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Distribution.
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<bibRefCitation id="EFEF33FBFF60FF46FBD7FFD3342EFB25" author="Endrodi, S." box="[1119,1362,1213,1237]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="171" pageNumber="170" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 209</bibRefCitation>
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H. W.
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<bibRefCitation id="EFEF33FBFF60FF46FD7BFE7332D6FAC5" author="Maes" box="[755,938,1309,1333]" firstAuthor="Maes" pageId="171" pageNumber="170" pagination="1 - 44" refId="ref213604" refString="Maes, J. - M. 1994. Los Dynastinae (Coleoptera: Scarabaeidae) de Nicaragua. Revista Nicaraguense de Entomologia 30: 1 - 44." type="journal article" year="1994">Maes 1994: 11</bibRefCitation>
;
<bibRefCitation id="EFEF33FBFF60FF46FC32FE73359AFAC5" author="Moore" box="[954,1254,1309,1333]" etAl="et al." firstAuthor="Moore" pageId="171" pageNumber="170" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 181</bibRefCitation>
.
<emphasis id="B90A9218FF60FF46FB7DFE723431FAC4" bold="true" box="[1269,1357,1308,1332]" pageId="171" pageNumber="170">Other:</emphasis>
H. W.
<bibRefCitation id="EFEF33FBFF60FF46FF48FE5330F5FAA5" author="Bates, H. W." bookContentInfo="432 p." box="[192,393,1341,1365]" journalOrPublisher="Frederick Ducane Godman and Osbert Salvin; London" pageId="171" pageNumber="170" pagination="161 - 336" refId="ref193515" refString="Bates, H. W. 1888. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. p. 161 - 336. In: Biologia Centrali-Americana. Frederick Ducane Godman and Osbert Salvin; London. 432 p." title="Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia" type="book chapter" volumeTitle="Biologia Centrali-Americana" year="1888">Bates 1888: 308</bibRefCitation>
(Guatemala, Mexico, Nicaragua, Panama);
<bibRefCitation id="EFEF33FBFF60FF46FC3BFE5335FFFAA5" author="Arrow, G. J." bookContentInfo="124 p." box="[947,1155,1341,1365]" editor="S. Schenkling" journalOrPublisher="Junk; The Hague, Netherlands" pageId="171" pageNumber="170" pagination="3 - 124" refId="ref192146" refString="Arrow, G. J. 1937 b. Scarabaeidae: Dynastinae. p. 3 - 124. In: S. Schenkling. Coleopterorum Catalogus Pars 156. W. Junk; The Hague, Netherlands. 124 p." title="Scarabaeidae: Dynastinae" type="book chapter" volumeTitle="Coleopterorum Catalogus Pars 156. W" year="1937">Arrow 1937b: 11</bibRefCitation>
(Mexico);
<bibRefCitation id="EFEF33FBFF60FF46FA80FE533045FA85" author="Blackwelder, R. E." journalOrPublisher="United States National Museum Bulletin" pageId="171" pageNumber="170" pagination="189 - 341" part="185" refId="ref193872" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341." title="Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2" type="journal article" year="1944">Blackwelder 1944: 251</bibRefCitation>
(Guatemala, Mexico, Nicaragua, Panama);
<bibRefCitation id="EFEF33FBFF60FF46FCEAFE333524FA85" author="Endrodi, S." box="[866,1112,1373,1397]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="171" pageNumber="170" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi 1966a: 209</bibRefCitation>
(Bolivia, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama), 1985a: 88, 125 (BoliviaMexico);
<bibRefCitation id="EFEF33FBFF60FF46FF48FEF3300DFA45" author="Maes" box="[192,369,1437,1461]" firstAuthor="Maes" pageId="171" pageNumber="170" pagination="1 - 44" refId="ref213604" refString="Maes, J. - M. 1994. Los Dynastinae (Coleoptera: Scarabaeidae) de Nicaragua. Revista Nicaraguense de Entomologia 30: 1 - 44." type="journal article" year="1994">Maes 1994: 11</bibRefCitation>
(Bolivia, Colombia, Ecuador, Guatemala, Mexico, Nicaragua, Panama);
<bibRefCitation id="EFEF33FBFF60FF46FB66FEF33190FA25" author="Ratcliffe" firstAuthor="Ratcliffe" pageId="171" pageNumber="170" pagination="1 - 506" refId="ref221631" refString="Ratcliffe, B. C. 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." type="journal article" year="2003">Ratcliffe 2003: 132</bibRefCitation>
(Brazil/BoliviaMexico);
<bibRefCitation id="EFEF33FBFF60FF46FD97FED333BCFA25" author="Ponchel" box="[543,704,1469,1493]" firstAuthor="Ponchel" pageId="171" pageNumber="170" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel 2011</bibRefCitation>
(not mentioned);
<bibRefCitation id="EFEF33FBFF60FF46FC1DFED33541FA25" author="Dupuis, F." box="[917,1085,1469,1493]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="171" pageNumber="170" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis 2016b</bibRefCitation>
(not mentioned);
<bibRefCitation id="EFEF33FBFF60FF46FA99FED3303EFA05" author="Moore" etAl="et al." firstAuthor="Moore" pageId="171" pageNumber="170" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. 2018b: 181</bibRefCitation>
(Belize, Bolivia, Colombia, Costa Rica, Ecuador, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Paraguay).
</paragraph>
<paragraph id="8BC14E0AFF60FF46FF48FD4B3235F96C" blockId="171.[192,1440,1572,1693]" pageId="171" pageNumber="170">
<emphasis id="B90A9218FF60FF46FF48FD4B3061F9CC" bold="true" box="[192,285,1572,1597]" pageId="171" pageNumber="170">Note 1.</emphasis>
<bibRefCitation id="EFEF33FBFF60FF46FEABFD4B336DF9CD" author="Endrodi, S." box="[291,529,1573,1597]" journalOrPublisher="Folia Entomologica Hungarica (N. S.)" pageId="171" pageNumber="170" pagination="433 - 470" part="17" refId="ref200586" refString="Endrodi, S. 1964. Eine Reihe von neuen Cyclocephala - Arten (Col., Melolonthidae, Dynastinae). Folia Entomologica Hungarica (N. S.) 17 (31): 433 - 470." title="Eine Reihe von neuen Cyclocephala - Arten (Col., Melolonthidae, Dynastinae)" type="journal article" year="1964">Endrödi (1964: 457)</bibRefCitation>
describes
<taxonomicName id="4C7E3589FF60FF46FD05FD4A32C3F9CC" authorityName="Endrodi" authorityYear="1964" box="[653,959,1572,1596]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="171" pageNumber="170" phylum="Arthropoda" rank="species" species="meinanderi">
<emphasis id="B90A9218FF60FF46FD05FD4A32C3F9CC" box="[653,959,1572,1596]" italics="true" pageId="171" pageNumber="170">Cyclocephala meinanderi</emphasis>
</taxonomicName>
. The species is reduced to an aberration of
<taxonomicName id="4C7E3589FF60FF46FF56FD2B3314F9AD" authority="in Endrodi (1967 b: 90)" authorityName="in Endrodi" authorityPageNumber="90" authorityYear="1967" box="[222,616,1605,1629]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="171" pageNumber="170" phylum="Arthropoda" rank="species" species="gravis">
<emphasis id="B90A9218FF60FF46FF56FD2B3030F9AC" box="[222,332,1605,1628]" italics="true" pageId="171" pageNumber="170">C. gravis</emphasis>
in
<bibRefCitation id="EFEF33FBFF60FF46FEFBFD2B3314F9AD" author="Endrodi, S." box="[371,616,1605,1629]" journalOrPublisher="Acta Zoologica Academiae Scientiarum Hungaricae" pageId="171" pageNumber="170" pagination="83 - 91" part="13" refId="ref200735" refString="Endrodi, S. 1967 b. Erganzungen zu meiner Monographie der Dynastinae: Cyclocephalini (Coleoptera). Acta Zoologica Academiae Scientiarum Hungaricae 13 (1 - 2): 83 - 91." title="Erganzungen zu meiner Monographie der Dynastinae: Cyclocephalini (Coleoptera)" type="journal article" year="1967">Endrödi (1967b: 90)</bibRefCitation>
</taxonomicName>
, and this is upheld in
<bibRefCitation id="EFEF33FBFF60FF46FCF0FD2B3511F9AD" author="Endrodi, S." bookContentInfo="800 p." box="[888,1133,1605,1629]" journalOrPublisher="Junk; The Hague, Netherlands and Akademiai Kiado; Budapest" pageId="171" pageNumber="170" refId="ref201383" refString="Endrodi, S. 1985 a. The Dynastinae of the world. Series Entomologica 28. Dr W. Junk; The Hague, Netherlands and Akademiai Kiado; Budapest. 800 p." title="The Dynastinae of the world. Series Entomologica 28. Dr W" type="book" year="1985">Endrödi (1985a: 88)</bibRefCitation>
.
<bibRefCitation id="EFEF33FBFF60FF46FBF1FD2B34E3F9AD" author="Dechambre, R. - P." box="[1145,1439,1605,1629]" journalOrPublisher="Revue francaise d'Entomologie (N. S.)" pageId="171" pageNumber="170" pagination="160 - 168" part="1" refId="ref198072" refString="Dechambre, R. - P. 1979 c. Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]. Revue francaise d'Entomologie (N. S.) 1 (4): 160 - 168." title="Missions entomologiques en Guyane et au Bresil [Coleoptera Dynastidae]" type="journal article" year="1979">Dechambre (1979c: 164)</bibRefCitation>
reinstates it as a valid species and explains this decision. Ratcliffe (B. Ratcliffe, pers. comm.) and
<bibRefCitation id="EFEF33FBFF60FF46FADBFD0B30E7F96D" author="Moore" etAl="et al." firstAuthor="Moore" pageId="171" pageNumber="170" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. (2018b: 181)</bibRefCitation>
still consider it a form of
<taxonomicName id="4C7E3589FF60FF46FD5DFDEB323EF96C" authorityName="in Endrodi" authorityYear="1967" box="[725,834,1669,1692]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="171" pageNumber="170" phylum="Arthropoda" rank="species" species="gravis">
<emphasis id="B90A9218FF60FF46FD5DFDEB323EF96C" box="[725,834,1669,1692]" italics="true" pageId="171" pageNumber="170">C. gravis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC14E0AFF60FF41FF48FDC3355BFEE1" blockId="171.[192,1440,1708,1893]" lastBlockId="172.[192,1440,217,273]" lastPageId="172" lastPageNumber="171" pageId="171" pageNumber="170">
<emphasis id="B90A9218FF60FF46FF48FDC33062F934" bold="true" box="[192,286,1708,1733]" pageId="171" pageNumber="170">Note 2.</emphasis>
As far as we know,
<taxonomicName id="4C7E3589FF60FF46FD98FDC2327FF934" authorityName="in Endrodi" authorityYear="1967" box="[528,771,1708,1732]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="171" pageNumber="170" phylum="Arthropoda" rank="species" species="gravis">
<emphasis id="B90A9218FF60FF46FD98FDC2327FF934" box="[528,771,1708,1732]" italics="true" pageId="171" pageNumber="170">Cyclocephala gravis</emphasis>
</taxonomicName>
is only reported for the research area by
<bibRefCitation id="EFEF33FBFF60FF46FB7EFDC33189F915" author="Bodkin, G. E." journalOrPublisher="The Entomologist's Monthly Magazine" pageId="171" pageNumber="170" pagination="210 - 219" part="55" refId="ref194211" refString="Bodkin, G. E. 1919. Notes on the Coleoptera of British Guiana. The Entomologist's Monthly Magazine 55: 210 - 219." title="Notes on the Coleoptera of British Guiana" type="journal article" year="1919">Bodkin (1919: 215)</bibRefCitation>
who records it for
<collectingCountry id="F3690E9AFF60FF46FE5DFDA33345F915" box="[469,569,1741,1765]" name="Guyana" pageId="171" pageNumber="170">Guyana</collectingCountry>
, by
<bibRefCitation id="EFEF33FBFF60FF46FDEEFDA33219F915" author="Endrodi, S." box="[614,869,1741,1765]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="171" pageNumber="170" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi (1966a: 209)</bibRefCitation>
who records it for
<collectingCountry id="F3690E9AFF60FF46FBCDFDA335DAF915" box="[1093,1190,1741,1765]" name="Guyana" pageId="171" pageNumber="170">Guyana</collectingCountry>
and
<collectingCountry id="F3690E9AFF60FF46FB69FDA334E0F915" box="[1249,1436,1741,1765]" name="French Guiana" pageId="171" pageNumber="170">French Guiana</collectingCountry>
, and by
<bibRefCitation id="EFEF33FBFF60FF46FE90FD8330ABF8F5" author="Maes" box="[280,471,1773,1797]" firstAuthor="Maes" pageId="171" pageNumber="170" pagination="1 - 44" refId="ref213604" refString="Maes, J. - M. 1994. Los Dynastinae (Coleoptera: Scarabaeidae) de Nicaragua. Revista Nicaraguense de Entomologia 30: 1 - 44." type="journal article" year="1994">Maes (1994: 11)</bibRefCitation>
who appears to have copied a record for
<collectingCountry id="F3690E9AFF60FF46FC48FD833545F8F5" box="[960,1081,1773,1797]" name="Suriname" pageId="171" pageNumber="170">Suriname</collectingCountry>
from an unknown source and mentions
<collectingCountry id="F3690E9AFF60FF46FEB1FC6330E6F8D5" box="[313,410,1805,1829]" name="Guyana" pageId="171" pageNumber="170">Guyana</collectingCountry>
and
<collectingCountry id="F3690E9AFF60FF46FE5FFC6333ECF8D5" box="[471,656,1805,1829]" name="French Guiana" pageId="171" pageNumber="170">French Guiana</collectingCountry>
based on
<bibRefCitation id="EFEF33FBFF60FF46FC82FC6332ACF8D5" author="Endrodi, S." box="[778,976,1805,1829]" journalOrPublisher="Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden" pageId="171" pageNumber="170" pagination="1 - 457" part="33" refId="ref200626" refString="Endrodi, S. 1966 a. Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil. Entomologische Abhandlungen Staatliches Museum fur Tierkunde in Dresden 33: 1 - 457." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). 1. Teil" type="journal article" year="1966">Endrödi (1966a)</bibRefCitation>
(and is in turn quoted by
<bibRefCitation id="EFEF33FBFF60FF46FA87FC633019F8B5" author="Moore" etAl="et al." firstAuthor="Moore" pageId="171" pageNumber="170" pagination="101 - 378" refId="ref215803" refString="Moore, M. R., R. D. Cave, and M. A. Branham. 2018 b. Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini). ZooKeys 745: 101 - 378." type="journal article" year="2018" yearSuffix="b">Moore et al. (2018b: 181))</bibRefCitation>
.
<bibRefCitation id="EFEF33FBFF60FF46FEF9FC433310F8B5" author="Duranton, M." box="[369,620,1837,1861]" journalOrPublisher="Coleopteres, numero special" pageId="171" pageNumber="170" pagination="1 - 31" part="1" refId="ref200092" refString="Duranton, M. 2011. Chorologie et ecologie des dynastides de Guyane francaise. Coleopteres, numero special 1: 1 - 31." title="Chorologie et ecologie des dynastides de Guyane francaise" type="journal article" year="2011">Duranton (2011: 13)</bibRefCitation>
notes he saw no definite specimens from
<collectingCountry id="F3690E9AFF60FF46FBFEFC433448F8B5" box="[1142,1332,1837,1861]" name="French Guiana" pageId="171" pageNumber="170">French Guiana</collectingCountry>
, and the species is not mentioned in the checklists for
<collectingCountry id="F3690E9AFF60FF46FD69FC2332E5F895" box="[737,921,1869,1893]" name="French Guiana" pageId="171" pageNumber="170">French Guiana</collectingCountry>
by
<bibRefCitation id="EFEF33FBFF60FF46FC4AFC23350AF895" author="Ponchel" box="[962,1142,1869,1893]" firstAuthor="Ponchel" pageId="171" pageNumber="170" pagination="60 - 61" refId="ref220051" refString="Ponchel, Y. 2011. Liste actualisee des Dynastinae de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 4: 60 - 61. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." type="journal article" year="2011">Ponchel (2011)</bibRefCitation>
and
<bibRefCitation id="EFEF33FBFF60FF46FB39FC233413F895" author="Dupuis, F." box="[1201,1391,1869,1893]" journalOrPublisher="Contribution a l'etude des coleopteres de Guyane" pageId="171" pageNumber="170" pagination="101 - 111" part="10" refId="ref199700" refString="Dupuis, F. 2016 b. Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes. Contribution a l'etude des coleopteres de Guyane 10: 101 - 111. [Supplement au Bulletin de liaison d'ACOREP-France &quot; Le Coleopteriste &quot;]." title="Contribution a la connaissance des Dynastidae de Guyane. Modifications nomen- claturales, signalements nouveaux et actualisation de la liste des especes" type="journal article" year="2016">Dupuis (2016b)</bibRefCitation>
. We have not seen any specimens from
<collectingCountry id="F3690E9AFF67FF41FDECFBB7339CFF01" box="[612,736,217,241]" name="Suriname" pageId="172" pageNumber="171">Suriname</collectingCountry>
, and because of the absence of any recent records from the research area, we consider its occurrence there in need of confirmation.
</paragraph>
</subSubSection>
</treatment>
</document>