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<document id="B4650420EB18758F51A14B310509A4F6" ID-CLB-Dataset="8721" ID-DOI="10.11646/zootaxa.4823.1.1" ID-GBIF-Dataset="a43d0118-9df5-4970-a316-0dae5c9caebf" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401916" ID-ZooBank="3DF06F26-FD35-452E-9056-1D05B49946EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609340622775" checkinUser="plazi" docAuthor="Dombrow, Holger E. &amp; Colville, Jonathan F." docDate="2020" docId="03DFBE05E03FFFD1FF64D551FC32F818" docLanguage="en" docName="zootaxa.4823.1.1.pdf" docOrigin="Zootaxa 4823" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Beckhoplia pulchra Dombrow 2020, new species" docType="treatment" docUuid="AB9FCC8F-A5DA-4004-8143-DE08345C578B" docUuidSource="ZooBank" docVersion="9" lastPageNumber="37" masterDocId="FFE6C67DE01CFFF5FFF3D41BFFBDFF8D" masterDocTitle="Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini)" masterLastPageNumber="79" masterPageNumber="1" pageNumber="36" updateTime="1698849823920" updateUser="ExternalLinkService">
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<mods:title id="AF036BCD96FFFE7EC04F3F2D9032FFEE">Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini)</mods:title>
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<mods:namePart id="1E9E5BD50301D693A222C6FF4E61F834">Dombrow, Holger E.</mods:namePart>
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<mods:namePart id="51038DE4FE71AB3FEAF8C99EDF7F9626">Colville, Jonathan F.</mods:namePart>
<mods:nameIdentifier id="0505E8DFB43EB082A001B7BFB96AD678" type="ORCID">0000-0003-2176-3077</mods:nameIdentifier>
<mods:nameIdentifier id="0F432206477FE9BD8BABBA28E2D5002D" type="email">jonathan.colville@gmail.com</mods:nameIdentifier>
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<mods:title id="6B82E81E9925763A48847D84CB141A71">Zootaxa</mods:title>
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<mods:date id="AE6AB5724835F8ED90E1205B11EB960A">2020</mods:date>
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<mods:number id="4509FAAD02AE41AD51070D65CF338F21">2020-08-10</mods:number>
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<treatment id="03DFBE05E03FFFD1FF64D551FC32F818" ID-DOI="http://doi.org/10.5281/zenodo.4455287" ID-GBIF-Taxon="177013516" ID-Zenodo-Dep="4455287" LSID="urn:lsid:zoobank.org:act:AB9FCC8F-A5DA-4004-8143-DE08345C578B" httpUri="http://treatment.plazi.org/id/03DFBE05E03FFFD1FF64D551FC32F818" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<subSubSection id="C36C5C98E03FFFD6FF64D551FD1AFEE9" box="[151,679,330,357]" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="8BC90F13E03FFFD6FF64D551FD1AFEE9" blockId="35.[151,874,330,428]" box="[151,679,330,357]" pageId="35" pageNumber="36">
<heading id="D081B87FE03FFFD6FF64D551FD1AFEE9" bold="true" box="[151,679,330,357]" fontSize="11" level="2" pageId="35" pageNumber="36" reason="4">
<emphasis id="B902D301E03FFFD6FF64D551FD1AFEE9" bold="true" box="[151,679,330,357]" pageId="35" pageNumber="36">
<taxonomicName id="4C767490E03FFFD6FF64D551FDB7FEE8" authority="Dombrow" authorityName="Dombrow" authorityYear="2020" box="[151,522,330,357]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pulchra" status="sp. nov.">
<emphasis id="B902D301E03FFFD6FF64D551FE3BFEE8" bold="true" box="[151,390,330,357]" italics="true" pageId="35" pageNumber="36">Beckhoplia pulchra</emphasis>
Dombrow
</taxonomicName>
,
<taxonomicNameLabel id="A2316E7AE03FFFD6FDE4D550FD1AFEE9" box="[535,679,330,357]" pageId="35" pageNumber="36" rank="species">new species</taxonomicNameLabel>
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</heading>
</paragraph>
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<subSubSection id="C36C5C98E03FFFD6FF64D574FCD7FE21" pageId="35" pageNumber="36" type="description">
<paragraph id="8BC90F13E03FFFD6FF64D574FE2DFE07" blockId="35.[151,874,330,428]" box="[151,400,366,394]" pageId="35" pageNumber="36">
(
<figureCitation id="134D1396E03FFFD6FF6CD574FF40FE04" box="[159,253,366,393]" captionStart="FIGURES 5054" captionStartId="24.[151,264,678,702]" captionTargetBox="[212,1375,186,616]" captionTargetId="figure-25@24.[208,1380,181,622]" captionTargetPageId="24" captionText="FIGURES 5054. General drawing of the types of prothoracic pretarsomere of Beckhoplia Dombrow, 2005 species, in lateral view, with roundish cutting ventrally, ♂♂. 50, distinctly thickened, smooth ventrally; 51, slightly thickened, with a shortened smooth ridge ventrally; 52, distinctly thickened, with a median tooth ventrally; 53, slightly thickened, with two median teeth consecutive ventrally; 54, distinctly thickened, with a shortened granulate ridge ventrally. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4401948" httpUri="https://zenodo.org/record/4401948/files/figure.png" pageId="35" pageNumber="36">Figs. 53</figureCitation>
,
<figureCitation id="134D1396E03FFFD6FEFAD575FEECFE04" box="[265,337,366,393]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="35" pageNumber="36">7475</figureCitation>
,
<figureCitation id="134D1396E03FFFD6FEAED574FE35FE07" box="[349,392,367,394]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="35" pageNumber="36">106</figureCitation>
)
</paragraph>
<paragraph id="8BC90F13E03FFFD6FF64D58FFCD7FE21" blockId="35.[151,874,330,428]" box="[151,874,404,428]" pageId="35" pageNumber="36">
http://zoobank.org/
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<subSubSection id="C36C5C98E03FFFD6FF64D5C7FA3CFC9B" pageId="35" pageNumber="36" type="materials_examined">
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<materialsCitation id="3B1E054EE03FFFD6FF64D5C7FC81FE7B" ID-GBIF-Occurrence="3022772319" box="[151,828,476,503]" collectionCode="NCP" country="South Africa" location="Farm Papkuilsfontein." pageId="35" pageNumber="36" specimenCount="1" typeStatus="holotype">
<emphasis id="B902D301E03FFFD6FF64D5C7FE8BFE7B" bold="true" box="[151,310,476,502]" pageId="35" pageNumber="36">
<typeStatus id="54CDB1B1E03FFFD6FF64D5C7FF6FFE7B" box="[151,210,476,502]" pageId="35" pageNumber="36">Type</typeStatus>
locality.
