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<mods:title id="E914A359378730891B17B210A9B908DA">An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas</mods:title>
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<treatment id="03D7FF1CFFC1FFE6FF48FD48332DFEC5" ID-DOI="http://doi.org/10.5281/zenodo.3684119" ID-GBIF-Taxon="162245817" ID-Zenodo-Dep="3684119" LSID="urn:lsid:plazi:treatment:03D7FF1CFFC1FFE6FF48FD48332DFEC5" httpUri="http://treatment.plazi.org/id/03D7FF1CFFC1FFE6FF48FD48332DFEC5" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9">
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<paragraph id="8BC14E0AFFC1FFE7FF48FD4832F7F9CE" blockId="10.[192,907,1574,1598]" box="[192,907,1574,1598]" pageId="10" pageNumber="9">
<heading id="D089F966FFC1FFE7FF48FD4832F7F9CE" bold="true" box="[192,907,1574,1598]" fontSize="10" level="2" pageId="10" pageNumber="9" reason="3">
<taxonomicName id="4C7E3589FFC1FFE7FF48FD4832F7F9CE" authority="(Klug, 1845)" baseAuthorityName="Klug" baseAuthorityYear="1845" box="[192,907,1574,1598]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="lanuginosus" subGenus="Neoathyreus">
<emphasis id="B90A9218FFC1FFE7FF48FD4832F7F9CE" bold="true" box="[192,907,1574,1598]" pageId="10" pageNumber="9">
<emphasis id="B90A9218FFC1FFE7FF48FD483017F9CE" bold="true" box="[192,363,1574,1598]" italics="true" pageId="10" pageNumber="9">Neoathyreus</emphasis>
(
<emphasis id="B90A9218FFC1FFE7FEF4FD48335BF9CE" bold="true" box="[380,551,1574,1598]" italics="true" pageId="10" pageNumber="9">Neoathyreus</emphasis>
)
<emphasis id="B90A9218FFC1FFE7FDB1FD48339DF9CE" bold="true" box="[569,737,1574,1598]" italics="true" pageId="10" pageNumber="9">lanuginosus</emphasis>
(
<bibRefCitation id="EFEF33FBFFC1FFE7FD79FD4832FEF9CE" author="Klug" box="[753,898,1574,1598]" firstAuthor="Klug" pageId="10" pageNumber="9" pagination="21 - 57" refId="ref210635" refString="Klug, J. C. F. 1845 (1843). Die Coleopteren-Gattungen Athyreus und Bolboceras, dargestellt nach den in der Sammlung hiesiger Konigl. Universitat davon vorhandenen Arten. Abhandlungen der Koniglichen Akademie der Wissenschaften 1843 (1845): 21 - 57." type="journal article" year="1845">Klug, 1845</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3641D81FFC1FFE7FF60FD2232D5F993" box="[232,937,1612,1635]" pageId="10" pageNumber="9" type="reference_group">
<paragraph id="8BC14E0AFFC1FFE7FF60FD2232D5F993" blockId="10.[232,937,1612,1635]" box="[232,937,1612,1635]" pageId="10" pageNumber="9">
<treatmentCitationGroup id="AB6E6924FFC1FFE7FF60FD2232D5F993" box="[232,937,1612,1635]" pageId="10" pageNumber="9">
<treatmentCitation id="0ADF681BFFC1FFE7FF60FD223306F992" author="Klug" box="[232,634,1612,1635]" page="28" pageId="10" pageNumber="9" year="1845">
<taxonomicName id="4C7E3589FFC1FFE7FF60FD223306F992" authority="Klug 1845: 28" authorityName="Klug" authorityPageNumber="28" authorityYear="1845" box="[232,634,1612,1635]" class="Insecta" family="Bolboceratidae" genus="Athyreus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="lanuginosus">
<emphasis id="B90A9218FFC1FFE7FF60FD2230A4F991" box="[232,472,1612,1633]" italics="true" pageId="10" pageNumber="9">Athyreus lanuginosus</emphasis>
<bibRefCitation id="EFEF33FBFFC1FFE7FE57FD233306F992" author="Klug" box="[479,634,1613,1635]" firstAuthor="Klug" pageId="10" pageNumber="9" pagination="21 - 57" refId="ref210635" refString="Klug, J. C. F. 1845 (1843). Die Coleopteren-Gattungen Athyreus und Bolboceras, dargestellt nach den in der Sammlung hiesiger Konigl. Universitat davon vorhandenen Arten. Abhandlungen der Koniglichen Akademie der Wissenschaften 1843 (1845): 21 - 57." type="journal article" year="1845">Klug 1845: 28</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<collectingCountry id="F3690E9AFFC1FFE7FD0FFD233393F993" box="[647,751,1613,1635]" name="Colombia" pageId="10" pageNumber="9">Colombia</collectingCountry>
-
<typeStatus id="54C5F0A8FFC1FFE7FC8DFD2332DFF993" box="[773,931,1613,1635]" pageId="10" pageNumber="9" type="lectotype">LECTOTYPE</typeStatus>
)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3641D81FFC1FFE6FF48FD1D332DFEC5" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9" type="distribution">
<paragraph id="8BC14E0AFFC1FFE7FF48FD1D30A3F8BB" blockId="10.[192,1440,1651,1867]" pageId="10" pageNumber="9">
<emphasis id="B90A9218FFC1FFE7FF48FD1D3368F97B" bold="true" box="[192,532,1651,1675]" pageId="10" pageNumber="9">
Distribution.
<collectingCountry id="F3690E9AFFC1FFE7FE0BFD1D3373F97B" box="[387,527,1651,1675]" name="Suriname" pageId="10" pageNumber="9">Suriname</collectingCountry>
:
</emphasis>
MHCN -
<emphasis id="B90A9218FFC1FFE7FD11FD1D3572F97B" bold="true" box="[665,1038,1651,1675]" pageId="10" pageNumber="9">NEW COUNTRY RECORD</emphasis>
(Suriname,
<collectingRegion id="49BA80E8FFC1FFE7FB39FD1D3449F97B" box="[1201,1333,1651,1675]" country="Suriname" name="Sipaliwini" pageId="10" pageNumber="9">Sipaliwini</collectingRegion>
, Puketi, (
<geoCoordinate id="EE4A28CDFFC1FFE7FF40FDFC3039F95B" box="[200,325,1682,1707]" degrees="4" direction="north" minutes="07" orientation="latitude" pageId="10" pageNumber="9" precision="15" seconds="30" value="4.125">4°0730″N</geoCoordinate>
,
<geoCoordinate id="EE4A28CDFFC1FFE7FEDBFDFC3098F95B" box="[339,484,1682,1707]" degrees="54" direction="west" minutes="37" orientation="longitude" pageId="10" pageNumber="9" precision="15" seconds="30" value="-54.625">54°3730″W</geoCoordinate>
). 16/
<date id="FFC068CAFFC1FFE7FD96FDFD33CFF95B" box="[542,691,1683,1707]" pageId="10" pageNumber="9" value="2013-07-21">21-VII-2013</date>
. A. J. Hielkema. In FIT in forest.); NZCS.
<emphasis id="B90A9218FFC1FFE7FB4FFDFD34DCF95B" bold="true" box="[1223,1440,1683,1707]" pageId="10" pageNumber="9">
<collectingCountry id="F3690E9AFFC1FFE7FB4FFDFD34E7F95B" box="[1223,1435,1683,1707]" name="French Guiana" pageId="10" pageNumber="9">French Guiana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFFC1FFE7FF48FDDD30EFF93B" author="Howden" box="[192,403,1715,1739]" firstAuthor="Howden" journalOrPublisher="Contributions of the American Entomological Institute" pageId="10" pageNumber="9" pagination="1 - 95" part="21" refId="ref208469" refString="Howden, H. F. 1985. A revision of the South American species in the genus Neoathyreus Howden and Martinez (Coleoptera, Scarabaeidae, Geotrupinae). Contributions of the American Entomological Institute 21 (4): 1 - 95." title="A revision of the South American species in the genus Neoathyreus Howden and Martinez (Coleoptera, Scarabaeidae, Geotrupinae)" type="journal article" year="1985">Howden 1985: 58</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFFC1FFE7FE17FDDD3318F93B" author="Boilly" box="[415,612,1715,1739]" firstAuthor="Boilly" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="10" pageNumber="9" pagination="19 - 31" part="4" refId="ref194293" refString="Boilly, O. 2011 b. Les Bolboceratidae de Guyane (Coleoptera, Scarabaeoidea). Contribution a l'etude des coleopteres de Guyane 4: 19 - 31. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Les Bolboceratidae de Guyane (Coleoptera, Scarabaeoidea)" type="journal article" year="2011" yearSuffix="b">Boilly 2011b: 30</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFFC1FFE7FDE7FDDD32FEF93B" author="Brule" bookContentInfo="120 p." box="[623,898,1715,1739]" etAl="et al." firstAuthor="Brule" journalOrPublisher="Rapport de la Societe entomologique Antilles-Guyane, SEAG, ONF" pageId="10" pageNumber="9" refId="ref195373" refString="Brule, S., J. Touroult, and P. - H. Dalens (coord.). 2011 a. Resultats de l'inventaire entomologique du site de Saut Parare, reserve des Nouragues (Guyane), 2009 - 2010. Rapport de la Societe entomologique Antilles-Guyane, SEAG, ONF. 120 p. + annexes." title="Resultats de l'inventaire entomologique du site de Saut Parare, reserve des Nouragues (Guyane)" type="book" year="2011" yearSuffix="a">Brûlé et al. 2011a: 183</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFFC1FFE7FC05FDDD3579F93B" author="Brule" bookContentInfo="144 p." box="[909,1029,1715,1739]" firstAuthor="Brule" journalOrPublisher="SEAG, PAG" pageId="10" pageNumber="9" refId="ref195498" refString="Brule, S., J. Touroult, E. Poirier, and P. - H. Dalens. 2014. Resultats de l'etude-inventaire entomologique du site de la Montagne Pelee, Saul (Guyane), 2010 - 2012. Rapport de la Societe entomologique Antilles-Guyane. SEAG, PAG. 144 p. + annexes." title="Resultats de l'etude-inventaire entomologique du site de la Montagne Pelee, Saul (Guyane), 2010 - 2012. Rapport de la Societe entomologique Antilles-Guyane" type="book" year="2014">2014: 174</bibRefCitation>
; MHCN.
<emphasis id="B90A9218FFC1FFE7FBFDFDDD346AF93B" bold="true" box="[1141,1302,1715,1739]" pageId="10" pageNumber="9">
<collectingCountry id="F3690E9AFFC1FFE7FBFDFDDD35B5F93B" box="[1141,1225,1715,1739]" name="Brazil" pageId="10" pageNumber="9">Brazil</collectingCountry>
(NA):
</emphasis>
<bibRefCitation id="EFEF33FBFFC1FFE7FA94FDDD31A1F91B" author="Klug" firstAuthor="Klug" pageId="10" pageNumber="9" pagination="21 - 57" refId="ref210635" refString="Klug, J. C. F. 1845 (1843). Die Coleopteren-Gattungen Athyreus und Bolboceras, dargestellt nach den in der Sammlung hiesiger Konigl. Universitat davon vorhandenen Arten. Abhandlungen der Koniglichen Akademie der Wissenschaften 1843 (1845): 21 - 57." type="journal article" year="1845">Klug 1845: 28</bibRefCitation>
(as
<taxonomicName id="4C7E3589FFC1FFE7FE85FDBD3369F91B" authorityName="Klug" authorityYear="1845" box="[269,533,1747,1771]" class="Insecta" family="Bolboceratidae" genus="Athyreus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="lanuginosus">
<emphasis id="B90A9218FFC1FFE7FE85FDBD3369F91B" box="[269,533,1747,1771]" italics="true" pageId="10" pageNumber="9">Athyreus lanuginosus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFFC1FFE7FDA3FDBD3387F91B" author="Howden" box="[555,763,1747,1771]" firstAuthor="Howden" journalOrPublisher="Contributions of the American Entomological Institute" pageId="10" pageNumber="9" pagination="1 - 95" part="21" refId="ref208469" refString="Howden, H. F. 1985. A revision of the South American species in the genus Neoathyreus Howden and Martinez (Coleoptera, Scarabaeidae, Geotrupinae). Contributions of the American Entomological Institute 21 (4): 1 - 95." title="A revision of the South American species in the genus Neoathyreus Howden and Martinez (Coleoptera, Scarabaeidae, Geotrupinae)" type="journal article" year="1985">Howden 1985: 58</bibRefCitation>
(NA).
<emphasis id="B90A9218FFC1FFE7FCC2FDBD3559F91B" bold="true" box="[842,1061,1747,1771]" pageId="10" pageNumber="9">
<collectingCountry id="F3690E9AFFC1FFE7FCC2FDBD32A4F91B" box="[842,984,1747,1771]" name="Venezuela" pageId="10" pageNumber="9">Venezuela</collectingCountry>
(EO):
</emphasis>
<bibRefCitation id="EFEF33FBFFC1FFE7FBA2FDBD3587F91B" author="Howden" box="[1066,1275,1747,1771]" firstAuthor="Howden" journalOrPublisher="Contributions of the American Entomological Institute" pageId="10" pageNumber="9" pagination="1 - 95" part="21" refId="ref208469" refString="Howden, H. F. 1985. A revision of the South American species in the genus Neoathyreus Howden and Martinez (Coleoptera, Scarabaeidae, Geotrupinae). Contributions of the American Entomological Institute 21 (4): 1 - 95." title="A revision of the South American species in the genus Neoathyreus Howden and Martinez (Coleoptera, Scarabaeidae, Geotrupinae)" type="journal article" year="1985">Howden 1985: 58</bibRefCitation>
(EO).
<emphasis id="B90A9218FFC1FFE7FAC0FDBD34DCF91B" bold="true" box="[1352,1440,1747,1771]" pageId="10" pageNumber="9">Other:</emphasis>
<bibRefCitation id="EFEF33FBFFC1FFE7FF48FD9D3018F8FB" author="Klug" box="[192,356,1779,1803]" firstAuthor="Klug" pageId="10" pageNumber="9" pagination="21 - 57" refId="ref210635" refString="Klug, J. C. F. 1845 (1843). Die Coleopteren-Gattungen Athyreus und Bolboceras, dargestellt nach den in der Sammlung hiesiger Konigl. Universitat davon vorhandenen Arten. Abhandlungen der Koniglichen Akademie der Wissenschaften 1843 (1845): 21 - 57." type="journal article" year="1845">Klug 1845: 28</bibRefCitation>
(Colombia - as
<taxonomicName id="4C7E3589FFC1FFE7FD91FD9D3261F8FB" authorityName="Klug" authorityYear="1845" box="[537,797,1779,1803]" class="Insecta" family="Bolboceratidae" genus="Athyreus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="lanuginosus">
<emphasis id="B90A9218FFC1FFE7FD91FD9D3261F8FB" box="[537,797,1779,1803]" italics="true" pageId="10" pageNumber="9">Athyreus lanuginosus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFFC1FFE7FCBBFD9D357DF8FB" author="Howden" box="[819,1025,1779,1803]" firstAuthor="Howden" journalOrPublisher="Contributions of the American Entomological Institute" pageId="10" pageNumber="9" pagination="1 - 95" part="21" refId="ref208469" refString="Howden, H. F. 1985. A revision of the South American species in the genus Neoathyreus Howden and Martinez (Coleoptera, Scarabaeidae, Geotrupinae). Contributions of the American Entomological Institute 21 (4): 1 - 95." title="A revision of the South American species in the genus Neoathyreus Howden and Martinez (Coleoptera, Scarabaeidae, Geotrupinae)" type="journal article" year="1985">Howden 1985: 58</bibRefCitation>
(Argentina, Bolivia, Nov. Valencia [= Colombia] - designation
<typeStatus id="54C5F0A8FFC1FFE7FD88FC7D33DBF8DB" box="[512,679,1811,1835]" pageId="10" pageNumber="9" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="4C7E3589FFC1FFE7FD24FC7D32CEF8DB" authorityName="Klug" authorityYear="1845" box="[684,946,1811,1835]" class="Insecta" family="Bolboceratidae" genus="Athyreus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="lanuginosus">
<emphasis id="B90A9218FFC1FFE7FD24FC7D32CEF8DB" box="[684,946,1811,1835]" italics="true" pageId="10" pageNumber="9">Athyreus lanuginosus</emphasis>
</taxonomicName>
, Ecuador, Paraguay, Peru);
<bibRefCitation id="EFEF33FBFFC1FFE7FA8BFC7D31A1F8BB" author="Boilly" firstAuthor="Boilly" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="10" pageNumber="9" pagination="19 - 31" part="4" refId="ref194293" refString="Boilly, O. 2011 b. Les Bolboceratidae de Guyane (Coleoptera, Scarabaeoidea). Contribution a l'etude des coleopteres de Guyane 4: 19 - 31. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Les Bolboceratidae de Guyane (Coleoptera, Scarabaeoidea)" type="journal article" year="2011" yearSuffix="b">Boilly 2011b: 30</bibRefCitation>
(Argentina, Mexico).
</paragraph>
<paragraph id="8BC14E0AFFC0FFE6FF48FBB3332DFEC5" blockId="11.[192,1440,221,309]" pageId="11" pageNumber="10">
<emphasis id="B90A9218FFC0FFE6FF48FBB33074FF05" bold="true" box="[192,264,221,245]" pageId="11" pageNumber="10">Note.</emphasis>
In
<bibRefCitation id="EFEF33FBFFC0FFE6FEA5FBB0335EFF05" author="Boucomont" bookContentInfo="47 p." box="[301,546,221,245]" editor="S. Schenkling" firstAuthor="Boucomont" journalOrPublisher="Junk; Berlin" pageId="11" pageNumber="10" pagination="3 - 47" refId="ref194974" refString="Boucomont, A. 1912. Scarabaeidae: Taurocerastinae, Geotrupinae. p. 3 - 47. In: S. Schenkling. Coleopterorum Catalogus Pars 46. W. Junk; Berlin. 47 p." title="Scarabaeidae: Taurocerastinae, Geotrupinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 46. W" year="1912">Boucomont (1912: 5)</bibRefCitation>
and
<bibRefCitation id="EFEF33FBFFC0FFE6FDD2FBB33201FF05" author="Blackwelder" box="[602,893,221,245]" firstAuthor="Blackwelder" journalOrPublisher="United States National Museum Bulletin" pageId="11" pageNumber="10" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder (1944: 219)</bibRefCitation>
, the present species
<taxonomicName id="4C7E3589FFC0FFE6FBF8FBB334DCFF05" baseAuthorityName="Klug" baseAuthorityYear="1845" box="[1136,1440,221,245]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="lanuginosus">
<emphasis id="B90A9218FFC0FFE6FBF8FBB334DCFF05" box="[1136,1440,221,245]" italics="true" pageId="11" pageNumber="10">Neoathyreus lanuginosus</emphasis>
</taxonomicName>
is treated as a synonym of
<taxonomicName id="4C7E3589FFC0FFE6FD88FB933395FEE4" authorityName="Laporte" authorityYear="1840" box="[512,745,253,277]" class="Insecta" family="Bolboceratidae" genus="Athyreus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B90A9218FFC0FFE6FD88FB933395FEE4" box="[512,745,253,277]" italics="true" pageId="11" pageNumber="10">Athyreus excavatus</emphasis>
</taxonomicName>
(now
<taxonomicName id="4C7E3589FFC0FFE6FCB9FB933539FEE4" baseAuthorityName="Laporte" baseAuthorityYear="1840" box="[817,1093,253,277]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B90A9218FFC0FFE6FCB9FB933539FEE4" box="[817,1093,253,277]" italics="true" pageId="11" pageNumber="10">Neoathyreus excavatus</emphasis>
</taxonomicName>
). The records are mentioned in the treatment of
<taxonomicName id="4C7E3589FFC0FFE6FE27FA733336FEC4" baseAuthorityName="Laporte" baseAuthorityYear="1840" box="[431,586,285,308]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="B90A9218FFC0FFE6FE27FA733336FEC4" box="[431,586,285,308]" italics="true" pageId="11" pageNumber="10">N. excavatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>