418 lines
52 KiB
XML
418 lines
52 KiB
XML
<document id="8075CD73001C22BDF4B3E6C8E44EA101" ID-CLB-Dataset="31171" ID-DOI="10.11646/zootaxa.4363.4.13" ID-GBIF-Dataset="42b57df3-f00d-4c0f-93cd-7b82649ef59a" ID-ISSN="1175-5326" ID-Zenodo-Dep="1114841" ID-ZooBank="77801C93-5D28-4AA6-94B8-F1003E9B938F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1513152368772" checkinUser="plazi" docAuthor="Chen, Chong, Watanabe, Hiromi Kayama & Araya, Juan Francisco" docDate="2017" docId="03EC87AC38249D58FF54DB5E974F96BA" docLanguage="en" docName="zootaxa.4363.4.12.pdf" docOrigin="Zootaxa 4363 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Columbellidae Swainson 1840" docType="treatment" docVersion="7" lastPageNumber="594" masterDocId="FFD5FFD438259D5AFFC3DA5E9219942E" masterDocTitle="First columbellid species (Gastropoda: Buccinoidea) from deep-sea hydrothermal vents, discovered in Okinawa Trough, Japan" masterLastPageNumber="596" masterPageNumber="592" pageNumber="593" updateTime="1698509680753" updateUser="ExternalLinkService">
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<mods:titleInfo id="527F362DF772F6F9CC424B43ADE18E4B">
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<mods:title id="F4B154C7F35C86832A06CE3B78BF3E5C">First columbellid species (Gastropoda: Buccinoidea) from deep-sea hydrothermal vents, discovered in Okinawa Trough, Japan</mods:title>
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<mods:name id="582F2742378071933C6FAD33F45F24EE" type="personal">
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<mods:namePart id="156E369AC9A81F78AB177020DD83AC10">Chen, Chong</mods:namePart>
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<mods:namePart id="D169B5E241CED539933609456C5EEBB7">Watanabe, Hiromi Kayama</mods:namePart>
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</mods:name>
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<mods:name id="B969BE6F82F88803AD07461E5310901E" type="personal">
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<mods:roleTerm id="89A8EBD5DAD474C6E08FF12704A37E5C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="240C441A7CCD60DE1A4B95017B017E22">Araya, Juan Francisco</mods:namePart>
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<mods:typeOfResource id="7CECBF400F7E50C29823652ED781D951">text</mods:typeOfResource>
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<mods:title id="4CE8C28440FF4F654D17F909C034E8D9">Zootaxa</mods:title>
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<mods:part id="1F69C09063756FC22A8771D0E3733931">
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<mods:date id="87A218354397867F844899875EAD42C5">2017</mods:date>
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<mods:number id="D85B0E0A9295F115AE50670906B26D0A">2017-12-13</mods:number>
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<treatment id="03EC87AC38249D58FF54DB5E974F96BA" ID-DOI="http://doi.org/10.5281/zenodo.6022939" ID-GBIF-Taxon="137679991" ID-Zenodo-Dep="6022939" LSID="urn:lsid:plazi:treatment:03EC87AC38249D58FF54DB5E974F96BA" httpUri="http://treatment.plazi.org/id/03EC87AC38249D58FF54DB5E974F96BA" lastPageId="2" lastPageNumber="594" pageId="1" pageNumber="593">
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<subSubSection id="C35F653138249D5BFF54DB5E90AB9512" pageId="1" pageNumber="593" type="nomenclature">
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<paragraph id="8BFA36BA38249D5BFF54DB5E90DF9534" blockId="1.[151,1436,152,2012]" box="[151,710,256,282]" pageId="1" pageNumber="593">
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<heading id="D0B281D638249D5BFF54DB5E90DF9534" bold="true" box="[151,710,256,282]" fontSize="11" level="1" pageId="1" pageNumber="593" reason="1">
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<materialsCitation id="3B2D3CE738249D5BFF54DB5E90DF9534" ID-GBIF-Occurrence="1705513546" box="[151,710,256,282]" location="Swainson" pageId="1" pageNumber="593" specimenCount="1" stateProvince="Swainson">
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<emphasis id="B931EAA838249D5BFF54DB5E90DF9534" bold="true" box="[151,710,256,282]" pageId="1" pageNumber="593">
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Family
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<taxonomicName id="4C454D3938249D5BFF35DB5E90DF9534" ID-CoL="7NGWV" authority="Swainson, 1840" authorityName="Swainson" authorityYear="1840" box="[246,710,256,282]" class="Gastropoda" family="Columbellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="593" phylum="Mollusca" rank="family">
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COLUMBELLIDAE
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<collectingRegion id="4981F85838249D5BFDC6DB5E90679534" box="[517,638,256,282]" country="American Samoa" name="Swains Island" pageId="1" pageNumber="593">Swainson</collectingRegion>
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, 1840
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</taxonomicName>
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</emphasis>
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</materialsCitation>
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</heading>
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</paragraph>
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<paragraph id="8BFA36BA38249D5BFF54DB7C90AB9512" blockId="1.[151,1436,152,2012]" box="[151,690,290,316]" pageId="1" pageNumber="593">
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<emphasis id="B931EAA838249D5BFF54DB7C90AB9512" bold="true" box="[151,690,290,316]" pageId="1" pageNumber="593">
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Genus
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<taxonomicName id="4C454D3938249D5BFF2DDB7C90AB9512" authority="H. Adams & A. Adams, 1853" authorityName="Chen & Watanabe & Araya" authorityYear="2017" box="[238,690,290,316]" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="593" phylum="Mollusca" rank="genus" status="sp. nov.">
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<emphasis id="B931EAA838249D5BFF2DDB7C93589512" bold="true" box="[238,321,290,316]" italics="true" pageId="1" pageNumber="593">Astyris</emphasis>
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H. Adams & A. Adams, 1853
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</taxonomicName>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F653138249D5BFF54DB1A967B9551" pageId="1" pageNumber="593" type="description">
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<paragraph id="8BFA36BA38249D5BFF54DB1A93C59571" blockId="1.[151,1436,152,2012]" box="[151,476,324,351]" pageId="1" pageNumber="593">
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<emphasis id="B931EAA838249D5BFF54DB1A93C59571" bold="true" box="[151,476,324,351]" pageId="1" pageNumber="593">
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<taxonomicName id="4C454D3938249D5BFF54DB1A938B9570" authority="Chen & Watanabe & Araya, 2017" authorityName="Chen & Watanabe & Araya" authorityYear="2017" box="[151,402,324,350]" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="593" phylum="Mollusca" rank="species" species="thermophilus" status="sp. nov.">
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<emphasis id="B931EAA838249D5BFF54DB1A938B9570" bold="true" box="[151,402,324,350]" italics="true" pageId="1" pageNumber="593">Astyris thermophilus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20257D338249D5BFE59DB1B93C59571" box="[410,476,325,351]" pageId="1" pageNumber="593" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="8BFA36BA38249D5BFF54DB36967B9551" blockId="1.[151,1436,152,2012]" box="[151,1122,360,383]" pageId="1" pageNumber="593">http://zoobank.org/urn:lsid:zoobank.org:act:43FA9CFA-3756-4CBC-9861-41FAF66397B9</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35F653138249D5BFF54DBD49731976F" pageId="1" pageNumber="593" type="materials_examined">
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<paragraph id="8BFA36BA38249D5BFF54DBD4974D9661" blockId="1.[151,1436,152,2012]" pageId="1" pageNumber="593">
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<emphasis id="B931EAA838249D5BFF54DBD49349958F" bold="true" box="[151,336,394,417]" pageId="1" pageNumber="593">Type specimens.</emphasis>
