210 lines
44 KiB
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210 lines
44 KiB
XML
<document id="276E5BE2408B16F102EE777316C5F9ED" ID-DOI="10.11646/phytotaxa.336.3.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13720104" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725658603287" checkinUser="felipe" docAuthor="Pennesi, Chiara, Majewska, Roksana, Sterrenburg, Frithjof A. S., Totti, Cecilia & Stefano, Mario De" docDate="2018" docId="03AC0834FFC38366FF6442E7867CAD0A" docLanguage="en" docName="phytotaxa.336.3.1.pdf" docOrigin="Phytotaxa 336 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.336.3.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Anorthoneis arthus-bertrandii De Stefano & C. Pennesi 2018, sp. nov." docType="treatment" docVersion="3" lastPageNumber="232" masterDocId="FF95704CFFDC8379FFEC42798569A827" masterDocTitle="Taxonomic revision and morphological cladistics analysis of the diatom genus Anorthoneis (Cocconeidaceae), with description of Anorthoneis arthus-bertrandii sp. nov." masterLastPageNumber="238" masterPageNumber="201" pageNumber="232" updateTime="1725670146395" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="6DAAEF9840921E88C35BC97686D662CA">
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<mods:title id="BEC1F3C43F936C9761A959ABA8135AF4">Taxonomic revision and morphological cladistics analysis of the diatom genus Anorthoneis (Cocconeidaceae), with description of Anorthoneis arthus-bertrandii sp. nov.</mods:title>
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<mods:name id="CED3F879DD1B4E2015AC631AD90A3911" type="personal">
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<mods:namePart id="C0109D8846D419E7A590CAA0118D8A4E">Pennesi, Chiara</mods:namePart>
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<mods:affiliation id="6335A58E49700A5D74A04157A4B7A8F6">Dipartimento di Scienze della Vita e dell’Ambiente, Università Politecnica delle Marche, via Brecce Bianche, 60131 Ancona, Italy</mods:affiliation>
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<mods:roleTerm id="8CEC96B2DE8B2673E46329E5073E8725">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="01284CB780B5F0A1F924CE87D7F29C60">Majewska, Roksana</mods:namePart>
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<mods:affiliation id="B37A52B064C91F6F810B0743D689A920">Unit for Environmental Sciences and Management, North-West University, Potchefstroom 2520, South Africa & Dipartimento di Scienze e Tecnologie Ambientali Biologiche e Farmaceutiche, Seconda Università di Napoli, 81100 Caserta, Italy</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="E6DA0C21CE5084A032F1D7D2BB4CAB23">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EBCE238DA18293B7A0BAB61ADCF7A59C">Sterrenburg, Frithjof A. S.</mods:namePart>
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<mods:affiliation id="C8231F52C4ABF17F2909BB8D102071EE">National Natural History Museum “ Naturalis ”, Leiden, Stationsweg 1581852 LN Heiloo, The Netherlands</mods:affiliation>
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</mods:name>
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<mods:namePart id="36A2235A9238C094E5ED78AAD9DA0235">Totti, Cecilia</mods:namePart>
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<mods:affiliation id="EDEAE53D2E00DE9FA75B5A352B607FB3">Dipartimento di Scienze della Vita e dell’Ambiente, Università Politecnica delle Marche, via Brecce Bianche, 60131 Ancona, Italy</mods:affiliation>
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<mods:namePart id="07B1B411906476F01A35046600926F60">Stefano, Mario De</mods:namePart>
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<mods:affiliation id="107F80209BCA7E79A97B39B9EA94188C">Dipartimento di Scienze e Tecnologie Ambientali Biologiche e Farmaceutiche, Seconda Università di Napoli, 81100 Caserta, Italy</mods:affiliation>
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<mods:typeOfResource id="838262465228289935BAACF9F99844E4">text</mods:typeOfResource>
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<mods:title id="F35F7B9FA2052E476383A76FF22B1B98">Phytotaxa</mods:title>
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<mods:part id="D89CA12C5BB99FDCAC8D0E86C5009D84">
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<mods:date id="1DB834D5B1C5E480F3D1C559072D4252">2018</mods:date>
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<mods:number id="5B3C32B781BDDA9D3D8175885CD1A66E">2018-02-02</mods:number>
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<mods:number id="B97B1D124DE688F68101558B0A27F357">336</mods:number>
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<mods:number id="300E2CF1FCF4E5348623BFE8605BFE13">3</mods:number>
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<mods:start id="08C4AA99A7DEEF1D5F1CBB18DBF88BDB">201</mods:start>
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<mods:url id="2BD43727AB154CB20D8FAEA3B8D1886F">http://dx.doi.org/10.11646/phytotaxa.336.3.1</mods:url>
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<mods:classification id="DA773B44BE61D0520D7DBEF3A868125A">journal article</mods:classification>
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<mods:identifier id="32DD3AADE9A813853A94AD6738A80C8F" type="DOI">10.11646/phytotaxa.336.3.1</mods:identifier>
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<mods:identifier id="3CBA227840942C89FEA522280F7C3E9A" type="ISSN">1179-3163</mods:identifier>
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<treatment id="03AC0834FFC38366FF6442E7867CAD0A" ID-DOI="http://doi.org/10.5281/zenodo.13720382" ID-Zenodo-Dep="13720382" LSID="urn:lsid:plazi:treatment:03AC0834FFC38366FF6442E7867CAD0A" httpUri="http://treatment.plazi.org/id/03AC0834FFC38366FF6442E7867CAD0A" lastPageNumber="232" pageId="31" pageNumber="232">
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<subSubSection id="C31FEAA9FFC38366FF6442E78163A89E" box="[136,1034,158,185]" pageId="31" pageNumber="232" type="nomenclature">
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<paragraph id="8BBAB922FFC38366FF6442E78163A89E" blockId="31.[136,1034,158,185]" box="[136,1034,158,185]" pageId="31" pageNumber="232">
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<heading id="D0F20E4EFFC38366FF6442E78163A89E" box="[136,1034,158,185]" fontSize="11" level="2" pageId="31" pageNumber="232" reason="3">
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<emphasis id="B9716530FFC38366FF6442E7863EA89E" bold="true" box="[136,855,158,185]" pageId="31" pageNumber="232">
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<taxonomicName id="4C05C2A1FFC38366FF6442E7879BA89E" authority="De Stefano & Pennesi" authorityName="De Stefano & C. Pennesi" authorityYear="2018" box="[136,754,158,185]" class="Bacillariophyceae" family="Cocconeidaceae" genus="Anorthoneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="31" pageNumber="232" phylum="Ochrophyta" rank="species" species="arthus-bertrandii" status="sp. nov.">
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||
<emphasis id="B9716530FFC38366FF6442E7848CA89E" bold="true" box="[136,485,158,185]" italics="true" pageId="31" pageNumber="232">Anorthoneis arthus-bertrandii</emphasis>
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De Stefano & Pennesi
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</taxonomicName>
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||
,
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<taxonomicNameLabel id="A242D84BFFC38366FD1142D9863EA89E" box="[765,855,160,185]" pageId="31" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
(
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||
<figureCitation id="133EA5A7FFC38366FC8A42D9816BA89E" box="[870,1026,160,185]" captionStart-0="FIGURES 130–135" captionStart-1="FIGURES 136–142" captionStart-2="FIGURE 143" captionStartId-0="29.[136,243,1748,1769]" captionStartId-1="30.[136,243,1851,1872]" captionStartId-2="31.[136,229,1993,2014]" captionTargetBox-0="[162,1430,201,1714]" captionTargetBox-1="[224,1362,178,1826]" captionTargetBox-2="[179,1403,1341,1963]" captionTargetId-0="figure-14@29.[151,1436,190,1723]" captionTargetId-1="figure-15@30.[224,1363,175,1826]" captionTargetId-2="figure-538@31.[177,1411,1333,1968]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionTargetPageId-2="31" captionText-0="FIGURES 130–135. Anorthoneis arthus-bertrandii De Stefano & C. Pennesi, sp. nov., SEM images. (130) External view of sternum valve; (131) Detail of sternum valve showing the presence of vestigial perforations in hyaline-like central area; (132). Detail of the sternum valve mantle showing biseriate striae close to the valve margin; (133) Internal view of sternum valve; (134) Detail of the sternum valve (internally) showing the eccentric central area with vestigial perforations; (135) Detail of circular areolae internally occluded by rota-like vela. Figs 130, 133 = 5 μm; Fig. 134 = 2 μm; Figs 131, 132, 135 = 1 μm." captionText-1="FIGURES 136–142. Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov., SEM images. (136) External raphe-sternum valve with asymmetric raphe branches; (137) Detail of coaxial central raphe endings (externally); (138) Detail of external distal raphe ending terminating in a roundish submarginal hyaline area (externally); (139) Internal view of raphe-sternum valve showing the thickening of sternum and small central nodule; (140) Detail of the internal, uncinated proximal raphe endings converging to the central nodule; (141) Detail of distal raphe ending terminating in a vestigial helictoglossa (internally). (142) Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov. from slide MDS 9 (Holotype), LM images. Scale bars: Fig. 140 = 10 μm; Figs 139, 142 = 5 μm; Fig. 137 = 2 μm; Figs 136, 138, 141 = 1 μm." captionText-2="FIGURE 143. Relationship tree of the genus Anorthoneis generated using maximum parsimony method (Cladistic analysis). The black circles indicate probable homology changes, and the white circles represent probable homoplasy or ancestrally-derived homology changes. Circle number, top = character number (Table 3); circle number, bottom = character state (Table 3)." figureDoi-0="http://doi.org/10.5281/zenodo.13720166" figureDoi-1="http://doi.org/10.5281/zenodo.13720168" figureDoi-2="http://doi.org/10.5281/zenodo.13720172" httpUri-0="https://zenodo.org/record/13720166/files/figure.png" httpUri-1="https://zenodo.org/record/13720168/files/figure.png" httpUri-2="https://zenodo.org/record/13720172/files/figure.png" pageId="31" pageNumber="232">Figs 130–143</figureCitation>
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||
)
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31FEAA9FFC38366FF6442BC8466A98D" pageId="31" pageNumber="232" type="diagnosis">
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||
<paragraph id="8BBAB922FFC38366FF6442BC8466A98D" blockId="31.[136,1452,197,566]" pageId="31" pageNumber="232">
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||
<emphasis id="B9716530FFC38366FF6442BC8462A8FD" bold="true" box="[136,267,197,218]" pageId="31" pageNumber="232">Diagnosis:—</emphasis>
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<emphasis id="B9716530FFC38366FEE742BC8466A98D" italics="true" pageId="31" pageNumber="232">
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valvae late ellipticae, 16–22 μm longae, 13–18 μm latae. Areovalva concava in area axiali et leviter convexa in area marginali. Sternum excentrico, lineare cum nodulo centrali conspicuo et eccentrico. Areovalva habet in valvae lato structuram striis cum monoseriatis (
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<quantity id="4CFD14C7FFC38366FE434373876FA938" box="[431,518,266,287]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" metricValueMax="4.572" metricValueMin="3.556" pageId="31" pageNumber="232" unit="in" value="16.0" valueMax="18.0" valueMin="14.0">14–18 in</quantity>
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10 μm) quae constant ex areolis oblongis (
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<quantity id="4CFD14C7FFC38366FC5C43738161A938" box="[944,1032,266,287]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" metricValueMax="3.556" metricValueMin="2.54" pageId="31" pageNumber="232" unit="in" value="12.0" valueMax="14.0" valueMin="10.0">10–14 in</quantity>
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10 μm). Implicatae rotae partes exteriores areolarum occludunt. Areovalva habet in limbo structuram striis biseriatis quae constant ex poroidis qui occlusi sunt hymenibus rimis cum inaequalibus. Raphovalva habet raphem rectam cum nodulo centrali et nodulis terminalibus oblongis. Striae monoseriatae (
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||
<quantity id="4CFD14C7FFC38366FF28430B8473A9A0" box="[196,282,370,391]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" metricValueMax="5.08" metricValueMin="4.572" pageId="31" pageNumber="232" unit="in" value="19.0" valueMax="20.0" valueMin="18.0">18–20 in</quantity>
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10 μm) quae constant ex poroidis qui occlusi sunt hymenibus rimis cum inaequalibus. Utraque valvocopula secta et caret fimbriis.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31FEAA9FFC38366FF6443C18106AA11" pageId="31" pageNumber="232" type="materials_examined">
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<paragraph id="8BBAB922FFC38366FF6443C184CFA9C8" blockId="31.[136,1452,197,566]" pageId="31" pageNumber="232">
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<materialsCitation id="3B6DB37FFFC38366FF6443C184CFA9C8" country="Italy" location="Italy" pageId="31" pageNumber="232" specimenCount="2" typeStatus="holotype">
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<emphasis id="B9716530FFC38366FF6443C1846CA9EA" bold="true" box="[136,261,440,461]" pageId="31" pageNumber="232">
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<typeStatus id="54BE0780FFC38366FF6443C18581A9EA" box="[136,232,440,461]" pageId="31" pageNumber="232" type="holotype">Holotype</typeStatus>
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:—
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</emphasis>
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slide MDS 9 deposited at the Herbarium of the Stazione Zoologica “Anton Dohrn” of Naples,
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<collectingCountry id="F312F9B2FFC38366FB9B43C181CCA9EA" box="[1143,1189,440,461]" name="Italy" pageId="31" pageNumber="232">Italy</collectingCountry>
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.
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<figureCitation id="133EA5A7FFC38366FB4743C18077A9EA" box="[1195,1310,440,461]" captionStart="FIGURES 136–142" captionStartId="30.[136,243,1851,1872]" captionTargetBox="[224,1362,178,1826]" captionTargetId="figure-15@30.[224,1363,175,1826]" captionTargetPageId="30" captionText="FIGURES 136–142. Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov., SEM images. (136) External raphe-sternum valve with asymmetric raphe branches; (137) Detail of coaxial central raphe endings (externally); (138) Detail of external distal raphe ending terminating in a roundish submarginal hyaline area (externally); (139) Internal view of raphe-sternum valve showing the thickening of sternum and small central nodule; (140) Detail of the internal, uncinated proximal raphe endings converging to the central nodule; (141) Detail of distal raphe ending terminating in a vestigial helictoglossa (internally). (142) Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov. from slide MDS 9 (Holotype), LM images. Scale bars: Fig. 140 = 10 μm; Figs 139, 142 = 5 μm; Fig. 137 = 2 μm; Figs 136, 138, 141 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720168" httpUri="https://zenodo.org/record/13720168/files/figure.png" pageId="31" pageNumber="232">Figures 142</figureCitation>
|
||
,
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||
<figureCitation id="133EA5A7FFC38366FACA43C18023A9EA" box="[1318,1354,440,461]" captionStart="FIGURE 143" captionStartId="31.[136,229,1993,2014]" captionTargetBox="[179,1403,1341,1963]" captionTargetId="figure-538@31.[177,1411,1333,1968]" captionTargetPageId="31" captionText="FIGURE 143. Relationship tree of the genus Anorthoneis generated using maximum parsimony method (Cladistic analysis). The black circles indicate probable homology changes, and the white circles represent probable homoplasy or ancestrally-derived homology changes. Circle number, top = character number (Table 3); circle number, bottom = character state (Table 3)." figureDoi="http://doi.org/10.5281/zenodo.13720172" httpUri="https://zenodo.org/record/13720172/files/figure.png" pageId="31" pageNumber="232">143</figureCitation>
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illustrates the
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<typeStatus id="54BE0780FFC38366FF0D43A3845CA9C8" box="[225,309,474,495]" pageId="31" pageNumber="232" type="holotype">holotype</typeStatus>
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specimens.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BBAB922FFC38366FF6443848106AA11" blockId="31.[136,1452,197,566]" pageId="31" pageNumber="232">
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||
<emphasis id="B9716530FFC38366FF6443848444AA35" bold="true" box="[136,301,509,530]" pageId="31" pageNumber="232">Type locality:—</emphasis>
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<collectingCountry id="F312F9B2FFC38366FEC1438484DAAA35" box="[301,435,509,530]" name="Indonesia" pageId="31" pageNumber="232">INDONESIA</collectingCountry>
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. Epiphytic on seagrasses leaves collected in
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<date id="FFBB9FE2FFC38366FC8643848686AA35" box="[874,1007,509,530]" pageId="31" pageNumber="232" value="2005-10">October 2005</date>
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along the coasts of Siladen Island,
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<collectingRegion id="49C177C0FFC38366FAA3438480C1AA35" box="[1359,1448,509,530]" country="Indonesia" name="Sulawesi Utara" pageId="31" pageNumber="232">Sulawesi</collectingRegion>
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, Indonesia, (
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<geoCoordinate id="EE31DFE5FFC38366FEDC405884AEAA11" box="[304,455,545,566]" degrees="1" direction="north" minutes="38" orientation="latitude" pageId="31" pageNumber="232" precision="1" seconds="25.23" value="1.6403416">1° 38′ 25.23″ N</geoCoordinate>
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,
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<geoCoordinate id="EE31DFE5FFC38366FE3D405887E9AA11" box="[465,640,545,566]" degrees="124" direction="east" minutes="48" orientation="longitude" pageId="31" pageNumber="232" precision="1" seconds="06.61" value="124.80184">124° 48′ 06.61″ E</geoCoordinate>
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). Collected by Dr. Francesca Azzini,
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<date id="FFBB9FE2FFC38366FC0940588102AA11" box="[997,1131,545,566]" pageId="31" pageNumber="232" value="2005-10">October 2005</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31FEAA9FFC38366FF644027866FAABD" pageId="31" pageNumber="232" type="etymology">
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<paragraph id="8BBAB922FFC38366FF644027866FAABD" blockId="31.[136,1452,606,1325]" pageId="31" pageNumber="232">
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<emphasis id="B9716530FFC38366FF6440278447AA50" bold="true" box="[136,302,606,631]" pageId="31" pageNumber="232">Etymology:—</emphasis>
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the specific epithet refers to Yann Arthus-Bertrand in recognition of his outstanding service in the promotion of scientific knowledge and its dissemination.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31FEAA9FFC38366FF5140DD867CAD0A" pageId="31" pageNumber="232" type="description">
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<paragraph id="8BBAB922FFC38366FF5140DD8754AAF8" blockId="31.[136,1452,606,1325]" pageId="31" pageNumber="232">
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<emphasis id="B9716530FFC38366FF5140DD8404AA9A" bold="true" box="[189,365,676,701]" pageId="31" pageNumber="232">Description:—</emphasis>
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Valves broadly elliptical to ovoid in outline (
|
||
<figureCitation id="133EA5A7FFC38366FC8A40DD816DAA9A" box="[870,1028,676,701]" captionStart="FIGURES 130–135" captionStartId="29.[136,243,1748,1769]" captionTargetBox="[162,1430,201,1714]" captionTargetId="figure-14@29.[151,1436,190,1723]" captionTargetPageId="29" captionText="FIGURES 130–135. Anorthoneis arthus-bertrandii De Stefano & C. Pennesi, sp. nov., SEM images. (130) External view of sternum valve; (131) Detail of sternum valve showing the presence of vestigial perforations in hyaline-like central area; (132). Detail of the sternum valve mantle showing biseriate striae close to the valve margin; (133) Internal view of sternum valve; (134) Detail of the sternum valve (internally) showing the eccentric central area with vestigial perforations; (135) Detail of circular areolae internally occluded by rota-like vela. Figs 130, 133 = 5 μm; Fig. 134 = 2 μm; Figs 131, 132, 135 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720166" httpUri="https://zenodo.org/record/13720166/files/figure.png" pageId="31" pageNumber="232">Figs 130, 133</figureCitation>
|
||
,
|
||
<figureCitation id="133EA5A7FFC38366FBFD40DD81C5AA9A" box="[1041,1196,676,701]" captionStart="FIGURES 136–142" captionStartId="30.[136,243,1851,1872]" captionTargetBox="[224,1362,178,1826]" captionTargetId="figure-15@30.[224,1363,175,1826]" captionTargetPageId="30" captionText="FIGURES 136–142. Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov., SEM images. (136) External raphe-sternum valve with asymmetric raphe branches; (137) Detail of coaxial central raphe endings (externally); (138) Detail of external distal raphe ending terminating in a roundish submarginal hyaline area (externally); (139) Internal view of raphe-sternum valve showing the thickening of sternum and small central nodule; (140) Detail of the internal, uncinated proximal raphe endings converging to the central nodule; (141) Detail of distal raphe ending terminating in a vestigial helictoglossa (internally). (142) Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov. from slide MDS 9 (Holotype), LM images. Scale bars: Fig. 140 = 10 μm; Figs 139, 142 = 5 μm; Fig. 137 = 2 μm; Figs 136, 138, 141 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720168" httpUri="https://zenodo.org/record/13720168/files/figure.png" pageId="31" pageNumber="232">136, 139, 142</figureCitation>
|
||
and
|
||
<figureCitation id="133EA5A7FFC38366FB0840DD807BAA9A" box="[1252,1298,676,701]" captionStart="FIGURE 143" captionStartId="31.[136,229,1993,2014]" captionTargetBox="[179,1403,1341,1963]" captionTargetId="figure-538@31.[177,1411,1333,1968]" captionTargetPageId="31" captionText="FIGURE 143. Relationship tree of the genus Anorthoneis generated using maximum parsimony method (Cladistic analysis). The black circles indicate probable homology changes, and the white circles represent probable homoplasy or ancestrally-derived homology changes. Circle number, top = character number (Table 3); circle number, bottom = character state (Table 3)." figureDoi="http://doi.org/10.5281/zenodo.13720172" httpUri="https://zenodo.org/record/13720172/files/figure.png" pageId="31" pageNumber="232">143</figureCitation>
|
||
). Apical axis: 16–22 μm, transapical axis: 13–18 μm.
|
||
</paragraph>
|
||
<paragraph id="8BBAB922FFC38366FF51409E84B9ABF5" blockId="31.[136,1452,606,1325]" pageId="31" pageNumber="232">
|
||
<emphasis id="B9716530FFC38366FF51409E846CAB25" bold="true" box="[189,261,743,770]" pageId="31" pageNumber="232">
|
||
<emphasis id="B9716530FFC38366FF51409E85B6AB26" bold="true" box="[189,223,743,769]" italics="true" pageId="31" pageNumber="232">SV</emphasis>
|
||
:—
|
||
</emphasis>
|
||
Externally, valve face slightly concave with a convex mantle, ending in a small marginal area covered by a row of very small pores (
|
||
<figureCitation id="133EA5A7FFC38366FE5741758777AB02" box="[443,542,780,805]" captionStart="FIGURES 130–135" captionStartId="29.[136,243,1748,1769]" captionTargetBox="[162,1430,201,1714]" captionTargetId="figure-14@29.[151,1436,190,1723]" captionTargetPageId="29" captionText="FIGURES 130–135. Anorthoneis arthus-bertrandii De Stefano & C. Pennesi, sp. nov., SEM images. (130) External view of sternum valve; (131) Detail of sternum valve showing the presence of vestigial perforations in hyaline-like central area; (132). Detail of the sternum valve mantle showing biseriate striae close to the valve margin; (133) Internal view of sternum valve; (134) Detail of the sternum valve (internally) showing the eccentric central area with vestigial perforations; (135) Detail of circular areolae internally occluded by rota-like vela. Figs 130, 133 = 5 μm; Fig. 134 = 2 μm; Figs 131, 132, 135 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720166" httpUri="https://zenodo.org/record/13720166/files/figure.png" pageId="31" pageNumber="232">Fig. 130</figureCitation>
|
||
). The eccentric sternum narrow, linear, developed entirely within the valve face, terminating close to the valve face-mantle junction (
|
||
<figureCitation id="133EA5A7FFC38366FD384157865FAB60" box="[724,822,814,839]" captionStart="FIGURES 130–135" captionStartId="29.[136,243,1748,1769]" captionTargetBox="[162,1430,201,1714]" captionTargetId="figure-14@29.[151,1436,190,1723]" captionTargetPageId="29" captionText="FIGURES 130–135. Anorthoneis arthus-bertrandii De Stefano & C. Pennesi, sp. nov., SEM images. (130) External view of sternum valve; (131) Detail of sternum valve showing the presence of vestigial perforations in hyaline-like central area; (132). Detail of the sternum valve mantle showing biseriate striae close to the valve margin; (133) Internal view of sternum valve; (134) Detail of the sternum valve (internally) showing the eccentric central area with vestigial perforations; (135) Detail of circular areolae internally occluded by rota-like vela. Figs 130, 133 = 5 μm; Fig. 134 = 2 μm; Figs 131, 132, 135 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720166" httpUri="https://zenodo.org/record/13720166/files/figure.png" pageId="31" pageNumber="232">Fig. 130</figureCitation>
|
||
). Central area small, circular, with very small, irregular, imperforated pores (
|
||
<figureCitation id="133EA5A7FFC38366FE864128876FAB4D" box="[362,518,849,874]" captionStart="FIGURES 130–135" captionStartId="29.[136,243,1748,1769]" captionTargetBox="[162,1430,201,1714]" captionTargetId="figure-14@29.[151,1436,190,1723]" captionTargetPageId="29" captionText="FIGURES 130–135. Anorthoneis arthus-bertrandii De Stefano & C. Pennesi, sp. nov., SEM images. (130) External view of sternum valve; (131) Detail of sternum valve showing the presence of vestigial perforations in hyaline-like central area; (132). Detail of the sternum valve mantle showing biseriate striae close to the valve margin; (133) Internal view of sternum valve; (134) Detail of the sternum valve (internally) showing the eccentric central area with vestigial perforations; (135) Detail of circular areolae internally occluded by rota-like vela. Figs 130, 133 = 5 μm; Fig. 134 = 2 μm; Figs 131, 132, 135 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720166" httpUri="https://zenodo.org/record/13720166/files/figure.png" pageId="31" pageNumber="232">Figs 130, 131</figureCitation>
|
||
). Transapical striae strongly radiate,
|
||
<quantity id="4CFD14C7FFC38366FC4C4128816BAB4D" box="[928,1026,849,874]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" metricValueMax="4.572" metricValueMin="3.556" pageId="31" pageNumber="232" unit="in" value="16.0" valueMax="18.0" valueMin="14.0">14–18 in</quantity>
|
||
10 μm, uniseriate, becoming biseriate in the last row of the mantle striae (
|
||
<figureCitation id="133EA5A7FFC38366FDCA410D87E3ABAA" box="[550,650,884,909]" captionStart="FIGURES 130–135" captionStartId="29.[136,243,1748,1769]" captionTargetBox="[162,1430,201,1714]" captionTargetId="figure-14@29.[151,1436,190,1723]" captionTargetPageId="29" captionText="FIGURES 130–135. Anorthoneis arthus-bertrandii De Stefano & C. Pennesi, sp. nov., SEM images. (130) External view of sternum valve; (131) Detail of sternum valve showing the presence of vestigial perforations in hyaline-like central area; (132). Detail of the sternum valve mantle showing biseriate striae close to the valve margin; (133) Internal view of sternum valve; (134) Detail of the sternum valve (internally) showing the eccentric central area with vestigial perforations; (135) Detail of circular areolae internally occluded by rota-like vela. Figs 130, 133 = 5 μm; Fig. 134 = 2 μm; Figs 131, 132, 135 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720166" httpUri="https://zenodo.org/record/13720166/files/figure.png" pageId="31" pageNumber="232">Fig. 132</figureCitation>
|
||
). Areolae circular to squared,
|
||
<quantity id="4CFD14C7FFC38366FC08410D8120ABAA" box="[996,1097,884,909]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" metricValueMax="3.556" metricValueMin="2.54" pageId="31" pageNumber="232" unit="in" value="12.0" valueMax="14.0" valueMin="10.0">10–14 in</quantity>
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10 μm, internally occluded by rota-like vela (
|
||
<figureCitation id="133EA5A7FFC38366FEC641EF84A0AB88" box="[298,457,918,943]" captionStart="FIGURES 130–135" captionStartId="29.[136,243,1748,1769]" captionTargetBox="[162,1430,201,1714]" captionTargetId="figure-14@29.[151,1436,190,1723]" captionTargetPageId="29" captionText="FIGURES 130–135. Anorthoneis arthus-bertrandii De Stefano & C. Pennesi, sp. nov., SEM images. (130) External view of sternum valve; (131) Detail of sternum valve showing the presence of vestigial perforations in hyaline-like central area; (132). Detail of the sternum valve mantle showing biseriate striae close to the valve margin; (133) Internal view of sternum valve; (134) Detail of the sternum valve (internally) showing the eccentric central area with vestigial perforations; (135) Detail of circular areolae internally occluded by rota-like vela. Figs 130, 133 = 5 μm; Fig. 134 = 2 μm; Figs 131, 132, 135 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720166" httpUri="https://zenodo.org/record/13720166/files/figure.png" pageId="31" pageNumber="232">Figs 131–135</figureCitation>
|
||
). Internally, the valve surface structures appear similar to the external ones, except for the larger central area (
|
||
<figureCitation id="133EA5A7FFC38366FE8D41C084AAABF5" box="[353,451,953,978]" captionStart="FIGURES 130–135" captionStartId="29.[136,243,1748,1769]" captionTargetBox="[162,1430,201,1714]" captionTargetId="figure-14@29.[151,1436,190,1723]" captionTargetPageId="29" captionText="FIGURES 130–135. Anorthoneis arthus-bertrandii De Stefano & C. Pennesi, sp. nov., SEM images. (130) External view of sternum valve; (131) Detail of sternum valve showing the presence of vestigial perforations in hyaline-like central area; (132). Detail of the sternum valve mantle showing biseriate striae close to the valve margin; (133) Internal view of sternum valve; (134) Detail of the sternum valve (internally) showing the eccentric central area with vestigial perforations; (135) Detail of circular areolae internally occluded by rota-like vela. Figs 130, 133 = 5 μm; Fig. 134 = 2 μm; Figs 131, 132, 135 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720166" httpUri="https://zenodo.org/record/13720166/files/figure.png" pageId="31" pageNumber="232">Fig. 133</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBAB922FFC38366FF5141A3867CAD0A" blockId="31.[136,1452,606,1325]" pageId="31" pageNumber="232">
|
||
<emphasis id="B9716530FFC38366FF5141A38471ABD2" bold="true" box="[189,280,986,1013]" pageId="31" pageNumber="232">
|
||
<emphasis id="B9716530FFC38366FF5141A3859BABD3" bold="true" box="[189,242,986,1012]" italics="true" pageId="31" pageNumber="232">RSV</emphasis>
|
||
:—
|
||
</emphasis>
|
||
Externally, valve concave to convex close to the margin. The margin is covered by two irregular rows of very small pores (
|
||
<figureCitation id="133EA5A7FFC38366FE9741878489AC30" box="[379,480,1022,1047]" captionStart="FIGURES 136–142" captionStartId="30.[136,243,1851,1872]" captionTargetBox="[224,1362,178,1826]" captionTargetId="figure-15@30.[224,1363,175,1826]" captionTargetPageId="30" captionText="FIGURES 136–142. Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov., SEM images. (136) External raphe-sternum valve with asymmetric raphe branches; (137) Detail of coaxial central raphe endings (externally); (138) Detail of external distal raphe ending terminating in a roundish submarginal hyaline area (externally); (139) Internal view of raphe-sternum valve showing the thickening of sternum and small central nodule; (140) Detail of the internal, uncinated proximal raphe endings converging to the central nodule; (141) Detail of distal raphe ending terminating in a vestigial helictoglossa (internally). (142) Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov. from slide MDS 9 (Holotype), LM images. Scale bars: Fig. 140 = 10 μm; Figs 139, 142 = 5 μm; Fig. 137 = 2 μm; Figs 136, 138, 141 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720168" httpUri="https://zenodo.org/record/13720168/files/figure.png" pageId="31" pageNumber="232">Fig. 136</figureCitation>
|
||
). Raphe-sternum asymmetric, narrow and linear developed entirely within the valve face (
|
||
<figureCitation id="133EA5A7FFC38366FF2B46588442AC1D" box="[199,299,1057,1082]" captionStart="FIGURES 136–142" captionStartId="30.[136,243,1851,1872]" captionTargetBox="[224,1362,178,1826]" captionTargetId="figure-15@30.[224,1363,175,1826]" captionTargetPageId="30" captionText="FIGURES 136–142. Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov., SEM images. (136) External raphe-sternum valve with asymmetric raphe branches; (137) Detail of coaxial central raphe endings (externally); (138) Detail of external distal raphe ending terminating in a roundish submarginal hyaline area (externally); (139) Internal view of raphe-sternum valve showing the thickening of sternum and small central nodule; (140) Detail of the internal, uncinated proximal raphe endings converging to the central nodule; (141) Detail of distal raphe ending terminating in a vestigial helictoglossa (internally). (142) Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov. from slide MDS 9 (Holotype), LM images. Scale bars: Fig. 140 = 10 μm; Figs 139, 142 = 5 μm; Fig. 137 = 2 μm; Figs 136, 138, 141 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720168" httpUri="https://zenodo.org/record/13720168/files/figure.png" pageId="31" pageNumber="232">Fig. 136</figureCitation>
|
||
). Proximal raphe endings slightly expanded, coaxial, converging within a small, roundish and clearly asymmetrical central area (
|
||
<figureCitation id="133EA5A7FFC38366FE57463D8734AC7A" box="[443,605,1092,1117]" captionStart="FIGURES 136–142" captionStartId="30.[136,243,1851,1872]" captionTargetBox="[224,1362,178,1826]" captionTargetId="figure-15@30.[224,1363,175,1826]" captionTargetPageId="30" captionText="FIGURES 136–142. Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov., SEM images. (136) External raphe-sternum valve with asymmetric raphe branches; (137) Detail of coaxial central raphe endings (externally); (138) Detail of external distal raphe ending terminating in a roundish submarginal hyaline area (externally); (139) Internal view of raphe-sternum valve showing the thickening of sternum and small central nodule; (140) Detail of the internal, uncinated proximal raphe endings converging to the central nodule; (141) Detail of distal raphe ending terminating in a vestigial helictoglossa (internally). (142) Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov. from slide MDS 9 (Holotype), LM images. Scale bars: Fig. 140 = 10 μm; Figs 139, 142 = 5 μm; Fig. 137 = 2 μm; Figs 136, 138, 141 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720168" httpUri="https://zenodo.org/record/13720168/files/figure.png" pageId="31" pageNumber="232">Figs 136, 137</figureCitation>
|
||
). Distal raphe endings similar in shape, terminate in roundish submarginal areas (
|
||
<figureCitation id="133EA5A7FFC38366FF38461F8413AC58" box="[212,378,1126,1151]" captionStart="FIGURES 136–142" captionStartId="30.[136,243,1851,1872]" captionTargetBox="[224,1362,178,1826]" captionTargetId="figure-15@30.[224,1363,175,1826]" captionTargetPageId="30" captionText="FIGURES 136–142. Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov., SEM images. (136) External raphe-sternum valve with asymmetric raphe branches; (137) Detail of coaxial central raphe endings (externally); (138) Detail of external distal raphe ending terminating in a roundish submarginal hyaline area (externally); (139) Internal view of raphe-sternum valve showing the thickening of sternum and small central nodule; (140) Detail of the internal, uncinated proximal raphe endings converging to the central nodule; (141) Detail of distal raphe ending terminating in a vestigial helictoglossa (internally). (142) Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov. from slide MDS 9 (Holotype), LM images. Scale bars: Fig. 140 = 10 μm; Figs 139, 142 = 5 μm; Fig. 137 = 2 μm; Figs 136, 138, 141 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720168" httpUri="https://zenodo.org/record/13720168/files/figure.png" pageId="31" pageNumber="232">Figs 136, 138</figureCitation>
|
||
). Transapical striae strongly radiate,
|
||
<quantity id="4CFD14C7FFC38366FCC4461F86E6AC58" box="[808,911,1126,1151]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" metricValueMax="5.08" metricValueMin="4.572" pageId="31" pageNumber="232" unit="in" value="19.0" valueMax="20.0" valueMin="18.0">18–20 in</quantity>
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||
10 μm, uniseriate in the valve face becoming biseriate in the mantle (
|
||
<figureCitation id="133EA5A7FFC38366FE6646F08482AC85" box="[394,491,1161,1186]" captionStart="FIGURES 136–142" captionStartId="30.[136,243,1851,1872]" captionTargetBox="[224,1362,178,1826]" captionTargetId="figure-15@30.[224,1363,175,1826]" captionTargetPageId="30" captionText="FIGURES 136–142. Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov., SEM images. (136) External raphe-sternum valve with asymmetric raphe branches; (137) Detail of coaxial central raphe endings (externally); (138) Detail of external distal raphe ending terminating in a roundish submarginal hyaline area (externally); (139) Internal view of raphe-sternum valve showing the thickening of sternum and small central nodule; (140) Detail of the internal, uncinated proximal raphe endings converging to the central nodule; (141) Detail of distal raphe ending terminating in a vestigial helictoglossa (internally). (142) Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov. from slide MDS 9 (Holotype), LM images. Scale bars: Fig. 140 = 10 μm; Figs 139, 142 = 5 μm; Fig. 137 = 2 μm; Figs 136, 138, 141 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720168" httpUri="https://zenodo.org/record/13720168/files/figure.png" pageId="31" pageNumber="232">Fig. 136</figureCitation>
|
||
). Areolae small, rounded,
|
||
<quantity id="4CFD14C7FFC38366FCE046F08607AC85" box="[780,878,1161,1186]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" metricValueMax="6.096" metricValueMin="5.588" pageId="31" pageNumber="232" unit="in" value="23.0" valueMax="24.0" valueMin="22.0">22–24 in</quantity>
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||
10 μm, internally occluded by perforated vela (
|
||
<figureCitation id="133EA5A7FFC38366FA9646F0844DACE2" captionStart="FIGURES 136–142" captionStartId="30.[136,243,1851,1872]" captionTargetBox="[224,1362,178,1826]" captionTargetId="figure-15@30.[224,1363,175,1826]" captionTargetPageId="30" captionText="FIGURES 136–142. Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov., SEM images. (136) External raphe-sternum valve with asymmetric raphe branches; (137) Detail of coaxial central raphe endings (externally); (138) Detail of external distal raphe ending terminating in a roundish submarginal hyaline area (externally); (139) Internal view of raphe-sternum valve showing the thickening of sternum and small central nodule; (140) Detail of the internal, uncinated proximal raphe endings converging to the central nodule; (141) Detail of distal raphe ending terminating in a vestigial helictoglossa (internally). (142) Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov. from slide MDS 9 (Holotype), LM images. Scale bars: Fig. 140 = 10 μm; Figs 139, 142 = 5 μm; Fig. 137 = 2 μm; Figs 136, 138, 141 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720168" httpUri="https://zenodo.org/record/13720168/files/figure.png" pageId="31" pageNumber="232">Figs 139, 140, 141</figureCitation>
|
||
). On the mantle margin, at the end of each biseriate striae, one smaller areola is present (
|
||
<figureCitation id="133EA5A7FFC38366FAED46D580F6ACE2" box="[1281,1439,1196,1221]" captionStart="FIGURES 136–142" captionStartId="30.[136,243,1851,1872]" captionTargetBox="[224,1362,178,1826]" captionTargetId="figure-15@30.[224,1363,175,1826]" captionTargetPageId="30" captionText="FIGURES 136–142. Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov., SEM images. (136) External raphe-sternum valve with asymmetric raphe branches; (137) Detail of coaxial central raphe endings (externally); (138) Detail of external distal raphe ending terminating in a roundish submarginal hyaline area (externally); (139) Internal view of raphe-sternum valve showing the thickening of sternum and small central nodule; (140) Detail of the internal, uncinated proximal raphe endings converging to the central nodule; (141) Detail of distal raphe ending terminating in a vestigial helictoglossa (internally). (142) Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov. from slide MDS 9 (Holotype), LM images. Scale bars: Fig. 140 = 10 μm; Figs 139, 142 = 5 μm; Fig. 137 = 2 μm; Figs 136, 138, 141 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720168" httpUri="https://zenodo.org/record/13720168/files/figure.png" pageId="31" pageNumber="232">Figs 136, 138</figureCitation>
|
||
). Internally, both the raphe-sternum and the small central nodule only slightly thickened (
|
||
<figureCitation id="133EA5A7FFC38366FBBE46B78187ACC0" box="[1106,1262,1230,1255]" captionStart="FIGURES 136–142" captionStartId="30.[136,243,1851,1872]" captionTargetBox="[224,1362,178,1826]" captionTargetId="figure-15@30.[224,1363,175,1826]" captionTargetPageId="30" captionText="FIGURES 136–142. Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov., SEM images. (136) External raphe-sternum valve with asymmetric raphe branches; (137) Detail of coaxial central raphe endings (externally); (138) Detail of external distal raphe ending terminating in a roundish submarginal hyaline area (externally); (139) Internal view of raphe-sternum valve showing the thickening of sternum and small central nodule; (140) Detail of the internal, uncinated proximal raphe endings converging to the central nodule; (141) Detail of distal raphe ending terminating in a vestigial helictoglossa (internally). (142) Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov. from slide MDS 9 (Holotype), LM images. Scale bars: Fig. 140 = 10 μm; Figs 139, 142 = 5 μm; Fig. 137 = 2 μm; Figs 136, 138, 141 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720168" httpUri="https://zenodo.org/record/13720168/files/figure.png" pageId="31" pageNumber="232">Figs 139, 140</figureCitation>
|
||
). Proximal raphe endings simple, deflected in opposite directions (
|
||
<figureCitation id="133EA5A7FFC38366FD4746888620AD2D" box="[683,841,1265,1290]" captionStart="FIGURES 136–142" captionStartId="30.[136,243,1851,1872]" captionTargetBox="[224,1362,178,1826]" captionTargetId="figure-15@30.[224,1363,175,1826]" captionTargetPageId="30" captionText="FIGURES 136–142. Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov., SEM images. (136) External raphe-sternum valve with asymmetric raphe branches; (137) Detail of coaxial central raphe endings (externally); (138) Detail of external distal raphe ending terminating in a roundish submarginal hyaline area (externally); (139) Internal view of raphe-sternum valve showing the thickening of sternum and small central nodule; (140) Detail of the internal, uncinated proximal raphe endings converging to the central nodule; (141) Detail of distal raphe ending terminating in a vestigial helictoglossa (internally). (142) Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov. from slide MDS 9 (Holotype), LM images. Scale bars: Fig. 140 = 10 μm; Figs 139, 142 = 5 μm; Fig. 137 = 2 μm; Figs 136, 138, 141 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720168" httpUri="https://zenodo.org/record/13720168/files/figure.png" pageId="31" pageNumber="232">Figs 139, 140</figureCitation>
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); distal ones terminate in vestigial helictoglossae (
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<figureCitation id="133EA5A7FFC38366FA9646888584AD0A" captionStart="FIGURES 136–142" captionStartId="30.[136,243,1851,1872]" captionTargetBox="[224,1362,178,1826]" captionTargetId="figure-15@30.[224,1363,175,1826]" captionTargetPageId="30" captionText="FIGURES 136–142. Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov., SEM images. (136) External raphe-sternum valve with asymmetric raphe branches; (137) Detail of coaxial central raphe endings (externally); (138) Detail of external distal raphe ending terminating in a roundish submarginal hyaline area (externally); (139) Internal view of raphe-sternum valve showing the thickening of sternum and small central nodule; (140) Detail of the internal, uncinated proximal raphe endings converging to the central nodule; (141) Detail of distal raphe ending terminating in a vestigial helictoglossa (internally). (142) Anorthoneis arthus-bertrandii De Stefano & Pennesi, sp. nov. from slide MDS 9 (Holotype), LM images. Scale bars: Fig. 140 = 10 μm; Figs 139, 142 = 5 μm; Fig. 137 = 2 μm; Figs 136, 138, 141 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720168" httpUri="https://zenodo.org/record/13720168/files/figure.png" pageId="31" pageNumber="232">Figs 139, 141</figureCitation>
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). Valvocopula open, with a smooth inner margin.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |