698 lines
89 KiB
XML
698 lines
89 KiB
XML
<document id="66221CC141346CCE9DD4A7EF35633BD1" ID-CLB-Dataset="8721" ID-DOI="10.11646/zootaxa.4823.1.1" ID-GBIF-Dataset="a43d0118-9df5-4970-a316-0dae5c9caebf" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401916" ID-ZooBank="3DF06F26-FD35-452E-9056-1D05B49946EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609340622775" checkinUser="plazi" docAuthor="Dombrow, Holger E. & Colville, Jonathan F." docDate="2020" docId="03DFBE05E03CFFD6FF64D551FCFAFF74" docLanguage="en" docName="zootaxa.4823.1.1.pdf" docOrigin="Zootaxa 4823" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Beckhoplia colvillei Dombrow 2005" docType="treatment" docVersion="9" lastPageNumber="36" masterDocId="FFE6C67DE01CFFF5FFF3D41BFFBDFF8D" masterDocTitle="Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini)" masterLastPageNumber="79" masterPageNumber="1" pageNumber="33" updateTime="1698849823920" updateUser="ExternalLinkService">
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<mods:title id="5E4C50A936A7DA101E77BBFAFEDCE910">Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini)</mods:title>
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<mods:name id="5919CB4FBEEF9F9559E6006E3E17B4FC" type="personal">
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<mods:namePart id="5B5C6556057AFDD4C87D0502EDBB5E0E">Dombrow, Holger E.</mods:namePart>
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<mods:namePart id="1AD0BC9B50E2B37BE22CDEFFBA94AAED">Colville, Jonathan F.</mods:namePart>
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<mods:nameIdentifier id="B5B718A634FA42F81D27B7E2C6ECC706" type="email">jonathan.colville@gmail.com</mods:nameIdentifier>
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<mods:title id="0F1A73C6F2861D9E94CD077A6D8F795E">Zootaxa</mods:title>
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<mods:date id="C78A6A628738161450703612351D532F">2020</mods:date>
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<mods:number id="96D6BFCCF35A9D6B35B26531D8A1DF3B">2020-08-10</mods:number>
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<mods:identifier id="5739152C0A2BA0017585921BF6A626B5" type="DOI">10.11646/zootaxa.4823.1.1</mods:identifier>
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<treatment id="03DFBE05E03CFFD6FF64D551FCFAFF74" ID-DOI="http://doi.org/10.5281/zenodo.4455285" ID-GBIF-Taxon="177013519" ID-Zenodo-Dep="4455285" LSID="urn:lsid:plazi:treatment:03DFBE05E03CFFD6FF64D551FCFAFF74" httpUri="http://treatment.plazi.org/id/03DFBE05E03CFFD6FF64D551FCFAFF74" lastPageId="35" lastPageNumber="36" pageId="32" pageNumber="33">
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<subSubSection id="C36C5C98E03CFFD5FF64D551FDEFFEE9" box="[151,594,330,357]" pageId="32" pageNumber="33" type="nomenclature">
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<paragraph id="8BC90F13E03CFFD5FF64D551FDEFFEE9" blockId="32.[151,861,330,428]" box="[151,594,330,357]" pageId="32" pageNumber="33">
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<heading id="D081B87FE03CFFD5FF64D551FDEFFEE9" bold="true" box="[151,594,330,357]" fontSize="11" level="2" pageId="32" pageNumber="33" reason="4">
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<taxonomicName id="4C767490E03CFFD5FF64D551FDEFFEE9" authority="Dombrow, 2005" authorityName="Dombrow" authorityYear="2005" box="[151,594,330,357]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="colvillei">
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<emphasis id="B902D301E03CFFD5FF64D551FDEFFEE9" bold="true" box="[151,594,330,357]" pageId="32" pageNumber="33">
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<emphasis id="B902D301E03CFFD5FF64D551FE3BFEE8" bold="true" box="[151,390,330,357]" italics="true" pageId="32" pageNumber="33">Beckhoplia colvillei</emphasis>
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<bibRefCitation id="EFE772E2E03CFFD5FE7ED551FDEFFEE9" author="Dombrow, H." box="[397,594,330,356]" pageId="32" pageNumber="33" pagination="125 - 130" refId="ref61191" refString="Dombrow, H. (2005) Beckhoplia a new genus of Hopliini from the Northern Cape of the Republic of South Africa. (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Coleoptera, 9, 125 - 130." type="journal article" year="2005">Dombrow, 2005</bibRefCitation>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C36C5C98E03CFFD5FF64D575FCE0FE21" pageId="32" pageNumber="33" type="description">
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<paragraph id="8BC90F13E03CFFD5FF64D575FCE0FE21" blockId="32.[151,861,330,428]" pageId="32" pageNumber="33">
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(
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<figureCitation id="134D1396E03CFFD5FF6CD575FF50FE05" box="[159,237,366,392]" captionStart="FIGURES 1–2" captionStartId="10.[151,264,456,480]" captionTargetBox="[156,1429,183,426]" captionTargetId="figure-25@10.[151,1436,179,431]" captionTargetPageId="10" captionText="FIGURES 1–2. General shape of the clypeus of Beckhoplia Dombrow, 2005 and Anisochelus Burmeister, 1844 species, ♂♂. 1, B. colvillei Dombrow, 2005; 2, A. inornatus Burmeister, 1844. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4401918" httpUri="https://zenodo.org/record/4401918/files/figure.png" pageId="32" pageNumber="33">Figs. 1</figureCitation>
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,
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<figureCitation id="134D1396E03CFFD5FF08D575FE9AFE04" box="[251,295,366,393]" captionStart="FIGURES 3–5" captionStartId="10.[151,264,808,832]" captionTargetBox="[156,1428,582,779]" captionTargetId="figure-59@10.[151,1436,577,785]" captionTargetPageId="10" captionText="FIGURES 3–5. General drawings of the type of incision of the anterior margin of Beckhoplia colvillei Dombrow, 2005 in anterior view, ♂♂. 3, concave incised medially; 4, curly bracket-shaped incised medially; 5, broad v-shaped incised medially. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4401920" httpUri="https://zenodo.org/record/4401920/files/figure.png" pageId="32" pageNumber="33">3–5</figureCitation>
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,
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<figureCitation id="134D1396E03CFFD5FEC0D575FEFFFE05" box="[307,322,366,392]" captionStart="FIGURES 8–10" captionStartId="10.[151,264,1454,1478]" captionTargetBox="[155,1430,1277,1425]" captionTargetId="figure-142@10.[151,1436,1273,1430]" captionTargetPageId="10" captionText="FIGURES 8–10. General drawings of the position of the eye of Beckhoplia Dombrow, 2005 species, in lateral view. 8, eye below head level, ♀♀; 9, eye on head level, ♀♀; 10, eye above head level, ♂♂. Scale bar 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.4401924" httpUri="https://zenodo.org/record/4401924/files/figure.png" pageId="32" pageNumber="33">8</figureCitation>
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,
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<figureCitation id="134D1396E03CFFD5FEBCD575FED0FE04" box="[335,365,366,393]" captionStart="FIGURES 15–18" captionStartId="16.[151,264,509,533]" captionTargetBox="[199,1387,184,446]" captionTargetId="figure-25@16.[195,1392,181,449]" captionTargetPageId="16" captionText="FIGURES 15–18. General shape of the antenna and clavus of Beckhoplia Dombrow, 2005 species, ♂♂, in lateral view, other- wise indicated. 15, nine segmented antenna: a, scapus; b, pedicellus; c, funicle I–IV = funiculum; d, three segmented lamellar clavus. Scale bar 1 mm. 16, clavus short elliptic, ♀♀; 17, clavus elliptic; 18, clavus long elliptic. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4401928" httpUri="https://zenodo.org/record/4401928/files/figure.png" pageId="32" pageNumber="33">15</figureCitation>
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,
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<figureCitation id="134D1396E03CFFD5FE8AD575FE7CFE05" box="[377,449,366,392]" captionStart="FIGURES 19–22" captionStartId="16.[151,264,1056,1080]" captionTargetBox="[199,1385,669,992]" captionTargetId="figure-88@16.[195,1392,665,996]" captionTargetPageId="16" captionText="FIGURES 19–22. General drawings of the mouth parts of Beckhoplia Dombrow, 2005 species, in ventral view, ♂♂. 19, labrum with epipharynx; 20, left mandible; 21, left maxillae with palpus maxillaris; 22, labium with palpus labialis. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4401930" httpUri="https://zenodo.org/record/4401930/files/figure.png" pageId="32" pageNumber="33">19–22</figureCitation>
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,
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<figureCitation id="134D1396E03CFFD5FE3ED575FDA8FE04" box="[461,533,366,393]" captionStart="FIGURES 34–35" captionStartId="17.[151,264,1858,1882]" captionTargetBox="[206,1381,1509,1794]" captionTargetId="figure-132@17.[202,1386,1504,1799]" captionTargetPageId="17" captionText="FIGURES 34–35. General drawings of the pygidium and abdomen of Beckhoplia Dombrow, 2005 species, in posterior view. 34, abdomen not compressed laterally, ♂♂; 35, abdomen distinctly pulvinate, ♀♀. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401940" httpUri="https://zenodo.org/record/4401940/files/figure.png" pageId="32" pageNumber="33">34–35</figureCitation>
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,
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<figureCitation id="134D1396E03CFFD5FDD2D575FD2EFE04" box="[545,659,366,393]" captionStart="FIGURES 41–46" captionStartId="23.[151,264,1293,1317]" captionTargetBox="[208,1379,833,1240]" captionTargetId="figure-110@23.[203,1384,830,1246]" captionTargetPageId="23" captionText="FIGURES 41–46. General drawing of the prothoracic tibia of Beckhoplia Dombrow, 2005 species, in dorsal view, otherwise indicated, ♂♂. 41, slender, basal tooth very small, interspace between teeth almost equal, shape of interspace concave, with a dense longitudinal row of deep, elongate roundish punctures; 42, basal tooth rudimentary, interspace between teeth almost equal, interspace between anterior and median tooth concave and between median and basal tooth linear; 43, interspace between anterior and median tooth u-shaped and between median and basal tooth concave; 44, with a medio-ventral ridge; 45, with a medio-ventral ridge, cross-section: a, internal ridge; b, external ridge; c, medio-ventral ridge; 46, smooth ventrally, cross-section: a, internal ridge; b, external ridge. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401944" httpUri="https://zenodo.org/record/4401944/files/figure.png" pageId="32" pageNumber="33">41, 44–45</figureCitation>
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,
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<figureCitation id="134D1396E03CFFD5FD6CD575FD00FE04" box="[671,701,366,393]" captionStart="FIGURES 50–54" captionStartId="24.[151,264,678,702]" captionTargetBox="[212,1375,186,616]" captionTargetId="figure-25@24.[208,1380,181,622]" captionTargetPageId="24" captionText="FIGURES 50–54. General drawing of the types of prothoracic pretarsomere of Beckhoplia Dombrow, 2005 species, in lateral view, with roundish cutting ventrally, ♂♂. 50, distinctly thickened, smooth ventrally; 51, slightly thickened, with a shortened smooth ridge ventrally; 52, distinctly thickened, with a median tooth ventrally; 53, slightly thickened, with two median teeth consecutive ventrally; 54, distinctly thickened, with a shortened granulate ridge ventrally. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4401948" httpUri="https://zenodo.org/record/4401948/files/figure.png" pageId="32" pageNumber="33">51</figureCitation>
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,
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<figureCitation id="134D1396E03CFFD5FD3AD575FCACFE05" box="[713,785,366,392]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="32" pageNumber="33">72–73</figureCitation>
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,
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<figureCitation id="134D1396E03CFFD5FCEED575FCF5FE04" box="[797,840,366,393]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="32" pageNumber="33">105</figureCitation>
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) http://zoobank.org/
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<subSubSection id="C36C5C98E03CFFD5FF64D5C1FDFBFE7F" box="[151,582,474,498]" pageId="32" pageNumber="33" type="reference_group">
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<paragraph id="8BC90F13E03CFFD5FF64D5C1FDFBFE7F" blockId="32.[151,582,474,498]" box="[151,582,474,498]" pageId="32" pageNumber="33">
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<treatmentCitation id="0AD72902E03CFFD5FF64D5C1FDFFFE7F" author="Dombrow, H." box="[151,578,474,498]" page="128" pageId="32" pageNumber="33" year="2005">
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<taxonomicName id="4C767490E03CFFD5FF64D5C1FDFFFE7F" authority="Dombrow, 2005: 128" authorityName="Dombrow" authorityPageNumber="128" authorityYear="2005" box="[151,578,474,498]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="colvillei">
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<emphasis id="B902D301E03CFFD5FF64D5C1FEDCFE7F" box="[151,353,474,498]" italics="true" pageId="32" pageNumber="33">Beckhoplia colvillei</emphasis>
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<bibRefCitation id="EFE772E2E03CFFD5FE94D5C1FDFFFE7F" author="Dombrow, H." box="[359,578,474,498]" pageId="32" pageNumber="33" pagination="125 - 130" refId="ref61191" refString="Dombrow, H. (2005) Beckhoplia a new genus of Hopliini from the Northern Cape of the Republic of South Africa. (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Coleoptera, 9, 125 - 130." type="journal article" year="2005">Dombrow, 2005: 128</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C5C98E03CFFD5FF64D63BFBD8FB82" pageId="32" pageNumber="33" type="materials_examined">
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<paragraph id="8BC90F13E03CFFD5FF64D63BFBE5FDB6" blockId="32.[151,1112,544,571]" box="[151,1112,544,571]" pageId="32" pageNumber="33">
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<materialsCitation id="3B1E054EE03CFFD5FF64D63BFBEEFDB6" ID-GBIF-Occurrence="3022772323" box="[151,1107,544,571]" collectionCode="NC" country="South Africa" location="RSA" pageId="32" pageNumber="33" specimenCount="1" typeStatus="holotype">
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<emphasis id="B902D301E03CFFD5FF64D63BFE8BFDB7" bold="true" box="[151,310,544,570]" pageId="32" pageNumber="33">
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<typeStatus id="54CDB1B1E03CFFD5FF64D63BFF6FFDB7" box="[151,210,544,570]" pageId="32" pageNumber="33">Type</typeStatus>
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locality.
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</emphasis>
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[
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<collectingCountry id="F3614F83E03CFFD5FEB0D63BFEC2FDB7" box="[323,383,544,570]" name="South Africa" pageId="32" pageNumber="33">RSA</collectingCountry>
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,
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<collectionCode id="ED6797D6E03CFFD5FE79D63BFE0AFDB6" box="[394,439,544,571]" pageId="32" pageNumber="33">NC</collectionCode>
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]: Farm Glenlyon [now Hantam National Botanical Garden]
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BC90F13E03CFFD5FF64D673FE33FC2F" blockId="32.[151,1437,616,930]" pageId="32" pageNumber="33">
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<emphasis id="B902D301E03CFFD5FF64D673FEFAFD0F" bold="true" box="[151,327,616,642]" pageId="32" pageNumber="33">Type material.</emphasis>
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[
|
||
<materialsCitation id="3B1E054EE03CFFD5FEA4D673FDB4FC9F" ID-GBIF-Occurrence="3022772316" collectingDate="1998-08-22" collectionCode="NCP, SAMC" collectorName="J. Colville & Dombrow" country="South Africa" determinerName="Dombrow" latitude="-31.24" location="Farm Glenlyon" longLatPrecision="731" longitude="19.08" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
45]
|
||
<emphasis id="B902D301E03CFFD5FE74D673FE49FD0F" bold="true" box="[391,500,616,642]" pageId="32" pageNumber="33">
|
||
<typeStatus id="54CDB1B1E03CFFD5FE74D673FE49FD0F" box="[391,500,616,642]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<specimenCount id="9D70C49AE03CFFD5FE0ED672FDA8FD0E" box="[509,533,617,643]" pageId="32" pageNumber="33" type="male">♂</specimenCount>
|
||
: white label: pr «
|
||
<collectingCountry id="F3614F83E03CFFD5FD2ED673FCA5FD0F" box="[733,792,616,642]" name="South Africa" pageId="32" pageNumber="33">RSA</collectingCountry>
|
||
,
|
||
<collectionCode id="ED6797D6E03CFFD5FCD7D673FCE7FD0F" box="[804,858,616,642]" country="United Kingdom" name="National Collection of Passiflora" pageId="32" pageNumber="33" type="Herbarium">NCP</collectionCode>
|
||
, Nieuwoudtv., |
|
||
<location id="8EA959C8E03CFFD5FBE6D672FB79FD0F" LSID="urn:lsid:plazi:treatment:03DFBE05E03CFFD6FF64D551FCFAFF74:8EA959C8E03CFFD5FBE6D672FB79FD0F" box="[1045,1220,616,643]" country="South Africa" latitude="-31.24" longLatPrecision="731" longitude="19.08" name="Farm Glenlyon" pageId="32" pageNumber="33">Farm Glenlyon</location>
|
||
,
|
||
<geoCoordinate id="EE4269D4E03CFFD5FB22D673FA93FD0F" box="[1233,1326,616,642]" degrees="31.24" direction="south" orientation="latitude" pageId="32" pageNumber="33" precision="555" value="-31.24">31.24S–</geoCoordinate>
|
||
|
|
||
<geoCoordinate id="EE4269D4E03CFFD5FAB6D673FA25FD0F" box="[1349,1432,616,642]" degrees="19.08" direction="east" orientation="longitude" pageId="32" pageNumber="33" precision="555" value="19.08">19.08E</geoCoordinate>
|
||
,
|
||
<date id="FFC829D3E03CFFD5FF64D697FEA7FD2B" box="[151,282,652,678]" pageId="32" pageNumber="33" value="1998-08-22">
|
||
<collectingDate id="EF8CD03BE03CFFD5FF64D697FEA7FD2B" box="[151,282,652,678]" pageId="32" pageNumber="33" value="1998-08-22">22/08/1998</collectingDate>
|
||
</date>
|
||
, | leg.:
|
||
<collectorName id="26836AC5E03CFFD5FE82D696FE54FD2B" box="[369,489,652,679]" pageId="32" pageNumber="33">J. Colville</collectorName>
|
||
» || yellow label hw «10168» | pr «coll.
|
||
<collectorName id="26836AC5E03CFFD5FC49D697FB93FD2B" box="[954,1070,652,678]" pageId="32" pageNumber="33">Dombrow</collectorName>
|
||
» || green label pr «Host plant: |
|
||
<taxonomicName id="4C767490E03CFFD5FF64D6AAFEFCFD47" authorityName="Roessler" authorityYear="1959" baseAuthorityName="Burm. fil." box="[151,321,688,715]" class="Magnoliopsida" family="Asteraceae" genus="Gazania" kingdom="Plantae" order="Asterales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B902D301E03CFFD5FF64D6AAFEFCFD47" box="[151,321,688,715]" italics="true" pageId="32" pageNumber="33">Gazania rigida</emphasis>
|
||
</taxonomicName>
|
||
» || blue label pr «Bioregion: | Shale Renosterveld | Vegetation Type: FRs2 – | Nieuwoudtville Shale Renosterveld» || yellow label pr «SAM–COL | A071226» || red type label pr «
|
||
<typeStatus id="54CDB1B1E03CFFD5FC0BD6CFFBDFFD63" box="[1016,1122,724,750]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
|
||
|
|
||
<taxonomicName id="4C767490E03CFFD5FB86D6CFFB4EFD63" authorityName="Dombrow" authorityYear="2005" box="[1141,1267,724,750]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902D301E03CFFD5FB86D6CFFB4EFD63" box="[1141,1267,724,750]" italics="true" pageId="32" pageNumber="33">Beckhoplia</emphasis>
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C767490E03CFFD5FAF5D6CFFADDFD63" authority="Dombrow & Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[1286,1376,724,750]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="colvillei" status="sp. nov.">
|
||
<emphasis id="B902D301E03CFFD5FAF5D6CFFADDFD63" box="[1286,1376,724,750]" italics="true" pageId="32" pageNumber="33">colvillei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2316E7AE03CFFD5FA94D6CEFA21FD62" box="[1383,1436,725,751]" pageId="32" pageNumber="33" rank="species">sp.n.</taxonomicNameLabel>
|
||
| det.:
|
||
<determinerName id="7E15E3D7E03CFFD5FF2ED6E3FEF2FC9F" box="[221,335,760,786]" pageId="32" pageNumber="33">Dombrow</determinerName>
|
||
2003» (
|
||
<collectionCode id="ED6797D6E03CFFD5FE5FD6E3FDBFFC9F" box="[428,514,760,786]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="32" pageNumber="33" type="Museum">SAMC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
. –
|
||
<materialsCitation id="3B1E054EE03CFFD5FDD9D6E3FD11FCD7" ID-GBIF-Occurrence="3022772325" collectionCode="SAMC" collectorName="Dombrow" country="South Africa" determinerName="Dombrow" pageId="32" pageNumber="33" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
|
||
<emphasis id="B902D301E03CFFD5FDD9D6E3FD1FFC9F" bold="true" box="[554,674,760,786]" pageId="32" pageNumber="33">
|
||
<typeStatus id="54CDB1B1E03CFFD5FDD9D6E3FD1FFC9F" box="[554,674,760,786]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<specimenCount id="9D70C49AE03CFFD5FD5AD6E3FD6BFC9E" box="[681,726,760,787]" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D70C49AE03CFFD5FD10D6E3FC99FC9E" box="[739,804,760,787]" pageId="32" pageNumber="33" type="female">2 ♀♀</specimenCount>
|
||
: same labels as
|
||
<typeStatus id="54CDB1B1E03CFFD5FC28D6E3FB80FC9F" box="[987,1085,760,786]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
|
||
but || yellow label hw «18», pr «.811,.817,.848 | coll.