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Holotype (
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<figureCitation id="137E2A3F38249D5BFE09DBD49026958F" box="[458,575,394,417]" captionStart="FIGURE 1" captionStartId="2.[151,244,1917,1939]" captionTargetBox="[151,1436,714,1896]" captionTargetId="figure@2.[151,1436,714,1896]" captionTargetPageId="2" captionText="FIGURE 1. Astyris thermophilus n. sp. A–D. Holotype (UMUT RM32644). E. Paratype #1 (NSMT Mo 78990). F. Paratype #2 (UMUT RM32645), periostracum removed to show spiral striae. G. Protoconch of paratype #6, a juvenile specimen (UMUT RM 32646). H. Operculum of paratype #2 (UMUT RM32645). I. Radula of paratype #2, UMUT RM32645). Scale bars = 2 mm (A–F), 500 µm(G, H), 20µm (I)." httpUri="https://zenodo.org/record/1114843/files/figure.png" pageId="1" pageNumber="593">Fig. 1A–D</figureCitation>
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) [SH
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<quantity id="4CBD9B5F38249D5BFD43DBD490CC958F" box="[640,725,394,417]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="593" unit="mm" value="8.5">8.5 mm</quantity>
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, SD
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<quantity id="4CBD9B5F38249D5BFCCADBD49147958F" box="[777,862,394,417]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="1" pageNumber="593" unit="mm" value="4.3">4.3 mm</quantity>
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, LW
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||
<quantity id="4CBD9B5F38249D5BFC5BDBD491F6958F" box="[920,1007,394,417]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="593" unit="mm" value="4.1">4.1 mm</quantity>
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], UMUT RM32644. Paratypes. #1 (
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<figureCitation id="137E2A3F38249D5BFAB3DBD492AD95EA" captionStart="FIGURE 1" captionStartId="2.[151,244,1917,1939]" captionTargetBox="[151,1436,714,1896]" captionTargetId="figure@2.[151,1436,714,1896]" captionTargetPageId="2" captionText="FIGURE 1. Astyris thermophilus n. sp. A–D. Holotype (UMUT RM32644). E. Paratype #1 (NSMT Mo 78990). F. Paratype #2 (UMUT RM32645), periostracum removed to show spiral striae. G. Protoconch of paratype #6, a juvenile specimen (UMUT RM 32646). H. Operculum of paratype #2 (UMUT RM32645). I. Radula of paratype #2, UMUT RM32645). Scale bars = 2 mm (A–F), 500 µm(G, H), 20µm (I)." httpUri="https://zenodo.org/record/1114843/files/figure.png" pageId="1" pageNumber="593">Fig. 1E</figureCitation>
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) [SH
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<quantity id="4CBD9B5F38249D5BFF39DBF3935695EA" box="[250,335,429,452]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="1" pageNumber="593" unit="mm" value="8.3">8.3 mm</quantity>
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, SD
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<quantity id="4CBD9B5F38249D5BFE44DBF393C495EA" box="[391,477,429,452]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="593" unit="mm" value="4.5">4.5 mm</quantity>
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, LW
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<quantity id="4CBD9B5F38249D5BFDD9DBF3906A95EA" box="[538,627,429,452]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="593" unit="mm" value="4.6">4.6 mm</quantity>
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], NSMT Mo 78990. #2 (
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<figureCitation id="137E2A3F38249D5BFC40DBF391CA95EA" box="[899,979,429,452]" captionStart="FIGURE 1" captionStartId="2.[151,244,1917,1939]" captionTargetBox="[151,1436,714,1896]" captionTargetId="figure@2.[151,1436,714,1896]" captionTargetPageId="2" captionText="FIGURE 1. Astyris thermophilus n. sp. A–D. Holotype (UMUT RM32644). E. Paratype #1 (NSMT Mo 78990). F. Paratype #2 (UMUT RM32645), periostracum removed to show spiral striae. G. Protoconch of paratype #6, a juvenile specimen (UMUT RM 32646). H. Operculum of paratype #2 (UMUT RM32645). I. Radula of paratype #2, UMUT RM32645). Scale bars = 2 mm (A–F), 500 µm(G, H), 20µm (I)." httpUri="https://zenodo.org/record/1114843/files/figure.png" pageId="1" pageNumber="593">Fig. 1F</figureCitation>
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) [SH
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<quantity id="4CBD9B5F38249D5BFBDBDBF3967795EA" box="[1048,1134,429,452]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="593" unit="mm" value="7.4">7.4 mm</quantity>
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, SD
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<quantity id="4CBD9B5F38249D5BFB65DBF396E595EA" box="[1190,1276,429,452]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="593" unit="mm" value="4.2">4.2 mm</quantity>
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, LW
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<quantity id="4CBD9B5F38249D5BFAFADBF3978895EA" box="[1337,1425,429,452]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="593" unit="mm" value="4.1">4.1 mm</quantity>
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], periostracum removed with diluted bleach to reveal fine sculpture, operculum and radula removed for SEM, UMUT RM32645. #3 [SH
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<quantity id="4CBD9B5F38249D5BFEABDBAC93A79627" box="[360,446,498,521]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="1" pageNumber="593" unit="mm" value="7.1">7.1 mm</quantity>
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, SD 4.0 mm, LW 4.0 mm], NSMT Mo 78991. #4 [SH
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<quantity id="4CBD9B5F38249D5BFBD9DBAC96699627" box="[1050,1136,498,521]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="1" pageNumber="593" unit="mm" value="7.7">7.7 mm</quantity>
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||
, SD
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||
<quantity id="4CBD9B5F38249D5BFB64DBAC96E59627" box="[1191,1276,498,521]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="593" unit="mm" value="4.1">4.1 mm</quantity>
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||
, LW
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||
<quantity id="4CBD9B5F38249D5BFAFADBAC97889627" box="[1337,1425,498,521]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="1" pageNumber="593" unit="mm" value="4.3">4.3 mm</quantity>
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||
], NSMT Mo 78992. #5 [SH
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||
<quantity id="4CBD9B5F38249D5BFE70D84B901E9602" box="[435,519,533,556]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="593" unit="mm" value="6.5">6.5 mm</quantity>
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||
, SD
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||
<quantity id="4CBD9B5F38249D5BFDFAD84B90949602" box="[569,653,533,556]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="593" unit="mm" value="3.7">3.7 mm</quantity>
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||
, LW
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||
<quantity id="4CBD9B5F38249D5BFD07D84B91039602" box="[708,794,533,556]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="1" pageNumber="593" unit="mm" value="3.9">3.9 mm</quantity>
|
||
], JAMSTEC 1140053525. #6 [SH 3.0 mm, SD
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||
<quantity id="4CBD9B5F38249D5BFAD5D84B97739602" box="[1302,1386,533,556]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="593" unit="mm" value="1.7">1.7 mm</quantity>
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||
, LW
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||
<quantity id="4CBD9B5F38249D5BFF54D86692F49661" box="[151,237,568,591]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="593" unit="mm" value="1.6">1.6 mm</quantity>
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||
], juvenile with intact protoconch, UMUT RM32646. All type specimens fixed and stored in 99% ethanol.
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</paragraph>
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<paragraph id="8BFA36BA38249D5BFF04D80490509699" blockId="1.[151,1436,152,2012]" pageId="1" pageNumber="593">
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<emphasis id="B931EAA838249D5BFF04D8049373965F" bold="true" box="[199,362,602,625]" pageId="1" pageNumber="593">
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||
<typeStatus id="54FE881838249D5BFF04D80492E6965F" box="[199,255,602,625]" pageId="1" pageNumber="593">Type</typeStatus>
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locality.
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</emphasis>
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Natsu hydrothermal vent site, Iheya North field,
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<collectingRegion id="4981F85838249D5BFC1AD8049620965F" box="[985,1081,602,625]" country="Japan" name="Okinawa" pageId="1" pageNumber="593">Okinawa</collectingRegion>
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Trough,
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<collectingCountry id="F352762A38249D5BFB70D80496EA965F" box="[1203,1267,602,625]" name="Japan" pageId="1" pageNumber="593">Japan</collectingCountry>
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;
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,
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<geoCoordinate id="EE71507D38249D5BFF54D823933696BA" box="[151,303,637,660]" direction="east" orientation="longitude" pageId="1" pageNumber="593" precision="1" value="126.9004">126°54.024’E</geoCoordinate>
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,
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<quantity id="4CBD9B5F38249D5BFEFAD823939096BA" box="[313,393,637,660]" metricMagnitude="3" metricUnit="m" metricValue="1.074" pageId="1" pageNumber="593" unit="m" value="1074.0">1074 m</quantity>
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deep;
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<date id="FFFB107A38249D5BFE13D823904C96BA" box="[464,597,637,660]" pageId="1" pageNumber="593" value="2014-01-24">2014/Jan/24</date>
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, collected by slurp gun,
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<collectionCode id="ED54AE7F38249D5BFC9BD823918996BA" box="[856,912,637,660]" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15413" name="Museo Civico di Rovereto" pageId="1" pageNumber="593">ROV</collectionCode>
|
||
<collectionCode id="ED54AE7F38249D5BFC55D823965396BD" box="[918,1098,637,660]" pageId="1" pageNumber="593">HYPER- DOLPHIN</collectionCode>
|
||
Dive #1614, R/V
|
||
<collectionCode id="ED54AE7F38249D5BFACED823974F96BD" box="[1293,1366,637,660]" pageId="1" pageNumber="593">KAIYO</collectionCode>
|
||
cruise KY14-01, principal scientist: Ken Takai.
|
||
</paragraph>
|
||
<paragraph id="8BFA36BA38249D5BFF04D89C9731976F" blockId="1.[151,1436,152,2012]" pageId="1" pageNumber="593">
|
||
<emphasis id="B931EAA838249D5BFF04D89C903396F7" bold="true" box="[199,554,706,729]" pageId="1" pageNumber="593">Additional material examined.</emphasis>
|
||
Two specimens from
|
||
<typeStatus id="54FE881838249D5BFCE5D89C914D96F7" box="[806,852,706,729]" pageId="1" pageNumber="593">type</typeStatus>
|
||
locality, fixed and stored in 99 % ethanol, used for genetic barcoding (tissue dissolved). Five specimens, live collected, fixed and stored in 99% ethanol. Aki hydrothermal vent site, Iheya North field,
|
||
<collectingRegion id="4981F85838249D5BFE05D956903F9731" box="[454,550,776,799]" country="Japan" name="Okinawa" pageId="1" pageNumber="593">Okinawa</collectingRegion>
|
||
Trough,
|
||
<collectingCountry id="F352762A38249D5BFD4AD95690D09731" box="[649,713,776,799]" name="Japan" pageId="1" pageNumber="593">Japan</collectingCountry>
|
||
;
|
||
<geoCoordinate id="EE71507D38249D5BFD16D956917A9731" box="[725,867,776,799]" direction="north" orientation="latitude" pageId="1" pageNumber="593" precision="1" value="27.768833">27°46.130’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE71507D38249D5BFCADD956961F9731" box="[878,1030,776,799]" direction="east" orientation="longitude" pageId="1" pageNumber="593" precision="1" value="126.90265">126°54.159’E</geoCoordinate>
|
||
,
|
||
<quantity id="4CBD9B5F38249D5BFBD2D956967B9731" box="[1041,1122,776,799]" metricMagnitude="3" metricUnit="m" metricValue="1.087" pageId="1" pageNumber="593" unit="m" value="1087.0">1087 m</quantity>
|
||
deep;
|
||
<date id="FFFB107A38249D5BFB68D95697299731" box="[1195,1328,776,799]" pageId="1" pageNumber="593" value="2014-01-25">2014/Jan/25</date>
|
||
, collected by slurp gun,
|
||
<collectionCode id="ED54AE7F38249D5BFEEAD9749378976F" box="[297,353,810,833]" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15413" name="Museo Civico di Rovereto" pageId="1" pageNumber="593">ROV</collectionCode>
|
||
<collectionCode id="ED54AE7F38249D5BFEABD9749005976F" box="[360,540,810,833]" pageId="1" pageNumber="593">HYPER- DOLPHIN</collectionCode>
|
||
Dive #1614, R/V
|
||
<collectionCode id="ED54AE7F38249D5BFD23D9749131976F" box="[736,808,810,833]" pageId="1" pageNumber="593">KAIYO</collectionCode>
|
||
cruise KY14-01, principal scientist: Ken Takai.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F653138249D5BFF04D913961E9787" pageId="1" pageNumber="593" type="etymology">
|
||
<paragraph id="8BFA36BA38249D5BFF04D91390E697A9" blockId="1.[151,1436,152,2012]" pageId="1" pageNumber="593">
|
||
<emphasis id="B931EAA838249D5BFF04D913935E974A" bold="true" box="[199,327,845,868]" pageId="1" pageNumber="593">Etymology.</emphasis>
|
||
From ‘
|
||
<emphasis id="B931EAA838249D5BFE5BD91393EC974A" box="[408,501,845,868]" italics="true" pageId="1" pageNumber="593">Thermós</emphasis>
|
||
’ (Greek), warm or hot, and ‘
|
||
<emphasis id="B931EAA838249D5BFCEFD9139173974A" box="[812,874,845,868]" italics="true" pageId="1" pageNumber="593">philíā</emphasis>
|
||
’ (Greek), love or affection; combined to mean heatloving, referring to its habitat in hydrothermal vent fields.
|
||
</paragraph>
|
||
<paragraph id="8BFA36BA38249D5BFF04D9CC961E9787" blockId="1.[151,1436,152,2012]" box="[199,1031,914,937]" pageId="1" pageNumber="593">
|
||
<emphasis id="B931EAA838249D5BFF04D9CC93669787" bold="true" box="[199,383,914,937]" pageId="1" pageNumber="593">Japanese Name.</emphasis>
|
||
‘
|
||
<emphasis id="B931EAA838249D5BFE4DD9CC90499787" box="[398,592,914,937]" italics="true" pageId="1" pageNumber="593">Yomotsu-mugi-gai</emphasis>
|
||
’, meaning ‘mitrid from the underworld’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F653138249D5BFF04D9EB9365901A" pageId="1" pageNumber="593" type="diagnosis">
|
||
<paragraph id="8BFA36BA38249D5BFF04D9EB9365901A" blockId="1.[151,1436,152,2012]" pageId="1" pageNumber="593">
|
||
<emphasis id="B931EAA838249D5BFF04D9EB932597E2" bold="true" box="[199,316,949,972]" pageId="1" pageNumber="593">Diagnosis.</emphasis>
|
||
A moderate-sized (up to SH
|
||
<quantity id="4CBD9B5F38249D5BFD55D9EB90F797E2" box="[662,750,949,972]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="593" unit="mm" value="8.5">8.5 mm</quantity>
|
||
)
|
||
<taxonomicName id="4C454D3938249D5BFCC0D9EB915497E2" authorityName="Chen & Watanabe & Araya" authorityYear="2017" box="[771,845,949,972]" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="593" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B931EAA838249D5BFCC0D9EB915497E2" box="[771,845,949,972]" italics="true" pageId="1" pageNumber="593">Astyris</emphasis>
|
||
</taxonomicName>
|
||
with rather tall-spired, uniformly white, thin, semitransparent shell; smooth including base, except 25 to 30 very fine spiral striae. Columella lacking columellar fold. Periostracum thin, greyish brown. Protoconch paucispiral. Radula stenoglossate with acuspate centre plate, three sharp cusps on each lateral.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F653138249D5BFF04DE1E970092AF" pageId="1" pageNumber="593" type="description">
|
||
<paragraph id="8BFA36BA38249D5BFF04DE1E90FB91FA" blockId="1.[151,1436,152,2012]" pageId="1" pageNumber="593">
|
||
<emphasis id="B931EAA838249D5BFF04DE1E934B9079" bold="true" box="[199,338,1088,1111]" pageId="1" pageNumber="593">Description.</emphasis>
|
||
Shell (
|
||
<figureCitation id="137E2A3F38249D5BFE6BDE1E90059079" box="[424,540,1088,1111]" captionStart="FIGURE 1" captionStartId="2.[151,244,1917,1939]" captionTargetBox="[151,1436,714,1896]" captionTargetId="figure@2.[151,1436,714,1896]" captionTargetPageId="2" captionText="FIGURE 1. Astyris thermophilus n. sp. A–D. Holotype (UMUT RM32644). E. Paratype #1 (NSMT Mo 78990). F. Paratype #2 (UMUT RM32645), periostracum removed to show spiral striae. G. Protoconch of paratype #6, a juvenile specimen (UMUT RM 32646). H. Operculum of paratype #2 (UMUT RM32645). I. Radula of paratype #2, UMUT RM32645). Scale bars = 2 mm (A–F), 500 µm(G, H), 20µm (I)." httpUri="https://zenodo.org/record/1114843/files/figure.png" pageId="1" pageNumber="593">Fig. 1A–F</figureCitation>
|
||
) moderate-sized for genus (up to SH
|
||
<quantity id="4CBD9B5F38249D5BFC12DE1E96329079" box="[977,1067,1088,1111]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="593" unit="mm" value="8.5">8.5 mm</quantity>
|
||
), rather tall-spired. Apex always decollate in adults, leaving at most three whorls remaining. Teleoconch thin, semi-transparent, uniformly white in colouration. Periostracum thin, greyish brown, earlier whorls slightly darker coloured. Often covered by further sulfide deposits. Teleoconch largely smooth except approximately 25 to 30 very fine, shallow, evenly spaced spiral striae present across entire whorl (see
|
||
<figureCitation id="137E2A3F38249D5BFE5FDE9493F290CF" box="[412,491,1226,1249]" captionStart="FIGURE 1" captionStartId="2.[151,244,1917,1939]" captionTargetBox="[151,1436,714,1896]" captionTargetId="figure@2.[151,1436,714,1896]" captionTargetPageId="2" captionText="FIGURE 1. Astyris thermophilus n. sp. A–D. Holotype (UMUT RM32644). E. Paratype #1 (NSMT Mo 78990). F. Paratype #2 (UMUT RM32645), periostracum removed to show spiral striae. G. Protoconch of paratype #6, a juvenile specimen (UMUT RM 32646). H. Operculum of paratype #2 (UMUT RM32645). I. Radula of paratype #2, UMUT RM32645). Scale bars = 2 mm (A–F), 500 µm(G, H), 20µm (I)." httpUri="https://zenodo.org/record/1114843/files/figure.png" pageId="1" pageNumber="593">Fig. 1F</figureCitation>
|
||
), increasing in strength anteriorly towards siphonal canal. Striations on posterior half of shell usually too fine to detect when covered by periostracum. Whorls elevated, slowly expanding, slightly convex but not angulated. Suture distinct, shallowly constricted. Aperture entirely lacking dentition, semi-circular in shape, siphonate, approximately twice as tall as wide, posteriorly acuminate. Outer lip simple, not thickened, completely smooth on inside. Columella straight, simple, with slightly thickened callus. Siphonal canal short with weakly constricted but distinct siphonal notch. Protoconch (
|
||
<figureCitation id="137E2A3F38249D5BFD05DF26910391A1" box="[710,794,1400,1423]" captionStart="FIGURE 1" captionStartId="2.[151,244,1917,1939]" captionTargetBox="[151,1436,714,1896]" captionTargetId="figure@2.[151,1436,714,1896]" captionTargetPageId="2" captionText="FIGURE 1. Astyris thermophilus n. sp. A–D. Holotype (UMUT RM32644). E. Paratype #1 (NSMT Mo 78990). F. Paratype #2 (UMUT RM32645), periostracum removed to show spiral striae. G. Protoconch of paratype #6, a juvenile specimen (UMUT RM 32646). H. Operculum of paratype #2 (UMUT RM32645). I. Radula of paratype #2, UMUT RM32645). Scale bars = 2 mm (A–F), 500 µm(G, H), 20µm (I)." httpUri="https://zenodo.org/record/1114843/files/figure.png" pageId="1" pageNumber="593">Fig. 1G</figureCitation>
|
||
) paucispiral, about 1.5 whorls, smooth, lacking velar sinus, sculpture or distinct growth lines. Transition edge between protoconch and teleoconch clearly marked by a varix. Suture of protoconch shallow, slightly higher than teleoconch.
|
||
</paragraph>
|
||
<paragraph id="8BFA36BA38249D5BFF04DFBE93789237" blockId="1.[151,1436,152,2012]" pageId="1" pageNumber="593">
|
||
Operculum (
|
||
<figureCitation id="137E2A3F38249D5BFE8CDFBE93BB91D9" box="[335,418,1504,1527]" captionStart="FIGURE 1" captionStartId="2.[151,244,1917,1939]" captionTargetBox="[151,1436,714,1896]" captionTargetId="figure@2.[151,1436,714,1896]" captionTargetPageId="2" captionText="FIGURE 1. Astyris thermophilus n. sp. A–D. Holotype (UMUT RM32644). E. Paratype #1 (NSMT Mo 78990). F. Paratype #2 (UMUT RM32645), periostracum removed to show spiral striae. G. Protoconch of paratype #6, a juvenile specimen (UMUT RM 32646). H. Operculum of paratype #2 (UMUT RM32645). I. Radula of paratype #2, UMUT RM32645). Scale bars = 2 mm (A–F), 500 µm(G, H), 20µm (I)." httpUri="https://zenodo.org/record/1114843/files/figure.png" pageId="1" pageNumber="593">Fig. 1H</figureCitation>
|
||
) corneous, small, length about half of aperture height. Lamellate with lateral nucleus on right often eroded away.
|
||
</paragraph>
|
||
<paragraph id="8BFA36BA38249D5BFF04DC7B970092AF" blockId="1.[151,1436,152,2012]" pageId="1" pageNumber="593">
|
||
Radula (
|
||
<figureCitation id="137E2A3F38249D5BFEE2DC7B93709212" box="[289,361,1573,1596]" captionStart="FIGURE 1" captionStartId="2.[151,244,1917,1939]" captionTargetBox="[151,1436,714,1896]" captionTargetId="figure@2.[151,1436,714,1896]" captionTargetPageId="2" captionText="FIGURE 1. Astyris thermophilus n. sp. A–D. Holotype (UMUT RM32644). E. Paratype #1 (NSMT Mo 78990). F. Paratype #2 (UMUT RM32645), periostracum removed to show spiral striae. G. Protoconch of paratype #6, a juvenile specimen (UMUT RM 32646). H. Operculum of paratype #2 (UMUT RM32645). I. Radula of paratype #2, UMUT RM32645). Scale bars = 2 mm (A–F), 500 µm(G, H), 20µm (I)." httpUri="https://zenodo.org/record/1114843/files/figure.png" pageId="1" pageNumber="593">
|
||
Fig.
|
||
<date id="FFFB107A38249D5BFE90DC7B93709212" box="[339,361,1573,1596]" pageId="1" pageNumber="593">1I</date>
|
||
</figureCitation>
|
||
) stenoglossate, typical of columbellids with one lateral on each side separated by an acuspate centre plate instead of rachidian. Laterals sigmoid, well-supported, with three strong cusps. Two distal cusps sharper, longer, closer spaced compared to basal cusp. Centre plate rectangular, slightly wider posteriorly, without sculpture.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F653138249D5BFF04DCD3974492C7" pageId="1" pageNumber="593" type="distribution">
|
||
<paragraph id="8BFA36BA38249D5BFF04DCD3974492C7" blockId="1.[151,1436,152,2012]" pageId="1" pageNumber="593">
|
||
<emphasis id="B931EAA838249D5BFF04DCD393FC928A" bold="true" box="[199,485,1677,1700]" pageId="1" pageNumber="593">Distribution and ecology.</emphasis>
|
||
Only known from Natsu and Aki sites of the Iheya North hydrothermal field (
|
||
<bibRefCitation id="EFD44B4B38249D5BFAE8DCD3931692E9" author="Nakamura" pageId="1" pageNumber="593" refString="Nakamura, K., Kawagucci, S., Kitada, K., Kumagai, H., Takai, K. & Okino, K. (2015) Water column imaging with multibeam echosounding in the mid-Okinawa Trough: Implications for distribution of deep-sea hydrothermal vent sites and the cause of acoustic water column anomaly. Geochemical Journal, 49, 579 - 596. https: // doi. org / 10.2343 / geochemj. 2.0387" type="journal article" year="2015">
|
||
Nakamura
|
||
<emphasis id="B931EAA838249D5BFF54DCEE92DE92E9" box="[151,199,1712,1735]" italics="true" pageId="1" pageNumber="593">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
), mid-Okinawa Trough,
|
||
<collectingCountry id="F352762A38249D5BFDD8DCEE904292E9" box="[539,603,1712,1735]" name="Japan" pageId="1" pageNumber="593">Japan</collectingCountry>
|
||
. Found in tubeworm bushes attached on the tubes, presumably a predator of other animals inhabiting the same habitat or alternatively it may be ovophagous and feed on eggs of other animals.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F653138249D58FF04DCAB90309539" lastPageId="2" lastPageNumber="594" pageId="1" pageNumber="593" type="discussion">
|
||
<paragraph id="8BFA36BA38249D58FF04DCAB90309539" blockId="1.[151,1436,152,2012]" lastBlockId="2.[151,1436,152,660]" lastPageId="2" lastPageNumber="594" pageId="1" pageNumber="593">
|
||
<emphasis id="B931EAA838249D5BFF04DCAB932C9322" bold="true" box="[199,309,1781,1804]" pageId="1" pageNumber="593">Remarks.</emphasis>
|
||
The present new species is assigned to genus
|
||
<taxonomicName id="4C454D3938249D5BFCEADCAB916A9322" authorityName="Chen & Watanabe & Araya" authorityYear="2017" box="[809,883,1781,1804]" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="593" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B931EAA838249D5BFCEADCAB916A9322" box="[809,883,1781,1804]" italics="true" pageId="1" pageNumber="593">Astyris</emphasis>
|
||
</taxonomicName>
|
||
as it matches well with the diagnosis for the genus given by
|
||
<bibRefCitation id="EFD44B4B38249D5BFEC1DD46902D9301" author="McLean" box="[258,564,1816,1839]" pageId="1" pageNumber="593" refString="McLean, J. H. & Gosliner, T. M. (1996) Taxonomic Atlas of the Benthic Fauna of the Santa Maria Basin and the Western Santa Barbara Channel. Vol. 9. Mollusca. Part 2. Gastropoda). Santa Barbara Museum of Natural History, Santa Barbara, CA, 228 pp." type="book" year="1996">McLean & Gosliner (1996)</bibRefCitation>
|
||
, most notably by having a small, high-spired shell with smooth surface and a paucispiral protoconch (
|
||
<bibRefCitation id="EFD44B4B38249D5BFE58DD64903F937F" author="Garcia" box="[411,550,1850,1873]" pageId="1" pageNumber="593" refString="Garcia, E. F. (2009) A new Astyris species (Gastropoda: Columbellidae) from the Gulf of Mexico, with notes on the genus. Novapex, 10, 5 - 8." type="journal article" year="2009">Garcia 2009</bibRefCitation>
|
||
). Of those species currently in
|
||
<taxonomicName id="4C454D3938249D5BFCBBDD6491DB937F" authorityName="Chen & Watanabe & Araya" authorityYear="2017" box="[888,962,1850,1873]" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="593" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B931EAA838249D5BFCBBDD6491DB937F" box="[888,962,1850,1873]" italics="true" pageId="1" pageNumber="593">Astyris</emphasis>
|
||
</taxonomicName>
|
||
, the new species most closely resembles
|
||
<taxonomicName id="4C454D3938249D5BFA46DD6493BA935A" authority="(Dall, 1890)" authorityName="Dall" authorityYear="1890" baseAuthorityName="Dall" baseAuthorityYear="1890" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="593" phylum="Mollusca" rank="species" species="permodesta">
|
||
<emphasis id="B931EAA838249D5BFA46DD64930A935A" italics="true" pageId="1" pageNumber="593">A. permodesta</emphasis>
|
||
(Dall, 1890)
|
||
</taxonomicName>
|
||
from methane seeps at Monterey Canyon and whale falls in California (
|
||
<bibRefCitation id="EFD44B4B38249D5BFB13DD039781935A" author="Smith" box="[1232,1432,1885,1908]" pageId="1" pageNumber="593" refString="Smith, C. R., Kukert, H., Wheatcroft, R. A., Jumars, P. A. & Deming, J. W. (1989) Vent fauna on whale remains. Nature, 341, 27 - 28. https: // doi. org / 10.1038 / 341027 a 0" type="journal article" year="1989">
|
||
Smith
|
||
<emphasis id="B931EAA838249D5BFADFDD039757935A" box="[1308,1358,1885,1908]" italics="true" pageId="1" pageNumber="593">et al</emphasis>
|
||
. 1989
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD44B4B38249D5BFF54DDDE937193B9" author="Bennett" box="[151,360,1920,1943]" pageId="1" pageNumber="593" refString="Bennett, B. A., Smith, C. R., Glaser, B. & Maybaum, H. L. (1994) Faunal community structure of a chemotrophic assemblage on whale bones in the deep northeast Pacific Ocean. Marine Ecology Progress Series, 108, 205 - 223. https: // doi. org / 10.3354 / meps 108205" type="journal article" year="1994">
|
||
Bennett
|
||
<emphasis id="B931EAA838249D5BFF31DDDE933B93B9" box="[242,290,1920,1943]" italics="true" pageId="1" pageNumber="593">et al</emphasis>
|
||
. 1994
|
||
</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="4C454D3938249D5BFE65DDDE91BA93B9" authority="Araya, Catalan & Aliaga, 2016" authorityName="Araya, Catalan & Aliaga" authorityYear="2016" box="[422,931,1920,1943]" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="593" phylum="Mollusca" rank="species" species="atacamensis">
|
||
<emphasis id="B931EAA838249D5BFE65DDDE905093B9" box="[422,585,1920,1943]" italics="true" pageId="1" pageNumber="593">A. atacamensis</emphasis>
|
||
<bibRefCitation id="EFD44B4B38249D5BFD91DDDE91BA93B9" author="Araya" box="[594,931,1920,1943]" pageId="1" pageNumber="593" refString="Araya, J. F., Catalan, R. & Aliaga, J. A. (2016) A new deep-water Astyris species (Buccinoidea: Columbellidae) from the southeastern Pacific. Zootaxa, 4139 (1), 140 - 144. https: // doi. org / 10.11646 / zootaxa. 4139.1.11" type="journal article" year="2016">Araya, Catalán & Aliaga, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
from northern
|
||
<collectingCountry id="F352762A38249D5BFB89DDDE969E93B9" box="[1098,1159,1920,1943]" name="Chile" pageId="1" pageNumber="593">Chile</collectingCountry>
|
||
. Although
|
||
<taxonomicName id="4C454D3938249D5BFAC3DDDE978293B9" baseAuthorityName="Dall" baseAuthorityYear="1890" box="[1280,1435,1920,1943]" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="593" phylum="Mollusca" rank="species" species="permodesta">
|
||
<emphasis id="B931EAA838249D5BFAC3DDDE978293B9" box="[1280,1435,1920,1943]" italics="true" pageId="1" pageNumber="593">A. permodesta</emphasis>
|
||
</taxonomicName>
|
||
has been reported also from off
|
||
<collectingRegion id="4981F85838249D5BFE30DDFC90229397" box="[499,571,1954,1977]" country="Peru" name="Callao" pageId="1" pageNumber="593">Callao</collectingRegion>
|
||
,
|
||
<collectingCountry id="F352762A38249D5BFD8BDDFC90609397" box="[584,633,1954,1977]" name="Peru" pageId="1" pageNumber="593">Perú</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFD44B4B38249D5BFD4BDDFC91519397" author="Levin" box="[648,840,1954,1977]" pageId="1" pageNumber="593" refString="Levin, L., Gutierrez, D., Rathburn, A., Neira, C., Sellanes, J., Munoz, P., Gallardo, V. & Salamanca, M. (2002) Benthic processes on the Peru margin: a transect across the oxygen minimum zone during the 1997 - 98 El Nino. Progress in Oceanography, 53, 1 - 27. https: // doi. org / 10.1016 / S 0079 - 6611 (02) 00022 - 8" type="journal article" year="2002">
|
||
Levin
|
||
<emphasis id="B931EAA838249D5BFD13DDFC911E9397" box="[720,775,1954,1977]" italics="true" pageId="1" pageNumber="593">et al.</emphasis>
|
||
2002
|
||
</bibRefCitation>
|
||
), this is likely in fact another record of
|
||
<taxonomicName id="4C454D3938249D5BFB3ADDFC935A93F2" authority="(Sellanes 2017)" authorityName="Sellanes" authorityYear="2017" baseAuthorityName="Sellanes" baseAuthorityYear="2017" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="593" phylum="Mollusca" rank="species" species="atacamensis">
|
||
<emphasis id="B931EAA838249D5BFB3ADDFC97859397" box="[1273,1436,1954,1977]" italics="true" pageId="1" pageNumber="593">A. atacamensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD44B4B38249D5BFF5DDD9B932293F2" author="Sellanes" box="[158,315,1989,2012]" pageId="1" pageNumber="593" refString="Sellanes, J. (2017) Comments on " A new deep-water Astyris species (Buccinoidea: Columbellidae) from the southeastern Pacific " by Araya et al. 2016. Zootaxa, 4247 (1), 55 - 56. https: // doi. org / 10.11646 / zootaxa. 4247.1.5" type="journal article" year="2017">Sellanes 2017</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Both of these species are easily separable from
|
||
<taxonomicName id="4C454D3938249D5BFC96DD9B961993F2" authority="Chen & Watanabe & Araya, 2017" authorityName="Chen & Watanabe & Araya" authorityYear="2017" box="[853,1024,1989,2012]" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="593" phylum="Mollusca" rank="species" species="thermophilus" status="sp. nov.">
|
||
<emphasis id="B931EAA838249D5BFC96DD9B961993F2" box="[853,1024,1989,2012]" italics="true" pageId="1" pageNumber="593">A. thermophilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20257D338249D5BFBCADD9B965E93F2" box="[1033,1095,1989,2012]" pageId="1" pageNumber="593" rank="species">
|
||
<emphasis id="B931EAA838249D5BFBCADD9B965E93F2" bold="true" box="[1033,1095,1989,2012]" pageId="1" pageNumber="593">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
as they have a thicker, broader shell with wider aperture and less constricted siphonal canal, as well as having spiral grooves on the base. The pronounced columellar fold seen in
|
||
<taxonomicName id="4C454D3938279D58FDD7DAE490AD94FF" baseAuthorityName="Sellanes" baseAuthorityYear="2017" box="[532,692,186,209]" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="594" phylum="Mollusca" rank="species" species="atacamensis">
|
||
<emphasis id="B931EAA838279D58FDD7DAE490AD94FF" box="[532,692,186,209]" italics="true" pageId="2" pageNumber="594">A. atacamensis</emphasis>
|
||
</taxonomicName>
|
||
and the lirae inside the outer lip are also lacking in
|
||
<taxonomicName id="4C454D3938279D58FB15DAE4979994FF" authority="Chen & Watanabe & Araya, 2017" authorityName="Chen & Watanabe & Araya" authorityYear="2017" box="[1238,1408,186,209]" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="594" phylum="Mollusca" rank="species" species="thermophilus" status="sp. nov.">
|
||
<emphasis id="B931EAA838279D58FB15DAE4979994FF" box="[1238,1408,186,209]" italics="true" pageId="2" pageNumber="594">A. thermophilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20257D338279D58FA45DAE492AE94DA" pageId="2" pageNumber="594" rank="species">
|
||
<emphasis id="B931EAA838279D58FA45DAE492AE94DA" bold="true" pageId="2" pageNumber="594">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Until the discovery of
|
||
<taxonomicName id="4C454D3938279D58FE68DA83904C94DA" authority="Chen & Watanabe & Araya, 2017" authorityName="Chen & Watanabe & Araya" authorityYear="2017" box="[427,597,221,244]" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="594" phylum="Mollusca" rank="species" species="thermophilus" status="sp. nov.">
|
||
<emphasis id="B931EAA838279D58FE68DA83904C94DA" box="[427,597,221,244]" italics="true" pageId="2" pageNumber="594">A. thermophilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20257D338279D58FD9FDA83908E94DA" box="[604,663,221,244]" pageId="2" pageNumber="594" rank="species">
|
||
<emphasis id="B931EAA838279D58FD9FDA83908E94DA" bold="true" box="[604,663,221,244]" pageId="2" pageNumber="594">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C454D3938279D58FD67DA83916994DA" baseAuthorityName="Dall" baseAuthorityYear="1890" box="[676,880,221,244]" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="594" phylum="Mollusca" rank="species" species="permodesta">
|
||
<emphasis id="B931EAA838279D58FD67DA83916994DA" box="[676,880,221,244]" italics="true" pageId="2" pageNumber="594">Astyris permodesta</emphasis>
|
||
</taxonomicName>
|
||
was the only other columbellid species known from deep-sea chemosynthetic ecosystems.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F653138279D58FF04DB7C974F96BA" pageId="2" pageNumber="594" type="reference_group">
|
||
<paragraph id="8BFA36BA38279D58FF04DB7C90B89661" blockId="2.[151,1436,152,660]" pageId="2" pageNumber="594">
|
||
<taxonomicName id="4C454D3938279D58FF04DB7C93BD9517" ID-CoL="67VF5" authority="Chen & Watanabe & Araya, 2017" authorityName="Chen & Watanabe & Araya" authorityYear="2017" box="[199,420,290,313]" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="594" phylum="Mollusca" rank="species" species="thermophilus" status="sp. nov.">
|
||
<emphasis id="B931EAA838279D58FF04DB7C93BD9517" box="[199,420,290,313]" italics="true" pageId="2" pageNumber="594">Astyris thermophilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20257D338279D58FE68DB7C93FE9517" box="[427,487,290,313]" pageId="2" pageNumber="594" rank="species">
|
||
<emphasis id="B931EAA838279D58FE68DB7C93FE9517" bold="true" box="[427,487,290,313]" pageId="2" pageNumber="594">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is conchologically similar to
|
||
<taxonomicName id="4C454D3938279D58FCE7DB7C96719517" ID-CoL="5D29L" authority="Okutani, 1964" authorityName="Okutani" authorityYear="1964" box="[804,1128,290,313]" class="Gastropoda" family="Columbellidae" genus="Zemitrella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="594" phylum="Mollusca" rank="species" species="cera">
|
||
<emphasis id="B931EAA838279D58FCE7DB7C91D09517" box="[804,969,290,313]" italics="true" pageId="2" pageNumber="594">Zemitrella cera</emphasis>
|
||
<bibRefCitation id="EFD44B4B38279D58FC13DB7C96719517" author="Okutani" box="[976,1128,290,313]" pageId="2" pageNumber="594" refString="Okutani, T. (1964) Report on the archibenthal and abyssal gastropod Mollusca mainly collected from Sagami Bay and adjacent waters by the R. V. Soyo-Maru during the years 1955 - 1963. Journal of the Faculty of Science, University of Tokyo, II, 15, 371 - 447." type="journal article" year="1964">Okutani, 1964</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<quantity id="4CBD9B5F38279D58FB69DB7C97259517" box="[1194,1340,290,313]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="2" pageNumber="594" unit="m" value="2000.0" valueMax="3000.0" valueMin="1000.0">1000–3000 m</quantity>
|
||
deep off Izu Islands,
|
||
<collectingCountry id="F352762A38279D58FEE0DB1B93469572" box="[291,351,325,348]" name="Japan" pageId="2" pageNumber="594">Japan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFD44B4B38279D58FEB1DB1B900D9572" author="Okutani" box="[370,532,325,348]" pageId="2" pageNumber="594" refString="Okutani, T. (1964) Report on the archibenthal and abyssal gastropod Mollusca mainly collected from Sagami Bay and adjacent waters by the R. V. Soyo-Maru during the years 1955 - 1963. Journal of the Faculty of Science, University of Tokyo, II, 15, 371 - 447." type="journal article" year="1964">Okutani, 1964</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD44B4B38279D58FDE7DB1B90449572" author="Okutani" box="[548,605,325,348]" pageId="2" pageNumber="594" refString="Okutani, T. (2017) Marine Mollusks in Japan. 2 nd Edition. Tokai University Press, Tokyo, 1382 pp." type="book" year="2017">2017</bibRefCitation>
|
||
), but differs by having spiral striation across the entire teleoconch whorl (sculpture is only present on the base in
|
||
<taxonomicName id="4C454D3938279D58FD97DB3690B89551" ID-CoL="5D29L" authorityName="Okutani" authorityYear="1964" box="[596,673,360,383]" class="Gastropoda" family="Columbellidae" genus="Zemitrella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="594" phylum="Mollusca" rank="species" species="cera">
|
||
<emphasis id="B931EAA838279D58FD97DB3690B89551" box="[596,673,360,383]" italics="true" pageId="2" pageNumber="594">Z. cera</emphasis>
|
||
</taxonomicName>
|
||
) and a slightly longer siphonal canal. As the protoconch of
|
||
<taxonomicName id="4C454D3938279D58FAF7DB3697989551" ID-CoL="5D29L" authorityName="Okutani" authorityYear="1964" box="[1332,1409,360,383]" class="Gastropoda" family="Columbellidae" genus="Zemitrella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="594" phylum="Mollusca" rank="species" species="cera">
|
||
<emphasis id="B931EAA838279D58FAF7DB3697989551" box="[1332,1409,360,383]" italics="true" pageId="2" pageNumber="594">Z. cera</emphasis>
|
||
</taxonomicName>
|
||
is unknown, it is not transferred to
|
||
<taxonomicName id="4C454D3938279D58FDD7DBD49047958F" ID-CoL="7NPZ9" authorityName="Chen & Watanabe & Araya" authorityYear="2017" box="[532,606,394,417]" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="594" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B931EAA838279D58FDD7DBD49047958F" box="[532,606,394,417]" italics="true" pageId="2" pageNumber="594">Astyris</emphasis>
|
||
</taxonomicName>
|
||
at this point. Furthermore, the new species is also very similar to a yet undescribed species reported from deep-water trawls and dredges carried out in northern Pacific coast of Honshu,
|
||
<collectingCountry id="F352762A38279D58FA9DDBF3978295EA" box="[1374,1435,429,452]" name="Japan" pageId="2" pageNumber="594">Japan</collectingCountry>
|
||
(
|
||
<taxonomicName id="4C454D3938279D58FF63DB8E92EF95C9" ID-CoL="7P94L" authorityName="Risso" authorityYear="1826" box="[160,246,464,487]" class="Gastropoda" family="Columbellidae" genus="Mitrella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="594" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B931EAA838279D58FF63DB8E92EF95C9" box="[160,246,464,487]" italics="true" pageId="2" pageNumber="594">Mitrella</emphasis>
|
||
</taxonomicName>
|
||
sp. 1
|
||
<emphasis id="B931EAA838279D58FEF4DB8E936A95C9" box="[311,371,464,487]" italics="true" pageId="2" pageNumber="594">sensu</emphasis>
|
||
<bibRefCitation id="EFD44B4B38279D58FEB9DB8E903795C9" author="Hasegawa" box="[378,558,464,487]" pageId="2" pageNumber="594" refString="Hasegawa, K. (2009) Upper bathyal gastropods of the Pacific Coast of northern Honshu, Japan, chiefly collected by R / V Wakataka- Maru. In: Fujita, T. (Ed.), Deep-sea Fauna and Pollutants off Pacific Coast of Northern Japan. National Museum of Nature and Science Monographs, 39, pp. 225 - 383." type="journal article" year="2009">Hasegawa, 2009</bibRefCitation>
|
||
: figs. 162–166), but differs from that species in the spiral striae on the teleoconch being less distinct and that the columella is always straight. At this point we cannot entirely dismiss the possibility that
|
||
<taxonomicName id="4C454D3938279D58FF54D84B92F79602" ID-CoL="7P94L" authorityName="Risso" authorityYear="1826" box="[151,238,533,556]" class="Gastropoda" family="Columbellidae" genus="Mitrella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="594" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B931EAA838279D58FF54D84B92F79602" box="[151,238,533,556]" italics="true" pageId="2" pageNumber="594">Mitrella</emphasis>
|
||
</taxonomicName>
|
||
sp. 1 (
|
||
<emphasis id="B931EAA838279D58FE81D84B93679602" box="[322,382,533,556]" italics="true" pageId="2" pageNumber="594">sensu</emphasis>
|
||
<bibRefCitation id="EFD44B4B38279D58FE4BD84B90279602" author="Hasegawa" box="[392,574,533,556]" pageId="2" pageNumber="594" refString="Hasegawa, K. (2009) Upper bathyal gastropods of the Pacific Coast of northern Honshu, Japan, chiefly collected by R / V Wakataka- Maru. In: Fujita, T. (Ed.), Deep-sea Fauna and Pollutants off Pacific Coast of Northern Japan. National Museum of Nature and Science Monographs, 39, pp. 225 - 383." type="journal article" year="2009">Hasegawa, 2009</bibRefCitation>
|
||
) may be conspecific with
|
||
<taxonomicName id="4C454D3938279D58FCAFD84B96549602" ID-CoL="67VF5" authority="Chen & Watanabe & Araya, 2017" authorityName="Chen & Watanabe & Araya" authorityYear="2017" box="[876,1101,533,556]" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="594" phylum="Mollusca" rank="species" species="thermophilus" status="sp. nov.">
|
||
<emphasis id="B931EAA838279D58FCAFD84B96549602" box="[876,1101,533,556]" italics="true" pageId="2" pageNumber="594">Astyris thermophilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20257D338279D58FB9BD84B96809602" box="[1112,1177,533,556]" pageId="2" pageNumber="594" rank="species">
|
||
<emphasis id="B931EAA838279D58FB9BD84B96809602" bold="true" box="[1112,1177,533,556]" pageId="2" pageNumber="594">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and the morphological differences may be due to environmental factors.
|
||
</paragraph>
|
||
<paragraph id="8BFA36BA38279D58FF04D804974F96BA" blockId="2.[151,1436,152,660]" pageId="2" pageNumber="594">
|
||
The phylogenetic reconstruction (
|
||
<figureCitation id="137E2A3F38279D58FDECD8049076965F" box="[559,623,602,625]" captionStart="FIGURE 2" captionStartId="3.[151,244,1124,1146]" captionTargetBox="[159,1420,203,1091]" captionTargetId="figure@3.[151,1436,191,1103]" captionTargetPageId="3" captionText="FIGURE 2. Phylogenetic reconstruction using the COI gene (618 bp) showing Astyris thermophilus n. sp. nested within Columbellidae. Node values indicate Bayesian posterior probability, those lower than 0.7 are omitted. The DDBJ/EMBL/GenBank accession numbers for each sequence used are shown inside brackets (LC329027 and LC329028 for the two sequenced specimens of A.thermophilus n. sp.)." httpUri="https://zenodo.org/record/1114845/files/figure.png" pageId="2" pageNumber="594">Fig. 2</figureCitation>
|
||
) confirmed the placement of the present new species within
|
||
<taxonomicName id="4C454D3938279D58FB3BD8049781965F" ID-CoL="7NGWV" authorityName="Swainson" authorityYear="1840" box="[1272,1432,602,625]" class="Gastropoda" family="Columbellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="594" phylum="Mollusca" rank="family">Columbellidae</taxonomicName>
|
||
. Its generic placement cannot be tested at present because sequences from other
|
||
<taxonomicName id="4C454D3938279D58FC24D823962896BA" ID-CoL="7NPZ9" authorityName="Chen & Watanabe & Araya" authorityYear="2017" box="[999,1073,637,660]" class="Gastropoda" family="Columbellidae" genus="Astyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="594" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B931EAA838279D58FC24D823962896BA" box="[999,1073,637,660]" italics="true" pageId="2" pageNumber="594">Astyris</emphasis>
|
||
</taxonomicName>
|
||
species were not available.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |