218 lines
41 KiB
XML
218 lines
41 KiB
XML
<document id="878229A91D246BC5D6E3AEFDAA6E63F6" ID-CLB-Dataset="24935" ID-DOI="10.11646/zootaxa.4695.3.1" ID-GBIF-Dataset="256591b2-6d7b-465d-a818-9a158017ca09" ID-ISSN="1175-5326" ID-Zenodo-Dep="3532515" ID-ZooBank="32E17629-F29A-47C8-ACCB-69A32C17B147" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573197328991" checkinUser="plazi" docAuthor="Gunter, Nicole L. & Weir, Thomas A." docDate="2019" docId="03ED0167FF86FFBEFF1DF1A0FA01F968" docLanguage="en" docName="zootaxa.4695.3.1.pdf" docOrigin="Zootaxa 4695 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepanus australis Matthews 1974" docType="key" docVersion="9" lastPageNumber="204" masterDocId="FFD4791FFF85FFBDFF8AF379FF9DFFA8" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups" masterLastPageNumber="252" masterPageNumber="201" pageNumber="204" updateTime="1698764404900" updateUser="ExternalLinkService">
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<mods:title id="C8D59E7E48BF23321454D0458A8AD69B">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups</mods:title>
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<mods:namePart id="1DE9E3ADC050E7568A3E0B7697E575B4">Gunter, Nicole L.</mods:namePart>
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<mods:namePart id="425037F5A1D90E209D1141AEE48C0D5B">Weir, Thomas A.</mods:namePart>
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<mods:title id="D3B1BF3426D6BBBF1AE65C00780F8158">Zootaxa</mods:title>
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<mods:date id="2BBAD155090C42142F09EC314737FCC9">2019</mods:date>
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<mods:number id="EF579CF5E0A8441D5D49B1158D93E24B">2019-11-08</mods:number>
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<mods:number id="607AF19248D794FB56C934E672C6D960">4695</mods:number>
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<mods:identifier id="3ACD38CCE983238192C379D52D213432" type="DOI">10.11646/zootaxa.4695.3.1</mods:identifier>
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<treatment id="03ED0167FF86FFBEFF1DF1A0FA01F968" ID-GBIF-Taxon="160218009" LSID="urn:lsid:plazi:treatment:03ED0167FF86FFBEFF1DF1A0FA01F968" httpUri="http://treatment.plazi.org/id/03ED0167FF86FFBEFF1DF1A0FA01F968" lastPageNumber="204" pageId="3" pageNumber="204">
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<subSubSection id="C35EE3FAFF86FFBEFF1DF1A0FD60FD5B" box="[151,765,729,756]" pageId="3" pageNumber="204" type="nomenclature">
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<paragraph id="8BFBB071FF86FFBEFF1DF1A0FD60FD5B" blockId="3.[151,765,729,756]" box="[151,765,729,756]" pageId="3" pageNumber="204">
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<heading id="D0B3071DFF86FFBEFF1DF1A0FD60FD5B" bold="true" box="[151,765,729,756]" fontSize="11" level="1" pageId="3" pageNumber="204" reason="1">
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<emphasis id="B9306C63FF86FFBEFF1DF1A0FD60FD5B" bold="true" box="[151,765,729,756]" pageId="3" pageNumber="204">
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Key to species of
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<taxonomicName id="4C44CBF2FF86FFBEFEE4F1A0FDDBFD5C" ID-CoL="6P7WD" authorityName="Matthews" authorityYear="1974" box="[366,582,729,756]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="australis">
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<emphasis id="B9306C63FF86FFBEFEE4F1A0FDDBFD5C" bold="true" box="[366,582,729,756]" italics="true" pageId="3" pageNumber="204">Lepanus australis</emphasis>
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</taxonomicName>
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species group.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35EE3FAFF86FFBEFF1DF067FA01F968" pageId="3" pageNumber="204" type="key">
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<keyStep id="30B0A8D4FF86FFBEFF1DF067FA01FC00" pageId="3" pageNumber="204">
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<paragraph id="8BFBB071FF86FFBEFF1DF067FA01FCC6" blockId="3.[151,1437,797,1729]" pageId="3" pageNumber="204">
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<keyLead id="30B51344FF86FFBEFF1DF067FA01FCC6" pageId="3" pageNumber="204">
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1. Front edge of protibiae truncate (
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<figureCitation id="137FACF4FF86FFBEFD91F064FDCDFC9C" box="[539,592,797,820]" captionStart="FIGURE 1" captionStartId="9.[151,250,188,213]" captionTargetId="figure@8.[242,1345,181,1957]" captionTargetPageId="8" captionText="FIGURE 1. Some diagnostic characters of the Lepanus australis species group. A, Male protibia of Lepanus eungella dotted line depicts triangular apical digit (grey arrow) and truncate front edge; B, female protibia of Lepanus eungella dotted line depicts truncate front edge, black arrow points to apical spur; C, male protibia of Lepanus australis dotted line depicts elongate apical digit (grey arrow) and deeply angulate front edge; D, female protibia of Lepanus australis dotted line depicts shortened apical digit (grey arrow) and deeply angulate front edge, black arrow points to apical spur; E, lateral view of Lepanus dukungarri white arrow points to row of punctures on metanepisternum; F, lateral view of Lepanus australis, dotted line indicates anteriorly straight inner border of lateral lobe of metaventrite, note the absence of puntures on metanepisternum and straight lower edge of elytral epipleura; G, lateral view of Lepanus oxleyi white arrow points to row of fine punctures on metanepisternum, grey arrow inducates curved lower edge of elytral epipleura; H, male metafemora of Lepanus dukungarri showing medial expansion; I, Head of Lepanus oxleyi white arrow points to interrupted basal carina; J, lateral view of Lepanus latheticus dotted line indicates anteriorly curved inner border of lateral lobe of metaventrite; K, Lepanus australis at bird droppings (image: Geoff Williams)." figureDoi="http://doi.org/10.5281/zenodo.3611235" httpUri="https://zenodo.org/record/3611235/files/figure.png" pageId="3" pageNumber="204">Fig. 1</figureCitation>
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A–B), apical digit short and triangular in males and absent in females, apical spur absent in males and present in females where it arises directly from the front edge; central
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<collectingRegion id="49807E93FF86FFBEFB8EF042FBE5FCF9" box="[1028,1144,827,849]" country="Australia" name="Queensland" pageId="3" pageNumber="204">Queensland</collectingRegion>
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, Eungella area..........................................................................
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<taxonomicName id="4C44CBF2FF86FFBEFC52F020FABDFCC6" ID-CoL="6P87Y" authority="Gunter & Weir" authorityName="Gunter & Weir" authorityYear="2019" box="[984,1312,856,878]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="eungella" status="sp. nov.">
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<emphasis id="B9306C63FF86FFBEFC52F020FB14FCC6" bold="true" box="[984,1161,856,878]" italics="true" pageId="3" pageNumber="204">Lepanus eungella</emphasis>
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Gunter & Weir
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</taxonomicName>
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,
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<taxonomicNameLabel id="A203D118FF86FFBEFAA3F020FA01FCC6" box="[1321,1436,856,878]" pageId="3" pageNumber="204" rank="species">new species</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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<paragraph id="8BFBB071FF86FFBEFF1DF00FFA01FC00" blockId="3.[151,1437,797,1729]" pageId="3" pageNumber="204">
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<keyLead id="30B51344FF86FFBEFF1DF00FFA01FC00" pageId="3" pageNumber="204">
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- Front edge of protibiae deeply angulate (
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<figureCitation id="137FACF4FF86FFBEFDFFF00CFD36FC24" box="[629,683,885,908]" captionStart="FIGURE 1" captionStartId="9.[151,250,188,213]" captionTargetId="figure@8.[242,1345,181,1957]" captionTargetPageId="8" captionText="FIGURE 1. Some diagnostic characters of the Lepanus australis species group. A, Male protibia of Lepanus eungella dotted line depicts triangular apical digit (grey arrow) and truncate front edge; B, female protibia of Lepanus eungella dotted line depicts truncate front edge, black arrow points to apical spur; C, male protibia of Lepanus australis dotted line depicts elongate apical digit (grey arrow) and deeply angulate front edge; D, female protibia of Lepanus australis dotted line depicts shortened apical digit (grey arrow) and deeply angulate front edge, black arrow points to apical spur; E, lateral view of Lepanus dukungarri white arrow points to row of punctures on metanepisternum; F, lateral view of Lepanus australis, dotted line indicates anteriorly straight inner border of lateral lobe of metaventrite, note the absence of puntures on metanepisternum and straight lower edge of elytral epipleura; G, lateral view of Lepanus oxleyi white arrow points to row of fine punctures on metanepisternum, grey arrow inducates curved lower edge of elytral epipleura; H, male metafemora of Lepanus dukungarri showing medial expansion; I, Head of Lepanus oxleyi white arrow points to interrupted basal carina; J, lateral view of Lepanus latheticus dotted line indicates anteriorly curved inner border of lateral lobe of metaventrite; K, Lepanus australis at bird droppings (image: Geoff Williams)." figureDoi="http://doi.org/10.5281/zenodo.3611235" httpUri="https://zenodo.org/record/3611235/files/figure.png" pageId="3" pageNumber="204">Fig. 1</figureCitation>
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C–D), apical digit elongate in both males and females, apical spur absent in males and present in females where it arises from apical digit..................................................
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<emphasis id="B9306C63FF86FFBEFA1AF0EAFA01FC00" bold="true" box="[1424,1436,915,936]" pageId="3" pageNumber="204">2</emphasis>
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</keyLead>
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</paragraph>
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||
</keyStep>
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||
<keyStep id="30B0A8D4FF86FFBEFF1DF0C9FA01FBDD" pageId="3" pageNumber="204">
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||
<paragraph id="8BFBB071FF86FFBEFF1DF0C9FA01FBB6" blockId="3.[151,1437,797,1729]" pageId="3" pageNumber="204">
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<keyLead id="30B51344FF86FFBEFF1DF0C9FA01FBB6" pageId="3" pageNumber="204">
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2. Elytra with striae 1–7 impressed, appearing as double lines (
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<figureCitation id="137FACF4FF86FFBEFC94F0C9FCF4FC6E" box="[798,873,944,966]" captionStart="FIGURE 2" captionStartId="10.[151,250,1947,1972]" captionTargetBox="[311,1277,196,1910]" captionTargetId="figure@10.[283,1305,181,1922]" captionTargetPageId="10" captionText="FIGURE 2. Habitus of species within the Lepanus australis species group. A, Lepanus australis; B, Lepanus burnetti; C, Lepanus dukungarri; D, Lepanus eungella; E, Lepanus latheticus; F, Lepanus oxleyi. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532517" httpUri="https://zenodo.org/record/3532517/files/figure.png" pageId="3" pageNumber="204">Fig. 2C</figureCitation>
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); head and disc of pronotum with obvious dense punctures; metanepisternum reticulate and with a row of obvious punctures (
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<figureCitation id="137FACF4FF86FFBEFCE3F0B4FC28FC4C" box="[873,949,973,996]" captionStart="FIGURE 1" captionStartId="9.[151,250,188,213]" captionTargetId="figure@8.[242,1345,181,1957]" captionTargetPageId="8" captionText="FIGURE 1. Some diagnostic characters of the Lepanus australis species group. A, Male protibia of Lepanus eungella dotted line depicts triangular apical digit (grey arrow) and truncate front edge; B, female protibia of Lepanus eungella dotted line depicts truncate front edge, black arrow points to apical spur; C, male protibia of Lepanus australis dotted line depicts elongate apical digit (grey arrow) and deeply angulate front edge; D, female protibia of Lepanus australis dotted line depicts shortened apical digit (grey arrow) and deeply angulate front edge, black arrow points to apical spur; E, lateral view of Lepanus dukungarri white arrow points to row of punctures on metanepisternum; F, lateral view of Lepanus australis, dotted line indicates anteriorly straight inner border of lateral lobe of metaventrite, note the absence of puntures on metanepisternum and straight lower edge of elytral epipleura; G, lateral view of Lepanus oxleyi white arrow points to row of fine punctures on metanepisternum, grey arrow inducates curved lower edge of elytral epipleura; H, male metafemora of Lepanus dukungarri showing medial expansion; I, Head of Lepanus oxleyi white arrow points to interrupted basal carina; J, lateral view of Lepanus latheticus dotted line indicates anteriorly curved inner border of lateral lobe of metaventrite; K, Lepanus australis at bird droppings (image: Geoff Williams)." figureDoi="http://doi.org/10.5281/zenodo.3611235" httpUri="https://zenodo.org/record/3611235/files/figure.png" pageId="3" pageNumber="204">Fig. 1E</figureCitation>
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); male metafemora expanded medially (
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<figureCitation id="137FACF4FF86FFBEFACBF0B4FA12FC4C" box="[1345,1423,973,996]" captionStart="FIGURE 1" captionStartId="9.[151,250,188,213]" captionTargetId="figure@8.[242,1345,181,1957]" captionTargetPageId="8" captionText="FIGURE 1. Some diagnostic characters of the Lepanus australis species group. A, Male protibia of Lepanus eungella dotted line depicts triangular apical digit (grey arrow) and truncate front edge; B, female protibia of Lepanus eungella dotted line depicts truncate front edge, black arrow points to apical spur; C, male protibia of Lepanus australis dotted line depicts elongate apical digit (grey arrow) and deeply angulate front edge; D, female protibia of Lepanus australis dotted line depicts shortened apical digit (grey arrow) and deeply angulate front edge, black arrow points to apical spur; E, lateral view of Lepanus dukungarri white arrow points to row of punctures on metanepisternum; F, lateral view of Lepanus australis, dotted line indicates anteriorly straight inner border of lateral lobe of metaventrite, note the absence of puntures on metanepisternum and straight lower edge of elytral epipleura; G, lateral view of Lepanus oxleyi white arrow points to row of fine punctures on metanepisternum, grey arrow inducates curved lower edge of elytral epipleura; H, male metafemora of Lepanus dukungarri showing medial expansion; I, Head of Lepanus oxleyi white arrow points to interrupted basal carina; J, lateral view of Lepanus latheticus dotted line indicates anteriorly curved inner border of lateral lobe of metaventrite; K, Lepanus australis at bird droppings (image: Geoff Williams)." figureDoi="http://doi.org/10.5281/zenodo.3611235" httpUri="https://zenodo.org/record/3611235/files/figure.png" pageId="3" pageNumber="204">Fig. 1H</figureCitation>
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); metatibiae with inner edge curved, more so in male than female; aedeagus as in
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<figureCitation id="137FACF4FF86FFBEFC68F092FBB0FBA9" box="[994,1069,1003,1025]" captionStart="FIGURE 3" captionStartId="11.[151,250,1924,1949]" captionTargetBox="[264,1312,179,1899]" captionTargetId="figure@11.[262,1324,166,1904]" captionTargetPageId="11" captionText="FIGURE 3. Left, dorsal, and right view of aedeagi of species within the Lepanus australis species group. A, Lepanus australis; B, Lepanus burnetti; C, Lepanus dukungarri; D, Lepanus eungella; E, Lepanus latheticus; F, Lepanus oxleyi. All specimens in comparative scale. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3532519" httpUri="https://zenodo.org/record/3532519/files/figure.png" pageId="3" pageNumber="204">Fig. 3C</figureCitation>
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); northern
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<collectingRegion id="49807E93FF86FFBEFB1CF092FA97FBA9" box="[1174,1290,1003,1025]" country="Australia" name="Queensland" pageId="3" pageNumber="204">Queensland</collectingRegion>
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, Wet Tropics.............................................................
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<taxonomicName id="4C44CBF2FF86FFBEFC36F770FABDFBB6" ID-CoL="6P7VY" authority="Gunter & Weir" authorityName="Gunter & Weir" authorityYear="2019" box="[956,1312,1032,1054]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="dukungarri" status="sp. nov.">
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<emphasis id="B9306C63FF86FFBEFC36F770FB14FBB6" bold="true" box="[956,1161,1032,1054]" italics="true" pageId="3" pageNumber="204">Lepanus dukungarri</emphasis>
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Gunter & Weir
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</taxonomicName>
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,
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<taxonomicNameLabel id="A203D118FF86FFBEFAA3F770FA01FBB6" box="[1321,1436,1032,1054]" pageId="3" pageNumber="204" rank="species">new species</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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<paragraph id="8BFBB071FF86FFBEFF1DF75FFA01FBDD" blockId="3.[151,1437,797,1729]" pageId="3" pageNumber="204">
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<keyLead id="30B51344FF86FFBEFF1DF75FFA01FBDD" pageId="3" pageNumber="204">
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- Elytra with striae more superficial, striae 1–3 or 1–4 appearing as double lines, striae 5–7 appearing single; head and disc of pronotum with punctures less obvious and less dense; metanepisternum either reticulate or with a row of punctures, but never both together; metafemora not expanded medially; metatibiae with inner edge almost straight.........................
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<emphasis id="B9306C63FF86FFBEFA1AF719FA01FBDD" bold="true" box="[1424,1436,1120,1141]" pageId="3" pageNumber="204">3</emphasis>
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</keyLead>
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</paragraph>
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||
</keyStep>
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<keyStep id="30B0A8D4FF86FFBEFF1DF707FA01FAEA" pageId="3" pageNumber="204">
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<paragraph id="8BFBB071FF86FFBEFF1DF707FA01FB43" blockId="3.[151,1437,797,1729]" pageId="3" pageNumber="204">
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<keyLead id="30B51344FF86FFBEFF1DF707FA01FB43" pageId="3" pageNumber="204">
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3. Basal carina of head present, interrupted in the middle (
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<figureCitation id="137FACF4FF86FFBEFD76F704FCA3FB3C" box="[764,830,1149,1172]" captionStart="FIGURE 1" captionStartId="9.[151,250,188,213]" captionTargetId="figure@8.[242,1345,181,1957]" captionTargetPageId="8" captionText="FIGURE 1. Some diagnostic characters of the Lepanus australis species group. A, Male protibia of Lepanus eungella dotted line depicts triangular apical digit (grey arrow) and truncate front edge; B, female protibia of Lepanus eungella dotted line depicts truncate front edge, black arrow points to apical spur; C, male protibia of Lepanus australis dotted line depicts elongate apical digit (grey arrow) and deeply angulate front edge; D, female protibia of Lepanus australis dotted line depicts shortened apical digit (grey arrow) and deeply angulate front edge, black arrow points to apical spur; E, lateral view of Lepanus dukungarri white arrow points to row of punctures on metanepisternum; F, lateral view of Lepanus australis, dotted line indicates anteriorly straight inner border of lateral lobe of metaventrite, note the absence of puntures on metanepisternum and straight lower edge of elytral epipleura; G, lateral view of Lepanus oxleyi white arrow points to row of fine punctures on metanepisternum, grey arrow inducates curved lower edge of elytral epipleura; H, male metafemora of Lepanus dukungarri showing medial expansion; I, Head of Lepanus oxleyi white arrow points to interrupted basal carina; J, lateral view of Lepanus latheticus dotted line indicates anteriorly curved inner border of lateral lobe of metaventrite; K, Lepanus australis at bird droppings (image: Geoff Williams)." figureDoi="http://doi.org/10.5281/zenodo.3611235" httpUri="https://zenodo.org/record/3611235/files/figure.png" pageId="3" pageNumber="204">
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Fig.
|
||
<date id="FFFA96B1FF86FFBEFCA3F707FCA3FB3C" box="[809,830,1150,1172]" pageId="3" pageNumber="204">1I</date>
|
||
</figureCitation>
|
||
); eye canthus not dividing eye; elytral epipleura not reticulate, lower edge curving upwards to meet lateral carina of elytra basally (
|
||
<figureCitation id="137FACF4FF86FFBEFCE3F7E2FC28FB18" box="[873,949,1179,1201]" captionStart="FIGURE 1" captionStartId="9.[151,250,188,213]" captionTargetId="figure@8.[242,1345,181,1957]" captionTargetPageId="8" captionText="FIGURE 1. Some diagnostic characters of the Lepanus australis species group. A, Male protibia of Lepanus eungella dotted line depicts triangular apical digit (grey arrow) and truncate front edge; B, female protibia of Lepanus eungella dotted line depicts truncate front edge, black arrow points to apical spur; C, male protibia of Lepanus australis dotted line depicts elongate apical digit (grey arrow) and deeply angulate front edge; D, female protibia of Lepanus australis dotted line depicts shortened apical digit (grey arrow) and deeply angulate front edge, black arrow points to apical spur; E, lateral view of Lepanus dukungarri white arrow points to row of punctures on metanepisternum; F, lateral view of Lepanus australis, dotted line indicates anteriorly straight inner border of lateral lobe of metaventrite, note the absence of puntures on metanepisternum and straight lower edge of elytral epipleura; G, lateral view of Lepanus oxleyi white arrow points to row of fine punctures on metanepisternum, grey arrow inducates curved lower edge of elytral epipleura; H, male metafemora of Lepanus dukungarri showing medial expansion; I, Head of Lepanus oxleyi white arrow points to interrupted basal carina; J, lateral view of Lepanus latheticus dotted line indicates anteriorly curved inner border of lateral lobe of metaventrite; K, Lepanus australis at bird droppings (image: Geoff Williams)." figureDoi="http://doi.org/10.5281/zenodo.3611235" httpUri="https://zenodo.org/record/3611235/files/figure.png" pageId="3" pageNumber="204">Fig. 1G</figureCitation>
|
||
, grey arrow); metanepisternum smooth, with a row of very fine punctures (
|
||
<figureCitation id="137FACF4FF86FFBEFE48F7C1FD93FB65" box="[450,526,1208,1230]" captionStart="FIGURE 1" captionStartId="9.[151,250,188,213]" captionTargetId="figure@8.[242,1345,181,1957]" captionTargetPageId="8" captionText="FIGURE 1. Some diagnostic characters of the Lepanus australis species group. A, Male protibia of Lepanus eungella dotted line depicts triangular apical digit (grey arrow) and truncate front edge; B, female protibia of Lepanus eungella dotted line depicts truncate front edge, black arrow points to apical spur; C, male protibia of Lepanus australis dotted line depicts elongate apical digit (grey arrow) and deeply angulate front edge; D, female protibia of Lepanus australis dotted line depicts shortened apical digit (grey arrow) and deeply angulate front edge, black arrow points to apical spur; E, lateral view of Lepanus dukungarri white arrow points to row of punctures on metanepisternum; F, lateral view of Lepanus australis, dotted line indicates anteriorly straight inner border of lateral lobe of metaventrite, note the absence of puntures on metanepisternum and straight lower edge of elytral epipleura; G, lateral view of Lepanus oxleyi white arrow points to row of fine punctures on metanepisternum, grey arrow inducates curved lower edge of elytral epipleura; H, male metafemora of Lepanus dukungarri showing medial expansion; I, Head of Lepanus oxleyi white arrow points to interrupted basal carina; J, lateral view of Lepanus latheticus dotted line indicates anteriorly curved inner border of lateral lobe of metaventrite; K, Lepanus australis at bird droppings (image: Geoff Williams)." figureDoi="http://doi.org/10.5281/zenodo.3611235" httpUri="https://zenodo.org/record/3611235/files/figure.png" pageId="3" pageNumber="204">Fig. 1G</figureCitation>
|
||
, white arrow); abdominal ventrite 6 virtually impunctate; southeastern
|
||
<collectingRegion id="49807E93FF86FFBEFB30F7C1FAB6FB66" box="[1210,1323,1208,1230]" country="Australia" name="Queensland" pageId="3" pageNumber="204">Queensland</collectingRegion>
|
||
.........................................................................
|
||
<taxonomicName id="4C44CBF2FF86FFBEFC79F7AFFABDFB43" ID-CoL="3T8V8" authority="Gunter & Weir" authorityName="Gunter & Weir" authorityYear="2019" box="[1011,1312,1237,1259]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="oxleyi" status="sp. nov.">
|
||
<emphasis id="B9306C63FF86FFBEFC79F7AFFB14FB43" bold="true" box="[1011,1161,1237,1259]" italics="true" pageId="3" pageNumber="204">Lepanus oxleyi</emphasis>
|
||
Gunter & Weir
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A203D118FF86FFBEFAA3F7AFFA01FB43" box="[1321,1436,1237,1259]" pageId="3" pageNumber="204" rank="species">new species</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BFBB071FF86FFBEFF1DF78AFA01FAEA" blockId="3.[151,1437,797,1729]" pageId="3" pageNumber="204">
|
||
<keyLead id="30B51344FF86FFBEFF1DF78AFA01FAEA" pageId="3" pageNumber="204">
|
||
- Basal carina of head absent; eye canthus dividing eye; elytral epipleura reticulate throughout, lower edge not curving upwards to meet lateral carina of elytra basally (
|
||
<figureCitation id="137FACF4FF86FFBEFDD8F669FD04FA8E" box="[594,665,1296,1318]" captionStart="FIGURE 1" captionStartId="9.[151,250,188,213]" captionTargetId="figure@8.[242,1345,181,1957]" captionTargetPageId="8" captionText="FIGURE 1. Some diagnostic characters of the Lepanus australis species group. A, Male protibia of Lepanus eungella dotted line depicts triangular apical digit (grey arrow) and truncate front edge; B, female protibia of Lepanus eungella dotted line depicts truncate front edge, black arrow points to apical spur; C, male protibia of Lepanus australis dotted line depicts elongate apical digit (grey arrow) and deeply angulate front edge; D, female protibia of Lepanus australis dotted line depicts shortened apical digit (grey arrow) and deeply angulate front edge, black arrow points to apical spur; E, lateral view of Lepanus dukungarri white arrow points to row of punctures on metanepisternum; F, lateral view of Lepanus australis, dotted line indicates anteriorly straight inner border of lateral lobe of metaventrite, note the absence of puntures on metanepisternum and straight lower edge of elytral epipleura; G, lateral view of Lepanus oxleyi white arrow points to row of fine punctures on metanepisternum, grey arrow inducates curved lower edge of elytral epipleura; H, male metafemora of Lepanus dukungarri showing medial expansion; I, Head of Lepanus oxleyi white arrow points to interrupted basal carina; J, lateral view of Lepanus latheticus dotted line indicates anteriorly curved inner border of lateral lobe of metaventrite; K, Lepanus australis at bird droppings (image: Geoff Williams)." figureDoi="http://doi.org/10.5281/zenodo.3611235" httpUri="https://zenodo.org/record/3611235/files/figure.png" pageId="3" pageNumber="204">Fig. 1F</figureCitation>
|
||
); metanepisternum reticulate, not punctate; abdominal ventrite 6 with discernible punctures............................................................................................
|
||
<emphasis id="B9306C63FF86FFBEFA1AF654FA01FAEA" bold="true" box="[1424,1436,1325,1346]" pageId="3" pageNumber="204">4</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30B0A8D4FF86FFBEFF1DF632FA01FA5B" pageId="3" pageNumber="204">
|
||
<paragraph id="8BFBB071FF86FFBEFF1DF632FA01FA33" blockId="3.[151,1437,797,1729]" pageId="3" pageNumber="204">
|
||
<keyLead id="30B51344FF86FFBEFF1DF632FA01FA33" pageId="3" pageNumber="204">
|
||
4. Elytra black or dark brown with red humeri (
|
||
<figureCitation id="137FACF4FF86FFBEFD12F632FD78FAC9" box="[664,741,1355,1377]" captionStart="FIGURE 2" captionStartId="10.[151,250,1947,1972]" captionTargetBox="[311,1277,196,1910]" captionTargetId="figure@10.[283,1305,181,1922]" captionTargetPageId="10" captionText="FIGURE 2. Habitus of species within the Lepanus australis species group. A, Lepanus australis; B, Lepanus burnetti; C, Lepanus dukungarri; D, Lepanus eungella; E, Lepanus latheticus; F, Lepanus oxleyi. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532517" httpUri="https://zenodo.org/record/3532517/files/figure.png" pageId="3" pageNumber="204">Fig. 2B</figureCitation>
|
||
); pronotum with punctures along basal margin circular, similar to those adjacent on disc; elytra with neither striae 6 or 7 reaching basal margin, ending in the red humeral patch and parallel to each other at this point; southeastern
|
||
<collectingRegion id="49807E93FF86FFBEFD9FF6FCFD1BFA33" box="[533,646,1413,1435]" country="Australia" name="Queensland" pageId="3" pageNumber="204">Queensland</collectingRegion>
|
||
............................
|
||
<taxonomicName id="4C44CBF2FF86FFBEFC55F6FFFABDFA33" ID-CoL="6P88B" authority="Gunter & Weir" authorityName="Gunter & Weir" authorityYear="2019" box="[991,1312,1413,1435]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="burnetti" status="sp. nov.">
|
||
<emphasis id="B9306C63FF86FFBEFC55F6FFFB14FA33" bold="true" box="[991,1161,1413,1435]" italics="true" pageId="3" pageNumber="204">Lepanus burnetti</emphasis>
|
||
Gunter & Weir
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A203D118FF86FFBEFAA3F6FFFA01FA33" box="[1321,1436,1413,1435]" pageId="3" pageNumber="204" rank="species">new species</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BFBB071FF86FFBEFF1DF6DAFA01FA5B" blockId="3.[151,1437,797,1729]" pageId="3" pageNumber="204">
|
||
<keyLead id="30B51344FF86FFBEFF1DF6DAFA01FA5B" pageId="3" pageNumber="204">
|
||
- Elytra uniformly black or brown; pronotum with punctures along basal margin somewhat elongate, different to those adjacent on disc; elytra with stria 6 reaching basal margin, stria 7 not reaching basal margin and converging towards stria 6 basally... ...................................................................................................
|
||
<emphasis id="B9306C63FF86FFBEFA1AF6A7FA01FA5B" bold="true" box="[1424,1436,1502,1523]" pageId="3" pageNumber="204">5</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30B0A8D4FF86FFBEFF1DF682FA01F968" pageId="3" pageNumber="204">
|
||
<paragraph id="8BFBB071FF86FFBEFF1DF682FA01F9E2" blockId="3.[151,1437,797,1729]" pageId="3" pageNumber="204">
|
||
<keyLead id="30B51344FF86FFBEFF1DF682FA01F9E2" pageId="3" pageNumber="204">
|
||
5. Lateral lobe of metaventrite with inner border beside mesocoxae anteriorly curving to meet lateral border (
|
||
<figureCitation id="137FACF4FF86FFBEFB6CF682FAB4F9B9" box="[1254,1321,1531,1553]" captionStart="FIGURE 1" captionStartId="9.[151,250,188,213]" captionTargetId="figure@8.[242,1345,181,1957]" captionTargetPageId="8" captionText="FIGURE 1. Some diagnostic characters of the Lepanus australis species group. A, Male protibia of Lepanus eungella dotted line depicts triangular apical digit (grey arrow) and truncate front edge; B, female protibia of Lepanus eungella dotted line depicts truncate front edge, black arrow points to apical spur; C, male protibia of Lepanus australis dotted line depicts elongate apical digit (grey arrow) and deeply angulate front edge; D, female protibia of Lepanus australis dotted line depicts shortened apical digit (grey arrow) and deeply angulate front edge, black arrow points to apical spur; E, lateral view of Lepanus dukungarri white arrow points to row of punctures on metanepisternum; F, lateral view of Lepanus australis, dotted line indicates anteriorly straight inner border of lateral lobe of metaventrite, note the absence of puntures on metanepisternum and straight lower edge of elytral epipleura; G, lateral view of Lepanus oxleyi white arrow points to row of fine punctures on metanepisternum, grey arrow inducates curved lower edge of elytral epipleura; H, male metafemora of Lepanus dukungarri showing medial expansion; I, Head of Lepanus oxleyi white arrow points to interrupted basal carina; J, lateral view of Lepanus latheticus dotted line indicates anteriorly curved inner border of lateral lobe of metaventrite; K, Lepanus australis at bird droppings (image: Geoff Williams)." figureDoi="http://doi.org/10.5281/zenodo.3611235" httpUri="https://zenodo.org/record/3611235/files/figure.png" pageId="3" pageNumber="204">Fig. 1J</figureCitation>
|
||
); mesepimeron usually smooth; pygidium with minute unevenly distributed punctures, often appearing impunctate (
|
||
<figureCitation id="137FACF4FF86FFBEFB45F561FA87F986" box="[1231,1306,1560,1582]" captionStart="FIGURE 4" captionStartId="12.[151,250,1270,1295]" captionTargetBox="[165,1415,192,1221]" captionTargetId="figure@12.[153,1434,181,1245]" captionTargetPageId="12" captionText="FIGURE 4. Pygidial configuration of species within the Lepanus australis species group. A, Lepanus australis; B, Lepanus burnetti; C, Lepanus dukungarri; D, Lepanus eungella; E, Lepanus latheticus; F, Lepanus oxleyi. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532521" httpUri="https://zenodo.org/record/3532521/files/figure.png" pageId="3" pageNumber="204">Fig. 4E</figureCitation>
|
||
); aedeagus as in
|
||
<figureCitation id="137FACF4FF86FFBEFF71F54CFED5F9E3" box="[251,328,1589,1611]" captionStart="FIGURE 3" captionStartId="11.[151,250,1924,1949]" captionTargetBox="[264,1312,179,1899]" captionTargetId="figure@11.[262,1324,166,1904]" captionTargetPageId="11" captionText="FIGURE 3. Left, dorsal, and right view of aedeagi of species within the Lepanus australis species group. A, Lepanus australis; B, Lepanus burnetti; C, Lepanus dukungarri; D, Lepanus eungella; E, Lepanus latheticus; F, Lepanus oxleyi. All specimens in comparative scale. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3532519" httpUri="https://zenodo.org/record/3532519/files/figure.png" pageId="3" pageNumber="204">Figs 3E</figureCitation>
|
||
; northern
|
||
<collectingRegion id="49807E93FF86FFBEFE23F54CFD80F9E3" box="[425,541,1589,1611]" country="Australia" name="Queensland" pageId="3" pageNumber="204">Queensland</collectingRegion>
|
||
, Wet Tropics...................................
|
||
<taxonomicName id="4C44CBF2FF86FFBEFBB0F54FFA01F9E2" ID-CoL="6P7Y6" authority="Matthews, 1974" authorityName="Matthews" authorityYear="1974" box="[1082,1436,1589,1611]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="latheticus">
|
||
<emphasis id="B9306C63FF86FFBEFBB0F54FFA01F9E2" bold="true" box="[1082,1436,1589,1611]" pageId="3" pageNumber="204">
|
||
<emphasis id="B9306C63FF86FFBEFBB0F54FFB68F9E3" bold="true" box="[1082,1269,1589,1611]" italics="true" pageId="3" pageNumber="204">Lepanus latheticus</emphasis>
|
||
<bibRefCitation id="EFD5CD80FF86FFBEFB71F54FFA01F9E2" author="Matthews, E. G." box="[1275,1436,1589,1611]" pageId="3" pageNumber="204" pagination="1 - 211" refId="ref56271" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini, Australian Journal of Zoology, Supplementary Series, 24, 1 - 211. https: // doi. org / 10.1071 / AJZS 024" type="journal article" year="1974">Matthews, 1974</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BFBB071FF86FFBEFF1DF52AFA01F968" blockId="3.[151,1437,797,1729]" pageId="3" pageNumber="204">
|
||
<keyLead id="30B51344FF86FFBEFF1DF52AFA01F968" pageId="3" pageNumber="204">
|
||
- Lateral lobe of metaventrite with inner border next to mesocoxae anteriorly straight, lateral border virtually absent (
|
||
<figureCitation id="137FACF4FF86FFBEFACCF52AFA0DF9C1" box="[1350,1424,1619,1641]" captionStart="FIGURE 1" captionStartId="9.[151,250,188,213]" captionTargetId="figure@8.[242,1345,181,1957]" captionTargetPageId="8" captionText="FIGURE 1. Some diagnostic characters of the Lepanus australis species group. A, Male protibia of Lepanus eungella dotted line depicts triangular apical digit (grey arrow) and truncate front edge; B, female protibia of Lepanus eungella dotted line depicts truncate front edge, black arrow points to apical spur; C, male protibia of Lepanus australis dotted line depicts elongate apical digit (grey arrow) and deeply angulate front edge; D, female protibia of Lepanus australis dotted line depicts shortened apical digit (grey arrow) and deeply angulate front edge, black arrow points to apical spur; E, lateral view of Lepanus dukungarri white arrow points to row of punctures on metanepisternum; F, lateral view of Lepanus australis, dotted line indicates anteriorly straight inner border of lateral lobe of metaventrite, note the absence of puntures on metanepisternum and straight lower edge of elytral epipleura; G, lateral view of Lepanus oxleyi white arrow points to row of fine punctures on metanepisternum, grey arrow inducates curved lower edge of elytral epipleura; H, male metafemora of Lepanus dukungarri showing medial expansion; I, Head of Lepanus oxleyi white arrow points to interrupted basal carina; J, lateral view of Lepanus latheticus dotted line indicates anteriorly curved inner border of lateral lobe of metaventrite; K, Lepanus australis at bird droppings (image: Geoff Williams)." figureDoi="http://doi.org/10.5281/zenodo.3611235" httpUri="https://zenodo.org/record/3611235/files/figure.png" pageId="3" pageNumber="204">Fig. 1F</figureCitation>
|
||
); mesepimeron usually reticulate; pygidium with discernible punctures, evenly distributed (
|
||
<figureCitation id="137FACF4FF86FFBEFBC5F509FB02F92E" box="[1103,1183,1648,1670]" captionStart="FIGURE 4" captionStartId="12.[151,250,1270,1295]" captionTargetBox="[165,1415,192,1221]" captionTargetId="figure@12.[153,1434,181,1245]" captionTargetPageId="12" captionText="FIGURE 4. Pygidial configuration of species within the Lepanus australis species group. A, Lepanus australis; B, Lepanus burnetti; C, Lepanus dukungarri; D, Lepanus eungella; E, Lepanus latheticus; F, Lepanus oxleyi. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532521" httpUri="https://zenodo.org/record/3532521/files/figure.png" pageId="3" pageNumber="204">Fig. 4A</figureCitation>
|
||
); aedeagus as in
|
||
<figureCitation id="137FACF4FF86FFBEFACDF509FA05F92D" box="[1351,1432,1648,1670]" captionStart="FIGURE 3" captionStartId="11.[151,250,1924,1949]" captionTargetBox="[264,1312,179,1899]" captionTargetId="figure@11.[262,1324,166,1904]" captionTargetPageId="11" captionText="FIGURE 3. Left, dorsal, and right view of aedeagi of species within the Lepanus australis species group. A, Lepanus australis; B, Lepanus burnetti; C, Lepanus dukungarri; D, Lepanus eungella; E, Lepanus latheticus; F, Lepanus oxleyi. All specimens in comparative scale. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3532519" httpUri="https://zenodo.org/record/3532519/files/figure.png" pageId="3" pageNumber="204">Figs 3A</figureCitation>
|
||
; southeastern
|
||
<collectingCountry id="F353F0E1FF86FFBEFED4F5F4FE2BF90B" box="[350,438,1677,1699]" name="Australia" pageId="3" pageNumber="204">Australia</collectingCountry>
|
||
from central
|
||
<collectingRegion id="49807E93FF86FFBEFDB2F5F4FD34F90B" box="[568,681,1677,1699]" country="Australia" name="Queensland" pageId="3" pageNumber="204">Queensland</collectingRegion>
|
||
(St Lawrence) to
|
||
<collectingRegion id="49807E93FF86FFBEFCDDF5F4FC39F90B" box="[855,932,1677,1699]" country="Australia" name="Victoria" pageId="3" pageNumber="204">Victoria</collectingRegion>
|
||
(Mallacoota)......................................................................................................
|
||
<taxonomicName id="4C44CBF2FF86FFBEFBCEF5D2FA01F968" ID-CoL="6P7WD" authority="Matthews, 1974" authorityName="Matthews" authorityYear="1974" box="[1092,1436,1707,1729]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B9306C63FF86FFBEFBCEF5D2FA01F968" bold="true" box="[1092,1436,1707,1729]" pageId="3" pageNumber="204">
|
||
<emphasis id="B9306C63FF86FFBEFBCEF5D2FB68F969" bold="true" box="[1092,1269,1707,1729]" italics="true" pageId="3" pageNumber="204">Lepanus australis</emphasis>
|
||
<bibRefCitation id="EFD5CD80FF86FFBEFB71F5D2FA01F968" author="Matthews, E. G." box="[1275,1436,1707,1728]" pageId="3" pageNumber="204" pagination="1 - 211" refId="ref56271" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini, Australian Journal of Zoology, Supplementary Series, 24, 1 - 211. https: // doi. org / 10.1071 / AJZS 024" type="journal article" year="1974">Matthews, 1974</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |