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<taxonomicName id="4C7E3589FF7EFF58FF48F90833D5FD8E" authority="(Olivier, 1789)" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[192,681,614,638]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="species" species="rustica">
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<emphasis id="B90A9218FF7EFF58FF48F908309CFD8E" bold="true" box="[192,480,614,638]" italics="true" pageId="181" pageNumber="180">Cyclocephala rustica</emphasis>
(
<bibRefCitation id="EFEF33FBFF7EFF58FE78F90833DCFD8E" author="Olivier" box="[496,672,614,638]" firstAuthor="Olivier" pageId="181" pageNumber="180" refId="ref218493" refString="Olivier, G. A. 1789. Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Coleopteres. Tome premier. Baudouin; Paris. [Note. Each genus is separately paginated]." type="book" year="1789">Olivier, 1789</bibRefCitation>
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<taxonomicName id="4C7E3589FF7EFF58FF60F9E23313FD52" authority="Olivier 1789: 27" authorityName="Olivier" authorityPageNumber="27" authorityYear="1789" box="[232,623,652,675]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="species" species="rustica">
<emphasis id="B90A9218FF7EFF58FF60F9E230C5FD52" box="[232,441,652,674]" italics="true" pageId="181" pageNumber="180">Melolontha rustica</emphasis>
<bibRefCitation id="EFEF33FBFF7EFF58FE48F9E33313FD52" author="Olivier" box="[448,623,653,675]" firstAuthor="Olivier" pageId="181" pageNumber="180" refId="ref218493" refString="Olivier, G. A. 1789. Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Coleopteres. Tome premier. Baudouin; Paris. [Note. Each genus is separately paginated]." type="book" year="1789">Olivier 1789: 27</bibRefCitation>
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(
<collectingCountry id="F3690E9AFF7EFF58FDF5F9E3321BFD53" box="[637,871,653,675]" name="Trinidad and Tobago" pageId="181" pageNumber="180">Trinidad and Tobago</collectingCountry>
)
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<treatmentCitationGroup id="AB6E6924FF7EFF58FF60F9C4350AFD30" box="[232,1142,681,704]" pageId="181" pageNumber="180">
=
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<taxonomicName id="4C7E3589FF7EFF58FF75F9C73247FD4F" authority="Hohne 1923 b: 365" authorityName="Hohne" authorityPageNumber="365" authorityYear="1923" box="[253,827,681,703]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="municipalis">
<emphasis id="B90A9218FF7EFF58FF75F9C73311FD4E" box="[253,621,681,703]" italics="true" pageId="181" pageNumber="180">Cyclocephala rustica municipalis</emphasis>
<bibRefCitation id="EFEF33FBFF7EFF58FDFCF9C73247FD4F" author="Hohne, W." box="[628,827,681,703]" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="181" pageNumber="180" pagination="345 - 373" part="1923" refId="ref208153" refString="Hohne, W. 1923 b. Neue Cyclocephalen (Col. Dyn.). Deutsche Entomologische Zeitschrift 1923 (4): 345 - 373." title="Neue Cyclocephalen (Col. Dyn.)" type="journal article" year="1923">Höhne 1923b: 365</bibRefCitation>
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(
<collectingCountry id="F3690E9AFF7EFF58FCC1F9C732C0FD4F" box="[841,956,681,703]" name="Venezuela" pageId="181" pageNumber="180">Venezuela</collectingCountry>
-
<typeStatus id="54C5F0A8FF7EFF58FC59F9C43513FD30" box="[977,1135,682,704]" pageId="181" pageNumber="180" type="lectotype">LECTOTYPE</typeStatus>
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<emphasis id="B90A9218FF7EFF58FF48F9A1337BFD17" bold="true" box="[192,519,719,743]" pageId="181" pageNumber="180">
Distribution.
<collectingCountry id="F3690E9AFF7EFF58FEF1F9A1337DFD17" box="[377,513,719,743]" name="Suriname" pageId="181" pageNumber="180">Suriname</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF7EFF58FD85F9BE3231FD18" author="Uyttenboogaart" box="[525,845,720,744]" firstAuthor="Uyttenboogaart" pageId="181" pageNumber="180" pagination="112 - 121" refId="ref227872" refString="Uyttenboogaart, D. L. 1902. List of beetles collected in the months of August and September 1900 in Surinam (Dutch Guyana) and on the island of Barbadoes. Tijdschrift voor Entomologie 45 (1 - 2): 112 - 121." type="journal article" year="1902">Uyttenboogaart 1902: 116</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF7EFF58FCD0F9BE3508FD18" author="Moore" box="[856,1140,720,744]" etAl="et al." firstAuthor="Moore" pageId="181" pageNumber="180" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 242</bibRefCitation>
; MHCN; NZCS.
<emphasis id="B90A9218FF7EFF58FAB5F9BE3050FCF7" bold="true" pageId="181" pageNumber="180">
<collectingCountry id="F3690E9AFF7EFF58FAB5F9BE305AFCF7" name="French Guiana" pageId="181" pageNumber="180">French Guiana</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF7EFF58FEB9F99E3367FCF8" author="Endrodi, S." box="[305,539,752,776]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="181" pageNumber="180" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 299</bibRefCitation>
(as
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<emphasis id="B90A9218FF7EFF58FDC1F99E322FFCF8" box="[585,851,752,776]" italics="true" pageId="181" pageNumber="180">C. rustica municipalis</emphasis>
</taxonomicName>
; as
<taxonomicName id="4C7E3589FF7EFF58FC08F99E3531FCF7" authority="rustica" box="[896,1101,752,775]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="rustica">
<emphasis id="B90A9218FF7EFF58FC08F99E3531FCF7" box="[896,1101,752,775]" italics="true" pageId="181" pageNumber="180">C. rustica rustica</emphasis>
</taxonomicName>
(?)), 1985a: 35, 116 (Guyana [&gt; Fr. Guiana] - as
<taxonomicName id="4C7E3589FF7EFF58FE27F87E33BCFCD8" authorityName="Hohne" authorityYear="1923" box="[431,704,784,808]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="municipalis">
<emphasis id="B90A9218FF7EFF58FE27F87E33BCFCD8" box="[431,704,784,808]" italics="true" pageId="181" pageNumber="180">C. rustica municipalis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF7EFF58FD51F87E32BFFCD8" author="Duranton, M." box="[729,963,784,808]" journalOrPublisher="Coleopteres, numero special" pageId="181" pageNumber="180" pagination="1 - 31" part="1" refId="ref200092" refString="Duranton, M. 2011. Chorologie et ecologie des dynastides de Guyane francaise. Coleopteres, numero special 1: 1 - 31." title="Chorologie et ecologie des dynastides de Guyane francaise" type="journal article" year="2011">Duranton 2011: 15</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF7EFF58FC59F87E35D8FCD8" author="Ponchel" box="[977,1188,784,808]" firstAuthor="Ponchel" pageId="181" pageNumber="180" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel 2011: 60</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF7EFF58FB3AF87E34E7FCD8" author="Dupuis, F." box="[1202,1435,784,808]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="181" pageNumber="180" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis 2016b: 109</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF7EFF58FF48F85E30A4FCB8" author="Moore" box="[192,472,816,840]" etAl="et al." firstAuthor="Moore" pageId="181" pageNumber="180" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 242</bibRefCitation>
.
<emphasis id="B90A9218FF7EFF58FE69F841330FFCB7" bold="true" box="[481,627,815,839]" pageId="181" pageNumber="180">“Guianas”:</emphasis>
<bibRefCitation id="EFEF33FBFF7EFF58FDF0F85E3228FCB8" author="Hohne, W." box="[632,852,816,840]" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="181" pageNumber="180" pagination="345 - 373" part="1923" refId="ref208153" refString="Hohne, W. 1923 b. Neue Cyclocephalen (Col. Dyn.). Deutsche Entomologische Zeitschrift 1923 (4): 345 - 373." title="Neue Cyclocephalen (Col. Dyn.)" type="journal article" year="1923">Höhne 1923b: 365</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF7EFF58FCD5F85E3558FCB8" author="Arrow, G. J." bookContentInfo="124 p." box="[861,1060,816,840]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="181" pageNumber="180" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 15</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF7EFF58FBA7F85E3438FCB8" author="Blackwelder, R. E." box="[1071,1348,816,840]" journalOrPublisher="United States National Museum Bulletin" pageId="181" pageNumber="180" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 252</bibRefCitation>
.
<emphasis id="B90A9218FF7EFF58FAC4F841307AFC97" bold="true" pageId="181" pageNumber="180">
<collectingCountry id="F3690E9AFF7EFF58FAC4F84134DCFCB7" box="[1356,1440,815,839]" name="Brazil" pageId="181" pageNumber="180">Brazil</collectingCountry>
(NA):
</emphasis>
<bibRefCitation id="EFEF33FBFF7EFF58FE83F83E309FFC98" author="Hohne, W." box="[267,483,848,872]" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="181" pageNumber="180" pagination="345 - 373" part="1923" refId="ref208153" refString="Hohne, W. 1923 b. Neue Cyclocephalen (Col. Dyn.). Deutsche Entomologische Zeitschrift 1923 (4): 345 - 373." title="Neue Cyclocephalen (Col. Dyn.)" type="journal article" year="1923">Höhne 1923b: 365</bibRefCitation>
(as
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<emphasis id="B90A9218FF7EFF58FD99F83E3266FC98" box="[529,794,848,872]" italics="true" pageId="181" pageNumber="180">C. rustica municipalis</emphasis>
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);
<bibRefCitation id="EFEF33FBFF7EFF58FCA7F83E3565FC98" author="Endrodi, S." box="[815,1049,848,872]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="181" pageNumber="180" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 299</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF7EFF58FBCFF83E342CFC98" authorityName="Hohne" authorityYear="1923" box="[1095,1360,848,872]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="municipalis">
<emphasis id="B90A9218FF7EFF58FBCFF83E342CFC98" box="[1095,1360,848,872]" italics="true" pageId="181" pageNumber="180">C. rustica municipalis</emphasis>
</taxonomicName>
) (NA);
<bibRefCitation id="EFEF33FBFF7EFF58FF48F81E30DEFC78" author="Endrodi, S." bookContentInfo="800 p." box="[192,418,880,904]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="181" pageNumber="180" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">Endrödi 1985a: 35</bibRefCitation>
, 116 (as
<taxonomicName id="4C7E3589FF7EFF58FD84F81E3266FC78" authorityName="Hohne" authorityYear="1923" box="[524,794,880,904]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="municipalis">
<emphasis id="B90A9218FF7EFF58FD84F81E3266FC78" box="[524,794,880,904]" italics="true" pageId="181" pageNumber="180">C. rustica municipalis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF7EFF58FCB9F81E352CFC78" author="Moore" box="[817,1104,880,904]" etAl="et al." firstAuthor="Moore" pageId="181" pageNumber="180" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 242</bibRefCitation>
.
<emphasis id="B90A9218FF7EFF58FBD3F81E358CFC78" bold="true" box="[1115,1264,880,904]" pageId="181" pageNumber="180">
<collectingCountry id="F3690E9AFF7EFF58FBD3F81E3597FC78" box="[1115,1259,880,904]" name="Venezuela" pageId="181" pageNumber="180">Venezuela</collectingCountry>
:
</emphasis>
<bibRefCitation id="EFEF33FBFF7EFF58FB7FF81E3190FC58" author="Hohne, W." journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="181" pageNumber="180" pagination="345 - 373" part="1923" refId="ref208153" refString="Hohne, W. 1923 b. Neue Cyclocephalen (Col. Dyn.). Deutsche Entomologische Zeitschrift 1923 (4): 345 - 373." title="Neue Cyclocephalen (Col. Dyn.)" type="journal article" year="1923">Höhne 1923b: 365</bibRefCitation>
(as
<taxonomicName id="4C7E3589FF7EFF58FEABF8FE334BFC58" authorityName="Hohne" authorityYear="1923" box="[291,567,912,936]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="municipalis">
<emphasis id="B90A9218FF7EFF58FEABF8FE334BFC58" box="[291,567,912,936]" italics="true" pageId="181" pageNumber="180">C. rustica municipalis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF7EFF58FDD9F8FE3239FC58" author="Endrodi, S." box="[593,837,912,936]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="181" pageNumber="180" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 299</bibRefCitation>
(designation
<typeStatus id="54C5F0A8FF7EFF58FC78F8FE35E4FC58" box="[1008,1176,912,936]" pageId="181" pageNumber="180" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="4C7E3589FF7EFF58FB29F8FE302DFC38" authorityName="Hohne" authorityYear="1923" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="municipalis">
<emphasis id="B90A9218FF7EFF58FB29F8FE302DFC38" italics="true" pageId="181" pageNumber="180">Cyclocephala rustica municipalis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF7EFF58FEEFF8DE333AFC38" author="Endrodi, S." bookContentInfo="800 p." box="[359,582,944,968]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="181" pageNumber="180" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">Endrödi 1985a: 35</bibRefCitation>
, 116 (as
<taxonomicName id="4C7E3589FF7EFF58FD23F8DE32CAFC38" authorityName="Hohne" authorityYear="1923" box="[683,950,944,968]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="municipalis">
<emphasis id="B90A9218FF7EFF58FD23F8DE32CAFC38" box="[683,950,944,968]" italics="true" pageId="181" pageNumber="180">C. rustica municipalis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF7EFF58FC44F8DE3599FC38" author="Moore" box="[972,1253,944,968]" etAl="et al." firstAuthor="Moore" pageId="181" pageNumber="180" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 242</bibRefCitation>
.
<emphasis id="B90A9218FF7EFF58FB66F8C1343AFC37" bold="true" box="[1262,1350,943,967]" pageId="181" pageNumber="180">Other:</emphasis>
<bibRefCitation id="EFEF33FBFF7EFF58FAC3F8DE305BFC18" author="Olivier" firstAuthor="Olivier" pageId="181" pageNumber="180" refId="ref218493" refString="Olivier, G. A. 1789. Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Coleopteres. Tome premier. Baudouin; Paris. [Note. Each genus is separately paginated]." type="book" year="1789">Olivier 1789: 27</bibRefCitation>
(Guadeloupe);
<bibRefCitation id="EFEF33FBFF7EFF58FE69F8BE33C2FC18" author="Hohne, W." box="[481,702,976,1000]" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="181" pageNumber="180" pagination="345 - 373" part="1923" refId="ref208153" refString="Hohne, W. 1923 b. Neue Cyclocephalen (Col. Dyn.). Deutsche Entomologische Zeitschrift 1923 (4): 345 - 373." title="Neue Cyclocephalen (Col. Dyn.)" type="journal article" year="1923">Höhne 1923b: 365</bibRefCitation>
(Colombia, Mexico - as
<taxonomicName id="4C7E3589FF7EFF58FC69F8BE358CFC18" authorityName="Hohne" authorityYear="1923" box="[993,1264,976,1000]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="municipalis">
<emphasis id="B90A9218FF7EFF58FC69F8BE358CFC18" box="[993,1264,976,1000]" italics="true" pageId="181" pageNumber="180">C. rustica municipalis</emphasis>
</taxonomicName>
; Trinidad and Tobago - as
<taxonomicName id="4C7E3589FF7EFF58FED0F89E3353FBF7" authority="rustica" box="[344,559,1008,1031]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="rustica">
<emphasis id="B90A9218FF7EFF58FED0F89E3353FBF7" box="[344,559,1008,1031]" italics="true" pageId="181" pageNumber="180">C. rustica rustica</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF7EFF58FDC0F89E321DFBF8" author="Blackwelder, R. E." box="[584,865,1008,1032]" journalOrPublisher="United States National Museum Bulletin" pageId="181" pageNumber="180" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 252</bibRefCitation>
(Guadeloupe);
<bibRefCitation id="EFEF33FBFF7EFF58FBA8F89E346FFBF8" author="Endrodi, S." box="[1056,1299,1008,1032]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="181" pageNumber="180" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 299</bibRefCitation>
(Colombia, Mexico, Peru - as
<taxonomicName id="4C7E3589FF7EFF58FE17FF7E33CDFBD8" authorityName="Hohne" authorityYear="1923" box="[415,689,1040,1064]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="municipalis">
<emphasis id="B90A9218FF7EFF58FE17FF7E33CDFBD8" box="[415,689,1040,1064]" italics="true" pageId="181" pageNumber="180">C. rustica municipalis</emphasis>
</taxonomicName>
; Guadeloupe, Trinidad and Tobago - designation
<typeStatus id="54C5F0A8FF7EFF58FA91FF7E34DCFBD8" box="[1305,1440,1040,1064]" pageId="181" pageNumber="180" type="neotype">NEOTYPE</typeStatus>
<taxonomicName id="4C7E3589FF7EFF58FF48FF5E30DBFBB7" authorityName="Olivier" authorityYear="1789" box="[192,423,1072,1096]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="species" species="rustica">
<emphasis id="B90A9218FF7EFF58FF48FF5E30DBFBB7" box="[192,423,1072,1096]" italics="true" pageId="181" pageNumber="180">Melolontha rustica</emphasis>
</taxonomicName>
), 1985a: 35, 116 (Colombia, Mexico, Peru - as
<taxonomicName id="4C7E3589FF7EFF58FC6DFF5E358AFBB8" box="[997,1270,1072,1096]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="municipalis">
<emphasis id="B90A9218FF7EFF58FC6DFF5E358AFBB8" box="[997,1270,1072,1096]" italics="true" pageId="181" pageNumber="180">C. rustica municipalis</emphasis>
</taxonomicName>
; Guadeloupe, Trinidad and Tobago - as
<taxonomicName id="4C7E3589FF7EFF58FD8DFF3E33A0FB97" authority="rustica" box="[517,732,1104,1127]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="rustica">
<emphasis id="B90A9218FF7EFF58FD8DFF3E33A0FB97" box="[517,732,1104,1127]" italics="true" pageId="181" pageNumber="180">C. rustica rustica</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFEF33FBFF7EFF58FD7DFF3E3566FB98" author="Moore" box="[757,1050,1104,1128]" etAl="et al." firstAuthor="Moore" pageId="181" pageNumber="180" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 242</bibRefCitation>
(Colombia, Peru, Trinidad and Tobago).
</paragraph>
<paragraph id="8BC14E0AFF7EFF58FF48FFF6343BFB20" blockId="181.[192,1440,1176,1232]" pageId="181" pageNumber="180">
<emphasis id="B90A9218FF7EFF58FF48FFF63061FB40" bold="true" box="[192,285,1176,1200]" pageId="181" pageNumber="180">Note 1.</emphasis>
<bibRefCitation id="EFEF33FBFF7EFF58FEABFFF63330FB40" author="Moore" box="[291,588,1176,1200]" etAl="et al." firstAuthor="Moore" pageId="181" pageNumber="180" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. (2018b: 107)</bibRefCitation>
conclude that the
<typeStatus id="54C5F0A8FF7EFF58FCA0FFF632F4FB40" box="[808,904,1176,1200]" pageId="181" pageNumber="180" type="neotype">neotype</typeStatus>
of
<taxonomicName id="4C7E3589FF7EFF58FC21FFF635F2FB5F" authorityName="Olivier" authorityYear="1789" box="[937,1166,1176,1200]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="species" species="rustica">
<emphasis id="B90A9218FF7EFF58FC21FFF635F2FB5F" box="[937,1166,1176,1200]" italics="true" pageId="181" pageNumber="180">Melolontha rustica</emphasis>
</taxonomicName>
designated in
<bibRefCitation id="EFEF33FBFF7EFF58FAB5FFF6302AFB20" author="Endrodi, S." journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="181" pageNumber="180" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi (1966a: 299)</bibRefCitation>
is invalid because it violates several demands of Article 75 of the
<bibRefCitation id="EFEF33FBFF7EFF58FBF4FFD6343FFB20" author="ICZN" box="[1148,1347,1208,1232]" firstAuthor="ICZN" pageId="181" pageNumber="180" refId="ref209183" refString="ICZN [International Commission on Zoological Nomenclature]. 1964. International code of zoological nomenclature adopted by the XV International Congress of Zoology. Second edition. International Trust for Zoological Nomenclature; London. 176 p." type="book" year="1964">ICZN (1964: 81)</bibRefCitation>
.
</paragraph>
<paragraph id="8BC14E0AFF7EFF58FF48FF8E3248FAA8" blockId="181.[192,1440,1248,1368]" pageId="181" pageNumber="180">
<emphasis id="B90A9218FF7EFF58FF48FF8E3062FB08" bold="true" box="[192,286,1248,1272]" pageId="181" pageNumber="180">Note 2.</emphasis>
<bibRefCitation id="EFEF33FBFF7EFF58FEADFF8E3084FB08" author="Olivier" box="[293,504,1248,1272]" firstAuthor="Olivier" pageId="181" pageNumber="180" refId="ref218493" refString="Olivier, G. A. 1789. Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Coleopteres. Tome premier. Baudouin; Paris. [Note. Each genus is separately paginated]." type="book" year="1789">Olivier (1789: 27)</bibRefCitation>
describes
<taxonomicName id="4C7E3589FF7EFF58FDFFFF8E320FFB07" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[631,883,1248,1272]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="species" species="rustica">
<emphasis id="B90A9218FF7EFF58FDFFFF8E320FFB07" box="[631,883,1248,1272]" italics="true" pageId="181" pageNumber="180">Cyclocephala rustica</emphasis>
</taxonomicName>
from
<collectingCountry id="F3690E9AFF7EFF58FC32FF8E3533FB08" box="[954,1103,1248,1272]" name="Guadeloupe" pageId="181" pageNumber="180">Guadeloupe</collectingCountry>
, and this record is copied in
<bibRefCitation id="EFEF33FBFF7EFF58FF48FE6E309BFAE8" author="Blackwelder, R. E." box="[192,487,1280,1304]" journalOrPublisher="United States National Museum Bulletin" pageId="181" pageNumber="180" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder (1944: 252)</bibRefCitation>
and
<bibRefCitation id="EFEF33FBFF7EFF58FDABFE6E3262FAE8" author="Endrodi, S." box="[547,798,1280,1304]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="181" pageNumber="180" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi (1966a: 299</bibRefCitation>
,
<bibRefCitation id="EFEF33FBFF7EFF58FCA1FE6E32DEFAE8" author="Endrodi, S." bookContentInfo="800 p." box="[809,930,1280,1304]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="181" pageNumber="180" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">1985a: 35</bibRefCitation>
, 116). However,
<bibRefCitation id="EFEF33FBFF7EFF58FBE4FE6E3184FAC8" author="Ratcliffe and Cave" firstAuthor="Ratcliffe" pageId="181" pageNumber="180" pagination="1 - 343" refId="ref222299" refString="Ratcliffe, B. C., and R. D. Cave. 2015. The dynastine scarab beetles of the West Indies (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 28: 1 - 343." type="journal article" year="2015">Ratcliffe and Cave (2015: 291)</bibRefCitation>
, addressing just
<bibRefCitation id="EFEF33FBFF7EFF58FE4EFE4E33F3FAC8" author="Endrodi, S." box="[454,655,1312,1336]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="181" pageNumber="180" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi (1966a)</bibRefCitation>
, claim this record is incorrect, as
<taxonomicName id="4C7E3589FF7EFF58FBA1FE4E35DCFAC7" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[1065,1184,1312,1335]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="species" species="rustica">
<emphasis id="B90A9218FF7EFF58FBA1FE4E35DCFAC7" box="[1065,1184,1312,1335]" italics="true" pageId="181" pageNumber="180">C. rustica</emphasis>
</taxonomicName>
is a South American species. Oliviers
<typeStatus id="54C5F0A8FF7EFF58FE1BFE2E30B4FAA8" box="[403,456,1344,1368]" pageId="181" pageNumber="180">type</typeStatus>
specimen may be mislabeled.
</paragraph>
<paragraph id="8BC14E0AFF7EFF58FF48FE0634DCF9F0" blockId="181.[192,1440,1384,1536]" pageId="181" pageNumber="180">
<emphasis id="B90A9218FF7EFF58FF48FE063067FA70" bold="true" box="[192,283,1384,1408]" pageId="181" pageNumber="180">Note 3.</emphasis>
In
<bibRefCitation id="EFEF33FBFF7EFF58FEC8FE063346FA70" author="Endrodi, S." box="[320,570,1384,1408]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="181" pageNumber="180" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi (1966a: 299)</bibRefCitation>
, both
<taxonomicName id="4C7E3589FF7EFF58FDF6FE0632B7FA8F" authority="rustica" box="[638,971,1384,1408]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="rustica">
<emphasis id="B90A9218FF7EFF58FDF6FE0632B7FA8F" box="[638,971,1384,1408]" italics="true" pageId="181" pageNumber="180">Cyclocephala rustica rustica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7E3589FF7EFF58FB8BFE063474FA70" authorityName="Hohne" authorityYear="1923" box="[1027,1288,1384,1408]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="municipalis">
<emphasis id="B90A9218FF7EFF58FB8BFE063474FA70" box="[1027,1288,1384,1408]" italics="true" pageId="181" pageNumber="180">C. rustica municipalis</emphasis>
</taxonomicName>
are reported from “Guayana: Cayenne,” which is
<collectingCountry id="F3690E9AFF7EFF58FD0CFEE6323FFA50" box="[644,835,1416,1440]" name="French Guiana" pageId="181" pageNumber="180">French Guiana</collectingCountry>
. In
<bibRefCitation id="EFEF33FBFF7EFF58FCFCFEE63517FA50" author="Endrodi, S." bookContentInfo="800 p." box="[884,1131,1416,1440]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="181" pageNumber="180" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">Endrödi (1985a: 35)</bibRefCitation>
<collectingCountry id="F3690E9AFF7EFF58FBFCFEE635A4FA50" box="[1140,1240,1416,1440]" name="Guyana" pageId="181" pageNumber="180">Guyana</collectingCountry>
, but not
<collectingCountry id="F3690E9AFF7EFF58FAC1FEE63062FA30" name="French Guiana" pageId="181" pageNumber="180">French Guiana</collectingCountry>
, is mentioned for
<taxonomicName id="4C7E3589FF7EFF58FE74FEC63271FA30" authorityName="Hohne" authorityYear="1923" box="[508,781,1448,1472]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="municipalis">
<emphasis id="B90A9218FF7EFF58FE74FEC63271FA30" box="[508,781,1448,1472]" italics="true" pageId="181" pageNumber="180">C. rustica municipalis</emphasis>
</taxonomicName>
(with
<taxonomicName id="4C7E3589FF7EFF58FCD7FEC63548FA4F" authority="rustica" box="[863,1076,1448,1471]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="180" phylum="Arthropoda" rank="subSpecies" species="rustica" subSpecies="rustica">
<emphasis id="B90A9218FF7EFF58FCD7FEC63548FA4F" box="[863,1076,1448,1471]" italics="true" pageId="181" pageNumber="180">C. rustica rustica</emphasis>
</taxonomicName>
only being mentioned for the Antilles). As all more specific locations were omitted in the 1985 publication, we assume the remaining “Guayana” erroneously became
<collectingCountry id="F3690E9AFF7EFF58FDCAFE8633DAF9F0" box="[578,678,1512,1536]" name="Guyana" pageId="181" pageNumber="180">Guyana</collectingCountry>
. We are unaware of actual records of this species from
<collectingCountry id="F3690E9AFF7EFF58FAB0FE8634E0F9F0" box="[1336,1436,1512,1536]" name="Guyana" pageId="181" pageNumber="180">Guyana</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>