</emphasis>
[
<collectingCountry id="F3614F83E03FFFD6FEB0D5C7FEC2FE7B" box="[323,383,476,502]" name="South Africa" pageId="35" pageNumber="36">RSA</collectingCountry>
,
<collectionCode id="ED6797D6E03FFFD6FE79D5C7FE7AFE7A" box="[394,455,476,503]" country="United Kingdom" name="National Collection of Passiflora" pageId="35" pageNumber="36" type="Herbarium">NCP</collectionCode>
]:
<quantity id="4C8EA2F6E03FFFD6FE2AD5C7FDB6FE7B" box="[473,523,476,502]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="36" unit="km" value="2.0">2km</quantity>
SW
<location id="8EA959C8E03FFFD6FDB1D5C6FC81FE7B" LSID="urn:lsid:plazi:treatment:03DFBE05E03FFFD1FF64D551FC32F818:8EA959C8E03FFFD6FDB1D5C6FC81FE7B" box="[578,828,476,503]" country="South Africa" name="Farm Papkuilsfontein." pageId="35" pageNumber="36">Farm Papkuilsfontein.</location>
</materialsCitation>
</paragraph>
<paragraph id="8BC90F13E03FFFD6FF64D63FFE85FD43" blockId="35.[151,1437,548,718]" pageId="35" pageNumber="36">
<emphasis id="B902D301E03FFFD6FF64D63FFEF8FDB3" bold="true" box="[151,325,548,574]" pageId="35" pageNumber="36">Type material.</emphasis>
[
<materialsCitation id="3B1E054EE03FFFD6FEA6D63FFE85FD43" ID-GBIF-Occurrence="3022772320" collectingDate="1989-08-30" collectionCode="ZSM" collectorName="Fritz &amp; Dombrow" country="South Africa" determinerName="H. Dombrow" elevation="680" latitude="-31.5725" location="Farm Papkuilsfontein" longLatPrecision="20" longitude="19.18111" pageId="35" pageNumber="36" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="holotype">
1]
<emphasis id="B902D301E03FFFD6FE80D63FFE5DFDB3" bold="true" box="[371,480,548,574]" pageId="35" pageNumber="36">
<typeStatus id="54CDB1B1E03FFFD6FE80D63FFE5DFDB3" box="[371,480,548,574]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D70C49AE03FFFD6FE14D63EFE42FDB2" box="[487,511,549,575]" pageId="35" pageNumber="36" type="male"></specimenCount>
: white label pr «AFRIKA:
<collectingCountry id="F3614F83E03FFFD6FCC3D63FFC1DFDB3" box="[816,928,548,574]" name="South Africa" pageId="35" pageNumber="36">Südafrika</collectingCountry>
,
<collectingRegion id="49B2C1F1E03FFFD6FC59D63FFB93FDB3" box="[938,1070,548,574]" country="South Africa" name="Western Cape" pageId="35" pageNumber="36">Kapprovinz</collectingRegion>
|
<quantity id="4C8EA2F6E03FFFD6FBB1D63FFBCEFDB3" box="[1090,1139,548,574]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="36" unit="km" value="2.0">2km</quantity>
SW
<location id="8EA959C8E03FFFD6FB58D63EFA21FDB3" LSID="urn:lsid:plazi:treatment:03DFBE05E03FFFD1FF64D551FC32F818:8EA959C8E03FFFD6FB58D63EFA21FDB3" box="[1195,1436,548,575]" country="South Africa" latitude="-31.5725" longLatPrecision="20" longitude="19.18111" name="Farm Papkuilsfontein" pageId="35" pageNumber="36" stateProvince="Western Cape">Farm Papkuilsfontein</location>
|
<geoCoordinate id="EE4269D4E03FFFD6FF50D653FE97FDEF" box="[163,298,584,610]" degrees="31" direction="south" minutes="34" orientation="latitude" pageId="35" pageNumber="36" precision="15" seconds="21" value="-31.5725">31°3421”S</geoCoordinate>
,
<geoCoordinate id="EE4269D4E03FFFD6FEC6D653FE03FDEE" box="[309,446,584,611]" degrees="19" direction="east" minutes="10" orientation="longitude" pageId="35" pageNumber="36" precision="15" seconds="52" value="19.18111">19°1052”E</geoCoordinate>
,
<quantity id="4C8EA2F6E03FFFD6FE3AD653FDADFDEE" box="[457,528,584,611]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="35" pageNumber="36" unit="m" value="680.0">
<elevation id="005BE820E03FFFD6FE3AD653FDADFDEE" box="[457,528,584,611]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="35" pageNumber="36" unit="m" value="680.0">680 m</elevation>
</quantity>
|
<date id="FFC829D3E03FFFD6FDD1D653FD1EFDEF" box="[546,675,584,610]" pageId="35" pageNumber="36" value="1989-08-30">
<collectingDate id="EF8CD03BE03FFFD6FDD1D653FD1EFDEF" box="[546,675,584,610]" pageId="35" pageNumber="36" value="1989-08-30">30.08.1989</collectingDate>
</date>
, leg.
<collectorName id="26836AC5E03FFFD6FD2DD653FCA8FDEF" box="[734,789,584,610]" pageId="35" pageNumber="36">Fritz</collectorName>
» || green label pr «Host plant: |
<taxonomicName id="4C767490E03FFFD6FB86D653FA82FDEF" box="[1141,1343,584,610]" class="Liliopsida" family="Iridaceae" genus="Babiana" kingdom="Plantae" order="Asparagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="vanziliae">
<emphasis id="B902D301E03FFFD6FB86D653FA82FDEF" box="[1141,1343,584,610]" italics="true" pageId="35" pageNumber="36">Babiana vanziliae</emphasis>
</taxonomicName>
» || blue label pr «Bioregion: | Trans-Escarpment Succulent Karroo | Vegetation Type: SKt2 | Hantam Karroo» || yellow label hw «54», pr «.921 | coll.
<collectorName id="26836AC5E03FFFD6FE1DD68BFDDFFD27" box="[494,610,656,682]" pageId="35" pageNumber="36">Dombrow</collectorName>
» || red label pr «
<typeStatus id="54CDB1B1E03FFFD6FCEAD68BFC3EFD27" box="[793,899,656,682]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
|
<taxonomicName id="4C767490E03FFFD6FC64D68BFBA8FD27" authorityName="Dombrow" authorityYear="2005" box="[919,1045,656,682]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B902D301E03FFFD6FC64D68BFBA8FD27" box="[919,1045,656,682]" italics="true" pageId="35" pageNumber="36">Beckhoplia</emphasis>
</taxonomicName>
|
<taxonomicName id="4C767490E03FFFD6FBDAD68BFB3DFD27" authority="Dombrow &amp; Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[1065,1152,656,682]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pulchra" status="sp. nov.">
<emphasis id="B902D301E03FFFD6FBDAD68BFB3DFD27" box="[1065,1152,656,682]" italics="true" pageId="35" pageNumber="36">pulchra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2316E7AE03FFFD6FB74D68AFB01FD26" box="[1159,1212,657,683]" pageId="35" pageNumber="36" rank="species">sp.n.</taxonomicNameLabel>
| det.:
<determinerName id="7E15E3D7E03FFFD6FAFBD68AFA20FD27" box="[1288,1437,656,683]" pageId="35" pageNumber="36">H. Dombrow</determinerName>
2019» (
<collectionCode id="ED6797D6E03FFFD6FF1FD6AFFE96FD43" box="[236,299,692,718]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="35" pageNumber="36" type="Museum">ZSM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BC90F13E03FFFD6FF64D6E7FA3CFC9B" blockId="35.[151,1409,764,791]" box="[151,1409,764,791]" pageId="35" pageNumber="36">
<emphasis id="B902D301E03FFFD6FF64D6E7FE21FC9B" bold="true" box="[151,412,764,790]" pageId="35" pageNumber="36">
Condition of
<typeStatus id="54CDB1B1E03FFFD6FEC2D6E7FE2AFC9B" box="[305,407,764,790]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
.
</emphasis>
Excellent. Glued on card. Dissected, parameres and spiculum gastrale glued behind beetle.
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E03FFFD6FF64D75FFCB1FB9E" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="8BC90F13E03FFFD6FF64D75FFCB1FB9E" blockId="35.[151,1437,836,1043]" pageId="35" pageNumber="36">
<emphasis id="B902D301E03FFFD6FF64D75FFE23FCD3" bold="true" box="[151,414,836,862]" pageId="35" pageNumber="36">Differential diagnosis.</emphasis>
Differs from
<taxonomicName id="4C767490E03FFFD6FDCBD75EFCB2FCD3" authority=": Lateral" authorityName="Lateral" box="[568,783,836,863]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="colvillei">
<emphasis id="B902D301E03FFFD6FDCBD75EFD0CFCD3" box="[568,689,836,862]" italics="true" pageId="35" pageNumber="36">B. colvillei</emphasis>
: Lateral
</taxonomicName>
margin of pronotum of
<taxonomicName id="4C767490E03FFFD6FBEED75EFB2EFCD3" authorityName="Dombrow" authorityYear="2020" box="[1053,1171,836,862]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B902D301E03FFFD6FBEED75EFB2EFCD3" box="[1053,1171,836,862]" italics="true" pageId="35" pageNumber="36">B. pulchra</emphasis>
</taxonomicName>
widest at posterior corner. Lateral margin of pronotum of
<taxonomicName id="4C767490E03FFFD6FDD6D772FD23FC0F" box="[549,670,872,898]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="colvillei">
<emphasis id="B902D301E03FFFD6FDD6D772FD23FC0F" box="[549,670,872,898]" italics="true" pageId="35" pageNumber="36">B. colvillei</emphasis>
</taxonomicName>
widest in anterior half. Externo-posterior margin of parameres of
<taxonomicName id="4C767490E03FFFD6FA77D772FF53FC2B" authorityName="Dombrow" authorityYear="2020" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B902D301E03FFFD6FA77D772FF53FC2B" italics="true" pageId="35" pageNumber="36">B. pulchra</emphasis>
</taxonomicName>
undulate in dorsal view (
<figureCitation id="134D1396E03FFFD6FDF8D797FDD7FC2B" box="[523,618,908,934]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="35" pageNumber="36">Figs. 74</figureCitation>
,
<figureCitation id="134D1396E03FFFD6FD86D797FD1EFC2A" box="[629,675,908,935]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="35" pageNumber="36">106</figureCitation>
). Externo-posterior margin of parameres of
<taxonomicName id="4C767490E03FFFD6FB63D796FAB4FC2B" box="[1168,1289,908,934]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="colvillei">
<emphasis id="B902D301E03FFFD6FB63D796FAB4FC2B" box="[1168,1289,908,934]" italics="true" pageId="35" pageNumber="36">B. colvillei</emphasis>
</taxonomicName>
linear with a median convex extension in dorsal view (
<figureCitation id="134D1396E03FFFD6FD94D7ABFD78FC47" box="[615,709,944,970]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="35" pageNumber="36">Figs. 72</figureCitation>
,
<figureCitation id="134D1396E03FFFD6FD3CD7ABFD40FC46" box="[719,765,944,971]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="35" pageNumber="36">105</figureCitation>
). Interno-posterior margin of parameres of
<taxonomicName id="4C767490E03FFFD6FB13D7AAFAEBFC47" authorityName="Dombrow" authorityYear="2020" box="[1248,1366,944,970]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B902D301E03FFFD6FB13D7AAFAEBFC47" box="[1248,1366,944,970]" italics="true" pageId="35" pageNumber="36">B. pulchra</emphasis>
</taxonomicName>
broad curly bracket-shaped in dorsal view (
<figureCitation id="134D1396E03FFFD6FDCBD7CFFD2BFC63" box="[568,662,980,1006]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="35" pageNumber="36">Figs. 74</figureCitation>
,
<figureCitation id="134D1396E03FFFD6FD52D7CFFD72FC62" box="[673,719,980,1007]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="35" pageNumber="36">106</figureCitation>
). Interno-posterior margin of parameres of
<taxonomicName id="4C767490E03FFFD6FB45D7CEFA92FC63" box="[1206,1327,980,1006]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="colvillei">
<emphasis id="B902D301E03FFFD6FB45D7CEFA92FC63" box="[1206,1327,980,1006]" italics="true" pageId="35" pageNumber="36">B. colvillei</emphasis>
</taxonomicName>
very narrow curly bracket-shaped in dorsal view (
<figureCitation id="134D1396E03FFFD6FD9AD7E3FD7AFB9F" box="[617,711,1016,1042]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="35" pageNumber="36">Figs. 72</figureCitation>
,
<figureCitation id="134D1396E03FFFD6FD21D7E3FCBDFB9E" box="[722,768,1016,1043]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="35" pageNumber="36">105</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E03FFFD1FF64D05BFEDEF9F8" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="description">
<paragraph id="8BC90F13E03FFFD6FF64D05BFE9CFBD7" blockId="35.[151,1437,1088,2015]" box="[151,289,1088,1114]" pageId="35" pageNumber="36">
<emphasis id="B902D301E03FFFD6FF64D05BFE9CFBD7" bold="true" box="[151,289,1088,1114]" pageId="35" pageNumber="36">Description</emphasis>
</paragraph>
<paragraph id="8BC90F13E03FFFD6FF34D07EFD61FBF2" blockId="35.[151,1437,1088,2015]" box="[199,732,1124,1151]" pageId="35" pageNumber="36">
<emphasis id="B902D301E03FFFD6FF34D07EFE3DFBF2" bold="true" box="[199,384,1125,1151]" pageId="35" pageNumber="36">Measurements:</emphasis>
length
<quantity id="4C8EA2F6E03FFFD6FE27D07FFD90FBF2" box="[468,557,1124,1151]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="35" pageNumber="36" unit="mm" value="7.8">7.8 mm</quantity>
; width 4.0 mm.
</paragraph>
<paragraph id="8BC90F13E03FFFD6FF34D093FD5CFB2F" blockId="35.[151,1437,1088,2015]" box="[199,737,1160,1186]" pageId="35" pageNumber="36">
<emphasis id="B902D301E03FFFD6FF34D093FEE6FB2F" bold="true" box="[199,347,1160,1186]" pageId="35" pageNumber="36">Integument:</emphasis>
Black; elytra transparently brown.
</paragraph>
<paragraph id="8BC90F13E03FFFD1FF34D0B7FBBAFE20" blockId="35.[151,1437,1088,2015]" lastBlockId="36.[151,1437,151,1655]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<emphasis id="B902D301E03FFFD6FF34D0B7FEFAFB4B" bold="true" box="[199,327,1196,1222]" pageId="35" pageNumber="36">Tegument:</emphasis>
<emphasis id="B902D301E03FFFD6FEA3D0B7FE17FB4B" box="[336,426,1196,1222]" italics="true" pageId="35" pageNumber="36">Clypeus</emphasis>
bare; sparse setose with very short, erect, brown setae on the inner side of the vertically reflexed anterior and lateral margin.
<emphasis id="B902D301E03FFFD6FDC5D0CBFDCEFB67" box="[566,627,1232,1258]" italics="true" pageId="35" pageNumber="36">Head</emphasis>
loose setose with medium-long to long, erect, black setae.
<emphasis id="B902D301E03FFFD6FAE0D0CAFA21FB67" box="[1299,1436,1232,1258]" italics="true" pageId="35" pageNumber="36">Eye canthus</emphasis>
with a loose setose ciliary of medium-long, black setae.
<taxonomicName id="4C767490E03FFFD6FCE3D0EEFCE2FA82" authorityName="Norman" authorityYear="1875" box="[784,863,1269,1295]" genus="Scapus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Cnidaria" rank="genus">
<emphasis id="B902D301E03FFFD6FCE3D0EEFCE2FA82" box="[784,863,1269,1295]" italics="true" pageId="35" pageNumber="36">Scapus</emphasis>
</taxonomicName>
loose setose with medium-long, erect, black setae, brown ventrally, sparse and short on
<emphasis id="B902D301E03FFFD6FDB2D103FC88FABF" box="[577,821,1304,1330]" italics="true" pageId="35" pageNumber="36">pedicellus; funiculum</emphasis>
with few very short, erect, black setae;
<emphasis id="B902D301E03FFFD6FAF6D103FAF0FABF" box="[1285,1357,1304,1330]" italics="true" pageId="35" pageNumber="36">clavus</emphasis>
sparse setose with extremely short, semi-erect, brown setae.
<emphasis id="B902D301E03FFFD6FCF1D126FCCCFADB" box="[770,881,1341,1366]" italics="true" pageId="35" pageNumber="36">Pronotum</emphasis>
loose setose with medium-long and long, erect, black setae; with a loose setose ciliary of long to distinctly long, black setae on lateral margin.
<taxonomicName id="4C767490E03FFFD6FB1AD17BFAEAFAF7" box="[1257,1367,1376,1402]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B902D301E03FFFD6FB1AD17BFAEAFAF7" box="[1257,1367,1376,1402]" italics="true" pageId="35" pageNumber="36">Scutellum</emphasis>
</taxonomicName>
loose setose with short to medium-long, semi-erect, brown setae, with a longitudinal, median, bar strip in anterior half.
<emphasis id="B902D301E03FFFD6FF64D1B3FF60FA4F" box="[151,221,1448,1474]" italics="true" pageId="35" pageNumber="36">Elytra</emphasis>
without pattern; sparse setose with very short, reclinate, brownish setae; short to medium-long, semi-erect, black setae interspersed, lengthening towards anterior margin; reclinate, narrowly-lanceolate, white scales interspersed medially behind disk and around apical torus; with a sparse juxta-sutural row of robust, long, erect, black setae, from anterior margin, crossing scutellar torus towards apical torus. Suture with a loose row of short to long, robust, semi-erect, black setae.
<emphasis id="B902D301E03FFFD6FE09D223FDF6F9DF" box="[506,587,1592,1618]" italics="true" pageId="35" pageNumber="36">Thorax</emphasis>
and
<emphasis id="B902D301E03FFFD6FD71D223FD59F9DF" box="[642,740,1592,1618]" italics="true" pageId="35" pageNumber="36">pleurites</emphasis>
loose pilose with long, black hairs.
<emphasis id="B902D301E03FFFD6FB8FD222FABFF9DF" box="[1148,1282,1593,1618]" italics="true" pageId="35" pageNumber="36">Ventrite IV</emphasis>
sparse setose with short, semi-erect, white setae; medium-long to long, semi-erect, black setae interspersed.
<taxonomicName id="4C767490E03FFFD6FB4DD247FAF2F9FB" authorityName="Bocchino" authorityYear="1964" box="[1214,1359,1628,1654]" class="Actinopterygii" genus="Propygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="35" pageNumber="36" phylum="Chordata" rank="genus">
<emphasis id="B902D301E03FFFD6FB4DD247FAF2F9FB" box="[1214,1359,1628,1654]" italics="true" pageId="35" pageNumber="36">Propygidium</emphasis>
</taxonomicName>
sparse setose with medium-long, semi-erect, black setae; short, reclinate, white setae interspersed at posterior margin.
<taxonomicName id="4C767490E03FFFD6FA86D29AFF58F933" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="35" pageNumber="36" phylum="Chordata" rank="genus">
<emphasis id="B902D301E03FFFD6FA86D29AFF58F933" italics="true" pageId="35" pageNumber="36">Pygidium</emphasis>
</taxonomicName>
loose setose with medium-long and long, erect, black setae; tip with a sparse ciliary of medium-long, black setae.
<emphasis id="B902D301E03FFFD6FF2CD2D3FE12F96F" box="[223,431,1736,1762]" italics="true" pageId="35" pageNumber="36">Prothoracic femur</emphasis>
loose pilose with long, black hairs; bare internally.
<emphasis id="B902D301E03FFFD6FBF0D2D3FB78F96F" box="[1027,1221,1736,1762]" italics="true" pageId="35" pageNumber="36">Prothoracic tibia</emphasis>
sparse setose with short, semi-erect, offwhite setae, sparser ventrally; medium-long, semi-erect, brown setae interspersed, longer and erect ventrally; with a sparse setose longitudinal row of medium-long, erect, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; a setaceous ciliary of short, brown setae on anterior margin; a longer, downwards pointing setose ciliary of brown setae along anterior margin of apical tooth.
<emphasis id="B902D301E03FFFD6FB84D343FAE4F8FF" box="[1143,1369,1880,1906]" italics="true" pageId="35" pageNumber="36">Mesothoracic femur</emphasis>
loose pilose with long, black hairs, distinctly long and brown ventrally; bare internally.
<emphasis id="B902D301E03FFFD6FBDDD367FABEF81B" box="[1070,1283,1916,1942]" italics="true" pageId="35" pageNumber="36">Mesothoracic tibia</emphasis>
sparse setose with short, reclinate, whitish setae; medium-long, semi-erect, black, robust setae interspersed, sparser, finer, medium-long internally. Posterior margin with a bristled ciliary of short, black bristles, lengthening dorsally, missing internally.
<emphasis id="B902D301E038FFD1FEE2D48CFE4CFF3C" box="[273,497,151,177]" italics="true" pageId="36" pageNumber="37">Metathoracic femur</emphasis>
loose setose with long to distinctly long, erect, black setae; pale-brown ventrally; bare internally.
<emphasis id="B902D301E038FFD1FEB7D4A0FDA8FF58" box="[324,533,187,213]" italics="true" pageId="36" pageNumber="37">Metathoracic tibia</emphasis>
sparse setose with very short and short, reclinate, brown setae; medium-long and long, robust, semi-erect, black setae interspersed, finer and longer internally. Posterior margin with a loose bristled outer marginal ciliary of short and medium-long, brown bristles, lengthening dorsally and ventrally.
<emphasis id="B902D301E038FFD1FAD5D518FF74FECD" italics="true" pageId="36" pageNumber="37">Tarsomere IIV</emphasis>
of all thoracic legs sparse setose with extremely short, reclinate, black setae; with a setaceous, apical, marginal ciliary of short, black setae, lengthening towards
<emphasis id="B902D301E038FFD1FD4CD550FC69FEE9" box="[703,980,331,357]" italics="true" pageId="36" pageNumber="37">metathoracic tarsomeres</emphasis>
, fine dorsally and more robust ventrally.
<emphasis id="B902D301E038FFD1FF64D574FE92FE05" box="[151,303,367,392]" italics="true" pageId="36" pageNumber="37">Pretarsomere</emphasis>
of all thoracic legs sparse setose with very short, reclinate, pale-brown setae; few, medium-long to long, semi-erect, black setae at apical margin, shorter and red-brown ventrally.
</paragraph>
<paragraph id="8BC90F13E038FFD1FF34D5ACFC06FD94" blockId="36.[151,1437,151,1655]" pageId="36" pageNumber="37">
<emphasis id="B902D301E038FFD1FF34D5ACFEADFE5C" bold="true" box="[199,272,439,465]" pageId="36" pageNumber="37">Head:</emphasis>
Suture between head and clypeus linear, complanate. Head dense punctate with small, shallow, roundish, not merging punctures. Funicle III cylindrical, slightly thickened medially; funicle III asymmetric frustum-shaped; clavus elliptic, slightly longer than pedicellus and funiculum combined.
</paragraph>
<paragraph id="8BC90F13E038FFD1FF34D638FB33FD08" blockId="36.[151,1437,151,1655]" pageId="36" pageNumber="37">
<emphasis id="B902D301E038FFD1FF34D638FEF4FDB0" bold="true" box="[199,329,547,573]" pageId="36" pageNumber="37">Pronotum:</emphasis>
Widest at anterior two fifth; loose punctate with shallow, very small, roundish punctures, dense in posterior third of medial furrow. Anterior corner obtuse. Lateral margin linearly attenuate anteriorly, subparallel posteriorly. With an indistinct, complete, shallow medial furrow. Posterior corner rounded.
</paragraph>
<paragraph id="8BC90F13E038FFD1FF34D695FCC6FD7C" blockId="36.[151,1437,151,1655]" pageId="36" pageNumber="37">
<emphasis id="B902D301E038FFD1FF34D695FEF5FD25" bold="true" box="[199,328,654,680]" pageId="36" pageNumber="37">
<taxonomicName id="4C767490E038FFD1FF34D695FEFEFD25" box="[199,323,654,680]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="36" pageNumber="70" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
:
</emphasis>
Parabola-shaped; longer than broad; dense punctate with shallow, small to large, roundish punctures, micropunctate within; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating medially. Lateral margin slightly convex. Tip pointed.
</paragraph>
<paragraph id="8BC90F13E038FFD1FF34D6E0FE9CFC60" blockId="36.[151,1437,151,1655]" pageId="36" pageNumber="37">
<emphasis id="B902D301E038FFD1FF34D6E0FEA0FC98" bold="true" box="[199,285,763,789]" pageId="36" pageNumber="37">Elytra:</emphasis>
Ovate; broader than long; widest in anterior third; protruding
<taxonomicName id="4C767490E038FFD1FC07D6E0FB35FC98" authorityName="Bocchino" authorityYear="1964" box="[1012,1160,763,789]" class="Actinopterygii" genus="Propygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="36" pageNumber="70" phylum="Chordata" rank="genus">propygidium</taxonomicName>
. Lateral margin slightly convex attenuate apically. Sculpture indistinct; slightly pulvinate laterally; humerus indistinct; with a very broad roundish torus around
<taxonomicName id="4C767490E038FFD1FE62D758FDBFFCD0" box="[401,514,835,861]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="36" pageNumber="70" phylum="Arthropoda" rank="genus">scutellum</taxonomicName>
; lateral trench missing; complanate beside
<taxonomicName id="4C767490E038FFD1FC2FD758FBF4FCD0" box="[988,1097,835,861]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="36" pageNumber="70" phylum="Arthropoda" rank="genus">scutellum</taxonomicName>
and along suture. Loose punctate with very small, shallow, roundish punctures, micropunctures in-between; a sparse juxta-sutural row of small, shallow, roundish punctures from anterior margin, crossing scutellar torus towards apical torus; a sparse supra-humeral row of small, shallow, roundish punctures from anterior margin between humerus and scutellar torus towards apical torus.
</paragraph>
<paragraph id="8BC90F13E038FFD1FF34D7EDFB53FBD4" blockId="36.[151,1437,151,1655]" pageId="36" pageNumber="37">
<emphasis id="B902D301E038FFD1FF34D7EDFEF8FB9D" bold="true" box="[199,325,1014,1040]" pageId="36" pageNumber="37">Abdomen:</emphasis>
Ventrite I extremely shortened medially; ventrite V broadest, not constricted ventrally.
<taxonomicName id="4C767490E038FFD1FAF9D7ECFA21FB9C" authorityName="Bocchino" authorityYear="1964" box="[1290,1436,1015,1041]" class="Actinopterygii" genus="Propygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="36" pageNumber="70" phylum="Chordata" rank="genus">Propygidium</taxonomicName>
with an indistinct stoma at posterior corner.
<taxonomicName id="4C767490E038FFD1FD74D000FD49FBB8" box="[647,756,1051,1077]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="36" pageNumber="70" phylum="Chordata" rank="genus">Pygidium</taxonomicName>
slightly pulvinate longitudinally and laterally; upper margin slightly convex; lateral margin subparallel in upper quarter, linearly attenuate apically; tip rounded.
</paragraph>
<paragraph id="8BC90F13E038FFD1FF34D079FBC6FA80" blockId="36.[151,1437,151,1655]" pageId="36" pageNumber="37">
<emphasis id="B902D301E038FFD1FF34D079FE37FBF0" bold="true" box="[199,394,1122,1149]" pageId="36" pageNumber="37">Prothoracic leg:</emphasis>
Femur and tibia slender. Tibia smooth ventrally. Median tooth elongate. Basal tooth very small. Interspace between teeth almost equal. Shape of interspace between teeth concave. Tarsomere IIV combined extremely short; lengthening from tarsomere IIV;
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/
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in
</quantity>
length of tibia. Pretarsomere slightly thickened; with two small, triangular-shaped, median teeth consecutive ventrally; slightly more than
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/
<quantity id="4C8EA2F6E038FFD1FBD2D0FAFBF8FB64" box="[1057,1093,1231,1263]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="36" pageNumber="37" unit="in" value="5.0">
<subScript id="17F20D56E038FFD1FBD2D0FAFB94FB62" attach="left" box="[1057,1065,1249,1263]" fontSize="6" pageId="36" pageNumber="37">5</subScript>
in
</quantity>
length of tarsomere IIV combined. Internal claw as long as pretarsomere. External claw
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/
<quantity id="4C8EA2F6E038FFD1FCB6D11EFCCFFA80" box="[837,882,1267,1299]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="36" pageNumber="37" unit="in" value="10.0">
<subScript id="17F20D56E038FFD1FCB6D11EFCE8FA9E" attach="left" box="[837,853,1285,1299]" fontSize="6" pageId="36" pageNumber="37">10</subScript>
in
</quantity>
length of internal claw.
</paragraph>
<paragraph id="8BC90F13E038FFD1FF34D10DFD08FA10" blockId="36.[151,1437,151,1655]" pageId="36" pageNumber="37">
<emphasis id="B902D301E038FFD1FF34D10DFE26FABC" bold="true" box="[199,411,1302,1329]" pageId="36" pageNumber="37">Mesothoracic leg:</emphasis>
Tibia
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/
<quantity id="4C8EA2F6E038FFD1FE07D132FD9FFABC" box="[500,546,1303,1335]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="36" pageNumber="37" unit="in" value="10.0">
<subScript id="17F20D56E038FFD1FE07D132FDB9FABA" attach="left" box="[500,516,1321,1335]" fontSize="6" pageId="36" pageNumber="37">10</subScript>
in
</quantity>
length of femur. Tarsomere IIV combined very short; slightly more than
<superScript id="7C03A25BE038FFD1FA94D10DFAD2FAA9" attach="right" box="[1383,1391,1302,1316]" fontSize="6" pageId="36" pageNumber="37">2</superScript>
/
<quantity id="4C8EA2F6E038FFD1FA84D132FA21FABC" box="[1399,1436,1303,1335]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="36" pageNumber="37" unit="in" value="5.0">
<subScript id="17F20D56E038FFD1FA84D132FAC2FABA" attach="left" box="[1399,1407,1321,1335]" fontSize="6" pageId="36" pageNumber="37">5</subScript>
in
</quantity>
length of tibia. Pretarsomere with a shortened granulate ridge ventrally, followed by a roundish cutting apically;
<superScript id="7C03A25BE038FFD1FA8FD121FA39FAC5" attach="right" box="[1404,1412,1338,1352]" fontSize="6" pageId="36" pageNumber="37">9</superScript>
/
<quantity id="4C8EA2F6E038FFD1FA7FD156FF10FAF4" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="36" pageNumber="37" unit="in" value="10.0">
<subScript id="17F20D56E038FFD1FA7FD156FA21FAD6" attach="none" box="[1420,1436,1357,1371]" fontSize="6" pageId="36" pageNumber="37">10</subScript>
in
</quantity>
length of tarsomere IIV combined. External claw slightly less than
<superScript id="7C03A25BE038FFD1FC44D145FC02FAE1" attach="right" box="[951,959,1374,1388]" fontSize="6" pageId="36" pageNumber="37">9</superScript>
/
<quantity id="4C8EA2F6E038FFD1FC34D16AFC48FAF4" box="[967,1013,1375,1407]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="36" pageNumber="37" unit="in" value="10.0">
<subScript id="17F20D56E038FFD1FC34D16AFC6AFAF2" attach="left" box="[967,983,1393,1407]" fontSize="6" pageId="36" pageNumber="37">10</subScript>
in
</quantity>
length of pretarsomere. Internal claw slightly more than
<superScript id="7C03A25BE038FFD1FE99D199FECFFA1D" attach="right" box="[362,370,1410,1424]" fontSize="6" pageId="36" pageNumber="37">9</superScript>
/
<quantity id="4C8EA2F6E038FFD1FE89D18EFE1AFA10" box="[378,423,1411,1443]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="36" pageNumber="37" unit="in" value="10.0">
<subScript id="17F20D56E038FFD1FE89D18EFE37FA2E" attach="left" box="[378,394,1429,1443]" fontSize="6" pageId="36" pageNumber="37">10</subScript>
in
</quantity>
length of external claw.
</paragraph>
<paragraph id="8BC90F13E038FFD1FF34D1BDFD54F9A0" blockId="36.[151,1437,151,1655]" pageId="36" pageNumber="37">
<emphasis id="B902D301E038FFD1FF34D1BDFE27FA4C" bold="true" box="[199,410,1446,1473]" pageId="36" pageNumber="37">Metathoracic leg:</emphasis>
Tibia slightly more than
<superScript id="7C03A25BE038FFD1FD49D1BDFD7FFA39" attach="right" box="[698,706,1446,1460]" fontSize="6" pageId="36" pageNumber="37">9</superScript>
/
<quantity id="4C8EA2F6E038FFD1FD3AD1A2FD45FA4C" box="[713,760,1447,1479]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="36" pageNumber="37" unit="in" value="10.0">
<subScript id="17F20D56E038FFD1FD3AD1A2FD64FA4A" attach="left" box="[713,729,1465,1479]" fontSize="6" pageId="36" pageNumber="37">10</subScript>
in
</quantity>
length of femur. Tarsomere IIV combined distinctly short; slightly more than
<superScript id="7C03A25BE038FFD1FE9AD1D0FECCFA54" attach="right" box="[361,369,1483,1497]" fontSize="6" pageId="36" pageNumber="37">7</superScript>
/
<quantity id="4C8EA2F6E038FFD1FE8AD1C6FE1BFA68" box="[377,422,1483,1515]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="36" pageNumber="37" unit="in" value="10.0">
<subScript id="17F20D56E038FFD1FE8AD1C6FE34FA66" attach="left" box="[377,393,1501,1515]" fontSize="6" pageId="36" pageNumber="37">10</subScript>
in
</quantity>
length of tibia. Pretarsomere slender; with an undulate ridge ventrally; slightly less than
<superScript id="7C03A25BE038FFD1FA77D1D1FA31FA55" attach="right" box="[1412,1420,1482,1496]" fontSize="6" pageId="36" pageNumber="37">1</superScript>
/
<quantity id="4C8EA2F6E038FFD1FA67D1C6FF10F984" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="36" pageNumber="37" unit="in" value="2.0">
<subScript id="17F20D56E038FFD1FA67D1C6FA21FA66" attach="none" box="[1428,1436,1501,1515]" fontSize="6" pageId="36" pageNumber="37">2</subScript>
in
</quantity>
length of tarsomere IIV combined. External claw slightly more than
<superScript id="7C03A25BE038FFD1FC4AD1F5FC7CFA71" attach="right" box="[953,961,1518,1532]" fontSize="6" pageId="36" pageNumber="37">1</superScript>
/
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more in length of pretarsomere. Internal claw slightly more than
<superScript id="7C03A25BE038FFD1FE55D209FE13F9AD" attach="right" box="[422,430,1554,1568]" fontSize="6" pageId="36" pageNumber="37">4</superScript>
/
<quantity id="4C8EA2F6E038FFD1FE45D23EFE66F9A0" box="[438,475,1555,1587]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="36" pageNumber="37" unit="in" value="5.0">
<subScript id="17F20D56E038FFD1FE45D23EFE03F9BE" attach="left" box="[438,446,1573,1587]" fontSize="6" pageId="36" pageNumber="37">5</subScript>
in
</quantity>
length of external claw.
</paragraph>
<paragraph id="8BC90F13E038FFD1FF34D22CFD93F9DF" blockId="36.[151,1437,151,1655]" box="[199,558,1590,1618]" pageId="36" pageNumber="37">
<emphasis id="B902D301E038FFD1FF34D22CFEECF9DC" bold="true" box="[199,337,1591,1617]" pageId="36" pageNumber="37">Parameres:</emphasis>
(
<figureCitation id="134D1396E038FFD1FE93D22CFE55F9DC" box="[352,488,1590,1617]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="36" pageNumber="37">Figs. 7475</figureCitation>
,
<figureCitation id="134D1396E038FFD1FE07D22CFD9FF9DF" box="[500,546,1591,1618]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="36" pageNumber="37">106</figureCitation>
).
</paragraph>
<paragraph id="8BC90F13E038FFD1FF34D247FEDEF9F8" blockId="36.[151,1437,151,1655]" box="[199,355,1627,1655]" pageId="36" pageNumber="37">
<emphasis id="B902D301E038FFD1FF34D247FF58F9FA" bold="true" box="[199,229,1628,1655]" pageId="36" pageNumber="37">
<specimenCount id="9D70C49AE038FFD1FF34D247FF63F9FA" box="[199,222,1628,1655]" pageId="36" pageNumber="37" type="female"></specimenCount>
:
</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E038FFD1FF64D2B9FDC8F888" pageId="36" pageNumber="37" type="distribution">
<paragraph id="8BC90F13E038FFD1FF64D2B9FBEDF930" blockId="36.[151,1104,1698,1725]" box="[151,1104,1698,1725]" pageId="36" pageNumber="37">
<emphasis id="B902D301E038FFD1FF64D2B9FE8CF931" bold="true" box="[151,305,1698,1724]" pageId="36" pageNumber="37">Distribution.</emphasis>
<collectingRegion id="49B2C1F1E038FFD1FEC4D2B8FDF5F930" box="[311,584,1699,1725]" country="South Africa" name="Northern Cape" pageId="36" pageNumber="37">Northern Cape Province</collectingRegion>
(
<figureCitation id="134D1396E038FFD1FDA4D2B8FD05F930" box="[599,696,1699,1725]" captionStart="FIGURE 140" captionStartId="72.[151,250,1944,1968]" captionTargetBox="[195,1393,188,1910]" captionTargetId="figure-25@72.[185,1402,181,1920]" captionTargetPageId="72" captionText="FIGURE 140. Distributions of 18 Beckhoplia Dombrow, 2005 species, showing the concentration of species around the Northern Cape town of Nieuwoudtville." figureDoi="http://doi.org/10.5281/zenodo.4401964" httpUri="https://zenodo.org/record/4401964/files/figure.png" pageId="36" pageNumber="37">Fig. 140</figureCitation>
), only known from the
<typeStatus id="54CDB1B1E038FFD1FC4CD2B8FC52F930" box="[959,1007,1699,1725]" pageId="36" pageNumber="37">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BC90F13E038FFD1FF64D2F1FDC8F888" blockId="36.[151,629,1770,1797]" box="[151,629,1770,1797]" pageId="36" pageNumber="37">
<emphasis id="B902D301E038FFD1FF64D2F1FEE3F888" bold="true" box="[151,350,1770,1797]" pageId="36" pageNumber="37">
Vegetation
<typeStatus id="54CDB1B1E038FFD1FEEFD2F0FEE4F888" box="[284,345,1771,1797]" pageId="36" pageNumber="37">Type</typeStatus>
.
</emphasis>
Hantam Karroo (SKt 2).
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E038FFD1FF64D328FD63F8C0" box="[151,734,1843,1869]" pageId="36" pageNumber="37" type="biology_ecology">
<paragraph id="8BC90F13E038FFD1FF64D328FD63F8C0" blockId="36.[151,734,1843,1869]" box="[151,734,1843,1869]" pageId="36" pageNumber="37">
<emphasis id="B902D301E038FFD1FF64D328FE95F8C0" bold="true" box="[151,296,1843,1869]" pageId="36" pageNumber="37">Host Plants.</emphasis>
<taxonomicName id="4C767490E038FFD1FEDCD328FE27F8C0" box="[303,410,1843,1869]" class="Liliopsida" family="Iridaceae" kingdom="Plantae" order="Asparagales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="family">Iridaceae</taxonomicName>
:
<taxonomicName id="4C767490E038FFD1FE57D328FD63F8C0" authority="L. Bolus." authorityName="L. Bolus." box="[420,734,1843,1869]" class="Liliopsida" family="Iridaceae" genus="Babiana" kingdom="Plantae" order="Asparagales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="vanziliae">
<emphasis id="B902D301E038FFD1FE57D328FDD2F8C0" box="[420,623,1843,1869]" italics="true" pageId="36" pageNumber="37">Babiana vanziliae</emphasis>
L. Bolus.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E038FFD1FF64D360FC32F818" box="[151,911,1914,1941]" pageId="36" pageNumber="37" type="etymology">
<paragraph id="8BC90F13E038FFD1FF64D360FC32F818" blockId="36.[151,911,1914,1941]" box="[151,911,1914,1941]" pageId="36" pageNumber="37">
<emphasis id="B902D301E038FFD1FF64D360FEA3F818" bold="true" box="[151,286,1915,1941]" pageId="36" pageNumber="37">Etymology.</emphasis>
The name derives from the [Latin: pulchr- = beautiful].
</paragraph>
</subSubSection>
</treatment>
</document>