|
||
<collectorName id="26836AC5E03CFFD5FE6DD707FDAFFCBB" box="[414,530,796,822]" pageId="32" pageNumber="33">Dombrow</collectorName>
|
||
» || yellow label pr «SAM–COL | A071226» || red label pr «
|
||
<typeStatus id="54CDB1B1E03CFFD5FB54D706FAB6FCBA" box="[1191,1291,797,823]" pageId="32" pageNumber="33" type="paratype">Paratype</typeStatus>
|
||
|
|
||
<taxonomicName id="4C767490E03CFFD5FAEDD707FA21FCBB" authorityName="Dombrow" authorityYear="2005" box="[1310,1436,796,822]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902D301E03CFFD5FAEDD707FA21FCBB" box="[1310,1436,796,822]" italics="true" pageId="32" pageNumber="33">Beckhoplia</emphasis>
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C767490E03CFFD5FF50D75BFF40FCD7" authority="Dombrow & Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[163,253,832,858]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="colvillei" status="sp. nov.">
|
||
<emphasis id="B902D301E03CFFD5FF50D75BFF40FCD7" box="[163,253,832,858]" italics="true" pageId="32" pageNumber="33">colvillei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2316E7AE03CFFD5FEF7D75AFE84FCD6" box="[260,313,833,859]" pageId="32" pageNumber="33" rank="species">sp.n.</taxonomicNameLabel>
|
||
| det.:
|
||
<determinerName id="7E15E3D7E03CFFD5FE71D75BFE49FCD7" box="[386,500,832,858]" pageId="32" pageNumber="33">Dombrow</determinerName>
|
||
2003» (
|
||
<collectionCode id="ED6797D6E03CFFD5FDBDD75BFD1EFCD7" box="[590,675,832,858]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="32" pageNumber="33" type="Museum">SAMC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B1E054EE03CFFD5FD45D75BFE37FC2F" ID-GBIF-Occurrence="3022772317" collectionCode="ESDW" collectorName="Dombrow" country="South Africa" determinerName="Dombrow" pageId="32" pageNumber="33" specimenCount="41" specimenCount-female="5" specimenCount-male="36" typeStatus="holotype">
|
||
<specimenCount id="9D70C49AE03CFFD5FD45D75BFCB8FCD6" box="[694,773,832,859]" pageId="32" pageNumber="33" type="male">36 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D70C49AE03CFFD5FCE3D75BFCEDFCD6" box="[784,848,832,859]" pageId="32" pageNumber="33" type="female">5 ♀♀</specimenCount>
|
||
: same labels as
|
||
<typeStatus id="54CDB1B1E03CFFD5FBF1D75BFBD9FCD7" box="[1026,1124,832,858]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
|
||
but || yellow label hw «18», pr «.774,.776–.810,.812–.816 | coll.
|
||
<collectorName id="26836AC5E03CFFD5FDCCD77FFD0EFCF3" box="[575,691,868,894]" pageId="32" pageNumber="33">Dombrow</collectorName>
|
||
» || red label pr «
|
||
<typeStatus id="54CDB1B1E03CFFD5FC9FD77EFC6DFCF2" box="[876,976,869,895]" pageId="32" pageNumber="33" type="paratype">Paratype</typeStatus>
|
||
|
|
||
<taxonomicName id="4C767490E03CFFD5FC16D77FFBDEFCF3" authorityName="Dombrow" authorityYear="2005" box="[997,1123,868,894]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902D301E03CFFD5FC16D77FFBDEFCF3" box="[997,1123,868,894]" italics="true" pageId="32" pageNumber="33">Beckhoplia</emphasis>
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C767490E03CFFD5FB84D77FFB6CFCF3" authority="Dombrow & Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[1143,1233,868,894]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="colvillei" status="sp. nov.">
|
||
<emphasis id="B902D301E03CFFD5FB84D77FFB6CFCF3" box="[1143,1233,868,894]" italics="true" pageId="32" pageNumber="33">colvillei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2316E7AE03CFFD5FB2AD77EFAB3FCF2" box="[1241,1294,869,895]" pageId="32" pageNumber="33" rank="species">sp.n.</taxonomicNameLabel>
|
||
| det.:
|
||
<determinerName id="7E15E3D7E03CFFD5FAA8D77EFF6CFC2F" pageId="32" pageNumber="33">Dombrow</determinerName>
|
||
2003» (
|
||
<collectionCode id="ED6797D6E03CFFD5FEDED793FE3EFC2F" box="[301,387,904,930]" pageId="32" pageNumber="33">ESDW</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03CFFD5FF64D7CBFBD8FB82" blockId="32.[151,1436,976,1039]" pageId="32" pageNumber="33">
|
||
<emphasis id="B902D301E03CFFD5FF64D7CBFE18FC67" bold="true" box="[151,421,976,1002]" pageId="32" pageNumber="33">
|
||
Condition of
|
||
<typeStatus id="54CDB1B1E03CFFD5FEC4D7CBFE1DFC67" box="[311,416,976,1002]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Good. Pinned. Dissected, parameres and spiculum gastrale glued on separate card below beetle. Right prothoracic tarsus and right metathoracic tarsus missing after tarsomere I.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C5C98E03CFFD5FF64D027FC45FB6A" pageId="32" pageNumber="33" type="discussion">
|
||
<paragraph id="8BC90F13E03CFFD5FF64D027FC45FB6A" blockId="32.[151,1437,1084,1255]" pageId="32" pageNumber="33">
|
||
<emphasis id="B902D301E03CFFD5FF64D027FEBDFBDB" bold="true" box="[151,256,1084,1110]" pageId="32" pageNumber="33">Remark.</emphasis>
|
||
The following specimens from the original
|
||
<typeStatus id="54CDB1B1E03CFFD5FD08D026FC96FBDA" box="[763,811,1085,1111]" pageId="32" pageNumber="33">type</typeStatus>
|
||
series are transferred to several other species, because of the distinct morphological differences see descriptions of the target species: Specimens from the locality “Nieuwoudtville Wildflower Reserve” are transferred to
|
||
<taxonomicName id="4C767490E03CFFD5FD39D09EFCD7FB13" authority="Dombrow & Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[714,874,1157,1182]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nigrosetosa" status="sp. nov.">
|
||
<emphasis id="B902D301E03CFFD5FD39D09EFCD7FB13" box="[714,874,1157,1182]" italics="true" pageId="32" pageNumber="33">B. nigrosetosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2316E7AE03CFFD5FC83D09EFC1AFB12" box="[880,935,1157,1183]" pageId="32" pageNumber="33" rank="species">sp.n.</taxonomicNameLabel>
|
||
; specimens with the locality “Farm Glenlyon …#210” are transferred to
|
||
<taxonomicName id="4C767490E03CFFD5FE32D0B2FDFEFB4F" authorityName="Dombrow" authorityYear="2020" box="[449,579,1193,1218]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="castanea">
|
||
<emphasis id="B902D301E03CFFD5FE32D0B2FDFEFB4F" box="[449,579,1193,1218]" italics="true" pageId="32" pageNumber="33">B. castanea</emphasis>
|
||
</taxonomicName>
|
||
; specimens from the locality “Hantam, Nieuwoudtville” to
|
||
<taxonomicName id="4C767490E03CFFD5FB2AD0B2FAE4FB4F" authority="Dombrow & Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[1241,1369,1192,1218]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="suturalis" status="sp. nov.">
|
||
<emphasis id="B902D301E03CFFD5FB2AD0B2FAE4FB4F" box="[1241,1369,1192,1218]" italics="true" pageId="32" pageNumber="33">B. suturalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2316E7AE03CFFD5FA93D0B2FA2AFB4E" box="[1376,1431,1193,1219]" pageId="32" pageNumber="33" rank="species">sp.n.</taxonomicNameLabel>
|
||
; and specimens from the locality “Biedouw Valley” to
|
||
<taxonomicName id="4C767490E03CFFD5FD07D0D6FC00FB6B" authority="Dombrow & Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[756,957,1228,1254]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="dolichiocnemis" status="sp. nov.">
|
||
<emphasis id="B902D301E03CFFD5FD07D0D6FC00FB6B" box="[756,957,1228,1254]" italics="true" pageId="32" pageNumber="33">B. dolichiocnemis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2316E7AE03CFFD5FC30D0D6FC45FB6A" box="[963,1016,1229,1255]" pageId="32" pageNumber="33" rank="species">sp.n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C5C98E03CFFD5FF64D10FFC9CFA6E" pageId="32" pageNumber="33" type="diagnosis">
|
||
<paragraph id="8BC90F13E03CFFD5FF64D10FFC9CFA6E" blockId="32.[151,1437,1300,1507]" pageId="32" pageNumber="33">
|
||
<emphasis id="B902D301E03CFFD5FF64D10FFE23FAA3" bold="true" box="[151,414,1300,1326]" pageId="32" pageNumber="33">Differential diagnosis.</emphasis>
|
||
Differs from
|
||
<taxonomicName id="4C767490E03CFFD5FDCAD10EFCB3FAA3" authority=": Lateral" authorityName="Lateral" box="[569,782,1300,1327]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pulchra">
|
||
<emphasis id="B902D301E03CFFD5FDCAD10EFD12FAA3" box="[569,687,1300,1326]" italics="true" pageId="32" pageNumber="33">B. pulchra</emphasis>
|
||
: Lateral
|
||
</taxonomicName>
|
||
margin of pronotum of
|
||
<taxonomicName id="4C767490E03CFFD5FBEED10EFB2BFAA3" authorityName="Dombrow" authorityYear="2020" box="[1053,1174,1300,1326]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="colvillei">
|
||
<emphasis id="B902D301E03CFFD5FBEED10EFB2BFAA3" box="[1053,1174,1300,1326]" italics="true" pageId="32" pageNumber="33">B. colvillei</emphasis>
|
||
</taxonomicName>
|
||
widest in anterior half. Lateral margin of pronotum of
|
||
<taxonomicName id="4C767490E03CFFD5FE0AD122FDCDFADF" box="[505,624,1336,1362]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pulchra">
|
||
<emphasis id="B902D301E03CFFD5FE0AD122FDCDFADF" box="[505,624,1336,1362]" italics="true" pageId="32" pageNumber="33">B. pulchra</emphasis>
|
||
</taxonomicName>
|
||
widest at posterior corner. Externo-posterior margin of parameres of
|
||
<taxonomicName id="4C767490E03CFFD5FA77D122FF4CFAFB" authorityName="Dombrow" authorityYear="2020" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="colvillei">
|
||
<emphasis id="B902D301E03CFFD5FA77D122FF4CFAFB" italics="true" pageId="32" pageNumber="33">B. colvillei</emphasis>
|
||
</taxonomicName>
|
||
linear with a median convex extension in dorsal view (
|
||
<figureCitation id="134D1396E03CFFD5FC93D147FC03FAFB" box="[864,958,1372,1398]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="32" pageNumber="33">Figs. 72</figureCitation>
|
||
,
|
||
<figureCitation id="134D1396E03CFFD5FC39D147FC45FAFA" box="[970,1016,1372,1399]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="32" pageNumber="33">105</figureCitation>
|
||
). Externo-posterior margin of parameres of
|
||
<taxonomicName id="4C767490E03CFFD5FF1BD19AFEE0FA17" box="[232,349,1408,1434]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pulchra">
|
||
<emphasis id="B902D301E03CFFD5FF1BD19AFEE0FA17" box="[232,349,1408,1434]" italics="true" pageId="32" pageNumber="33">B. pulchra</emphasis>
|
||
</taxonomicName>
|
||
undulate in dorsal view (
|
||
<figureCitation id="134D1396E03CFFD5FD85D19BFD69FA17" box="[630,724,1408,1434]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="32" pageNumber="33">Figs. 74</figureCitation>
|
||
,
|
||
<figureCitation id="134D1396E03CFFD5FD2DD19BFCB1FA16" box="[734,780,1408,1435]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="32" pageNumber="33">106</figureCitation>
|
||
). Interno-posterior margin of parameres of
|
||
<taxonomicName id="4C767490E03CFFD5FB1FD19AFAD9FA17" authorityName="Dombrow" authorityYear="2020" box="[1260,1380,1408,1434]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="colvillei">
|
||
<emphasis id="B902D301E03CFFD5FB1FD19AFAD9FA17" box="[1260,1380,1408,1434]" italics="true" pageId="32" pageNumber="33">B. colvillei</emphasis>
|
||
</taxonomicName>
|
||
very narrow curly bracket-shaped in dorsal view (
|
||
<figureCitation id="134D1396E03CFFD5FD6BD1BFFD44FA33" box="[664,761,1444,1470]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="32" pageNumber="33">Figs. 72</figureCitation>
|
||
,
|
||
<figureCitation id="134D1396E03CFFD5FCF5D1BFFC89FA32" box="[774,820,1444,1471]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="32" pageNumber="33">105</figureCitation>
|
||
). Interno-posterior margin of parameres of
|
||
<taxonomicName id="4C767490E03CFFD5FAD7D1BEFA21FA33" box="[1316,1436,1444,1470]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pulchra">
|
||
<emphasis id="B902D301E03CFFD5FAD7D1BEFA21FA33" box="[1316,1436,1444,1470]" italics="true" pageId="32" pageNumber="33">B. pulchra</emphasis>
|
||
</taxonomicName>
|
||
broad curly bracket-shaped in dorsal view (
|
||
<figureCitation id="134D1396E03CFFD5FD8ED1D3FD66FA6F" box="[637,731,1480,1506]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="32" pageNumber="33">Figs. 74</figureCitation>
|
||
,
|
||
<figureCitation id="134D1396E03CFFD5FD15D1D3FCA9FA6E" box="[742,788,1480,1507]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="32" pageNumber="33">106</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C5C98E03CFFD7FF64D20BFBD7F819" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33" type="description">
|
||
<paragraph id="8BC90F13E03CFFD5FF64D20BFE80F9A7" blockId="32.[151,1437,1552,2011]" box="[151,317,1552,1578]" pageId="32" pageNumber="33">
|
||
<heading id="D081B87FE03CFFD5FF64D20BFE80F9A7" box="[151,317,1552,1578]" pageId="32" pageNumber="33">
|
||
<emphasis id="B902D301E03CFFD5FF64D20BFE80F9A7" bold="true" box="[151,317,1552,1578]" pageId="32" pageNumber="33">Redescription</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03CFFD5FF34D22FFC83F9C2" blockId="32.[151,1437,1552,2011]" box="[199,830,1588,1615]" pageId="32" pageNumber="33">
|
||
<emphasis id="B902D301E03CFFD5FF34D22FFE3DF9C3" bold="true" box="[199,384,1588,1614]" pageId="32" pageNumber="33">Measurements:</emphasis>
|
||
length
|
||
<quantity id="4C8EA2F6E03CFFD5FE27D22FFDE3F9C2" box="[468,606,1588,1615]" metricMagnitude="-3" metricUnit="m" metricValue="7.55" metricValueMax="8.4" metricValueMin="6.7" pageId="32" pageNumber="33" unit="mm" value="7.55" valueMax="8.4" valueMin="6.7">6.7–8.4 mm</quantity>
|
||
; width
|
||
<quantity id="4C8EA2F6E03CFFD5FD43D22FFC87F9C2" box="[688,826,1588,1615]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="32" pageNumber="33" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.5–4.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03CFFD5FF34D243FEADF91B" blockId="32.[151,1437,1552,2011]" pageId="32" pageNumber="33">
|
||
<emphasis id="B902D301E03CFFD5FF34D243FEE6F9FF" bold="true" box="[199,347,1624,1650]" pageId="32" pageNumber="33">Integument:</emphasis>
|
||
Black; funiculum pale-brown annulated; elytra transparently pale-brown, margins sometimes blackened.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03CFFD4FF34D2BBFB23FC28" blockId="32.[151,1437,1552,2011]" lastBlockId="33.[151,1437,151,2019]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
|
||
<emphasis id="B902D301E03CFFD5FF34D2BBFEFAF937" bold="true" box="[199,327,1696,1722]" pageId="32" pageNumber="33">Tegument:</emphasis>
|
||
<emphasis id="B902D301E03CFFD5FEBCD2BBFE14F937" box="[335,425,1696,1722]" italics="true" pageId="32" pageNumber="33">Clypeus</emphasis>
|
||
bare; loose setose with short, erect, brown setae on the inner side of the reflexed anterior and lateral margin.
|
||
<emphasis id="B902D301E03CFFD5FE82D2DFFE13F953" box="[369,430,1732,1758]" italics="true" pageId="32" pageNumber="33">Head</emphasis>
|
||
loose setose with long, erect, black setae.
|
||
<emphasis id="B902D301E03CFFD5FC7AD2DEFBACF953" box="[905,1041,1732,1758]" italics="true" pageId="32" pageNumber="33">Eye canthus</emphasis>
|
||
with a dense setose ciliary of long, black setae.
|
||
<emphasis id="B902D301E03CFFD5FED2D2F2FECDF88E" box="[289,368,1769,1795]" italics="true" pageId="32" pageNumber="33">Scapus</emphasis>
|
||
loose setose with medium-long, black setae, pale-brown ventrally, sparse and short on
|
||
<emphasis id="B902D301E03CFFD5FAB1D2F3FF05F8AB" italics="true" pageId="32" pageNumber="33">pedicellus</emphasis>
|
||
;
|
||
<emphasis id="B902D301E03CFFD5FF35D317FE8BF8AB" box="[198,310,1804,1830]" italics="true" pageId="32" pageNumber="33">funiculum</emphasis>
|
||
with few short, semi-erect, black setae; sparse and brown on
|
||
<emphasis id="B902D301E03CFFD5FC11D317FB97F8AB" box="[994,1066,1804,1830]" italics="true" pageId="32" pageNumber="33">clavus</emphasis>
|
||
.
|
||
<emphasis id="B902D301E03CFFD5FBC4D316FB1BF8AB" box="[1079,1190,1805,1830]" italics="true" pageId="32" pageNumber="33">Pronotum</emphasis>
|
||
loose setose with medium-long and long, erect, brown setae; with a loose setose ciliary of long and distinctly-long, black setae on lateral margin.
|
||
<emphasis id="B902D301E03CFFD5FF00D34FFEDCF8E3" box="[243,353,1876,1902]" italics="true" pageId="32" pageNumber="33">Scutellum</emphasis>
|
||
loose setose with short and medium-long, erect, black setae.
|
||
<emphasis id="B902D301E03CFFD5FBF9D34FFBEDF8E3" box="[1034,1104,1876,1902]" italics="true" pageId="32" pageNumber="33">Elytra</emphasis>
|
||
without pattern. Loose setose with short, semi-erect, brown setae; medium-long, semi-erect, black setae interspersed, long on humerus and along anterior margin; few short, semi-erect, whitish, squamiform setae interspersed around apical torus; with two sparse rows of medium-long, erect, black setae, slightly shorter in posterior half: a juxta-sutural row from anterior margin, crossing scutellar torus to apical torus; a juxta-humeral row from beside humerus to apical torus. Suture with a loose row of robust, medium-long to distinctly-long, semi-erect, black setae.
|
||
<emphasis id="B902D301E03DFFD4FC02D4A0FBFFFF58" box="[1009,1090,187,213]" italics="true" pageId="33" pageNumber="34">Thorax</emphasis>
|
||
and
|
||
<emphasis id="B902D301E03DFFD4FB84D4A0FB64FF58" box="[1143,1241,187,213]" italics="true" pageId="33" pageNumber="34">pleurites</emphasis>
|
||
loose pilose with long, black hairs.
|
||
<emphasis id="B902D301E03DFFD4FEAED4C4FE09FF75" box="[349,436,223,248]" italics="true" pageId="33" pageNumber="34">Ventrite</emphasis>
|
||
I–V loose setose with short, reclinate, brownish setae; medium-long and long, semi-erect, black setae interspersed.
|
||
<emphasis id="B902D301E03DFFD4FE5CD518FDFDFE90" box="[431,576,259,285]" italics="true" pageId="33" pageNumber="34">Propygidium</emphasis>
|
||
loose squamose with reclinate, narrowly-lanceolate, pale-brown and whitish scales; medium-long and long, semi-erect, black setae interspersed.
|
||
<emphasis id="B902D301E03DFFD4FC75D53CFC4CFECC" box="[902,1009,295,321]" italics="true" pageId="33" pageNumber="34">Pygidium</emphasis>
|
||
sparse setose with long and distinctlylong, erect, black setae; tip with a loose ciliary of medium-long and long, black setae.
|
||
<emphasis id="B902D301E03DFFD4FB75D550FAEAFEE8" box="[1158,1367,331,357]" italics="true" pageId="33" pageNumber="34">Prothoracic femur</emphasis>
|
||
loose pilose with long, black hairs; bare internally.
|
||
<emphasis id="B902D301E03DFFD4FD63D574FCEDFE04" box="[656,848,367,393]" italics="true" pageId="33" pageNumber="34">Prothoracic tibia</emphasis>
|
||
loose setose with short, reclinate, white setae; short, semi-erect, black setae interspersed dorsally; medium-long, erect, black setae interspersed ventrally; with a sparse setose longitudinal row of short and medium-long, erect, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; a setaceous ciliary of short, black setae on anterior margin, more robust towards internal ridge; a medium-long, downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth.
|
||
<emphasis id="B902D301E03DFFD4FF64D638FEC4FDB0" box="[151,377,547,573]" italics="true" pageId="33" pageNumber="34">Mesothoracic femur</emphasis>
|
||
loose pilose with long, black hairs; bare internally.
|
||
<emphasis id="B902D301E03DFFD4FC4FD638FB2DFDB0" box="[956,1168,547,573]" italics="true" pageId="33" pageNumber="34">Mesothoracic tibia</emphasis>
|
||
loose setose with short, reclinate, white setae, sparser internally; medium-long, semi-erect, robust, black setae interspersed, long internally. Posterior margin with a bristled ciliary of very short and short, black bristles; missing internally.
|
||
<emphasis id="B902D301E03DFFD4FB4DD670FA21FD08" box="[1214,1436,619,645]" italics="true" pageId="33" pageNumber="34">Metathoracic femur</emphasis>
|
||
loose setose with long, erect, black setae, long black hairs interspersed; bare internally.
|
||
<emphasis id="B902D301E03DFFD4FB90D694FA8EFD24" box="[1123,1331,655,681]" italics="true" pageId="33" pageNumber="34">Metathoracic tibia</emphasis>
|
||
loose setose with short, reclinate, brown setae; medium-long and long, semi-erect, black setae interspersed, denser and long internally. Posterior margin with a loose bristled outer marginal ciliary of short and medium-long, brown bristles, lengthening dorsally and ventrally, more robust ventrally.
|
||
<emphasis id="B902D301E03DFFD4FCEBD6E0FC23FC98" box="[792,926,763,789]" italics="true" pageId="33" pageNumber="34">Prothoracic</emphasis>
|
||
and
|
||
<emphasis id="B902D301E03DFFD4FC21D6E0FAAAFC99" box="[978,1303,763,789]" italics="true" pageId="33" pageNumber="34">mesothoracic tarsomere I–IV</emphasis>
|
||
bare.
|
||
<emphasis id="B902D301E03DFFD4FAAAD6E0FE1BFCB5" italics="true" pageId="33" pageNumber="34">Metathoracic tarsomere I–IV</emphasis>
|
||
sparse setose with short, reclinate, brown setae.
|
||
<emphasis id="B902D301E03DFFD4FC3AD704FBC5FCB5" box="[969,1144,799,824]" italics="true" pageId="33" pageNumber="34">Tarsomere I–IV</emphasis>
|
||
of all thoracic legs with a setaceous, apical, marginal ciliary of short, black setae, lengthening towards
|
||
<emphasis id="B902D301E03DFFD4FC1ED758FABFFCD1" box="[1005,1282,835,861]" italics="true" pageId="33" pageNumber="34">metathoracic tarsomeres</emphasis>
|
||
, fine dorsally and more robust ventrally.
|
||
<emphasis id="B902D301E03DFFD4FE36D77CFDE0FC0D" box="[453,605,871,896]" italics="true" pageId="33" pageNumber="34">Pretarsomere</emphasis>
|
||
of all thoracic legs sparse setose with short, semi-erect, pale-brown setae; few, medium-long, semi-erect, red-brown setae at apical margin, shorter and paler ventrally.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03DFFD4FF34D7B4FABCFB9C" blockId="33.[151,1437,151,2019]" pageId="33" pageNumber="34">
|
||
<emphasis id="B902D301E03DFFD4FF34D7B4FEADFC44" bold="true" box="[199,272,943,969]" pageId="33" pageNumber="34">Head:</emphasis>
|
||
Suture between head and clypeus undulate, crenated. Head dense punctate with small, shallow, roundish, not merging punctures, micropunctate within. Funicle I–II cylindrical, slightly thickened medially; funicle III short asymmetric frustum-shaped; clavus elliptic, slightly longer than pedicellus and funiculum combined.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03DFFD4FF34D000FD13FBF0" blockId="33.[151,1437,151,2019]" pageId="33" pageNumber="34">
|
||
<emphasis id="B902D301E03DFFD4FF34D000FEF4FBB8" bold="true" box="[199,329,1051,1077]" pageId="33" pageNumber="34">Pronotum:</emphasis>
|
||
Widest medially; dense punctate with small, shallow, roundish punctures, loose on disk. Anterior corner orthogonal. Lateral margin linearly attenuate anteriorly, subparallel posteriorly. With an indistinct, complete, shallow medial furrow. Posterior corner angled.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03DFFD4FF34D09DFAC0FB64" blockId="33.[151,1437,151,2019]" pageId="33" pageNumber="34">
|
||
<emphasis id="B902D301E03DFFD4FF34D09DFEF5FB2D" bold="true" box="[199,328,1158,1184]" pageId="33" pageNumber="34">Scutellum:</emphasis>
|
||
Buckler-shaped; longer than broad; dense punctate with small to large, shallow, roundish punctures, micropunctate within; with a shortened, median, longitudinal, very narrow, smooth stripe from anterior margin terminating in anterior third. Lateral margin linearly attenuate in anterior two thirds, stronger apically. Tip pointed.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03DFFD4FF34D0E8FE3DFA68" blockId="33.[151,1437,151,2019]" pageId="33" pageNumber="34">
|
||
<emphasis id="B902D301E03DFFD4FF34D0E8FEA0FA80" bold="true" box="[199,285,1267,1293]" pageId="33" pageNumber="34">Elytra:</emphasis>
|
||
Ovate; longer than broad; widest medially; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; slightly pulvinate laterally; humerus indistinct; with a broad roundish torus around scutellum; lateral trench missing; complanate beside scutellum and along suture. Loose punctate with extremely small to very small, shallow, roundish punctures, micropunctures in-between; a sparse juxta-sutural row of small, shallow, roundish punctures from anterior margin, crossing scutellar torus towards apical torus; a sparse supra-humeral row of small, shallow, roundish punctures from anterior margin between humerus and scutellar torus towards apical torus.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03DFFD4FF34D1F5FB91F9DC" blockId="33.[151,1437,151,2019]" pageId="33" pageNumber="34">
|
||
<emphasis id="B902D301E03DFFD4FF34D1F5FEF8F985" bold="true" box="[199,325,1518,1544]" pageId="33" pageNumber="34">Abdomen:</emphasis>
|
||
Ventrite I extremely shortened medially; ventrite V broadest, not constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium slightly pulvinate longitudinally and laterally; upper margin moderately convex; lateral margin linearly attenuate apically; tip rounded.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03DFFD4FF34D241FE93F8A4" blockId="33.[151,1437,151,2019]" pageId="33" pageNumber="34">
|
||
<emphasis id="B902D301E03DFFD4FF34D241FE3BF9F8" bold="true" box="[199,390,1626,1653]" pageId="33" pageNumber="34">Prothoracic leg:</emphasis>
|
||
Femur robust. Tibia slender; with a longitudinal, medio-ventral ridge. Median tooth elongate. Basal tooth very small. Interspace between teeth almost equal. Shape of interspace between teeth concave. Tarsomere I–IV combined extremely short; lengthening from tarsomere I–IV; slightly more than
|
||
<superScript id="7C03A25BE03DFFD4FB40D2B9FB06F93D" attach="right" box="[1203,1211,1698,1712]" fontSize="6" pageId="33" pageNumber="34">2</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03DFFD4FB30D2AEFB57F930" box="[1219,1258,1699,1731]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="33" pageNumber="34" unit="in" value="5.0">
|
||
<subScript id="17F20D56E03DFFD4FB30D2AEFB76F94E" attach="left" box="[1219,1227,1717,1731]" fontSize="6" pageId="33" pageNumber="34">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia. Pretarsomere slightly thickened; with a shortened smooth ridge ventrally; as long as tarsomere I–IV combined. Internal claw slightly less than
|
||
<superScript id="7C03A25BE03DFFD4FE04D2F1FE42F975" attach="right" box="[503,511,1770,1784]" fontSize="6" pageId="33" pageNumber="34">1</superScript>
|
||
/
|
||
<subScript id="17F20D56E03DFFD4FDF4D2E6FDAAF886" attach="left" box="[519,535,1789,1803]" fontSize="6" pageId="33" pageNumber="34">10</subScript>
|
||
more in length of pretarsomere. External claw slightly less than
|
||
<superScript id="7C03A25BE03DFFD4FB01D2F0FB47F974" attach="right" box="[1266,1274,1771,1785]" fontSize="6" pageId="33" pageNumber="34">7</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03DFFD4FAF1D2E6FA8DF888" box="[1282,1328,1771,1803]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="33" pageNumber="34" unit="in" value="10.0">
|
||
<subScript id="17F20D56E03DFFD4FAF1D2E6FAAFF886" attach="left" box="[1282,1298,1789,1803]" fontSize="6" pageId="33" pageNumber="34">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of internal claw.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03DFFD4FF34D329FCC0F834" blockId="33.[151,1437,151,2019]" pageId="33" pageNumber="34">
|
||
<emphasis id="B902D301E03DFFD4FF34D329FE21F8C0" bold="true" box="[199,412,1842,1869]" pageId="33" pageNumber="34">Mesothoracic leg:</emphasis>
|
||
Tibia slightly more than
|
||
<superScript id="7C03A25BE03DFFD4FD48D329FD7EF8CD" attach="right" box="[699,707,1842,1856]" fontSize="6" pageId="33" pageNumber="34">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03DFFD4FD38D35EFD4CF8C0" box="[715,753,1843,1875]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="33" pageNumber="34" unit="in" value="5.0">
|
||
<subScript id="17F20D56E03DFFD4FD38D35EFD6EF8DE" attach="left" box="[715,723,1861,1875]" fontSize="6" pageId="33" pageNumber="34">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of femur. Tarsomere I–IV combined extremely short; slightly less than
|
||
<superScript id="7C03A25BE03DFFD4FEABD34DFEDDF8E9" attach="right" box="[344,352,1878,1892]" fontSize="6" pageId="33" pageNumber="34">2</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03DFFD4FE9BD372FE30F8FC" box="[360,397,1879,1911]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="33" pageNumber="34" unit="in" value="5.0">
|
||
<subScript id="17F20D56E03DFFD4FE9BD372FECDF8FA" attach="left" box="[360,368,1897,1911]" fontSize="6" pageId="33" pageNumber="34">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia. Pretarsomere with a shortened smooth ridge ventrally, followed by a roundish cutting apically;
|
||
<superScript id="7C03A25BE03DFFD4FEA1D361FEE7F805" attach="right" box="[338,346,1914,1928]" fontSize="6" pageId="33" pageNumber="34">9</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03DFFD4FE91D396FE32F818" box="[354,399,1915,1947]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="33" pageNumber="34" unit="in" value="10.0">
|
||
<subScript id="17F20D56E03DFFD4FE91D396FECFF816" attach="left" box="[354,370,1933,1947]" fontSize="6" pageId="33" pageNumber="34">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of tarsomere I–IV combined. External claw slightly less than
|
||
<superScript id="7C03A25BE03DFFD4FB7FD361FB29F805" attach="right" box="[1164,1172,1914,1928]" fontSize="6" pageId="33" pageNumber="34">9</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03DFFD4FB6FD396FB74F818" box="[1180,1225,1915,1947]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="33" pageNumber="34" unit="in" value="10.0">
|
||
<subScript id="17F20D56E03DFFD4FB6FD396FB11F816" attach="left" box="[1180,1196,1933,1947]" fontSize="6" pageId="33" pageNumber="34">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of pretarsomere. Internal claw slightly less than
|
||
<superScript id="7C03A25BE03DFFD4FDC9D385FDFFF821" attach="right" box="[570,578,1950,1964]" fontSize="6" pageId="33" pageNumber="34">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03DFFD4FDB9D3AAFDD2F834" box="[586,623,1951,1983]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="33" pageNumber="34" unit="in" value="5.0">
|
||
<subScript id="17F20D56E03DFFD4FDB9D3AAFDEFF832" attach="left" box="[586,594,1969,1983]" fontSize="6" pageId="33" pageNumber="34">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of external claw.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03DFFD7FF34D3D9FCF2FF75" blockId="33.[151,1437,151,2019]" lastBlockId="34.[151,1437,150,1946]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
|
||
<emphasis id="B902D301E03DFFD4FF34D3D9FE27F850" bold="true" box="[199,410,1986,2013]" pageId="33" pageNumber="34">Metathoracic leg:</emphasis>
|
||
Tibia slightly more than
|
||
<superScript id="7C03A25BE03DFFD4FD49D3D9FD7FF85D" attach="right" box="[698,706,1986,2000]" fontSize="6" pageId="33" pageNumber="34">9</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03DFFD4FD3AD3CEFD45F850" box="[713,760,1987,2019]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="33" pageNumber="34" unit="in" value="10.0">
|
||
<subScript id="17F20D56E03DFFD4FD3AD3CEFD64F86E" attach="left" box="[713,729,2005,2019]" fontSize="6" pageId="33" pageNumber="34">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of femur. Tarsomere I–IV combined distinctly short; slightly more than
|
||
<superScript id="7C03A25BE03EFFD7FE9BD48DFECDFF29" attach="right" box="[360,368,150,164]" fontSize="6" pageId="34" pageNumber="35">7</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03EFFD7FE8BD4B2FE19FF3D" box="[376,420,150,183]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="34" pageNumber="35" unit="in" value="10.0">
|
||
<subScript id="17F20D56E03EFFD7FE8BD4B2FE35FF3A" attach="left" box="[376,392,169,183]" fontSize="6" pageId="34" pageNumber="35">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia. Pretarsomere slightly thickened; with a smooth ridge ventrally; slightly less than
|
||
<superScript id="7C03A25BE03EFFD7FF3CD4A1FF6AFF45" attach="right" box="[207,215,186,200]" fontSize="6" pageId="34" pageNumber="35">1</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03EFFD7FF2CD4D6FEBBFF59" box="[223,262,186,219]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="34" pageNumber="35" unit="in" value="2.0">
|
||
<subScript id="17F20D56E03EFFD7FF2CD4D6FF5AFF56" attach="left" box="[223,231,205,219]" fontSize="6" pageId="34" pageNumber="35">2</subScript>
|
||
in
|
||
</quantity>
|
||
length of tarsomere I–IV combined. External claw slightly less than
|
||
<superScript id="7C03A25BE03EFFD7FBE5D4A1FBA3FF45" attach="right" box="[1046,1054,186,200]" fontSize="6" pageId="34" pageNumber="35">1</superScript>
|
||
/
|
||
<subScript id="17F20D56E03EFFD7FBD5D4D7FB93FF57" attach="left" box="[1062,1070,204,218]" fontSize="6" pageId="34" pageNumber="35">5</subScript>
|
||
more in length of pretarsomere. Internal claw slightly more than
|
||
<superScript id="7C03A25BE03EFFD7FDF7D4C5FDB1FF61" attach="right" box="[516,524,222,236]" fontSize="6" pageId="34" pageNumber="35">7</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03EFFD7FDE7D4EAFDFCFF75" box="[532,577,222,255]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="34" pageNumber="35" unit="in" value="10.0">
|
||
<subScript id="17F20D56E03EFFD7FDE7D4EAFD99FF72" attach="left" box="[532,548,241,255]" fontSize="6" pageId="34" pageNumber="35">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of external claw.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03EFFD7FF34D519FD93FE90" blockId="34.[151,1437,150,1946]" box="[199,558,258,285]" pageId="34" pageNumber="35">
|
||
<emphasis id="B902D301E03EFFD7FF34D519FEECFE91" bold="true" box="[199,337,258,284]" pageId="34" pageNumber="35">Parameres:</emphasis>
|
||
(
|
||
<figureCitation id="134D1396E03EFFD7FE93D519FE55FE91" box="[352,488,258,284]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="34" pageNumber="35">Figs. 72–73</figureCitation>
|
||
,
|
||
<figureCitation id="134D1396E03EFFD7FE07D519FD9FFE90" box="[500,546,258,285]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="34" pageNumber="35">105</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03EFFD7FF34D533FCCBFECC" blockId="34.[151,1437,150,1946]" box="[199,886,294,323]" pageId="34" pageNumber="35">
|
||
<emphasis id="B902D301E03EFFD7FF34D533FE05FECD" bold="true" box="[199,440,294,323]" pageId="34" pageNumber="35">
|
||
<specimenCount id="9D70C49AE03EFFD7FF34D533FF4FFECE" box="[199,242,296,323]" pageId="34" pageNumber="35" type="female">♀♀</specimenCount>
|
||
. Measurements:
|
||
</emphasis>
|
||
length
|
||
<quantity id="4C8EA2F6E03EFFD7FDFFD53DFD2BFECC" box="[524,662,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" metricValueMax="9.3" metricValueMin="8.5" pageId="34" pageNumber="35" unit="mm" value="8.9" valueMax="9.3" valueMin="8.5">8.5–9.3 mm</quantity>
|
||
; width
|
||
<quantity id="4C8EA2F6E03EFFD7FD1BD53DFCCFFECC" box="[744,882,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="5.1" metricValueMin="4.2" pageId="34" pageNumber="35" unit="mm" value="4.65" valueMax="5.1" valueMin="4.2">4.2–5.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03EFFD7FF34D551FA25FEE9" blockId="34.[151,1437,150,1946]" box="[199,1432,330,356]" pageId="34" pageNumber="35">
|
||
<emphasis id="B902D301E03EFFD7FF34D551FEE6FEE9" bold="true" box="[199,347,330,356]" pageId="34" pageNumber="35">Integument:</emphasis>
|
||
Black; flagellum brown; thoracic legs dark brown; metathoracic tibia and all thoracic tarsi paler.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03EFFD7FF34D575FCD3FBD4" blockId="34.[151,1437,150,1946]" pageId="34" pageNumber="35">
|
||
<emphasis id="B902D301E03EFFD7FF34D575FEFAFE05" bold="true" box="[199,327,366,392]" pageId="34" pageNumber="35">Tegument:</emphasis>
|
||
<emphasis id="B902D301E03EFFD7FEBED575FE1AFE05" box="[333,423,366,392]" italics="true" pageId="34" pageNumber="35">Clypeus</emphasis>
|
||
loose setose with short, erect, brown setae, medium-long on
|
||
<emphasis id="B902D301E03EFFD7FBBBD575FBC3FE05" box="[1096,1150,366,392]" italics="true" pageId="34" pageNumber="35">head</emphasis>
|
||
.
|
||
<emphasis id="B902D301E03EFFD7FB78D574FAAFFE05" box="[1163,1298,366,392]" italics="true" pageId="34" pageNumber="35">Eye canthus</emphasis>
|
||
with a loose setose ciliary of medium-long, brown setae. Setae on
|
||
<emphasis id="B902D301E03EFFD7FD1DD589FCE3FE21" box="[750,862,402,428]" italics="true" pageId="34" pageNumber="35">funiculum</emphasis>
|
||
and
|
||
<emphasis id="B902D301E03EFFD7FC67D589FC61FE21" box="[916,988,402,428]" italics="true" pageId="34" pageNumber="35">clavus</emphasis>
|
||
extremely short.
|
||
<emphasis id="B902D301E03EFFD7FB6CD588FAB3FE21" box="[1183,1294,403,428]" italics="true" pageId="34" pageNumber="35">Pronotum</emphasis>
|
||
without pattern. Loose setose with short, reclinate and long, semi-erect, brown setae; with a loose setose ciliary of long, brown setae on lateral margin.
|
||
<emphasis id="B902D301E03EFFD7FE5BD5C1FDABFE79" box="[424,534,474,500]" italics="true" pageId="34" pageNumber="35">Scutellum</emphasis>
|
||
loose setose with short and medium-long, semi-erect, whitish and brown setae.
|
||
<emphasis id="B902D301E03EFFD7FF64D5E5FF60FD95" box="[151,221,510,536]" italics="true" pageId="34" pageNumber="35">Elytra</emphasis>
|
||
loose setose with very short and short, semi-erect, pale-brown setae; medium-long, semi-erect, brown setae interspersed; few long, semi-erect, pale-brown, squamiform setae within lateral trench and around apical torus. Suture with a loose row of short and medium-long, semi-erect, robust, brown setae.
|
||
<emphasis id="B902D301E03EFFD7FB9AD65DFB07FDED" box="[1129,1210,582,608]" italics="true" pageId="34" pageNumber="35">Thorax</emphasis>
|
||
and
|
||
<emphasis id="B902D301E03EFFD7FB06D65DFAEAFDED" box="[1269,1367,582,608]" italics="true" pageId="34" pageNumber="35">pleurites</emphasis>
|
||
loose pilose with long, pale-brown hairs.
|
||
<emphasis id="B902D301E03EFFD7FDDAD670FD0DFD09" box="[553,688,619,644]" italics="true" pageId="34" pageNumber="35">Ventrite I–V</emphasis>
|
||
loose setose with short, reclinate, pale-brown setae; medium-long and long, erect, brown setae interspersed.
|
||
<emphasis id="B902D301E03EFFD7FD9ED695FD43FD25" box="[621,766,654,680]" italics="true" pageId="34" pageNumber="35">Propygidium</emphasis>
|
||
and
|
||
<emphasis id="B902D301E03EFFD7FCC7D695FC21FD25" box="[820,924,654,680]" italics="true" pageId="34" pageNumber="35">pygidium</emphasis>
|
||
loose setose with short, reclinate, pale-brown setae; medium-long, semi-erect, brown setae interspersed; tip of pygidium with a loose ciliary of short and mediumlong, brown setae.
|
||
<emphasis id="B902D301E03EFFD7FE9BD6CDFD88FD7D" box="[360,565,726,752]" italics="true" pageId="34" pageNumber="35">Prothoracic femur</emphasis>
|
||
loose pilose with short and medium-long, brown hairs.
|
||
<emphasis id="B902D301E03EFFD7FB6FD6CDFAE6FD7D" box="[1180,1371,726,752]" italics="true" pageId="34" pageNumber="35">Prothoracic tibia</emphasis>
|
||
loose setose with short, reclinate, pale-brown setae dorsally, erect, medium-long and long ventrally; with a setaceous ciliary of short, pale-brown setae on anterior margin.
|
||
<emphasis id="B902D301E03EFFD7FD25D705FC07FCB5" box="[726,954,798,824]" italics="true" pageId="34" pageNumber="35">Mesothoracic femur</emphasis>
|
||
loose pilose with long, brown hairs; paler ventrally.
|
||
<emphasis id="B902D301E03EFFD7FEF4D759FE66FCD1" box="[263,475,834,860]" italics="true" pageId="34" pageNumber="35">Mesothoracic tibia</emphasis>
|
||
loose setose with short, reclinate, pale-brown setae; sparser internally; medium-long, semi-erect, robust, red-brown setae interspersed, longer internally. Posterior margin with a bristled ciliary of very short and short, red-brown bristles; lengthening dorsally and ventrally; missing internally.
|
||
<emphasis id="B902D301E03EFFD7FB8ED791FAE6FC29" box="[1149,1371,906,932]" italics="true" pageId="34" pageNumber="35">Metathoracic femur</emphasis>
|
||
loose setose with medium-long and long, whitish and brown setae.
|
||
<emphasis id="B902D301E03EFFD7FCC9D7B5FBB4FC45" box="[826,1033,942,968]" italics="true" pageId="34" pageNumber="35">Metathoracic tibia</emphasis>
|
||
sparse setose with short, semi-erect, white setae; medium-long and long, robust, semi-erect, brown setae interspersed, loose and finer internally. Posterior margin with a loose bristled outer marginal ciliary of very short, red-brown bristles, lengthening dorsally and ventrally, more robust ventrally. Ciliary of
|
||
<emphasis id="B902D301E03EFFD7FD82D000FCA4FBB9" box="[625,793,1051,1076]" italics="true" pageId="34" pageNumber="35">tarsomere I–IV</emphasis>
|
||
of thoracic legs red-brown.
|
||
<emphasis id="B902D301E03EFFD7FBA2D000FB54FBB9" box="[1105,1257,1051,1076]" italics="true" pageId="34" pageNumber="35">Pretarsomere</emphasis>
|
||
of
|
||
<emphasis id="B902D301E03EFFD7FAFFD001FAD4FBB9" box="[1292,1385,1050,1076]" italics="true" pageId="34" pageNumber="35">thoracic</emphasis>
|
||
legs sparse setose with extremely short, semi-erect, pale-brown setae.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03EFFD7FF34D079FE3AFB65" blockId="34.[151,1437,150,1946]" pageId="34" pageNumber="35">
|
||
<emphasis id="B902D301E03EFFD7FF34D079FEADFBF1" bold="true" box="[199,272,1122,1148]" pageId="34" pageNumber="35">Head:</emphasis>
|
||
Clypeus very slightly depressed laterally; dense punctate with large, shallow roundish punctures. Anterior margin of clypeus slightly vertically reflexed. Lateral margin very slightly reflexed. Head dense punctate with small to large, shallow, roundish, partly merging punctures. Eye below head level. Clavus as long as pedicellus and funiculum combined.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03EFFD7FF34D0E9FF46FABD" blockId="34.[151,1437,150,1946]" pageId="34" pageNumber="35">
|
||
<emphasis id="B902D301E03EFFD7FF34D0E9FEF4FA81" bold="true" box="[199,329,1266,1292]" pageId="34" pageNumber="35">Pronotum:</emphasis>
|
||
Moderate pulvinate laterally. Anterior corner orthogonal. Lateral margin convex. Posterior corner rounded.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03EFFD7FF34D121FE61FAF5" blockId="34.[151,1437,150,1946]" pageId="34" pageNumber="35">
|
||
<emphasis id="B902D301E03EFFD7FF34D121FEF5FAD9" bold="true" box="[199,328,1338,1364]" pageId="34" pageNumber="35">Scutellum:</emphasis>
|
||
Parabola-shaped; as broad as long; dense punctate with small, shallow, roundish punctures. Lateral margin convex. Tip rounded.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03EFFD7FF34D199FAF7FA4D" blockId="34.[151,1437,150,1946]" pageId="34" pageNumber="35">
|
||
<emphasis id="B902D301E03EFFD7FF34D199FEA0FA11" bold="true" box="[199,285,1410,1436]" pageId="34" pageNumber="35">Elytra:</emphasis>
|
||
Widest in anterior quarter; covering propygidium. Sculpture indistinct; scutellar torus missing; lateral trench indistinct. Dense punctate with very small, shallow, roundish punctures, micropunctures in-between.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03EFFD7FF34D1D1FBEFF985" blockId="34.[151,1437,150,1946]" pageId="34" pageNumber="35">
|
||
<emphasis id="B902D301E03EFFD7FF34D1D1FEF8FA69" bold="true" box="[199,325,1482,1508]" pageId="34" pageNumber="35">Abdomen:</emphasis>
|
||
Distinctly pulvinate; ventrite I distinctly shortened medially; ventrite II–V broadened medially; ventrite V not constricted ventrally. Pygidium slightly declined posteriorly; tip linear.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03EFFD7FF34D209FD83F915" blockId="34.[151,1437,150,1946]" pageId="34" pageNumber="35">
|
||
<emphasis id="B902D301E03EFFD7FF34D209FE3BF9A1" bold="true" box="[199,390,1554,1580]" pageId="34" pageNumber="35">Prothoracic leg:</emphasis>
|
||
Median tooth broad triangular-shaped. Tarsomere I–IV combined very short; equal in length;
|
||
<superScript id="7C03A25BE03EFFD7FF64D22DFF22F9C9" attach="none" box="[151,159,1590,1604]" fontSize="6" pageId="34" pageNumber="35">7</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03EFFD7FF54D252FF6EF9DD" box="[167,211,1590,1623]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="34" pageNumber="35" unit="in" value="10.0">
|
||
<subScript id="17F20D56E03EFFD7FF54D252FF0AF9DA" attach="left" box="[167,183,1609,1623]" fontSize="6" pageId="34" pageNumber="35">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia. Pretarsomere slender; smooth ventrally, roundish cutting missing; slightly less than
|
||
<superScript id="7C03A25BE03EFFD7FAEED22DFA98F9C9" attach="right" box="[1309,1317,1590,1604]" fontSize="6" pageId="34" pageNumber="35">3</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03EFFD7FADED253FAECF9DD" box="[1325,1361,1590,1622]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="34" pageNumber="35" unit="in" value="5.0">
|
||
<subScript id="17F20D56E03EFFD7FADED253FA88F9DB" attach="left" box="[1325,1333,1608,1622]" fontSize="6" pageId="34" pageNumber="35">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of tarsomere I–IV combined. Internal claw slender; not broadened; as long as pretarsomere. External claw slightly less than
|
||
<superScript id="7C03A25BE03EFFD7FF0CD265FEBAF901" attach="right" box="[255,263,1662,1676]" fontSize="6" pageId="34" pageNumber="35">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03EFFD7FEFCD28BFE89F915" box="[271,308,1662,1694]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="34" pageNumber="35" unit="in" value="5.0">
|
||
<subScript id="17F20D56E03EFFD7FEFCD28BFEAAF913" attach="left" box="[271,279,1680,1694]" fontSize="6" pageId="34" pageNumber="35">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of internal claw.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03EFFD7FF34D2B9FDB9F8A5" blockId="34.[151,1437,150,1946]" pageId="34" pageNumber="35">
|
||
<emphasis id="B902D301E03EFFD7FF34D2B9FE26F931" bold="true" box="[199,411,1698,1724]" pageId="34" pageNumber="35">Mesothoracic leg:</emphasis>
|
||
Tibia
|
||
<superScript id="7C03A25BE03EFFD7FE17D2B9FE51F93D" attach="right" box="[484,492,1698,1712]" fontSize="6" pageId="34" pageNumber="35">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03EFFD7FE07D2AFFDA4F931" box="[500,537,1698,1730]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="34" pageNumber="35" unit="in" value="5.0">
|
||
<subScript id="17F20D56E03EFFD7FE07D2AFFE41F94F" attach="left" box="[500,508,1716,1730]" fontSize="6" pageId="34" pageNumber="35">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of femur. Tarsomere I–IV combined distinctly short; slightly less than
|
||
<superScript id="7C03A25BE03EFFD7FA77D2B9FA31F93D" attach="right" box="[1412,1420,1698,1712]" fontSize="6" pageId="34" pageNumber="35">1</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03EFFD7FA67D2AEFF10F96D" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="34" pageNumber="35" unit="in" value="2.0">
|
||
<subScript id="17F20D56E03EFFD7FA67D2AEFA21F94E" attach="none" box="[1428,1436,1717,1731]" fontSize="6" pageId="34" pageNumber="35">2</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia. Pretarsomere slightly thickened; smooth ventrally, roundish cutting missing; slightly more than
|
||
<superScript id="7C03A25BE03EFFD7FF64D2F1FF22F975" attach="none" box="[151,159,1770,1784]" fontSize="6" pageId="34" pageNumber="35">7</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03EFFD7FF54D2E6FF68F889" box="[167,213,1770,1803]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="34" pageNumber="35" unit="in" value="10.0">
|
||
<subScript id="17F20D56E03EFFD7FF54D2E6FF0AF886" attach="left" box="[167,183,1789,1803]" fontSize="6" pageId="34" pageNumber="35">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of tarsomere I–IV combined. External claw slender; not broadened medially; as long as pretarsomere. Internal claw slightly less than
|
||
<superScript id="7C03A25BE03EFFD7FE07D315FE41F891" attach="right" box="[500,508,1806,1820]" fontSize="6" pageId="34" pageNumber="35">9</superScript>
|
||
/
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03EFFD7FDF7D33AFC82F8A5" blockId="34.[151,1437,150,1946]" box="[516,831,1806,1839]" pageId="34" pageNumber="35">10 in length of external claw.</paragraph>
|
||
<paragraph id="8BC90F13E03EFFD7FF34D329FBD7F819" blockId="34.[151,1437,150,1946]" pageId="34" pageNumber="35">
|
||
<emphasis id="B902D301E03EFFD7FF34D329FE26F8C1" bold="true" box="[199,411,1842,1868]" pageId="34" pageNumber="35">Metathoracic leg:</emphasis>
|
||
Femur slightly thickened. Tibia slightly extended dorso-apically; slightly more than
|
||
<superScript id="7C03A25BE03EFFD7FAADD329FADBF8CD" attach="right" box="[1374,1382,1842,1856]" fontSize="6" pageId="34" pageNumber="35">7</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03EFFD7FA9DD35EFA21F8C1" box="[1390,1436,1842,1875]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="34" pageNumber="35" unit="in" value="10.0">
|
||
<subScript id="17F20D56E03EFFD7FA9DD35EFAC3F8DE" attach="left" box="[1390,1406,1861,1875]" fontSize="6" pageId="34" pageNumber="35">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of femur. Tarsomere I–IV combined short; slightly less than
|
||
<superScript id="7C03A25BE03EFFD7FC77D34DFC31F8E9" attach="right" box="[900,908,1878,1892]" fontSize="6" pageId="34" pageNumber="35">7</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03EFFD7FC67D372FC7CF8FD" box="[916,961,1878,1911]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="34" pageNumber="35" unit="in" value="10.0">
|
||
<subScript id="17F20D56E03EFFD7FC67D372FC19F8FA" attach="left" box="[916,932,1897,1911]" fontSize="6" pageId="34" pageNumber="35">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia. Pretarsomere slender; with a granulate ridge ventrally. Internal claw slightly less than
|
||
<superScript id="7C03A25BE03EFFD7FCD4D361FC92F805" attach="right" box="[807,815,1914,1928]" fontSize="6" pageId="34" pageNumber="35">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E03EFFD7FCC4D397FCE1F819" box="[823,860,1914,1946]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="34" pageNumber="35" unit="in" value="5.0">
|
||
<subScript id="17F20D56E03EFFD7FCC4D397FC82F817" attach="left" box="[823,831,1932,1946]" fontSize="6" pageId="34" pageNumber="35">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of external claw.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C5C98E03EFFD6FF64D3D9FCEBFF3C" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="distribution">
|
||
<paragraph id="8BC90F13E03EFFD7FF64D3D9FBEDF851" blockId="34.[151,1104,1986,2013]" box="[151,1104,1986,2013]" pageId="34" pageNumber="35">
|
||
<emphasis id="B902D301E03EFFD7FF64D3D9FE8CF851" bold="true" box="[151,305,1986,2012]" pageId="34" pageNumber="35">Distribution.</emphasis>
|
||
<collectingRegion id="49B2C1F1E03EFFD7FEC4D3D9FDF5F851" box="[311,584,1986,2012]" country="South Africa" name="Northern Cape" pageId="34" pageNumber="35">Northern Cape Province</collectingRegion>
|
||
(
|
||
<figureCitation id="134D1396E03EFFD7FDA4D3D9FD05F851" box="[599,696,1986,2012]" captionStart="FIGURE 140" captionStartId="72.[151,250,1944,1968]" captionTargetBox="[195,1393,188,1910]" captionTargetId="figure-25@72.[185,1402,181,1920]" captionTargetPageId="72" captionText="FIGURE 140. Distributions of 18 Beckhoplia Dombrow, 2005 species, showing the concentration of species around the Northern Cape town of Nieuwoudtville." figureDoi="http://doi.org/10.5281/zenodo.4401964" httpUri="https://zenodo.org/record/4401964/files/figure.png" pageId="34" pageNumber="35">Fig. 140</figureCitation>
|
||
), only known from the
|
||
<typeStatus id="54CDB1B1E03EFFD7FC4CD3D8FC52F850" box="[959,1007,1987,2013]" pageId="34" pageNumber="35">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E03FFFD6FF64D48DFCEBFF3C" blockId="35.[151,854,150,177]" box="[151,854,150,177]" pageId="35" pageNumber="36">
|
||
<emphasis id="B902D301E03FFFD6FF64D48DFEE3FF3C" bold="true" box="[151,350,150,177]" pageId="35" pageNumber="36">
|
||
Vegetation
|
||
<typeStatus id="54CDB1B1E03FFFD6FEEFD48CFEE4FF3C" box="[284,345,151,177]" pageId="35" pageNumber="36">Type</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Nieuwoudtville Shale Renosterveld (FRs 2).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C5C98E03FFFD6FF64D4C4FCFAFF74" box="[151,839,223,250]" pageId="35" pageNumber="36" type="biology_ecology">
|
||
<paragraph id="8BC90F13E03FFFD6FF64D4C4FCFAFF74" blockId="35.[151,839,223,250]" box="[151,839,223,250]" pageId="35" pageNumber="36">
|
||
<emphasis id="B902D301E03FFFD6FF64D4C4FE95FF74" bold="true" box="[151,296,223,249]" pageId="35" pageNumber="36">Host Plants.</emphasis>
|
||
<taxonomicName id="4C767490E03FFFD6FEDED4C4FE11FF74" box="[301,428,223,249]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C767490E03FFFD6FE44D4C4FDDCFF74" authorityName="Roessler" authorityYear="1959" baseAuthorityName="Burm. fil." box="[439,609,223,249]" class="Magnoliopsida" family="Asteraceae" genus="Gazania" kingdom="Plantae" order="Asterales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="rigida">
|
||
<emphasis id="B902D301E03FFFD6FE44D4C4FDDCFF74" box="[439,609,223,249]" italics="true" pageId="35" pageNumber="36">Gazania rigida</emphasis>
|
||
</taxonomicName>
|
||
(Burm. f.) Roessler.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |