381 lines
46 KiB
XML
381 lines
46 KiB
XML
<document id="D9515546DAA8E9C13CD475EC40E8E3A2" ID-DOI="10.11646/zootaxa.3946.3.3" ID-GBIF-Dataset="fcec1ba5-a256-4419-a228-ca2b529948bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="245390" ID-ZooBank="8D7575DB-636C-4473-91C3-A58C1E5F9E73" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461034695781" checkinUser="plazi" docAuthor="Zakrzewska, Marta & Giłka, Wojciech" docDate="2015" docId="03CD3E7A711EA71EFF60AFD0D3E9F655" docLanguage="en" docName="zt03946p360.pdf" docOrigin="Zootaxa 3946 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tanytarsus protogregarius Gilka et Zakrzewska, sp. nov." docType="treatment" docVersion="11" lastPageNumber="359" masterDocId="FFF446027117A712FFF7AC04D275F355" masterDocTitle="The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk" masterLastPageNumber="360" masterPageNumber="347" pageNumber="356" updateTime="1698574398835" updateUser="plazi">
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<mods:titleInfo id="88DDFE732E77726D98954B0CF3D7E8A8">
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<mods:title id="48F8508934E4F1238ABD21768F6CEFBB">The Tanytarsini (Diptera: Chironomidae) in the collection of the Museum of Amber Inclusions, University of Gdańsk</mods:title>
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</mods:titleInfo>
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<mods:name id="ACEE51D8E3A82D94D9051CA20F53E20D" type="personal">
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<mods:roleTerm id="29CC2E9449C37DAABEFE99B75092EA50">Author</mods:roleTerm>
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<mods:namePart id="FB044922451F8902E3CF89D5E98D5433">Zakrzewska, Marta</mods:namePart>
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</mods:name>
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<mods:name id="81237FA6521A36393A51898DFEF7C00C" type="personal">
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<mods:role id="BC7D468E92ED74907840CF430D8EB7D8">
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<mods:roleTerm id="A95DB7B07559E8BD59836868C515A635">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9B1C26D9BD817D2CFA3071700A767763">Giłka, Wojciech</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="B272FF1665B4446DEA42F8EE0B820390">text</mods:typeOfResource>
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<mods:title id="F7B6FFA7BD44ACA1B5C093CE8543D36B">Zootaxa</mods:title>
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<mods:part id="09D6280503199508D29B7A70477C71D9">
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<mods:date id="8908BC5941BF5792A88C72F48A20E639">2015</mods:date>
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<mods:detail id="23799BFB6B9F59C93350623E135DC88A" type="volume">
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<mods:number id="75C8C32E909F67C2F194A3A749B9AE5F">3946</mods:number>
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<mods:detail id="6C6A112E6493BC28FD9E4CA69EA8ABCA" type="issue">
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<mods:number id="4142DD6F4529F3343637573493F2C064">3</mods:number>
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<mods:classification id="E4D6A5D55249909DBF8EB5220DE8B503">journal article</mods:classification>
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<mods:identifier id="FE0B1A48DCC0D32877EE92B1DACA36B4" type="DOI">10.11646/zootaxa.3946.3.3</mods:identifier>
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<mods:identifier id="101DB71C0BDA3A3F27A76921A7A01496" type="GBIF-Dataset">fcec1ba5-a256-4419-a228-ca2b529948bd</mods:identifier>
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<mods:identifier id="DD3254A4F084BD1B63E8CC170CD07145" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="1A9704DD15DAF8A0AB8BC481E212C970" type="Zenodo-Dep">245390</mods:identifier>
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<mods:identifier id="0AB051EAB256BAFAC0E1C595322F1AD8" type="ZooBank">8D7575DB-636C-4473-91C3-A58C1E5F9E73</mods:identifier>
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<treatment id="03CD3E7A711EA71EFF60AFD0D3E9F655" ID-DOI="http://doi.org/10.5281/zenodo.5616330" ID-GBIF-Taxon="119608924" ID-Zenodo-Dep="5616330" LSID="urn:lsid:plazi:treatment:03CD3E7A711EA71EFF60AFD0D3E9F655" httpUri="http://treatment.plazi.org/id/03CD3E7A711EA71EFF60AFD0D3E9F655" lastPageId="12" lastPageNumber="359" pageId="9" pageNumber="356">
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<subSubSection id="C37EDCE7711EA71BFF60AFD0D13EF0BB" box="[151,843,980,1006]" pageId="9" pageNumber="356" type="nomenclature">
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<paragraph id="8BDB8F6C711EA71BFF60AFD0D13EF0BB" blockId="9.[151,843,980,1006]" box="[151,843,980,1006]" pageId="9" pageNumber="356">
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<heading id="D0933800711EA71BFF60AFD0D13EF0BB" bold="true" box="[151,843,980,1006]" fontSize="11" level="1" pageId="9" pageNumber="356" reason="1">
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<emphasis id="B910537E711EA71BFF60AFD0D13EF0BB" bold="true" box="[151,843,980,1006]" pageId="9" pageNumber="356">
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<taxonomicName id="4C64F4EF711EA71BFF60AFD0D094F0BB" ID-CoL="9HM8N" authority="Gilka et Zakrzewska" authorityName="Gilka et Zakrzewska" box="[151,737,980,1006]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="protogregarius" status="sp. nov.">
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<emphasis id="B910537E711EA71BFF60AFD0D3A0F0BB" bold="true" box="[151,469,980,1006]" italics="true" pageId="9" pageNumber="356">Tanytarsus protogregarius</emphasis>
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Giłka
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<emphasis id="B910537E711EA71BFDDDAFD0D04AF0BB" bold="true" box="[554,575,980,1006]" italics="true" pageId="9" pageNumber="356">et</emphasis>
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Zakrzewska
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</taxonomicName>
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,
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<taxonomicNameLabel id="A223EE05711EA71BFD19AFD0D13EF0BB" box="[750,843,980,1006]" pageId="9" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37EDCE7711EA71BFF60A81ED0A8F796" pageId="9" pageNumber="356" type="materials_examined">
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<paragraph id="8BDB8F6C711EA71BFF60A81ED0A8F796" blockId="9.[151,1437,1050,2020]" pageId="9" pageNumber="356">
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<emphasis id="B910537E711EA71BFF60A81ED330F766" bold="true" box="[151,325,1050,1075]" pageId="9" pageNumber="356">
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<typeStatus id="54DF31CE711EA71BFF60A81ED2A7F766" box="[151,210,1050,1075]" pageId="9" pageNumber="356">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54DF31CE711EA71BFEBBA81ED3CDF766" box="[332,440,1050,1075]" pageId="9" pageNumber="356" type="holotype">Holotype</typeStatus>
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. Adult male (tarsi of hind legs broken and separated) preserved in
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<date id="FFDAA9AC711EA71BFB50A81ED689F766" box="[1191,1276,1050,1075]" pageId="9" pageNumber="356" value="2013-10-15">15 x 13</date>
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||
x
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<quantity id="4C9C2289711EA71BFAEFA81ED72FF767" box="[1304,1370,1050,1075]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="356" unit="mm" value="5.0">5 mm</quantity>
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piece of amber (Eocene, ~45–40 Ma, Baltic amber, Gulf of Gdańsk; MAI-4325a;
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<figureCitation id="135F93E9711EA71BFBC5A83AD60EF702" box="[1074,1147,1086,1111]" captionStart="FIGURE 7" captionStartId="10.[151,250,1955,1977]" captionTargetBox="[151,1436,194,1932]" captionTargetId="figure@10.[151,1436,194,1933]" captionTargetPageId="10" captionText="FIGURE 7. Tanytarsus protogregarius sp. nov., adult male, holotype MAI- 4325 a (A, C – G), paratypes MAI- 4352 (B) (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A, B — inclusions; C — habitus; D — head; E — antenna (arrows: borders between flagellomeres); F, G — wing photographed in reflected (F) and transmitted light (G)." httpUri="https://zenodo.org/record/245397/files/figure.png" pageId="9" pageNumber="356">Fig. 7</figureCitation>
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A); animal syninclusion:
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<taxonomicName id="4C64F4EF711EA71BFF60A866D28AF72E" box="[151,255,1122,1147]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
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(1 ind., MAI-4325).
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<typeStatus id="54DF31CE711EA71BFE03A867D010F72E" box="[500,613,1123,1147]" pageId="9" pageNumber="356" type="paratype">Paratypes</typeStatus>
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. Two adult males (tarsi of all legs broken and separated and/or missing, wings poorly preserved), as syninclusions, slightly compressed, preserved in
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<date id="FFDAA9AC711EA71BFBE7A882D61FF7CA" box="[1040,1130,1158,1183]" pageId="9" pageNumber="356" value="1918-10-16">16 x 18</date>
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||
x
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<quantity id="4C9C2289711EA71BFB7BA883D6BAF7CB" box="[1164,1231,1159,1183]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="356" unit="mm" value="4.0">4 mm</quantity>
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piece of clarified amber (MAI-4352, same data as
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<typeStatus id="54DF31CE711EA71BFDF0A8AED019F796" box="[519,620,1194,1219]" pageId="9" pageNumber="356" type="holotype">holotype</typeStatus>
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;
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<figureCitation id="135F93E9711EA71BFD8FA8AED0CEF796" box="[632,699,1194,1219]" captionStart="FIGURE 7" captionStartId="10.[151,250,1955,1977]" captionTargetBox="[151,1436,194,1932]" captionTargetId="figure@10.[151,1436,194,1933]" captionTargetPageId="10" captionText="FIGURE 7. Tanytarsus protogregarius sp. nov., adult male, holotype MAI- 4325 a (A, C – G), paratypes MAI- 4352 (B) (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A, B — inclusions; C — habitus; D — head; E — antenna (arrows: borders between flagellomeres); F, G — wing photographed in reflected (F) and transmitted light (G)." httpUri="https://zenodo.org/record/245397/files/figure.png" pageId="9" pageNumber="356">Fig. 7</figureCitation>
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B).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37EDCE7711EA71BFF30A8CAD11FF65E" pageId="9" pageNumber="356" type="etymology">
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<paragraph id="8BDB8F6C711EA71BFF30A8CAD11FF65E" blockId="9.[151,1437,1050,2020]" pageId="9" pageNumber="356">
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<emphasis id="B910537E711EA71BFF30A8CAD3DCF7B2" bold="true" box="[199,425,1230,1255]" pageId="9" pageNumber="356">Derivatio nominis.</emphasis>
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Combination of two words: ‘
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<emphasis id="B910537E711EA71BFCF8A8CAD1EFF7B2" box="[783,922,1230,1255]" italics="true" pageId="9" pageNumber="356">protoplastus</emphasis>
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’ and ‘
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<taxonomicName id="4C64F4EF711EA71BFC19A8CBD62FF7B3" box="[1006,1114,1231,1254]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="gregarius">
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<emphasis id="B910537E711EA71BFC19A8CBD62FF7B3" box="[1006,1114,1231,1254]" italics="true" pageId="9" pageNumber="356">gregarius</emphasis>
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</taxonomicName>
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’. The species is the oldest known representative of the
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<taxonomicName id="4C64F4EF711EA71BFE22A8F0D0B7F65E" box="[469,706,1268,1291]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="species" species="gregarius">
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<emphasis id="B910537E711EA71BFE22A8F0D0B7F65E" box="[469,706,1268,1291]" italics="true" pageId="9" pageNumber="356">Tanytarsus gregarius</emphasis>
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</taxonomicName>
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species group.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C37EDCE7711EA71BFF30A912D1F0F6CE" pageId="9" pageNumber="356" type="diagnosis">
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||
<paragraph id="8BDB8F6C711EA71BFF30A912D1F0F6CE" blockId="9.[151,1437,1050,2020]" pageId="9" pageNumber="356">
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<emphasis id="B910537E711EA71BFF30A912D334F67A" bold="true" box="[199,321,1302,1327]" pageId="9" pageNumber="356">Diagnosis.</emphasis>
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Femur of mid leg longer than femur of hind leg. Gonostylus with subapical tooth-like process. Anal point slender, tapering to blunt apex, bearing spinulae arranged in row. Superior volsella posteriorly directed, with nipple-like extension on apex and 4 strong medially directed setae on median margin. Digitus not observed. Stem of median volsella long, straight, with 5–6 slender foliate lamellae.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37EDCE7711EA71EFF30A9A2D0B3F1B6" lastPageId="12" lastPageNumber="359" pageId="9" pageNumber="356" type="description">
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<paragraph id="8BDB8F6C711EA71BFF30A9A2D039F6EA" blockId="9.[151,1437,1050,2020]" box="[199,588,1446,1471]" pageId="9" pageNumber="356">
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<emphasis id="B910537E711EA71BFF30A9A2D324F6EA" bold="true" box="[199,337,1446,1471]" pageId="9" pageNumber="356">Description</emphasis>
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. Adult male (n = 1–3)
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</paragraph>
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<paragraph id="8BDB8F6C711EA71BFF30A9CED152F6B7" blockId="9.[151,1437,1050,2020]" box="[199,807,1482,1507]" pageId="9" pageNumber="356">
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Total length
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<quantity id="4C9C2289711EA71BFEA3A9CED3ABF6B7" box="[340,478,1482,1507]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="9" pageNumber="356" unit="mm" value="2.85" valueMax="3.1" valueMin="2.6">2.6–3.1 mm</quantity>
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; wing length
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<date id="FFDAA9AC711EA71BFD82A9CED086F6B6" box="[629,755,1482,1507]" pageId="9" pageNumber="356" value="1475" valueMax="1575">1475–1575</date>
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Μm.
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</paragraph>
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<paragraph id="8BDB8F6C711EA71BFF30A9EAD3F1F526" blockId="9.[151,1437,1050,2020]" pageId="9" pageNumber="356">
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<emphasis id="B910537E711EA71BFF30A9EAD371F552" box="[199,260,1518,1543]" italics="true" pageId="9" pageNumber="356">Head</emphasis>
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(
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<figureCitation id="135F93E9711EA71BFEE2A9EAD32FF553" box="[277,346,1518,1543]" captionStart="FIGURE 7" captionStartId="10.[151,250,1955,1977]" captionTargetBox="[151,1436,194,1932]" captionTargetId="figure@10.[151,1436,194,1933]" captionTargetPageId="10" captionText="FIGURE 7. Tanytarsus protogregarius sp. nov., adult male, holotype MAI- 4325 a (A, C – G), paratypes MAI- 4352 (B) (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A, B — inclusions; C — habitus; D — head; E — antenna (arrows: borders between flagellomeres); F, G — wing photographed in reflected (F) and transmitted light (G)." httpUri="https://zenodo.org/record/245397/files/figure.png" pageId="9" pageNumber="356">Fig. 7</figureCitation>
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D, E). Eyes bare, with well developed dorsomedian extensions. Frontal tubercles not observed. Antenna with 13 well discernible flagellomeres, AR 0.92–0.97, plume fully developed (
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<figureCitation id="135F93E9711EA71BFB45AA16D68EF57E" box="[1202,1275,1554,1579]" captionStart="FIGURE 7" captionStartId="10.[151,250,1955,1977]" captionTargetBox="[151,1436,194,1932]" captionTargetId="figure@10.[151,1436,194,1933]" captionTargetPageId="10" captionText="FIGURE 7. Tanytarsus protogregarius sp. nov., adult male, holotype MAI- 4325 a (A, C – G), paratypes MAI- 4352 (B) (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A, B — inclusions; C — habitus; D — head; E — antenna (arrows: borders between flagellomeres); F, G — wing photographed in reflected (F) and transmitted light (G)." httpUri="https://zenodo.org/record/245397/files/figure.png" pageId="9" pageNumber="356">Fig. 7</figureCitation>
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E). Length of palpomeres 2–5 (Μm): 40–48, 113–137, 115–125, 149–185. Clypeals present but poorly visible (at least
|
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<quantity id="4C9C2289711EA71BFAA5AA32D7E9F51A" box="[1362,1436,1590,1615]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.524" metricValueMin="1.016" pageId="9" pageNumber="356" unit="in" value="5.0" valueMax="6.0" valueMin="4.0">4–6 in</quantity>
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||
<typeStatus id="54DF31CE711EA71BFF60AA5FD282F526" box="[151,247,1627,1651]" pageId="9" pageNumber="356" type="paratype">paratype</typeStatus>
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||
specimens).
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||
</paragraph>
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||
<paragraph id="8BDB8F6C711EA71BFF30AA7AD676F5C2" blockId="9.[151,1437,1050,2020]" box="[199,1027,1662,1687]" pageId="9" pageNumber="356">
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||
<emphasis id="B910537E711EA71BFF30AA7AD36DF5C2" box="[199,280,1662,1687]" italics="true" pageId="9" pageNumber="356">Thorax</emphasis>
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. Ac at least 12, Dc 13 on each side, Pa 3 on each side, Scts 12–13.
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</paragraph>
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<paragraph id="8BDB8F6C711EA71BFF30AAA0D1A6F478" blockId="9.[151,1437,1050,2020]" pageId="9" pageNumber="356">
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||
<emphasis id="B910537E711EA71BFF30AAA0D28AF5EE" box="[199,255,1700,1723]" italics="true" pageId="9" pageNumber="356">Wing</emphasis>
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||
(
|
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<figureCitation id="135F93E9711EA71BFEE5AAA6D323F5EF" box="[274,342,1698,1723]" captionStart="FIGURE 7" captionStartId="10.[151,250,1955,1977]" captionTargetBox="[151,1436,194,1932]" captionTargetId="figure@10.[151,1436,194,1933]" captionTargetPageId="10" captionText="FIGURE 7. Tanytarsus protogregarius sp. nov., adult male, holotype MAI- 4325 a (A, C – G), paratypes MAI- 4352 (B) (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). A, B — inclusions; C — habitus; D — head; E — antenna (arrows: borders between flagellomeres); F, G — wing photographed in reflected (F) and transmitted light (G)." httpUri="https://zenodo.org/record/245397/files/figure.png" pageId="9" pageNumber="356">Fig. 7</figureCitation>
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F, G). Ellipse-shaped, with anal lobe weak, broadest at 2/3 length, width: 385–395 Μm, length/ width ratio 3.83–3.98. FCu placed well distally of RM; VR Cu 1.33. Veins ending as in most extant
|
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<taxonomicName id="4C64F4EF711EA71BFB2BAAC3D723F58B" box="[1244,1366,1735,1758]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="356" phylum="Arthropoda" rank="genus">
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<emphasis id="B910537E711EA71BFB2BAAC3D723F58B" box="[1244,1366,1735,1758]" italics="true" pageId="9" pageNumber="356">Tanytarsus</emphasis>
|
||
</taxonomicName>
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(from base to tip): An, Sc, Cu1, R1, R2+3, M3+4, R4+5, M1+2; distances between ends of R1–R2+3 and R2+3–R4+5 unequal (VR C 1.35). Almost whole wing (except base) covered with dense macrotrichia.
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</paragraph>
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<paragraph id="8BDB8F6C711EA71BFF30AB3DD326F4B6" blockId="9.[151,1437,1050,2020]" pageId="9" pageNumber="356">
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||
<emphasis id="B910537E711EA71BFF30AB3DD289F405" box="[199,252,1849,1872]" italics="true" pageId="9" pageNumber="356">Legs</emphasis>
|
||
. Tibia of fore leg with single, 28–30 Μm long spur (a vestigial comb composed of 2–3 teeth, observed on one leg of
|
||
<typeStatus id="54DF31CE711EA71BFEE6AB59D307F420" box="[273,370,1885,1909]" pageId="9" pageNumber="356" type="paratype">paratype</typeStatus>
|
||
specimen is recognized as artefact). Tibial combs of mid and hind legs separated, fan-shaped, teeth up to 20 Μm long (mid leg) and c. 20–24 Μm long (hind leg); each comb with slender spur, up to 30 Μm long (mid leg) and 36–48 Μm long (hind leg). Sensilla chaetica on ta1 of p2 not observed. Lengths of leg segments and leg ratios in
|
||
<tableCitation id="C6E6BAD7711EA71BFF0EABCFD33BF4B6" box="[249,334,1995,2020]" captionStart="TABLE 3" captionStartId="12.[151,239,152,175]" captionTargetBox="[159,1404,202,359]" captionTargetPageId="12" captionText="TABLE 3. Leg segment lengths (Μm) and leg ratios of male Tanytarsus protogregarius sp. nov." httpUri="http://table.plazi.org/id/DF1BDFE4711BA71EFF60AC9CD6E6F3FA" pageId="9" pageNumber="356" tableUuid="DF1BDFE4711BA71EFF60AC9CD6E6F3FA">Table 3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF1BDFE4711DA718FF60ABA7D631F4A3" httpUri="https://zenodo.org/record/245397/files/figure.png" pageId="10" pageNumber="357" targetBox="[151,1436,194,1932]" targetPageId="10">
|
||
<paragraph id="8BDB8F6C711DA718FF60ABA7D631F4A3" blockId="10.[151,1436,1955,2038]" pageId="10" pageNumber="357">
|
||
<emphasis id="B910537E711DA718FF60ABA7D36CF4ED" bold="true" box="[151,281,1955,1977]" pageId="10" pageNumber="357">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C64F4EF711DA718FED2ABA0D043F4EC" box="[293,566,1956,1977]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="357" phylum="Arthropoda" rank="species" species="protogregarius" status="sp. nov.">
|
||
<emphasis id="B910537E711DA718FED2ABA0D043F4EC" box="[293,566,1956,1977]" italics="true" pageId="10" pageNumber="357">Tanytarsus protogregarius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B910537E711DA718FDB4ABA7D0ECF4ED" bold="true" box="[579,665,1955,1976]" pageId="10" pageNumber="357">
|
||
<taxonomicNameLabel id="A223EE05711DA718FDB4ABA7D0ECF4ED" box="[579,665,1955,1976]" pageId="10" pageNumber="357" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, adult male, holotype MAI-4325a (
|
||
<emphasis id="B910537E711DA718FBD6ABA7D646F4ED" bold="true" box="[1057,1075,1955,1976]" pageId="10" pageNumber="357">A</emphasis>
|
||
,
|
||
<emphasis id="B910537E711DA718FBB1ABA7D60CF4EC" bold="true" box="[1094,1145,1955,1977]" pageId="10" pageNumber="357">C–G</emphasis>
|
||
), paratypes MAI-4352 (
|
||
<emphasis id="B910537E711DA718FA74ABA7D7E1F4ED" bold="true" box="[1411,1428,1955,1976]" pageId="10" pageNumber="357">B</emphasis>
|
||
) (Eocene, ~45–40 Ma; Baltic amber, Gulf of Gdańsk).
|
||
<emphasis id="B910537E711DA718FD32ABC5D0A2F483" bold="true" box="[709,727,1985,2006]" pageId="10" pageNumber="357">A</emphasis>
|
||
,
|
||
<emphasis id="B910537E711DA718FD12ABC6D083F482" bold="true" box="[741,758,1986,2007]" pageId="10" pageNumber="357">B</emphasis>
|
||
—inclusions;
|
||
<emphasis id="B910537E711DA718FC72ABC5D1E2F482" bold="true" box="[901,919,1985,2007]" pageId="10" pageNumber="357">C</emphasis>
|
||
—habitus;
|
||
<emphasis id="B910537E711DA718FBFEABC6D66EF482" bold="true" box="[1033,1051,1986,2007]" pageId="10" pageNumber="357">D</emphasis>
|
||
—head;
|
||
<emphasis id="B910537E711DA718FB84ABC6D6F1F482" bold="true" box="[1139,1156,1986,2007]" pageId="10" pageNumber="357">E</emphasis>
|
||
—antenna (arrows: borders between flagellomeres);
|
||
<emphasis id="B910537E711DA718FE66ABE4D3EBF4A0" bold="true" box="[401,414,2016,2037]" pageId="10" pageNumber="357">F</emphasis>
|
||
,
|
||
<emphasis id="B910537E711DA718FE5CABE4D3CAF4A3" bold="true" box="[427,447,2016,2038]" pageId="10" pageNumber="357">G</emphasis>
|
||
—wing photographed in reflected (
|
||
<emphasis id="B910537E711DA718FCD3ABE4D146F4A0" bold="true" box="[804,819,2016,2037]" pageId="10" pageNumber="357">F</emphasis>
|
||
) and transmitted light (
|
||
<emphasis id="B910537E711DA718FBD5ABE4D643F4A3" bold="true" box="[1058,1078,2016,2038]" pageId="10" pageNumber="357">G</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1BDFE4711CA719FF60AB70D1E9F4B3" httpUri="https://zenodo.org/record/245398/files/figure.png" pageId="11" pageNumber="358" targetBox="[165,1408,193,1851]" targetPageId="11">
|
||
<paragraph id="8BDB8F6C711CA719FF60AB70D1E9F4B3" blockId="11.[151,1436,1908,2022]" pageId="11" pageNumber="358">
|
||
<emphasis id="B910537E711CA719FF60AB70D367F4DC" bold="true" box="[151,274,1908,1930]" pageId="11" pageNumber="358">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C64F4EF711CA719FEEEAB71D051F4DF" box="[281,548,1909,1930]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="358" phylum="Arthropoda" rank="species" species="protogregarius" status="sp. nov.">
|
||
<emphasis id="B910537E711CA719FEEEAB71D051F4DF" box="[281,548,1909,1930]" italics="true" pageId="11" pageNumber="358">Tanytarsus protogregarius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A223EE05711CA719FDDCAB70D00EF4DC" box="[555,635,1908,1929]" pageId="11" pageNumber="358" rank="species">
|
||
<emphasis id="B910537E711CA719FDDCAB70D00EF4DC" bold="true" box="[555,635,1908,1929]" pageId="11" pageNumber="358">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult male, holotype MAI-4325a (
|
||
<emphasis id="B910537E711CA719FC13AB70D660F4DC" bold="true" box="[996,1045,1908,1930]" pageId="11" pageNumber="358">A–D</emphasis>
|
||
), paratype MAI–4352 (
|
||
<emphasis id="B910537E711CA719FAF0AB70D76DF4DC" bold="true" box="[1287,1304,1908,1929]" pageId="11" pageNumber="358">E</emphasis>
|
||
,
|
||
<emphasis id="B910537E711CA719FAD3AB70D746F4DC" bold="true" box="[1316,1331,1908,1929]" pageId="11" pageNumber="358">F</emphasis>
|
||
) (Eocene, ~45–40 Ma; Baltic amber, Gulf of Gdańsk). Hypopygium in dorsal (
|
||
<emphasis id="B910537E711CA719FCAAAB96D11AF4F2" bold="true" box="[861,879,1938,1959]" pageId="11" pageNumber="358">A</emphasis>
|
||
,
|
||
<emphasis id="B910537E711CA719FC89AB96D1E5F4FD" bold="true" box="[894,912,1938,1960]" pageId="11" pageNumber="358">C</emphasis>
|
||
) and ventral aspect (
|
||
<emphasis id="B910537E711CA719FB9BAB97D608F4FD" bold="true" box="[1132,1149,1939,1960]" pageId="11" pageNumber="358">B</emphasis>
|
||
); photographed in reflected light (
|
||
<emphasis id="B910537E711CA719FF23ABB5D293F493" bold="true" box="[212,230,1969,1990]" pageId="11" pageNumber="358">A</emphasis>
|
||
,
|
||
<emphasis id="B910537E711CA719FF04ABB5D371F493" bold="true" box="[243,260,1969,1990]" pageId="11" pageNumber="358">B</emphasis>
|
||
), and drawn (
|
||
<emphasis id="B910537E711CA719FE64ABB5D3D0F492" bold="true" box="[403,421,1969,1991]" pageId="11" pageNumber="358">C</emphasis>
|
||
);
|
||
<emphasis id="B910537E711CA719FE4CABB5D3B8F493" bold="true" box="[443,461,1969,1990]" pageId="11" pageNumber="358">D</emphasis>
|
||
—superior volsella;
|
||
<emphasis id="B910537E711CA719FD6BABB5D0D8F493" bold="true" box="[668,685,1969,1990]" pageId="11" pageNumber="358">E</emphasis>
|
||
,
|
||
<emphasis id="B910537E711CA719FD4DABB5D0BCF493" bold="true" box="[698,713,1969,1990]" pageId="11" pageNumber="358">F</emphasis>
|
||
—median volsella photographed in transmitted light (
|
||
<emphasis id="B910537E711CA719FB1CABB5D689F493" bold="true" box="[1259,1276,1969,1990]" pageId="11" pageNumber="358">E</emphasis>
|
||
) and drawn (
|
||
<emphasis id="B910537E711CA719FA73ABB5D7E6F493" bold="true" box="[1412,1427,1969,1990]" pageId="11" pageNumber="358">F</emphasis>
|
||
) (D—magnified c. 2 times and F—c. 5 times relative to hypopygium drawn).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1BDFE4711BA71EFF60AC9CD6E6F3FA" ID-Table-UUID="DF1BDFE4711BA71EFF60AC9CD6E6F3FA" box="[151,1171,152,175]" httpUri="http://table.plazi.org/id/DF1BDFE4711BA71EFF60AC9CD6E6F3FA" pageId="12" pageNumber="359" targetBox="[159,1404,202,359]" targetIsTable="true" targetPageId="12">
|
||
<paragraph id="8BDB8F6C711BA71EFF60AC9CD6E6F3FA" blockId="12.[151,1171,152,175]" box="[151,1171,152,175]" pageId="12" pageNumber="359">
|
||
<emphasis id="B910537E711BA71EFF60AC9CD37CF3FA" bold="true" box="[151,265,152,175]" pageId="12" pageNumber="359">TABLE 3.</emphasis>
|
||
Leg segment lengths (Μm) and leg ratios of male
|
||
<taxonomicName id="4C64F4EF711BA71EFCEAAC9CD643F3FA" box="[797,1078,152,175]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="protogregarius" status="sp. nov.">
|
||
<emphasis id="B910537E711BA71EFCEAAC9CD643F3FA" box="[797,1078,152,175]" italics="true" pageId="12" pageNumber="359">Tanytarsus protogregarius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A223EE05711BA71EFBCAAC9CD6E6F3FA" box="[1085,1171,152,175]" pageId="12" pageNumber="359" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDB8F6C711BA71EFEDAACCED737F231" pageId="12" pageNumber="359">
|
||
<table id="F9647DCC711B58EDFF68ACCED709F232" box="[159,1404,202,359]" gridcols="9" gridrows="4" pageId="12" pageNumber="359">
|
||
<tr id="35548D2E711B58EDFF68ACCED709F3B1" box="[159,1404,202,228]" gridrow="0" pageId="12" pageNumber="359" rowspan-0="1">
|
||
<th id="7685E452711B58EDFEDAACCED326F3B1" box="[301,339,202,228]" gridcol="1" gridrow="0" pageId="12" pageNumber="359">fe</th>
|
||
<th id="7685E452711B58EDFE4BACCED060F3B1" box="[444,533,202,228]" gridcol="2" gridrow="0" pageId="12" pageNumber="359">ti</th>
|
||
<th id="7685E452711B58EDFDBCACCED0D1F3B1" box="[587,676,202,228]" gridcol="3" gridrow="0" pageId="12" pageNumber="359">ta1</th>
|
||
<th id="7685E452711B58EDFD2DACCED146F3B1" box="[730,819,202,228]" gridcol="4" gridrow="0" pageId="12" pageNumber="359">ta2</th>
|
||
<th id="7685E452711B58EDFC9EACCED1B7F3B1" box="[873,962,202,228]" gridcol="5" gridrow="0" pageId="12" pageNumber="359">ta3</th>
|
||
<th id="7685E452711B58EDFC0FACCED624F3B1" box="[1016,1105,202,228]" gridcol="6" gridrow="0" pageId="12" pageNumber="359">ta4</th>
|
||
<th id="7685E452711B58EDFB70ACCED6AAF3B1" box="[1159,1247,202,228]" gridcol="7" gridrow="0" pageId="12" pageNumber="359">ta5</th>
|
||
<th id="7685E452711B58EDFAE1ACCED709F3B1" box="[1302,1404,202,228]" gridcol="8" gridrow="0" pageId="12" pageNumber="359">LR</th>
|
||
</tr>
|
||
<tr id="35548D2E711B58EDFF68ACF2D709F245" box="[159,1404,246,272]" gridrow="1" pageId="12" pageNumber="359">
|
||
<th id="7685E452711B58EDFF68ACF2D2C6F245" box="[159,179,246,272]" gridcol="0" gridrow="1" pageId="12" pageNumber="359">p1</th>
|
||
<td id="7685E452711B58EDFEDAACF2D326F245" box="[301,339,246,272]" gridcol="1" gridrow="1" pageId="12" pageNumber="359">850</td>
|
||
<td id="7685E452711B58EDFE4BACF2D060F245" box="[444,533,246,272]" gridcol="2" gridrow="1" pageId="12" pageNumber="359">560</td>
|
||
<td id="7685E452711B58EDFDBCACF2D0D1F245" box="[587,676,246,272]" gridcol="3" gridrow="1" pageId="12" pageNumber="359">715–765</td>
|
||
<td id="7685E452711B58EDFD2DACF2D146F245" box="[730,819,246,272]" gridcol="4" gridrow="1" pageId="12" pageNumber="359">445–470</td>
|
||
<td id="7685E452711B58EDFC9EACF2D1B7F245" box="[873,962,246,272]" gridcol="5" gridrow="1" pageId="12" pageNumber="359">355–380</td>
|
||
<td id="7685E452711B58EDFC0FACF2D624F245" box="[1016,1105,246,272]" gridcol="6" gridrow="1" pageId="12" pageNumber="359">250–270</td>
|
||
<td id="7685E452711B58EDFB70ACF2D6AAF245" box="[1159,1247,246,272]" gridcol="7" gridrow="1" pageId="12" pageNumber="359">110–120</td>
|
||
<td id="7685E452711B58EDFAE1ACF2D709F245" box="[1302,1404,246,272]" gridcol="8" gridrow="1" pageId="12" pageNumber="359">1.28–1.37</td>
|
||
</tr>
|
||
<tr id="35548D2E711B58EDFF68AD26D709F269" box="[159,1404,290,316]" gridrow="2" pageId="12" pageNumber="359">
|
||
<th id="7685E452711B58EDFF68AD26D2C6F269" box="[159,179,290,316]" gridcol="0" gridrow="2" pageId="12" pageNumber="359">p2</th>
|
||
<td id="7685E452711B58EDFEDAAD26D326F269" box="[301,339,290,316]" gridcol="1" gridrow="2" pageId="12" pageNumber="359">835</td>
|
||
<td id="7685E452711B58EDFE4BAD26D060F269" box="[444,533,290,316]" gridcol="2" gridrow="2" pageId="12" pageNumber="359">715–730</td>
|
||
<td id="7685E452711B58EDFDBCAD26D0D1F269" box="[587,676,290,316]" gridcol="3" gridrow="2" pageId="12" pageNumber="359">390–415</td>
|
||
<td id="7685E452711B58EDFD2DAD26D146F269" box="[730,819,290,316]" gridcol="4" gridrow="2" pageId="12" pageNumber="359">250–260</td>
|
||
<td id="7685E452711B58EDFC9EAD26D1B7F269" box="[873,962,290,316]" gridcol="5" gridrow="2" pageId="12" pageNumber="359">185–195</td>
|
||
<td id="7685E452711B58EDFC0FAD26D624F269" box="[1016,1105,290,316]" gridcol="6" gridrow="2" pageId="12" pageNumber="359">125</td>
|
||
<td id="7685E452711B58EDFB70AD26D6AAF269" box="[1159,1247,290,316]" gridcol="7" gridrow="2" pageId="12" pageNumber="359">80–85</td>
|
||
<td id="7685E452711B58EDFAE1AD26D709F269" box="[1302,1404,290,316]" gridcol="8" gridrow="2" pageId="12" pageNumber="359">0.55–0.58</td>
|
||
</tr>
|
||
<tr id="35548D2E711B58EDFF68AD4AD709F232" box="[159,1404,334,359]" gridrow="3" pageId="12" pageNumber="359">
|
||
<th id="7685E452711B58EDFF68AD4AD2C6F232" box="[159,179,334,359]" gridcol="0" gridrow="3" pageId="12" pageNumber="359">p3</th>
|
||
<td id="7685E452711B58EDFEDAAD4AD326F232" box="[301,339,334,359]" gridcol="1" gridrow="3" pageId="12" pageNumber="359">810</td>
|
||
<td id="7685E452711B58EDFE4BAD4AD060F232" box="[444,533,334,359]" gridcol="2" gridrow="3" pageId="12" pageNumber="359">835–945</td>
|
||
<td id="7685E452711B58EDFDBCAD4AD0D1F232" box="[587,676,334,359]" gridcol="3" gridrow="3" pageId="12" pageNumber="359">470</td>
|
||
<td id="7685E452711B58EDFD2DAD4AD146F232" box="[730,819,334,359]" gridcol="4" gridrow="3" pageId="12" pageNumber="359">310</td>
|
||
<td id="7685E452711B58EDFC9EAD4AD1B7F232" box="[873,962,334,359]" gridcol="5" gridrow="3" pageId="12" pageNumber="359">270–275</td>
|
||
<td id="7685E452711B58EDFC0FAD4AD624F232" box="[1016,1105,334,359]" gridcol="6" gridrow="3" pageId="12" pageNumber="359">155–165</td>
|
||
<td id="7685E452711B58EDFB70AD4AD6AAF232" box="[1159,1247,334,359]" gridcol="7" gridrow="3" pageId="12" pageNumber="359">95</td>
|
||
<td id="7685E452711B58EDFAE1AD4AD709F232" box="[1302,1404,334,359]" gridcol="8" gridrow="3" pageId="12" pageNumber="359">0.56</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BDB8F6C711BA71EFF30ADA8D0B3F1B6" blockId="12.[151,1437,427,1280]" pageId="12" pageNumber="359">
|
||
<emphasis id="B910537E711BA71EFF30ADA8D323F296" box="[199,342,428,451]" italics="true" pageId="12" pageNumber="359">Hypopygium</emphasis>
|
||
(
|
||
<figureCitation id="135F93E9711BA71EFE9AADAFD3C3F291" box="[365,438,427,452]" captionStart="FIGURE 8" captionStartId="11.[151,250,1908,1930]" captionTargetBox="[165,1408,193,1851]" captionTargetId="figure@11.[165,1409,193,1869]" captionTargetPageId="11" captionText="FIGURE 8. Tanytarsus protogregarius sp. nov., adult male, holotype MAI- 4325 a (A – D), paratype MAI – 4352 (E, F) (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). Hypopygium in dorsal (A, C) and ventral aspect (B); photographed in reflected light (A, B), and drawn (C); D — superior volsella; E, F — median volsella photographed in transmitted light (E) and drawn (F) (D—magnified c. 2 times and F—c. 5 times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/245398/files/figure.png" pageId="12" pageNumber="359">Fig. 8</figureCitation>
|
||
A–F). Gonostylus stout, slightly curved at mid length, c. 95–115 Μm, longer than gonocoxite, bearing several setae placed at median margin of distal half, with subapical anteromedially directed tooth-like process. Median setae on anal tergite not observed, at least 3 posterolateral setae on each side of anal point. Anal point slender, tapering to blunt apex, bearing at least 3 spinulae arranged in row, crests tapering towards anal point apex. Superior volsella posteriorly directed, oval, slightly elongated, with nipple-like extension on apex and 4 strong medially directed setae placed on distinct protuberances on median margin (
|
||
<figureCitation id="135F93E9711BA71EFB50AE5BD698F12D" box="[1191,1261,607,632]" captionStart="FIGURE 8" captionStartId="11.[151,250,1908,1930]" captionTargetBox="[165,1408,193,1851]" captionTargetId="figure@11.[165,1409,193,1869]" captionTargetPageId="11" captionText="FIGURE 8. Tanytarsus protogregarius sp. nov., adult male, holotype MAI- 4325 a (A – D), paratype MAI – 4352 (E, F) (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). Hypopygium in dorsal (A, C) and ventral aspect (B); photographed in reflected light (A, B), and drawn (C); D — superior volsella; E, F — median volsella photographed in transmitted light (E) and drawn (F) (D—magnified c. 2 times and F—c. 5 times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/245398/files/figure.png" pageId="12" pageNumber="359">Fig. 8</figureCitation>
|
||
D). Digitus not observed. Stem of median volsella c. 25 Μm long, straight, bearing 5–6 slender foliate lamellae (
|
||
<figureCitation id="135F93E9711BA71EFAE6AE87D722F1C9" box="[1297,1367,643,668]" captionStart="FIGURE 8" captionStartId="11.[151,250,1908,1930]" captionTargetBox="[165,1408,193,1851]" captionTargetId="figure@11.[165,1409,193,1869]" captionTargetPageId="11" captionText="FIGURE 8. Tanytarsus protogregarius sp. nov., adult male, holotype MAI- 4325 a (A – D), paratype MAI – 4352 (E, F) (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). Hypopygium in dorsal (A, C) and ventral aspect (B); photographed in reflected light (A, B), and drawn (C); D — superior volsella; E, F — median volsella photographed in transmitted light (E) and drawn (F) (D—magnified c. 2 times and F—c. 5 times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/245398/files/figure.png" pageId="12" pageNumber="359">Fig. 8</figureCitation>
|
||
E, F). Inferior volsella reaching 1/3 length of gonostylus at most, club-shaped, with posteromedially turned head-like apical part, armed with several stout curved setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37EDCE7711BA71EFF30AEEBD3E9F655" pageId="12" pageNumber="359" type="discussion">
|
||
<paragraph id="8BDB8F6C711BA71EFF30AEEBD3E9F655" blockId="12.[151,1437,427,1280]" pageId="12" pageNumber="359">
|
||
<emphasis id="B910537E711BA71EFF30AEEBD34EF05D" bold="true" box="[199,315,751,776]" pageId="12" pageNumber="359">Remarks.</emphasis>
|
||
<taxonomicName id="4C64F4EF711BA71EFEB4AEF4D019F052" box="[323,620,752,775]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="protogregarius">
|
||
<emphasis id="B910537E711BA71EFEB4AEF4D019F052" box="[323,620,752,775]" italics="true" pageId="12" pageNumber="359">Tanytarsus protogregarius</emphasis>
|
||
</taxonomicName>
|
||
is here proposed to be included in the
|
||
<taxonomicName id="4C64F4EF711BA71EFBDDAEF4D6E3F052" box="[1066,1174,752,775]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="gregarius">
|
||
<emphasis id="B910537E711BA71EFBDDAEF4D6E3F052" box="[1066,1174,752,775]" italics="true" pageId="12" pageNumber="359">gregarius</emphasis>
|
||
</taxonomicName>
|
||
species group due to a distinct similarity to the extant species—
|
||
<taxonomicName id="4C64F4EF711BA71EFD99AF10D677F079" authority="Kieffer, 1909" authorityName="Kieffer" authorityYear="1909" box="[622,1026,787,812]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="gregarius">
|
||
<emphasis id="B910537E711BA71EFD99AF10D12BF07E" box="[622,862,788,811]" italics="true" pageId="12" pageNumber="359">Tanytarsus gregarius</emphasis>
|
||
Kieffer, 1909
|
||
</taxonomicName>
|
||
. Both the species are similar in the shape of the anal point and the superior volsella that is posteriorly directed and armed with nipple-like apical extension (
|
||
<emphasis id="B910537E711BA71EFEE4AF58D35AF021" box="[275,303,860,884]" italics="true" pageId="12" pageNumber="359">cf.</emphasis>
|
||
<figureCitation id="135F93E9711BA71EFEC1AF5FD30DF021" box="[310,376,859,884]" captionStart="FIGURE 8" captionStartId="11.[151,250,1908,1930]" captionTargetBox="[165,1408,193,1851]" captionTargetId="figure@11.[165,1409,193,1869]" captionTargetPageId="11" captionText="FIGURE 8. Tanytarsus protogregarius sp. nov., adult male, holotype MAI- 4325 a (A – D), paratype MAI – 4352 (E, F) (Eocene, ~ 45 – 40 Ma; Baltic amber, Gulf of Gdańsk). Hypopygium in dorsal (A, C) and ventral aspect (B); photographed in reflected light (A, B), and drawn (C); D — superior volsella; E, F — median volsella photographed in transmitted light (E) and drawn (F) (D—magnified c. 2 times and F—c. 5 times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/245398/files/figure.png" pageId="12" pageNumber="359">Fig. 8</figureCitation>
|
||
and
|
||
<bibRefCitation id="EFF5F29D711BA71EFE58AF5FD0D7F021" author="Reiss" box="[431,674,859,884]" pageId="12" pageNumber="359" refString="Reiss, F. & Fittkau, E. J. (1971) Taxonomie und Okologie europaisch verbreiteter Tanytarsus - Arten (Chironomidae, Diptera). Archiv fur Hydrobiologie, supplement 40, 75 - 200." type="journal article" year="1971">Reiss & Fittkau 1971</bibRefCitation>
|
||
, fig. 26). The best characters separating the new species from other members of the group is the subapical tooth-like process of the gonostylus and the presence of four strong medially directed setae on median margin of the superior volsella (three strong setae are known from extant species). Moreover, in the adult male of
|
||
<taxonomicName id="4C64F4EF711BA71EFDE7AFCCD0ADF08A" box="[528,728,968,991]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="protogregarius">
|
||
<emphasis id="B910537E711BA71EFDE7AFCCD0ADF08A" box="[528,728,968,991]" italics="true" pageId="12" pageNumber="359">T. protogregarius</emphasis>
|
||
</taxonomicName>
|
||
we observed interesting proportions of leg segments. In all examined individuals femora of mid legs are longer than those of hind legs (
|
||
<tableCitation id="C6E6BAD7711BA71EFC1FAFEFD64BF751" box="[1000,1086,1003,1028]" captionStart="TABLE 3" captionStartId="12.[151,239,152,175]" captionTargetBox="[159,1404,202,359]" captionTargetPageId="12" captionText="TABLE 3. Leg segment lengths (Μm) and leg ratios of male Tanytarsus protogregarius sp. nov." httpUri="http://table.plazi.org/id/DF1BDFE4711BA71EFF60AC9CD6E6F3FA" pageId="12" pageNumber="359" tableUuid="DF1BDFE4711BA71EFF60AC9CD6E6F3FA">Table 3</tableCitation>
|
||
). Similar proportions we found in other fossil species—
|
||
<taxonomicName id="4C64F4EF711BA71EFE5FA814D188F77D" authority="Gilka et Zakrzewska, 2013" authorityName="Gilka et Zakrzewska" authorityYear="2013" box="[424,1021,1039,1064]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="congregabilis">
|
||
<emphasis id="B910537E711BA71EFE5FA814D0B1F77D" box="[424,708,1039,1064]" italics="true" pageId="12" pageNumber="359">Tanytarsus congregabilis</emphasis>
|
||
<bibRefCitation id="EFF5F29D711BA71EFD3BA80BD188F77D" author="Gilka" box="[716,1021,1039,1064]" pageId="12" pageNumber="359" refString="Gilka, W., Zakrzewska, M., Dominiak, P. & Urbanek, A. (2013 b) Non-biting midges of the tribe Tanytarsini in Eocene amber from the Rovno region (Ukraine): a pioneer systematic study with notes on the phylogeny (Diptera: Chironomidae). Zootaxa, 3736 (5), 569 - 586. http: // dx. doi. org / 10.11646 / zootaxa. 3736.5.8" type="journal article" year="2013">
|
||
Giłka
|
||
<emphasis id="B910537E711BA71EFCE5A814D153F772" box="[786,806,1040,1063]" italics="true" pageId="12" pageNumber="359">et</emphasis>
|
||
Zakrzewska, 2013
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
of the
|
||
<taxonomicName id="4C64F4EF711BA71EFBB8A80BD6EDF77D" box="[1103,1176,1039,1064]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B910537E711BA71EFBB8A80BD6EDF77D" box="[1103,1176,1039,1064]" italics="true" pageId="12" pageNumber="359">lugens</emphasis>
|
||
</taxonomicName>
|
||
group. It may indicate close relations between the two groups, which were previously proposed also to be treated as one—the concept based on adult male morphological characters of extant species (
|
||
<bibRefCitation id="EFF5F29D711BA71EFC6FA853D657F725" author="Gilka" box="[920,1058,1111,1136]" pageId="12" pageNumber="359" refString="Gilka, W. (2000) Rewizja systematyczna polskich ochotkowatych z plemienia Tanytarsini (Diptera: Chironomidae) [A systematic revision of Polish non-biting midges of the tribe Tanytarsini (Diptera: Chironomidae]. Doctoral dissertation. Department of Invertebrate Zoology, University of Gdansk, Gdynia, Poland; 154 pp. + 90 figure plates. [in Polish]" type="book" year="2000">Giłka 2000</bibRefCitation>
|
||
). The latter character discussed, however, is not known from any extant European species of these two groups [
|
||
<taxonomicName id="4C64F4EF711BA71EFBF1A878D6FBF7C6" box="[1030,1166,1148,1171]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="gregarius">
|
||
<emphasis id="B910537E711BA71EFBF1A878D6FBF7C6" box="[1030,1166,1148,1171]" italics="true" pageId="12" pageNumber="359">T. gregarius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C64F4EF711BA71EFB6CA878D2A7F7ED" authority="Lindeberg, 1970" authorityName="Lindeberg" authorityYear="1970" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="aberrans">
|
||
<emphasis id="B910537E711BA71EFB6CA878D76EF7C1" box="[1179,1307,1147,1172]" italics="true" pageId="12" pageNumber="359">T. aberrans</emphasis>
|
||
Lindeberg, 1970
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C64F4EF711BA71EFF29A8A4D027F7ED" authority="Goetghebuer, 1921" authorityName="Goetghebuer" authorityYear="1921" box="[222,594,1183,1208]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="inaequalis">
|
||
<emphasis id="B910537E711BA71EFF29A8A4D304F7ED" box="[222,369,1183,1208]" italics="true" pageId="12" pageNumber="359">T. inaequalis</emphasis>
|
||
Goetghebuer, 1921
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C64F4EF711BA71EFD66A8A4D1DEF7ED" authority="Kieffer, 1916" authorityName="Kieffer" authorityYear="1916" box="[657,939,1183,1208]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="lugens">
|
||
<emphasis id="B910537E711BA71EFD66A8A4D082F7ED" box="[657,759,1183,1208]" italics="true" pageId="12" pageNumber="359">T. lugens</emphasis>
|
||
(Kieffer, 1916)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C64F4EF711BA71EFC4EA8A4D68EF7ED" authority="Kieffer, 1911" authorityName="Kieffer" authorityYear="1911" box="[953,1275,1183,1208]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="bathophilus">
|
||
<emphasis id="B910537E711BA71EFC4EA8A4D62FF7ED" box="[953,1114,1183,1208]" italics="true" pageId="12" pageNumber="359">T. bathophilus</emphasis>
|
||
Kieffer, 1911
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C64F4EF711BA71EFAFFA8A4D337F789" authority="Edwards, 1941" authorityName="Edwards" authorityYear="1941" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="latiforceps">
|
||
<emphasis id="B910537E711BA71EFAFFA8A4D7E9F7ED" box="[1288,1436,1183,1208]" italics="true" pageId="12" pageNumber="359">T. latiforceps</emphasis>
|
||
Edwards, 1941
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C64F4EF711BA71EFEB8A8C0D0CEF789" authority="Gilka et Paasivirta, 2007" authorityName="Gilka et Paasivirta" authorityYear="2007" box="[335,699,1219,1244]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="359" phylum="Arthropoda" rank="species" species="trux">
|
||
<emphasis id="B910537E711BA71EFEB8A8C0D3E3F78E" box="[335,406,1220,1243]" italics="true" pageId="12" pageNumber="359">T. trux</emphasis>
|
||
Giłka
|
||
<emphasis id="B910537E711BA71EFE11A8C0D38FF78E" box="[486,506,1220,1243]" italics="true" pageId="12" pageNumber="359">et</emphasis>
|
||
Paasivirta, 2007
|
||
</taxonomicName>
|
||
; all examined], in which femora of mid legs are typically shorter than those of hind legs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |