613" box="[588,643,1010,1031]" gridcol="1" gridrow="3" pageId="7" pageNumber="48">9/10</td>
|
||
<td id="76A58177FFE369ACE4672488FCED7613" box="[804,859,1010,1031]" gridcol="2" gridrow="3" pageId="7" pageNumber="48">10/8</td>
|
||
<td id="76A58177FFE369ACE4BE2488FB857613" box="[1021,1075,1010,1031]" gridcol="3" gridrow="3" pageId="7" pageNumber="48">10/10</td>
|
||
<td id="76A58177FFE369ACE3962488FA257613" box="[1237,1427,1010,1031]" gridcol="4" gridrow="3" pageId="7" pageNumber="48">9/9</td>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D42360FA257658" box="[151,1427,1050,1100]" gridrow="4" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D42360FE1C7658" box="[151,426,1050,1100]" gridcol="0" gridrow="4" pageId="7" pageNumber="48">Crown Scales 7 between CO</th>
|
||
<td id="76A58177FFE369ACE50F2360FD357658" box="[588,643,1050,1100]" gridcol="1" gridrow="4" pageId="7" pageNumber="48">7</td>
|
||
<td id="76A58177FFE369ACE4672360FCED7658" box="[804,859,1050,1100]" gridcol="2" gridrow="4" pageId="7" pageNumber="48">7</td>
|
||
<td id="76A58177FFE369ACE4BE2360FB857658" box="[1021,1075,1050,1100]" gridcol="3" gridrow="4" pageId="7" pageNumber="48">8</td>
|
||
<td id="76A58177FFE369ACE3962360FA257658" box="[1237,1427,1050,1100]" gridcol="4" gridrow="4" pageId="7" pageNumber="48">6</td>
|
||
</tr>
|
||
<tr id="3574E80BFFE369ACE7D42325FA257686" box="[151,1427,1119,1170]" gridrow="5" pageId="7" pageNumber="48">
|
||
<th id="76A58177FFE369ACE7D42325FE1C7686" box="[151,426,1119,1170]" gridcol="0" gridrow="5" pageId="7" pageNumber="48">Maximum number 21 of crown scales</th>
|
||
<td id="76A58177FFE369ACE50F2325FD357686" box="[588,643,1119,1170]" gridcol="1" gridrow="5" pageId="7" pageNumber="48">22</td>
|
||
<td id="76A58177FFE369ACE4672325FCED7686" box="[804,859,1119,1170]" gridcol="2" gridrow="5" pageId="7" pageNumber="48">22</td>
|
||
<td id="76A58177FFE369ACE4BE2325FB857686" box="[1021,1075,1119,1170]" gridcol="3" gridrow="5" pageId="7" pageNumber="48">21</td>
|
||
<td id="76A58177FFE369ACE3962325FA257686" box="[1237,1427,1119,1170]" gridcol="4" gridrow="5" pageId="7" pageNumber="48">19</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BFBEA49FFE39654E78523A1FCA6748B" blockId="7.[151,1437,1243,2004]" pageId="7" pageNumber="48">
|
||
<emphasis id="B930365BFFE39654E78523A1FE8676E1" bold="true" box="[198,304,1243,1269]" pageId="7" pageNumber="48">Habitat:</emphasis>
|
||
All specimens were collected among leaf litter on the forest floor in evergreen mid-altitude wet forest, from approximately
|
||
<quantity id="4CBC47ACFFE39654E6992278FD857708" box="[474,563,1282,1308]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="48" unit="m" value="1000.0">1000 m</quantity>
|
||
a.s.l. on Mount Mabu (
|
||
<figureCitation id="137FF6CCFFE39654E40B2278FC387708" box="[840,910,1282,1308]" captionStart="FIGURE 8" captionStartId="11.[151,256,1006,1030]" captionTargetBox="[195,1399,266,982]" captionTargetId="figure@11.[195,1401,176,982]" captionTargetPageId="11" captionText="FIGURE 8. General view of closed-canopy, mid-altitude forest habitat on Mount Mabue, northern Mozambique. The central valley includes the main camp from whose surroundings the type series of Atheris mabuensis were collected (photograph J. Bayliss)." httpUri="https://zenodo.org/record/274885/files/figure.png" pageId="7" pageNumber="48">Fig. 8</figureCitation>
|
||
) and at
|
||
<quantity id="4CBC47ACFFE39654E4AD2278FBF17708" box="[1006,1095,1282,1308]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="7" pageNumber="48" unit="m" value="1550.0">1550 m</quantity>
|
||
a.s.l. on Mount Namuli. The presence of the species at a higher altitude on Mount Namuli may be due to the loss of mid-altitude forest habitat on the lower slopes as a result of subsistence farming. A similar displacement of forest birds to higher montane forests on Mount Namuli has been proposed by Dowesett- Lemaire (in Jackson
|
||
<emphasis id="B930365BFFE39654E388220DFABE7784" box="[1227,1288,1399,1424]" italics="true" pageId="7" pageNumber="48">et al.</emphasis>
|
||
, 2009). The habitat comprises closed-canopy forest (
|
||
<figureCitation id="137FF6CCFFE39654E53B22E7FD7777A3" box="[632,705,1437,1463]" captionStart="FIGURE 9" captionStartId="11.[151,255,1976,2000]" captionTargetBox="[197,1403,1153,1952]" captionTargetId="figure@11.[197,1403,1153,1953]" captionTargetPageId="11" captionText="FIGURE 9. Closed-canopy forest habitat around the main camp, Mount Mabu, northern Mozambique. The type series of Atheris mabuensis was collected in leaf litter on the forest floor (photograph J. Bayliss)." httpUri="https://zenodo.org/record/274886/files/figure.png" pageId="7" pageNumber="48">Fig. 9</figureCitation>
|
||
), except for small gaps caused by tree-falls and along stream gullies. Tall trees (up to
|
||
<quantity id="4CBC47ACFFE39654E6F822BEFD9277CA" box="[443,548,1476,1502]" metricMagnitude="1" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="7" pageNumber="48" unit="m" value="42.5" valueMax="45.0" valueMin="40.0">40–45 m</quantity>
|
||
height) emerge from the canopy, with
|
||
<taxonomicName id="4C4491CAFFE39654E4B622BEFB4777C9" box="[1013,1265,1476,1501]" class="Magnoliopsida" family="Strombosiaceae" genus="Strombosia" kingdom="Plantae" order="Santalales" pageId="7" pageNumber="48" phylum="Tracheophyta" rank="species" species="scheffleri">
|
||
<emphasis id="B930365BFFE39654E4B622BEFB4777C9" box="[1013,1265,1476,1501]" italics="true" pageId="7" pageNumber="48">Strombosia scheffleri</emphasis>
|
||
</taxonomicName>
|
||
dominant and others including
|
||
<taxonomicName id="4C4491CAFFE39654E6262291FE6F7410" box="[357,473,1515,1540]" class="Aves" family="Vangidae" genus="Newtonia" kingdom="Animalia" order="Passeriformes" pageId="7" pageNumber="48" phylum="Chordata" rank="genus">
|
||
<emphasis id="B930365BFFE39654E6262291FE6F7410" box="[357,473,1515,1540]" italics="true" pageId="7" pageNumber="48">Newtonia</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<taxonomicName id="4C4491CAFFE39654E5562291FC3D7410" box="[533,907,1515,1540]" class="Magnoliopsida" family="Sapotaceae" genus="Chrysophyllum" kingdom="Plantae" order="Ericales" pageId="7" pageNumber="48" phylum="Tracheophyta" rank="species" species="gorungosanum">
|
||
<emphasis id="B930365BFFE39654E5562291FC3D7410" box="[533,907,1515,1540]" italics="true" pageId="7" pageNumber="48">Chrysophyllum gorungosanum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B930365BFFE39654E4DF2291FED3743E" italics="true" pageId="7" pageNumber="48">
|
||
<taxonomicName id="4C4491CAFFE39654E4DF2291FAB97410" authority="Ficus" authorityName="Ficus" box="[924,1295,1515,1540]" class="Magnoliopsida" family="Chrysobalanaceae" genus="Maranthes" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="48" phylum="Tracheophyta" rank="species" species="goetzeniana">Maranthes goetzeniana, Ficus</taxonomicName>
|
||
thonningii, Blighia unijugata
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="4C4491CAFFE39654E6E4216BFD33743E" box="[423,645,1553,1578]" class="Magnoliopsida" family="Apocynaceae" genus="Funtumia" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="48" phylum="Tracheophyta" rank="species" species="africana">
|
||
<emphasis id="B930365BFFE39654E6E4216BFD33743E" box="[423,645,1553,1578]" italics="true" pageId="7" pageNumber="48">Funtumia africana</emphasis>
|
||
</taxonomicName>
|
||
. Smaller trees in the understorey include
|
||
<taxonomicName id="4C4491CAFFE39654E331216BFB59743E" box="[1138,1263,1553,1578]" class="Magnoliopsida" family="Sapindaceae" genus="Allophylus" kingdom="Animalia" order="Sapindales" pageId="7" pageNumber="48" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B930365BFFE39654E331216BFB59743E" box="[1138,1263,1553,1578]" italics="true" pageId="7" pageNumber="48">Allophylus</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<taxonomicName id="4C4491CAFFE39654E26B216BFA2A743E" box="[1320,1436,1553,1578]" class="Magnoliopsida" family="Rubiaceae" genus="Canthium" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="48" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B930365BFFE39654E26B216BFA2A743E" box="[1320,1436,1553,1578]" italics="true" pageId="7" pageNumber="48">Canthium</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<emphasis id="B930365BFFE39654E7842142FB207445" box="[199,1174,1592,1617]" italics="true" pageId="7" pageNumber="48">
|
||
<taxonomicName id="4C4491CAFFE39654E7842142FD887445" authority="Rawsonia" authorityName="Rawsonia" box="[199,574,1592,1617]" class="Magnoliopsida" family="Rubiaceae" genus="Oxyanthus" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="48" phylum="Tracheophyta" rank="species" species="speciosus">Oxyanthus speciosus, Rawsonia</taxonomicName>
|
||
lucida, Tabernaemontana ventricosa, Vepris stolzii
|
||
</emphasis>
|
||
, and small saplings of
|
||
<taxonomicName id="4C4491CAFFE39654E7D42125FE54746C" authority="Drypetes" authorityName="Drypetes" box="[151,482,1631,1656]" class="Insecta" family="Noctuidae" genus="Cola" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="greenwayi">
|
||
<emphasis id="B930365BFFE39654E7D42125FE54746C" box="[151,482,1631,1656]" italics="true" pageId="7" pageNumber="48">Cola greenwayi, Drypetes</emphasis>
|
||
</taxonomicName>
|
||
sp. and
|
||
<taxonomicName id="4C4491CAFFE39654E50D2125FD60746C" box="[590,726,1631,1656]" class="Magnoliopsida" family="Chrysobalanaceae" genus="Maranthes" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="48" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B930365BFFE39654E50D2125FD60746C" box="[590,726,1631,1656]" italics="true" pageId="7" pageNumber="48">Maranthes</emphasis>
|
||
</taxonomicName>
|
||
sp. Canopy lianas are dominated by
|
||
<taxonomicName id="4C4491CAFFE39654E3F32125FA2B746C" box="[1200,1437,1631,1656]" class="Globothalamea" family="Millettiidae" genus="Millettia" kingdom="Chromista" order="Rotaliida" pageId="7" pageNumber="48" phylum="Foraminifera" rank="species" species="lasiantha">
|
||
<emphasis id="B930365BFFE39654E3F32125FA2B746C" box="[1200,1437,1631,1656]" italics="true" pageId="7" pageNumber="48">Millettia lasiantha</emphasis>
|
||
</taxonomicName>
|
||
(Dowsett-Lemaire and Dowsett, unpubl. report 2008).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35EB9C2FFE3965BE78521D1FA90779D" lastPageId="8" lastPageNumber="49" pageId="7" pageNumber="48" type="description">
|
||
<paragraph id="8BFBEA49FFE39654E78521D1FBBD75C0" blockId="7.[151,1437,1243,2004]" pageId="7" pageNumber="48">
|
||
<emphasis id="B930365BFFE39654E78521D1FE2174D1" bold="true" box="[198,407,1707,1733]" pageId="7" pageNumber="48">Natural History:</emphasis>
|
||
The original specimen (PEM R17910) was maintained in captivity for over 37 months. The other specimens were kept for shorter periods (2–4 months). During this period they readily accepted small geckos (
|
||
<taxonomicName id="4C4491CAFFE39654E6062183FDE57506" box="[325,595,1785,1810]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis id="B930365BFFE39654E6062183FDE57506" box="[325,595,1785,1810]" italics="true" pageId="7" pageNumber="48">Lygodactylus capensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4491CAFFE39654E5CD2183FC157506" box="[654,931,1785,1810]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="mabouia">
|
||
<emphasis id="B930365BFFE39654E5CD2183FC157506" box="[654,931,1785,1810]" italics="true" pageId="7" pageNumber="48">Hemidactylus mabouia</emphasis>
|
||
</taxonomicName>
|
||
) as well as small amphibians (
|
||
<taxonomicName id="4C4491CAFFE39654E25B2183FA2A7506" box="[1304,1436,1785,1810]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="genus">
|
||
<emphasis id="B930365BFFE39654E25B2183FA2A7506" box="[1304,1436,1785,1810]" italics="true" pageId="7" pageNumber="48">Hyperolius</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<taxonomicName id="4C4491CAFFE39654E789205AFEDF752D" box="[202,361,1824,1849]" class="Amphibia" family="Pyxicephalidae" genus="Strongylopus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="genus">
|
||
<emphasis id="B930365BFFE39654E789205AFEDF752D" box="[202,361,1824,1849]" italics="true" pageId="7" pageNumber="48">Strongylopus</emphasis>
|
||
</taxonomicName>
|
||
sp., and
|
||
<taxonomicName id="4C4491CAFFE39654E699205AFD18752D" box="[474,686,1824,1849]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="genus">
|
||
<emphasis id="B930365BFFE39654E699205AFD18752D" box="[474,686,1824,1849]" italics="true" pageId="7" pageNumber="48">Phrynobatrachus</emphasis>
|
||
</taxonomicName>
|
||
sp.). While in captivity they were observed to use the bright yellow tail as a caudal lure to entice prey into striking distance. In a terrarium with leaf litter and small branches they either coiled partially concealed among large leaves on the floor, or in dead twigs or leaf clumps up to
|
||
<quantity id="4CBC47ACFFE39654E79D20EEFE9675BA" box="[222,288,1940,1966]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="48" unit="cm" value="20.0">20cm</quantity>
|
||
above the floor. The only adult female (PEM R17902) has thick muscular oviducts and small vitellogenic ovarian follicles indicating sexual maturity at only
|
||
<quantity id="4CBC47ACFFE39654E43D20C0FC6175C0" box="[894,983,1978,2004]" metricMagnitude="-1" metricUnit="m" metricValue="3.84" pageId="7" pageNumber="48" unit="mm" value="384.0">384mm</quantity>
|
||
TL.
|
||
</paragraph>
|
||
<caption id="DF3BBAC1FFEC965BE7D424ACFF687626" httpUri="https://zenodo.org/record/274882/files/figure.png" pageId="8" pageNumber="49" targetBox="[151,1435,194,957]" targetPageId="8">
|
||
<paragraph id="8BFBEA49FFEC965BE7D424ACFF687626" blockId="8.[151,1436,982,1074]" pageId="8" pageNumber="49">
|
||
<emphasis id="B930365BFFEC965BE7D424ACFEAF71FA" bold="true" box="[151,281,982,1006]" pageId="8" pageNumber="49">FIGURE 5.</emphasis>
|
||
Midbody, right lateral view of
|
||
<taxonomicName id="4C4491CAFFEC965BE52124ACFC9371F9" box="[610,805,982,1005]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="49" phylum="Chordata" rank="species" species="mabuensis" status="sp. nov.">
|
||
<emphasis id="B930365BFFEC965BE52124ACFC9371F9" box="[610,805,982,1005]" italics="true" pageId="8" pageNumber="49">Atheris mabuensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B930365BFFEC965BE46F24ACFCD171FA" bold="true" box="[812,871,982,1006]" pageId="8" pageNumber="49">
|
||
<taxonomicNameLabel id="A2038B20FFEC965BE46F24ACFCD171FA" box="[812,871,982,1006]" pageId="8" pageNumber="49" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Holotype PEM R17901 Mount Mabu): note oblique orientation of lateral scale rows 1–5, and the progressive asymmetry in the shape and lack of serrated keels on these scales.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFBEA49FFEC965BE785231BFA90779D" blockId="8.[151,1437,1121,1417]" pageId="8" pageNumber="49">
|
||
The species was locally common within the large tract of wet forest (>
|
||
<quantity id="4CBC47ACFFEC965BE4BB231BFBFA766F" box="[1016,1100,1121,1147]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="49" unit="m" value="1000.0">1000m</quantity>
|
||
) on Mount Mabu, and it was only observed within leaf litter on the forest floor (
|
||
<figureCitation id="137FF6CCFFEC965BE5AA23FDFC9876B5" box="[745,814,1159,1185]" captionStart="FIGURE 6" captionStartId="9.[151,258,1869,1893]" captionTargetBox="[152,1433,194,1846]" captionTargetId="figure@9.[152,1435,194,1846]" captionTargetPageId="9" captionText="FIGURE 6. Atheris mabuensis n. sp. (PEM R 17910): Top - small juvenile (<150 mm TL) photographed in situ on capture (25 January 2006, Julian Bayliss); note drab colouration imparting camouflage in habitat among leaf litter on forest floor. Bottom – same specimen (photographed 19 Nov 2008, after 34 months in captivity); note retention of juvenile colouration, particularly yellow tail tip, and development of gold tips to the keels on the head and forebody scales." httpUri="https://zenodo.org/record/274883/files/figure.png" pageId="8" pageNumber="49">Fig. 6</figureCitation>
|
||
top). Local hunters when interviewed confirmed this habitat preference, and also noted that the species remained small, growing to not much more than
|
||
<quantity id="4CBC47ACFFEC965BE27023D4FACB76DC" box="[1331,1405,1198,1224]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="49" unit="cm" value="35.0">35 cm</quantity>
|
||
in length. This observation was supported by the growth of the first specimen, which measured approximately
|
||
<quantity id="4CBC47ACFFEC965BE7D42381FF4A7701" box="[151,252,1275,1301]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="49" unit="mm" value="150.0">150 mm</quantity>
|
||
TL when first captured, and grew to only
|
||
<quantity id="4CBC47ACFFEC965BE44B2381FCD87701" box="[776,878,1275,1301]" metricMagnitude="-1" metricUnit="m" metricValue="3.67" pageId="8" pageNumber="49" unit="mm" value="367.0">367 mm</quantity>
|
||
TL after over three years in captivity despite regular feeding. Local hunters also noted that bites from the species were painful but not lethal. Caution is still advised because serious envenomation has followed bites from
|
||
<taxonomicName id="4C4491CAFFEC965BE4CA2233FB937776" box="[905,1061,1353,1378]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="49" phylum="Chordata" rank="species" species="chlorechis">
|
||
<emphasis id="B930365BFFEC965BE4CA2233FB937776" box="[905,1061,1353,1378]" italics="true" pageId="8" pageNumber="49">A. chlorechis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFD597B8FFEC965BE3772233FB7C7777" author="Rodel" box="[1076,1226,1353,1379]" pageId="8" pageNumber="54" refString="Rodel, M-O. (1999). Uber die Folgen eines Bisses von Atheris chlorechis (Pel, 1851). Herpetofauna, 21 (123), 21 - 24." type="journal article" year="1999">Rödel, 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD597B8FFEC965BE3942233FA257777" author="Top" box="[1239,1427,1353,1379]" pageId="8" pageNumber="54" refString="Top, L. J., Tulleken, J. E., Ligtenberg, J. J. M., Meertens, J. H. J. M., van der Werf, T. S. & Zijlstra, J. G. (2006). Serious envenomation after a snakebite by a Western bush viper (Atheris chlorechis) in the Netherlands: a case report. The Netherlands Journal of Medicine, 64 (5), 153 - 156." type="journal article" year="2006">
|
||
Top
|
||
<emphasis id="B930365BFFEC965BE24F2233FAF77776" box="[1292,1345,1353,1378]" italics="true" pageId="8" pageNumber="49">et al</emphasis>
|
||
., 2006
|
||
</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="4C4491CAFFEC965BE78B220AFDEB779D" authority="Mebs et al., 1998" authorityName="Mebs et al." authorityYear="1998" box="[200,605,1391,1417]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="49" phylum="Chordata" rank="species" species="squamigera">
|
||
<emphasis id="B930365BFFEC965BE78B220AFEC5779D" box="[200,371,1392,1417]" italics="true" pageId="8" pageNumber="49">A. squamigera</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD597B8FFEC965BE6C02215FDE2779D" author="Mebs" box="[387,596,1391,1417]" pageId="8" pageNumber="54" refString="Mebs, D., Holada, K., Kornalik, F., Simak, J., Vankova, H., Muller, D., Schoenemann, H., Lange, H. & Herrmann, H. (1998). Severe coagulopathy after a bite of a green bush viper (Atheris squamiger): case report and biochemical analysis of the venom. Toxicon, 36 (10), 1333 - 1340." type="journal article" year="1998">
|
||
Mebs
|
||
<emphasis id="B930365BFFEC965BE68E220AFDB1779D" box="[461,519,1392,1417]" italics="true" pageId="8" pageNumber="49">et al.</emphasis>
|
||
, 1998
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and even a death from the latter (
|
||
<bibRefCitation id="EFD597B8FFEC965BE4A82215FAAC779D" author="Lanoie" box="[1003,1306,1391,1417]" pageId="8" pageNumber="54" refString="Lanoie, L. O. and Branch, W. R. (1991). Atheris squamiger: Fatal Envenomation. Journal of the Herpetological Association of Africa, 39, 29." type="journal article" year="1991">Lanoie and Branch, 1991</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |