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<document id="8480B9D03578061154435A443BDD0749" ID-CLB-Dataset="24554" ID-DOI="10.11646/zootaxa.4712.1.6" ID-GBIF-Dataset="6be10c07-890a-41b6-80b9-55206dbf2fc5" ID-ISSN="1175-5326" ID-Zenodo-Dep="3586882" ID-ZooBank="63DD38EB-47D8-4010-912B-754C931C0C63" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576824309777" checkinUser="plazi" docAuthor="Song, Xiao-Bin, Zhou, De-Yao, Chen, Zhi-Bing, Lin, Ming-Zhen &amp; Zhu, Jian-Qing" docDate="2019" docId="039B87E15A002B6AFF00FDC0C42A0746" docLanguage="en" docName="zootaxa.4712.1.6.pdf" docOrigin="Zootaxa 4712 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laena hongqiao Zhou &amp; Song 2019, new species" docType="treatment" docUuid="0907CCF6-E359-4789-88ED-BA2C794FD45C" docUuidSource="ZooBank" docVersion="11" lastPageNumber="94" masterDocId="FFA2FF995A012B6EFF97FFADC6760418" masterDocTitle="First record of tribe Laenini (Coleoptera, Tenebrionidae, Lagriinae) from Shanghai, with description of a new species of the genus Laena Dejean" masterLastPageNumber="100" masterPageNumber="90" pageNumber="91" updateTime="1732905079956" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="3D3EC2BF209A7DB2D3894D31525BE664" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2366692599597651BD9A6B9394472795">
<mods:title id="F7F18384461A28A072FAB5875FD19012">First record of tribe Laenini (Coleoptera, Tenebrionidae, Lagriinae) from Shanghai, with description of a new species of the genus Laena Dejean</mods:title>
</mods:titleInfo>
<mods:name id="38981BAA31063CC1A117B67D88413211" type="personal">
<mods:role id="D7106E11593185324E9D35F09722C843">
<mods:roleTerm id="B9969341AE144D7D0C172249126AE25A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9134A723BF31097C2397D6AE42DDCE43">Song, Xiao-Bin</mods:namePart>
</mods:name>
<mods:name id="8F175ED153591E885B35F26349A7FA84" type="personal">
<mods:role id="A179CBA86C059815B1B56AA37964EA78">
<mods:roleTerm id="A7E7608B72C4799F7794FA0F740B7948">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7DF02CB86FBF07ED4A4C9319848014F0">Zhou, De-Yao</mods:namePart>
</mods:name>
<mods:name id="D28CA59431B2D409C29A050807C4CEA9" type="personal">
<mods:role id="81B9473C40FB8A681A0B957A7EF69A72">
<mods:roleTerm id="8D24C9E250968936A721EA8710A9DBB0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DFDFA82DA11308B6D2CB43E4D5406912">Chen, Zhi-Bing</mods:namePart>
</mods:name>
<mods:name id="C073392B72FC311D1518ABAB47528622" type="personal">
<mods:role id="33DAAE853725D1779A4579A6964BED4C">
<mods:roleTerm id="9B21DE0AA4DA8B5E6C9D3E4666C8E936">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="84CCDBC73EF32FA062C9EC9DC15BA8A3">Lin, Ming-Zhen</mods:namePart>
</mods:name>
<mods:name id="023510371A36B8269D86E50D8DCB6C5B" type="personal">
<mods:role id="D078A439E894C3A112DF547AFDEEEABF">
<mods:roleTerm id="4BBCD2F06624B7DF80613632B8A948FA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="72DBEFAF45F5B7D120E528E56AC65615">Zhu, Jian-Qing</mods:namePart>
</mods:name>
<mods:typeOfResource id="38ABB481A60F6A99ADE83154A0A7483C">text</mods:typeOfResource>
<mods:relatedItem id="FF8EBC11E33729248796778D70FB1911" type="host">
<mods:titleInfo id="62EDDFEE5E62884059E905E78E06C1D6">
<mods:title id="61CE927066D85D1A38939313C82504C1">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="94AD31C2057A4A94F69D6AEBE99224D8">
<mods:date id="583EBFCCD6B49DF871D29BDE189BEA0B">2019</mods:date>
<mods:detail id="BDCE610779F7C1DDA57EA9189ACD9384" type="pubDate">
<mods:number id="A159827A5EDED6489B0A23D4E8C4E65C">2019-12-18</mods:number>
</mods:detail>
<mods:detail id="7C998D5CE1510D70BA83B8657FA75B22" type="volume">
<mods:number id="6D9C9D2DB741449890106DF3C27C88E9">4712</mods:number>
</mods:detail>
<mods:detail id="6EA3C331ABD5F0F0D6E011F38AD550DB" type="issue">
<mods:number id="EC6200A68527E07C7D280AD1BD91F966">1</mods:number>
</mods:detail>
<mods:extent id="BEDE4726BAE67F536027176155AC5C9B" unit="page">
<mods:start id="08767D02557F49D9B24BD2FEDC4D894C">90</mods:start>
<mods:end id="A5E8B990E699D4AE58F1307E329EBAF7">100</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A953ABB50D0B5B74125F804DE719589D">journal article</mods:classification>
<mods:identifier id="D18663245FA417C1DFEA00670D47BF0C" type="CLB-Dataset">24554</mods:identifier>
<mods:identifier id="0E218321EB53B9BD036666FDB434F72D" type="DOI">10.11646/zootaxa.4712.1.6</mods:identifier>
<mods:identifier id="E8583BBCAAA03C4FDA5D52FEA5081D73" type="GBIF-Dataset">6be10c07-890a-41b6-80b9-55206dbf2fc5</mods:identifier>
<mods:identifier id="175E44D47D74F6E7049F0FCA087D2B70" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9EC07C5031A33F2F8F2EA3D0EF3E092D" type="Zenodo-Dep">3586882</mods:identifier>
<mods:identifier id="9440B76D1DC125414F9AB3DE7E4F47CD" type="ZooBank">63DD38EB-47D8-4010-912B-754C931C0C63</mods:identifier>
</mods:mods>
<treatment id="039B87E15A002B6AFF00FDC0C42A0746" ID-GBIF-Taxon="160797999" LSID="urn:lsid:zoobank.org:act:0907CCF6-E359-4789-88ED-BA2C794FD45C" httpUri="http://treatment.plazi.org/id/039B87E15A002B6AFF00FDC0C42A0746" lastPageId="4" lastPageNumber="94" pageId="1" pageNumber="91" scope_family="Tenebrionidae" scope_order="Coleoptera">
<subSubSection id="C328657C5A002B6FFF00FDC0C69D06B3" pageId="1" pageNumber="91" type="nomenclature">
<paragraph id="8B8D36F75A002B6FFF00FDC0C4DD069E" blockId="1.[151,683,620,683]" box="[151,683,620,647]" pageId="1" pageNumber="91">
<heading id="D0C5819B5A002B6FFF00FDC0C4DD069E" bold="true" box="[151,683,620,647]" fontSize="11" level="1" pageId="1" pageNumber="91" reason="1">
<emphasis id="B946EAE55A002B6FFF00FDC0C4DD069E" bold="true" box="[151,683,620,647]" pageId="1" pageNumber="91">
<taxonomicName id="4C324D745A002B6FFF00FDC0C478069E" authority="Zhou &amp; Song" authorityName="Zhou &amp; Song" authorityYear="2019" box="[151,526,620,647]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="species" species="hongqiao" status="sp. nov.">
<emphasis id="B946EAE55A002B6FFF00FDC0C72B069F" bold="true" box="[151,349,620,647]" italics="true" pageId="1" pageNumber="91">Laena hongqiao</emphasis>
Zhou &amp; Song
</taxonomicName>
,
<taxonomicNameLabel id="A275579E5A002B6FFD8CFDC0C4DD069E" box="[539,683,620,647]" pageId="1" pageNumber="91" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8D36F75A002B6FFF00FD3DC69D06B3" blockId="1.[151,683,620,683]" box="[151,235,656,683]" pageId="1" pageNumber="91">
(
<figureCitation id="13092A725A002B6FFF37FD3DC69606B3" box="[160,224,656,683]" captionStart="FIGURE" captionStartId="1.[151,251,1785,1810]" captionTargetBox="[181,1436,703,1757]" captionTargetId="figure@1.[151,1436,702,1757]" captionText="FIGURE. 1 Dorsal habitus of Laena hongqiao sp. nov., male (Holotype). B. Living L. hongqiao sifted from leaf litter. C. Habitat of L. hongqiao in Shanghai Zoo." figureDoi="http://doi.org/10.5281/zenodo.3688273" httpUri="https://zenodo.org/record/3688273/files/figure.png" pageId="1" pageNumber="91">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="DF4D667F5A002B6FFF00F954C474032D" ID-DOI="http://doi.org/10.5281/zenodo.3688273" ID-Zenodo-Dep="3688273" httpUri="https://zenodo.org/record/3688273/files/figure.png" pageId="1" pageNumber="91" startId="1.[151,251,1785,1810]" targetBox="[181,1436,703,1757]" targetPageId="1">
<paragraph id="8B8D36F75A002B6FFF00F954C474032D" blockId="1.[151,1437,1785,1846]" pageId="1" pageNumber="91">
<emphasis id="B946EAE55A002B6FFF00F954C765030A" bold="true" box="[151,275,1785,1810]" pageId="1" pageNumber="91">FIGURE. 1</emphasis>
Dorsal habitus of
<taxonomicName id="4C324D745A002B6FFE5CF957C4060309" authority="Song &amp; Zhou &amp; Chen &amp; Lin &amp; Zhu, 2019" authorityName="Song &amp; Zhou &amp; Chen &amp; Lin &amp; Zhu" authorityYear="2019" box="[459,624,1785,1809]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="species" species="hongqiao" status="sp. nov.">
<emphasis id="B946EAE55A002B6FFE5CF957C4060309" box="[459,624,1785,1809]" italics="true" pageId="1" pageNumber="91">Laena hongqiao</emphasis>
</taxonomicName>
<emphasis id="B946EAE55A002B6FFDE2F957C4B3030A" bold="true" box="[629,709,1786,1810]" pageId="1" pageNumber="91">
<taxonomicNameLabel id="A275579E5A002B6FFDE2F957C4B3030A" box="[629,709,1786,1810]" pageId="1" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (
<typeStatus id="548988555A002B6FFC87F954C502030A" box="[784,884,1785,1810]" pageId="1" pageNumber="91" type="holotype">Holotype</typeStatus>
).
<emphasis id="B946EAE55A002B6FFC13F957C5E3030A" bold="true" box="[900,917,1786,1810]" pageId="1" pageNumber="91">B</emphasis>
. Living
<taxonomicName id="4C324D745A002B6FFC7DF957C2120309" authorityName="Song &amp; Zhou &amp; Chen &amp; Lin &amp; Zhu" authorityYear="2019" box="[1002,1124,1785,1809]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="species" species="hongqiao">
<emphasis id="B946EAE55A002B6FFC7DF957C2120309" box="[1002,1124,1785,1809]" italics="true" pageId="1" pageNumber="91">L. hongqiao</emphasis>
</taxonomicName>
sifted from leaf litter.
<emphasis id="B946EAE55A002B6FFAD3F954C32B030A" bold="true" box="[1348,1373,1785,1810]" pageId="1" pageNumber="91">C.</emphasis>
Habitat of
<taxonomicName id="4C324D745A002B6FFF45F8B3C73B032D" authorityName="Song &amp; Zhou &amp; Chen &amp; Lin &amp; Zhu" authorityYear="2019" box="[210,333,1821,1845]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="91" phylum="Arthropoda" rank="species" species="hongqiao">
<emphasis id="B946EAE55A002B6FFF45F8B3C73B032D" box="[210,333,1821,1845]" italics="true" pageId="1" pageNumber="91">L. hongqiao</emphasis>
</taxonomicName>
in
<collectingRegion id="49F6F8155A002B6FFEFAF8B0C7BD032E" box="[365,459,1821,1846]" country="China" name="Shanghai" pageId="1" pageNumber="91">Shanghai</collectingRegion>
Zoo.
</paragraph>
</caption>
<subSubSection id="C328657C5A002B6FFF00F8C9C7830362" box="[151,501,1890,1918]" pageId="1" pageNumber="91" type="description">
<paragraph id="8B8D36F75A002B6FFF00F8C9C7830362" blockId="1.[151,501,1890,1918]" box="[151,501,1890,1918]" pageId="1" pageNumber="91">
<emphasis id="B946EAE55A002B6FFF00F8C9C70E0366" bold="true" box="[151,376,1892,1918]" pageId="1" pageNumber="91">Vernacular name:</emphasis>
ṘŘẌƤ
</paragraph>
</subSubSection>
<subSubSection id="C328657C5A002B6CFF00F801C5F90507" lastPageId="2" lastPageNumber="92" pageId="1" pageNumber="91" type="materials_examined">
<paragraph id="8B8D36F75A002B6FFF00F801C3EB03F2" blockId="1.[151,1437,1962,2026]" pageId="1" pageNumber="91">
<emphasis id="B946EAE55A002B6FFF00F801C73303DE" bold="true" box="[151,325,1963,1990]" pageId="1" pageNumber="91">Type material.</emphasis>
<materialsCitation id="3B5A3CAA5A002B6FFEDCF801C2C703F2" ID-GBIF-Occurrence="2521263243" collectingDate="2017-09-06" collectionCode="SNUC" collectorName="Zoo &amp; Song Xiao-Bin" country="China" location="Changning District" municipality="China" pageId="1" pageNumber="91" specimenCode="T 3" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Shanghai" typeStatus="holotype">
<specimenCount id="9D34FD7E5A002B6FFEDCF801C7FA03DE" box="[331,396,1964,1990]" pageId="1" pageNumber="91" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D34FD7E5A002B6FFE0FF801C7A103DE" box="[408,471,1964,1990]" pageId="1" pageNumber="91" type="female">3 ♀♀</specimenCount>
.
<emphasis id="B946EAE55A002B6FFE72F801C4BE03DE" bold="true" box="[485,712,1963,1990]" pageId="1" pageNumber="91">
<typeStatus id="548988555A002B6FFE72F801C42003DE" box="[485,598,1964,1990]" pageId="1" pageNumber="91" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F32576675A002B6FFDF5F806C4B203DE" box="[610,708,1963,1990]" name="China" pageId="1" pageNumber="91">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="9D34FD7E5A002B6FFD58F801C49203DE" box="[719,740,1964,1990]" pageId="1" pageNumber="91" type="male">♂</specimenCount>
(
<collectionCode id="ED23AE325A002B6FFD64F801C53503DE" box="[755,835,1964,1990]" pageId="1" pageNumber="91">SNUC</collectionCode>
), labeled ‘
<collectingMunicipality id="6BE9AC8D5A002B6FFC25F801C58803DE" box="[946,1022,1964,1990]" pageId="1" pageNumber="91">China</collectingMunicipality>
:
<collectingRegion id="49F6F8155A002B6FFB9FF801C20503DE" box="[1032,1139,1964,1990]" country="China" name="Shanghai" pageId="1" pageNumber="91">Shanghai</collectingRegion>
,
<location id="8EED602C5A002B6FFBE9F801C32203DE" LSID="urn:lsid:plazi:treatment:039B87E15A002B6AFF00FDC0C42A0746:8EED602C5A002B6FFBE9F801C32203DE" box="[1150,1364,1964,1990]" country="China" municipality="China" name="Changning District" pageId="1" pageNumber="91" stateProvince="Shanghai">Changning District</location>
(K
<specimenCode id="DB949E8C5A002B6FFA14F807C6C503FD" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="91">Ṫ ³</specimenCode>
),
<collectingRegion id="49F6F8155A002B6FFF5DF87DC74403F2" box="[202,306,2000,2026]" country="China" name="Shanghai" pageId="1" pageNumber="91">Shanghai</collectingRegion>
<collectorName id="26C753215A002B6FFEAFF87DC71303F2" box="[312,357,2000,2026]" pageId="1" pageNumber="91">Zoo</collectorName>
(±dzḿn'ẘ), artificial mixed forest, sifted,
<date id="FF8C10375A002B6FFCF5F87DC5B803F2" box="[866,974,2000,2026]" pageId="1" pageNumber="91" value="2017-09-06">
<collectingDate id="EFC8E9DF5A002B6FFCF5F87DC5B803F2" box="[866,974,2000,2026]" pageId="1" pageNumber="91" value="2017-09-06">6.ix.2017</collectingDate>
</date>
,
<collectorName id="26C753215A002B6FFC4EF87DC2F703F2" box="[985,1153,2000,2026]" pageId="1" pageNumber="91">Song Xiao-Bin</collectorName>
leg.
</materialsCitation>
/
<typeStatus id="548988555A002B6FFB52F87DC32F03F2" box="[1221,1369,2000,2026]" pageId="1" pageNumber="91" type="holotype">HOLOTYPE</typeStatus>
[red],
</paragraph>
<paragraph id="8B8D36F75A032B6CFF00FF37C5F90507" blockId="2.[151,1436,152,648]" pageId="2" pageNumber="92">
<taxonomicName id="4C324D745A032B6CFF00FF37C73904AB" authority="Song &amp; Zhou &amp; Chen &amp; Lin &amp; Zhu, 2019" authorityName="Song &amp; Zhou &amp; Chen &amp; Lin &amp; Zhu" authorityYear="2019" box="[151,335,153,179]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="hongqiao" status="sp. nov.">
<emphasis id="B946EAE55A032B6CFF00FF37C73904AB" box="[151,335,153,179]" italics="true" pageId="2" pageNumber="92">Laena hongqiao</emphasis>
</taxonomicName>
<emphasis id="B946EAE55A032B6CFEC1FF34C7C604AB" bold="true" box="[342,432,153,179]" pageId="2" pageNumber="92">
<taxonomicNameLabel id="A275579E5A032B6CFEC1FF34C7C604AB" box="[342,432,153,179]" pageId="2" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Zhou &amp; Song det.
<materialsCitation id="3B5A3CAA5A032B6CFD18FF34C7BE04CF" ID-GBIF-Occurrence="2521263244" collectionCode="SHZ, MHBU" country="China" location="CHINA" pageId="2" pageNumber="92" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
2018’
<emphasis id="B946EAE55A032B6CFD42FF34C5B204AB" bold="true" box="[725,964,152,179]" pageId="2" pageNumber="92">
<typeStatus id="548988555A032B6CFD42FF34C52704AB" box="[725,849,153,179]" pageId="2" pageNumber="92" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F32576675A032B6CFCC9FF35C5B604AB" box="[862,960,152,179]" name="China" pageId="2" pageNumber="92">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="9D34FD7E5A032B6CFC5BFF34C58C04AB" box="[972,1018,153,179]" pageId="2" pageNumber="92" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D34FD7E5A032B6CFB97FF34C25104AB" box="[1024,1063,153,179]" pageId="2" pageNumber="92" type="female">1 ♀</specimenCount>
(cSXB),
<specimenCount id="9D34FD7E5A032B6CFB04FF34C2DA04AB" box="[1171,1196,153,179]" pageId="2" pageNumber="92" type="female">1♀</specimenCount>
(
<collectionCode id="ED23AE325A032B6CFB22FF34C28004AB" box="[1205,1270,153,179]" pageId="2" pageNumber="92">SHZ</collectionCode>
),
<specimenCount id="9D34FD7E5A032B6CFA9BFF34C35204AB" box="[1292,1316,153,179]" pageId="2" pageNumber="92" type="female">1♀</specimenCount>
(
<collectionCode id="ED23AE325A032B6CFABBFF34C3F904AB" box="[1324,1423,153,179]" pageId="2" pageNumber="92">MHBU</collectionCode>
), same label data as
<typeStatus id="548988555A032B6CFEF4FF10C7BE04CF" box="[355,456,189,215]" pageId="2" pageNumber="92" type="holotype">holotype</typeStatus>
</materialsCitation>
, /
<typeStatus id="548988555A032B6CFE48FF10C41C04CF" box="[479,618,189,215]" pageId="2" pageNumber="92" type="paratype">PARATYPE</typeStatus>
[yellow],
<taxonomicName id="4C324D745A032B6CFD4CFF13C5E704CF" authority="Song &amp; Zhou &amp; Chen &amp; Lin &amp; Zhu, 2019" authorityName="Song &amp; Zhou &amp; Chen &amp; Lin &amp; Zhu" authorityYear="2019" box="[731,913,189,215]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="hongqiao" status="sp. nov.">
<emphasis id="B946EAE55A032B6CFD4CFF13C5E704CF" box="[731,913,189,215]" italics="true" pageId="2" pageNumber="92">Laena hongqiao</emphasis>
</taxonomicName>
<emphasis id="B946EAE55A032B6CFC01FF10C59804CF" bold="true" box="[918,1006,189,215]" pageId="2" pageNumber="92">
<taxonomicNameLabel id="A275579E5A032B6CFC01FF10C59804CF" box="[918,1006,189,215]" pageId="2" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<determinerName id="7E51DA335A032B6CFC6CFF10C24004CF" box="[1019,1078,189,215]" pageId="2" pageNumber="92">Zhou</determinerName>
&amp;
<determinerName id="7E51DA335A032B6CFBC0FF10C2E704CF" box="[1111,1169,189,215]" pageId="2" pageNumber="92">Song</determinerName>
det.
<materialsCitation id="3B5A3CAA5A032B6CFB53FF10C5360507" ID-GBIF-Occurrence="2521263241" collectingDate="2016-05-26" collectionCode="SNUC, PARA-, TYPE" collectorName="Tu" country="China" county="Peng Zhong" determinerName="Zhou &amp; Song" location="Zhou De-Yao" municipality="Wang Dan" pageId="2" pageNumber="92" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
2018;
<specimenCount id="9D34FD7E5A032B6CFA9DFF10C34404CF" box="[1290,1330,189,215]" pageId="2" pageNumber="92" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED23AE325A032B6CFAD7FF10C3E604CF" box="[1344,1424,189,215]" pageId="2" pageNumber="92">SNUC</collectionCode>
), same label data as
<typeStatus id="548988555A032B6CFEFDFF4CC7B904E3" box="[362,463,225,251]" pageId="2" pageNumber="92" type="holotype">holotype</typeStatus>
, except ‘
<date id="FF8C10375A032B6CFDB8FF4CC4D204E3" box="[559,676,225,251]" pageId="2" pageNumber="92" value="2016-05-26">
<collectingDate id="EFC8E9DF5A032B6CFDB8FF4CC4D204E3" box="[559,676,225,251]" pageId="2" pageNumber="92" value="2016-05-26">26.v.2016</collectingDate>
</date>
,
<location id="8EED602C5A032B6CFD27FF4CC53D04E3" LSID="urn:lsid:plazi:treatment:039B87E15A002B6AFF00FDC0C42A0746:8EED602C5A032B6CFD27FF4CC53D04E3" box="[688,843,225,251]" country="China" county="Peng Zhong" municipality="Wang Dan" name="Zhou De-Yao" pageId="2" pageNumber="92">Zhou De-Yao</location>
,
<collectingMunicipality id="6BE9AC8D5A032B6CFCC1FF4CC5B804E3" box="[854,974,225,251]" pageId="2" pageNumber="92">Wang Dan</collectingMunicipality>
,
<collectingCounty id="62EC4E7B5A032B6CFC4DFF4CC21504E3" box="[986,1123,225,251]" pageId="2" pageNumber="92">Peng Zhong</collectingCounty>
&amp;
<collectorName id="26C753215A032B6CFB11FF4CC2D204E3" box="[1158,1188,225,251]" pageId="2" pageNumber="92">Tu</collectorName>
Yue-Ye leg., /
<collectionCode id="ED23AE325A032B6CFADCFF4CC6AB0507" pageId="2" pageNumber="92">PARA- TYPE</collectionCode>
[yellow],
<taxonomicName id="4C324D745A032B6CFEC6FEABC47E0507" authority="Song &amp; Zhou &amp; Chen &amp; Lin &amp; Zhu, 2019" authorityName="Song &amp; Zhou &amp; Chen &amp; Lin &amp; Zhu" authorityYear="2019" box="[337,520,261,287]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="hongqiao" status="sp. nov.">
<emphasis id="B946EAE55A032B6CFEC6FEABC47E0507" box="[337,520,261,287]" italics="true" pageId="2" pageNumber="92">Laena hongqiao</emphasis>
</taxonomicName>
<emphasis id="B946EAE55A032B6CFD98FEA8C41E0507" bold="true" box="[527,616,261,287]" pageId="2" pageNumber="92">
<taxonomicNameLabel id="A275579E5A032B6CFD98FEA8C41E0507" box="[527,616,261,287]" pageId="2" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<determinerName id="7E51DA335A032B6CFDE1FEA8C4C70507" box="[630,689,261,287]" pageId="2" pageNumber="92">Zhou</determinerName>
&amp;
<determinerName id="7E51DA335A032B6CFD42FEA8C5790507" box="[725,783,261,287]" pageId="2" pageNumber="92">Song</determinerName>
det.
</materialsCitation>
2018’.
</paragraph>
</subSubSection>
<subSubSection id="C328657C5A032B6CFF50FE85C3050593" pageId="2" pageNumber="92" type="diagnosis">
<paragraph id="8B8D36F75A032B6CFF50FE85C3050593" blockId="2.[151,1436,152,648]" pageId="2" pageNumber="92">
<emphasis id="B946EAE55A032B6CFF50FE85C734055B" bold="true" box="[199,322,296,323]" pageId="2" pageNumber="92">Diagnosis.</emphasis>
Dark colored species, with all femora umarmed. Pronotum nearly rounded, bearing large and sparse punctures with its lateral margins bordered. Male anterior tibiae and middle tibiae each with a protruded inner apex, male anterior tarsi not widened. Parameres long, about 0.25 as long as the aedeagus length, with an acute apex.
</paragraph>
</subSubSection>
<subSubSection id="C328657C5A032B6AFF50FE39C52106EB" lastPageId="4" lastPageNumber="94" pageId="2" pageNumber="92" type="description">
<paragraph id="8B8D36F75A032B6CFF50FE39C44F05CB" blockId="2.[151,1436,152,648]" pageId="2" pageNumber="92">
<emphasis id="B946EAE55A032B6CFF50FE39C72F05B7" bold="true" box="[199,345,404,431]" pageId="2" pageNumber="92">Description.</emphasis>
Male (
<figureCitation id="13092A725A032B6CFE3AFE38C47E05B7" box="[429,520,405,431]" captionStart="FIGURE" captionStartId="1.[151,251,1785,1810]" captionTargetBox="[181,1436,703,1757]" captionTargetId="figure@1.[151,1436,702,1757]" captionText="FIGURE. 1 Dorsal habitus of Laena hongqiao sp. nov., male (Holotype). B. Living L. hongqiao sifted from leaf litter. C. Habitat of L. hongqiao in Shanghai Zoo." figureDoi="http://doi.org/10.5281/zenodo.3688273" httpUri="https://zenodo.org/record/3688273/files/figure.png" pageId="2" pageNumber="92">Fig. 1A</figureCitation>
). Body length
<quantity id="4CCA9B125A032B6CFD23FE39C52A05B7" box="[692,860,404,431]" metricMagnitude="-3" metricUnit="m" metricValue="4.5649999999999995" metricValueMax="5.03" metricValueMin="4.1" pageId="2" pageNumber="92" unit="mm" value="4.5649999999999995" valueMax="5.03" valueMin="4.1">4.10–5.03 mm</quantity>
; body black, antennae, femora, tibiae dark brown, maxillary palpi and tarsi light brown.
</paragraph>
<paragraph id="8B8D36F75A032B6CFF50FE70C5C6069F" blockId="2.[151,1436,152,648]" pageId="2" pageNumber="92">
Head (
<figureCitation id="13092A725A032B6CFE84FE70C71B05EF" box="[275,365,477,503]" captionStart="FIGURE" captionStartId="2.[151,251,1728,1753]" captionTargetBox="[151,1422,676,1702]" captionTargetId="figure@2.[151,1436,669,1703]" captionText="FIGURE. 2 Diagnostic features of Laena hongqiao A. Head in dorsal view. B. Pronotum in dorsal view. C. Left elytron in lateral view, showing setiferous umbilicate pores on intervals (arrowed). Abbreviations: i7, interval 7; i9, interval 9." figureDoi="http://doi.org/10.5281/zenodo.3688277" httpUri="https://zenodo.org/record/3688277/files/figure.png" pageId="2" pageNumber="92">Fig. 2A</figureCitation>
) wide, subhexagonal; clypeus with broad and shallow emargination; genae moderately raised; frons weekly convex in middle; frons covered with sharply marked and deep punctures, on sides separated by spaces as wide as puncture diameters, less dense in middle. Eyes round, prominent. Antennae (
<figureCitation id="13092A725A032B6CFBEBFD88C2A30627" box="[1148,1237,549,575]" captionStart="FIGURE" captionStartId="3.[151,251,1713,1738]" captionTargetBox="[177,1414,356,1682]" captionTargetId="figure@3.[164,1423,345,1689]" captionText="FIGURE. 3 Diagnostic features of Laena hongqiao A. Left antenna in dorsal view, B. Aedeagus in dorsal view. C. Aedeagus in lateral view. D. Apical portion of right anterior tibia of male in dorsal view. E. Apical portion of left middle tibia of male in dorsal view. F. Right anterior tibia and anterior femur in dorsal view. G. Left middle tibia and middle femur in dorsal view. H. Left posterior tibia and posterior femur in dorsal view. I. Sternite VII of male in ventral view. J. Sternite VII of female in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3688279" httpUri="https://zenodo.org/record/3688279/files/figure.png" pageId="2" pageNumber="92">Fig. 3A</figureCitation>
) extending to the base of elytra, relative ratio of the length (width) of antennomeres II–XI as follows: 7.5 (6.1): 9.2 (6.0): 7.6 (6.0): 8.4 (6.6): 8.9 (6.5): 9.5 (7.3): 9.5 (7.3): 9.5 (8.4): 9.6 (9.9): 15.4 (11.8).
</paragraph>
<caption id="DF4D667F5A032B6CFF00F96DC36402E5" ID-DOI="http://doi.org/10.5281/zenodo.3688277" ID-Zenodo-Dep="3688277" httpUri="https://zenodo.org/record/3688277/files/figure.png" pageId="2" pageNumber="92" startId="2.[151,251,1728,1753]" subCaptionStartIDs="2.[864,1012,1764,1789]" subCaptionStarts="Abbr" targetBox="[151,1422,676,1702]" targetPageId="2">
<paragraph id="8B8D36F75A032B6CFF00F96DC36402E5" blockId="2.[151,1436,1728,1789]" pageId="2" pageNumber="92">
<emphasis id="B946EAE55A032B6CFF00F96DC76502C1" bold="true" box="[151,275,1728,1753]" pageId="2" pageNumber="92">FIGURE. 2</emphasis>
Diagnostic features of
<taxonomicName id="4C324D745A032B6CFE68F96FC4D302C1" authorityName="Song &amp; Zhou &amp; Chen &amp; Lin &amp; Zhu" authorityYear="2019" box="[511,677,1729,1753]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="92" phylum="Arthropoda" rank="species" species="hongqiao">
<emphasis id="B946EAE55A032B6CFE68F96FC4D302C1" box="[511,677,1729,1753]" italics="true" pageId="2" pageNumber="92">Laena hongqiao</emphasis>
</taxonomicName>
<emphasis id="B946EAE55A032B6CFD3CF96DC4B202C1" bold="true" box="[683,708,1728,1753]" pageId="2" pageNumber="92">A.</emphasis>
Head in dorsal view.
<emphasis id="B946EAE55A032B6CFC37F96CC5CE02C1" bold="true" box="[928,952,1729,1753]" pageId="2" pageNumber="92">B.</emphasis>
Pronotum in dorsal view.
<emphasis id="B946EAE55A032B6CFB54F96DC2AD02C1" bold="true" box="[1219,1243,1728,1753]" pageId="2" pageNumber="92">C.</emphasis>
Left elytron in lateral view, showing setiferous umbilicate pores on intervals (arrowed). Abbreviations:
<date id="FF8C10375A032B6CFC6BF949C26402E5" box="[1020,1042,1764,1789]" pageId="2" pageNumber="92">i7</date>
, interval 7;
<date id="FF8C10375A032B6CFB1CF949C2D602E5" box="[1163,1184,1764,1789]" pageId="2" pageNumber="92">i9</date>
, interval 9.
</paragraph>
</caption>
<paragraph id="8B8D36F75A032B6CFF50F881C32603AA" blockId="2.[151,1436,1834,2042]" pageId="2" pageNumber="92">
Pronotum (
<figureCitation id="13092A725A032B6CFED0F886C7E8035D" box="[327,414,1835,1861]" captionStart="FIGURE" captionStartId="2.[151,251,1728,1753]" captionTargetBox="[151,1422,676,1702]" captionTargetId="figure@2.[151,1436,669,1703]" captionText="FIGURE. 2 Diagnostic features of Laena hongqiao A. Head in dorsal view. B. Pronotum in dorsal view. C. Left elytron in lateral view, showing setiferous umbilicate pores on intervals (arrowed). Abbreviations: i7, interval 7; i9, interval 9." figureDoi="http://doi.org/10.5281/zenodo.3688277" httpUri="https://zenodo.org/record/3688277/files/figure.png" pageId="2" pageNumber="92">Fig. 2B</figureCitation>
) with straight anterior margin, 1.1 times as wide as long, widest near anterior 2/5; lateral margins bordered, posterior margin not bordered and not bent downwards, posterior angles rounded. Punctation of disc distinct, composed of large and deep punctures, distance between punctures equal to 1–2 puncture diameters, most punctures with long and erect setae. Propleura sparsely bearing large punctures, each with a short seta.
</paragraph>
<paragraph id="8B8D36F75A032B6DFF50F816C43904E3" blockId="2.[151,1436,1834,2042]" lastBlockId="3.[151,1437,153,323]" lastPageId="3" lastPageNumber="93" pageId="2" pageNumber="92">
Elytra oblong, width across the humeri wider than posterior margin of pronotum, 1.6–1.7 times as long as wide, widest at just posterior to the middle; dorsum flattened, elytra with rows of punctures in striae, the punctures as large as those on pronotum, each bearing a long seta, intervals between rows with minute punctures, each bearing a short and erect seta, all the intervals flat and shagreened, interval IX with 6 setiferous umbilicate pore, interval VII with 2 setiferous umbilicate pores (
<figureCitation id="13092A725A022B6DFE7EFF4CC43404E3" box="[489,578,225,251]" captionStart="FIGURE" captionStartId="2.[151,251,1728,1753]" captionTargetBox="[151,1422,676,1702]" captionTargetId="figure@2.[151,1436,669,1703]" captionText="FIGURE. 2 Diagnostic features of Laena hongqiao A. Head in dorsal view. B. Pronotum in dorsal view. C. Left elytron in lateral view, showing setiferous umbilicate pores on intervals (arrowed). Abbreviations: i7, interval 7; i9, interval 9." figureDoi="http://doi.org/10.5281/zenodo.3688277" httpUri="https://zenodo.org/record/3688277/files/figure.png" pageId="3" pageNumber="93">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="8B8D36F75A022B6DFF50FEA8C6B1055B" blockId="3.[151,1437,153,323]" pageId="3" pageNumber="93">
All femora (
<figureCitation id="13092A725A022B6DFEC6FEA8C7E70507" box="[337,401,261,287]" captionStart="FIGURE" captionStartId="3.[151,251,1713,1738]" captionTargetBox="[177,1414,356,1682]" captionTargetId="figure@3.[164,1423,345,1689]" captionText="FIGURE. 3 Diagnostic features of Laena hongqiao A. Left antenna in dorsal view, B. Aedeagus in dorsal view. C. Aedeagus in lateral view. D. Apical portion of right anterior tibia of male in dorsal view. E. Apical portion of left middle tibia of male in dorsal view. F. Right anterior tibia and anterior femur in dorsal view. G. Left middle tibia and middle femur in dorsal view. H. Left posterior tibia and posterior femur in dorsal view. I. Sternite VII of male in ventral view. J. Sternite VII of female in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3688279" httpUri="https://zenodo.org/record/3688279/files/figure.png" pageId="3" pageNumber="93">Fig. 3</figureCitation>
F–H) without teeth. Inner apex of anterior tibiae (
<figureCitation id="13092A725A022B6DFC29FEA8C2630507" box="[958,1045,261,287]" captionStart="FIGURE" captionStartId="3.[151,251,1713,1738]" captionTargetBox="[177,1414,356,1682]" captionTargetId="figure@3.[164,1423,345,1689]" captionText="FIGURE. 3 Diagnostic features of Laena hongqiao A. Left antenna in dorsal view, B. Aedeagus in dorsal view. C. Aedeagus in lateral view. D. Apical portion of right anterior tibia of male in dorsal view. E. Apical portion of left middle tibia of male in dorsal view. F. Right anterior tibia and anterior femur in dorsal view. G. Left middle tibia and middle femur in dorsal view. H. Left posterior tibia and posterior femur in dorsal view. I. Sternite VII of male in ventral view. J. Sternite VII of female in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3688279" httpUri="https://zenodo.org/record/3688279/files/figure.png" pageId="3" pageNumber="93">Fig. 3D</figureCitation>
) and middle tibiae projecting (
<figureCitation id="13092A725A022B6DFAF8FEA8C6CC055B" captionStart="FIGURE" captionStartId="3.[151,251,1713,1738]" captionTargetBox="[177,1414,356,1682]" captionTargetId="figure@3.[164,1423,345,1689]" captionText="FIGURE. 3 Diagnostic features of Laena hongqiao A. Left antenna in dorsal view, B. Aedeagus in dorsal view. C. Aedeagus in lateral view. D. Apical portion of right anterior tibia of male in dorsal view. E. Apical portion of left middle tibia of male in dorsal view. F. Right anterior tibia and anterior femur in dorsal view. G. Left middle tibia and middle femur in dorsal view. H. Left posterior tibia and posterior femur in dorsal view. I. Sternite VII of male in ventral view. J. Sternite VII of female in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3688279" httpUri="https://zenodo.org/record/3688279/files/figure.png" pageId="3" pageNumber="93">Fig. 3E</figureCitation>
).
</paragraph>
<caption id="DF4D667F5A022B6DFF00F91CC6B80342" ID-DOI="http://doi.org/10.5281/zenodo.3688279" ID-Zenodo-Dep="3688279" httpUri="https://zenodo.org/record/3688279/files/figure.png" pageId="3" pageNumber="93" startId="3.[151,251,1713,1738]" targetBox="[177,1414,356,1682]" targetPageId="3">
<paragraph id="8B8D36F75A022B6DFF00F91CC6B80342" blockId="3.[151,1437,1713,1882]" pageId="3" pageNumber="93">
<emphasis id="B946EAE55A022B6DFF00F91CC76202D2" bold="true" box="[151,276,1713,1738]" pageId="3" pageNumber="93">FIGURE. 3</emphasis>
Diagnostic features of
<taxonomicName id="4C324D745A022B6DFD93F91EC48D02D2" authority="A. Left" authorityName="A. Left" box="[516,763,1713,1738]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="93" phylum="Arthropoda" rank="species" species="hongqiao">
<emphasis id="B946EAE55A022B6DFD93F91EC4DC02D2" box="[516,682,1714,1738]" italics="true" pageId="3" pageNumber="93">Laena hongqiao</emphasis>
<emphasis id="B946EAE55A022B6DFD26F91CC4BC02D2" bold="true" box="[689,714,1713,1738]" pageId="3" pageNumber="93">A.</emphasis>
Left
</taxonomicName>
antenna in dorsal view,
<emphasis id="B946EAE55A022B6DFC62F91FC27A02D2" bold="true" box="[1013,1036,1714,1738]" pageId="3" pageNumber="93">B.</emphasis>
Aedeagus in dorsal view.
<emphasis id="B946EAE55A022B6DFA8DF91CC34402D2" bold="true" box="[1306,1330,1713,1738]" pageId="3" pageNumber="93">C.</emphasis>
Aedeagus in lateral view.
<emphasis id="B946EAE55A022B6DFEA3F97BC73B02F6" bold="true" box="[308,333,1750,1774]" pageId="3" pageNumber="93">D.</emphasis>
Apical portion of right anterior tibia of male in dorsal view.
<emphasis id="B946EAE55A022B6DFC2FF97BC5A602F6" bold="true" box="[952,976,1750,1774]" pageId="3" pageNumber="93">E.</emphasis>
Apical portion of left middle tibia of male in dorsal view.
<emphasis id="B946EAE55A022B6DFE8FF957C75D030A" bold="true" box="[280,299,1786,1810]" pageId="3" pageNumber="93">F.</emphasis>
Right anterior tibia and anterior femur in dorsal view.
<emphasis id="B946EAE55A022B6DFCCEF954C502030A" bold="true" box="[857,884,1785,1810]" pageId="3" pageNumber="93">G.</emphasis>
Left middle tibia and middle femur in dorsal view.
<emphasis id="B946EAE55A022B6DFA15F957C3EA030A" bold="true" box="[1410,1436,1786,1810]" pageId="3" pageNumber="93">H.</emphasis>
Left posterior tibia and posterior femur in dorsal view.
<emphasis id="B946EAE55A022B6DFD2AF8B3C4BB032E" bold="true" box="[701,717,1822,1846]" pageId="3" pageNumber="93">I.</emphasis>
Sternite VII of male in ventral view.
<emphasis id="B946EAE55A022B6DFBD5F8B3C223032E" bold="true" box="[1090,1109,1822,1846]" pageId="3" pageNumber="93">J.</emphasis>
Sternite VII of female in ventral view.
</paragraph>
</caption>
<paragraph id="8B8D36F75A022B6DFF50F825C21903BB" blockId="3.[151,1436,1928,2027]" box="[199,1135,1928,1955]" pageId="3" pageNumber="93">
Sternite VII (
<figureCitation id="13092A725A022B6DFECCF825C7D103BB" box="[347,423,1928,1955]" captionStart="FIGURE" captionStartId="3.[151,251,1713,1738]" captionTargetBox="[177,1414,356,1682]" captionTargetId="figure@3.[164,1423,345,1689]" captionText="FIGURE. 3 Diagnostic features of Laena hongqiao A. Left antenna in dorsal view, B. Aedeagus in dorsal view. C. Aedeagus in lateral view. D. Apical portion of right anterior tibia of male in dorsal view. E. Apical portion of left middle tibia of male in dorsal view. F. Right anterior tibia and anterior femur in dorsal view. G. Left middle tibia and middle femur in dorsal view. H. Left posterior tibia and posterior femur in dorsal view. I. Sternite VII of male in ventral view. J. Sternite VII of female in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3688279" httpUri="https://zenodo.org/record/3688279/files/figure.png" pageId="3" pageNumber="93">
Fig.
<date id="FF8C10375A022B6DFE18F824C7D103BB" box="[399,423,1929,1955]" pageId="3" pageNumber="93">3I</date>
</figureCitation>
) somewhat rounded at apical margin. Aedeagus see
<figureCitation id="13092A725A022B6DFC64F825C24403BB" box="[1011,1074,1928,1955]" captionStart="FIGURE" captionStartId="3.[151,251,1713,1738]" captionTargetBox="[177,1414,356,1682]" captionTargetId="figure@3.[164,1423,345,1689]" captionText="FIGURE. 3 Diagnostic features of Laena hongqiao A. Left antenna in dorsal view, B. Aedeagus in dorsal view. C. Aedeagus in lateral view. D. Apical portion of right anterior tibia of male in dorsal view. E. Apical portion of left middle tibia of male in dorsal view. F. Right anterior tibia and anterior femur in dorsal view. G. Left middle tibia and middle femur in dorsal view. H. Left posterior tibia and posterior femur in dorsal view. I. Sternite VII of male in ventral view. J. Sternite VII of female in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3688279" httpUri="https://zenodo.org/record/3688279/files/figure.png" pageId="3" pageNumber="93">Fig. 3</figureCitation>
B–C.
</paragraph>
<paragraph id="8B8D36F75A022B6DFF50F801C27303F2" blockId="3.[151,1436,1928,2027]" pageId="3" pageNumber="93">
Females. Similar to male, but body larger, body length 5.00–
<quantity id="4CCA9B125A022B6DFCE2F801C5AD03DF" box="[885,987,1964,1991]" metricMagnitude="-3" metricUnit="m" metricValue="5.56" pageId="3" pageNumber="93" unit="mm" value="5.56">5.56 mm</quantity>
, with unmodified inner apex of anterior tibiae and middle tibiae; sternite VII (
<figureCitation id="13092A725A022B6DFDAAF87DC4FC03F3" box="[573,650,2000,2027]" captionStart="FIGURE" captionStartId="3.[151,251,1713,1738]" captionTargetBox="[177,1414,356,1682]" captionTargetId="figure@3.[164,1423,345,1689]" captionText="FIGURE. 3 Diagnostic features of Laena hongqiao A. Left antenna in dorsal view, B. Aedeagus in dorsal view. C. Aedeagus in lateral view. D. Apical portion of right anterior tibia of male in dorsal view. E. Apical portion of left middle tibia of male in dorsal view. F. Right anterior tibia and anterior femur in dorsal view. G. Left middle tibia and middle femur in dorsal view. H. Left posterior tibia and posterior femur in dorsal view. I. Sternite VII of male in ventral view. J. Sternite VII of female in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3688279" httpUri="https://zenodo.org/record/3688279/files/figure.png" pageId="3" pageNumber="93">Fig. 3J</figureCitation>
) nearly rounded at apical margin.
</paragraph>
<paragraph id="8B8D36F75A052B6AFF50FF35C7FD05B7" blockId="4.[151,1437,152,863]" pageId="4" pageNumber="94">
<emphasis id="B946EAE55A052B6AFF50FF35C7C404AB" bold="true" box="[199,434,152,179]" pageId="4" pageNumber="94">Comparative notes.</emphasis>
Within described Chinese, Japanese and Korean congeners,
<taxonomicName id="4C324D745A052B6AFBF0FF34C35604AA" box="[1127,1312,152,178]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="hongqiao">
<emphasis id="B946EAE55A052B6AFBF0FF34C35604AA" box="[1127,1312,152,178]" italics="true" pageId="4" pageNumber="94">Laena hongqiao</emphasis>
</taxonomicName>
shares unarmed femora with
<taxonomicName id="4C324D745A052B6AFEE2FF10C4AB04CF" authority="Schawaller, 2008" authorityName="Schawaller" authorityYear="2008" box="[373,733,188,215]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="moxica">
<emphasis id="B946EAE55A052B6AFEE2FF10C46204CE" box="[373,532,189,214]" italics="true" pageId="4" pageNumber="94">Laena moxica</emphasis>
<bibRefCitation id="EFA34B065A052B6AFD8BFF11C4AB04CF" author="Schawaller, W." box="[540,733,188,215]" pageId="4" pageNumber="94" pagination="387 - 411" refId="ref19622" refString="Schawaller, W. (2008) The genus Laena Latreille (Coleoptera: Tenebrionidae) in China (part 2), with descriptions of 30 new species and a new identification key. Stuttgarter Beitrage zur Naturkunde, Serie A (Biologie), 1, 387 - 411." type="journal article" year="2008">Schawaller, 2008</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="49F6F8155A052B6AFCB4FF11C5F604CE" box="[803,896,188,214]" country="China" name="Sichuan" pageId="4" pageNumber="94">Sichuan</collectingRegion>
,
<taxonomicName id="4C324D745A052B6AFC1BFF10C29904CF" authority="Schawaller, 2008" authorityName="Schawaller" authorityYear="2008" box="[908,1263,188,215]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="guizhouica">
<emphasis id="B946EAE55A052B6AFC1BFF10C25304CE" box="[908,1061,188,214]" italics="true" pageId="4" pageNumber="94">L. guizhouica</emphasis>
<bibRefCitation id="EFA34B065A052B6AFBBAFF11C29904CF" author="Schawaller, W." box="[1069,1263,188,215]" pageId="4" pageNumber="94" pagination="387 - 411" refId="ref19622" refString="Schawaller, W. (2008) The genus Laena Latreille (Coleoptera: Tenebrionidae) in China (part 2), with descriptions of 30 new species and a new identification key. Stuttgarter Beitrage zur Naturkunde, Serie A (Biologie), 1, 387 - 411." type="journal article" year="2008">Schawaller, 2008</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="49F6F8155A052B6AFAA2FF11C3EE04CE" box="[1333,1432,188,214]" country="China" name="Guizhou" pageId="4" pageNumber="94">Guizhou</collectingRegion>
,
<taxonomicName id="4C324D745A052B6AFF00FF4CC79104E3" authority="Schawaller, 2008" authorityName="Schawaller" authorityYear="2008" box="[151,487,224,251]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="jiangxica">
<emphasis id="B946EAE55A052B6AFF00FF4CC76B04E2" box="[151,285,225,250]" italics="true" pageId="4" pageNumber="94">L. jiangxica</emphasis>
<bibRefCitation id="EFA34B065A052B6AFEB2FF4DC79104E3" author="Schawaller, W." box="[293,487,224,251]" pageId="4" pageNumber="94" pagination="387 - 411" refId="ref19622" refString="Schawaller, W. (2008) The genus Laena Latreille (Coleoptera: Tenebrionidae) in China (part 2), with descriptions of 30 new species and a new identification key. Stuttgarter Beitrage zur Naturkunde, Serie A (Biologie), 1, 387 - 411." type="journal article" year="2008">Schawaller, 2008</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="49F6F8155A052B6AFDBBFF4DC4F604E2" box="[556,640,224,250]" country="China" name="Jiangxi" pageId="4" pageNumber="94">Jiangxi</collectingRegion>
,
<taxonomicName id="4C324D745A052B6AFD1CFF4CC59104E3" authority="Schawaller, 2008" authorityName="Schawaller" authorityYear="2008" box="[651,999,224,251]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="guangxica">
<emphasis id="B946EAE55A052B6AFD1CFF4CC56804E2" box="[651,798,225,250]" italics="true" pageId="4" pageNumber="94">L. guangxica</emphasis>
<bibRefCitation id="EFA34B065A052B6AFCB2FF4DC59104E3" author="Schawaller, W." box="[805,999,224,251]" pageId="4" pageNumber="94" pagination="387 - 411" refId="ref19622" refString="Schawaller, W. (2008) The genus Laena Latreille (Coleoptera: Tenebrionidae) in China (part 2), with descriptions of 30 new species and a new identification key. Stuttgarter Beitrage zur Naturkunde, Serie A (Biologie), 1, 387 - 411." type="journal article" year="2008">Schawaller, 2008</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="49F6F8155A052B6AFBBBFF4DC2F904E2" box="[1068,1167,224,250]" country="China" name="Guangxi" pageId="4" pageNumber="94">Guangxi</collectingRegion>
,
<taxonomicName id="4C324D745A052B6AFB0CFF4CC7570507" authority="Masumoto &amp; Yin, 1994" authorityName="Masumoto &amp; Yin" authorityYear="1994" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis id="B946EAE55A052B6AFB0CFF4CC36B04E2" box="[1179,1309,224,250]" italics="true" pageId="4" pageNumber="94">L. daliensis</emphasis>
<bibRefCitation id="EFA34B065A052B6AFAB3FF4CC7570507" author="Masumoto, K. &amp; Yin, W. Y." pageId="4" pageNumber="94" pagination="165 - 170" refId="ref19482" refString="Masumoto, K. &amp; Yin, W. Y. (1994) Four new Laena species (Coleoptera, Tenebrionidae) from Yunnan, Southwest China. Elytra, 22 (1), 165 - 170." type="journal article" year="1994">Masumoto &amp; Yin, 1994</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C324D745A052B6AFEC0FEA8C4810507" authority="Masumoto &amp; Yin, 1993" authorityName="Masumoto &amp; Yin" authorityYear="1993" box="[343,759,260,287]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="watanabei">
<emphasis id="B946EAE55A052B6AFEC0FEA8C79F0506" box="[343,489,260,286]" italics="true" pageId="4" pageNumber="94">L. watanabei</emphasis>
<bibRefCitation id="EFA34B065A052B6AFE78FEA8C4810507" author="Masumoto, K. &amp; Yin W. Y." box="[495,759,260,287]" pageId="4" pageNumber="94" pagination="239 - 243" refId="ref19438" refString="Masumoto, K. &amp; Yin W. Y. (1993) Two new Yunnanese beetles of the genera Laena (Tenebrionidae) and Sivacrypticus (Archeocrypticidae) (Coleoptera). Elytra, 21 (2), 239 - 243." type="journal article" year="1993">Masumoto &amp; Yin, 1993</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="49F6F8155A052B6AFCADFEA8C5E50507" box="[826,915,261,287]" country="China" name="Yunnan" pageId="4" pageNumber="94">Yunnan</collectingRegion>
,
<taxonomicName id="4C324D745A052B6AFC0AFEA8C2AF0507" authority="Zhao &amp; Ren, 2011" authorityName="Zhao &amp; Ren" authorityYear="2011" box="[925,1241,260,287]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="zhengi">
<emphasis id="B946EAE55A052B6AFC0AFEA8C2720506" box="[925,1028,260,286]" italics="true" pageId="4" pageNumber="94">L. zhengi</emphasis>
<bibRefCitation id="EFA34B065A052B6AFB9CFEA9C2AF0507" author="Zhao, X. L. &amp; Ren, G. D." box="[1035,1241,260,287]" pageId="4" pageNumber="94" pagination="568 - 570" refId="ref20040" refString="Zhao, X. L. &amp; Ren, G. D. (2011) A new species of the genus Laena Dejean from Xizang, China (Coleoptera, Tenebrionidae, Lagriinae). Acta Zootaxonomica Sinica, 36 (3), 568 - 570." type="journal article" year="2011">Zhao &amp; Ren, 2011</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C324D745A052B6AFA98FEA8C70C055B" authority="Zhao &amp; Ren, 2012 a" authorityName="Zhao &amp; Ren" authorityYear="2012" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="motogana">
<emphasis id="B946EAE55A052B6AFA98FEA8C3EA0506" box="[1295,1436,261,286]" italics="true" pageId="4" pageNumber="94">L. motogana</emphasis>
<bibRefCitation id="EFA34B065A052B6AFF00FE85C70C055B" author="Zhao, X. L. &amp; Ren, G. D." box="[151,378,296,323]" pageId="4" pageNumber="94" pagination="15 - 36" refId="ref20089" refString="Zhao, X. L. &amp; Ren, G. D. (2012 a) Six new species of the genus Laena Dejean from China (Coleoptera, Tenebrionidae, Lagriinae). ZooKeys, 177, 15 - 36. https: // doi. org / 10.3897 / zookeys. 177.2426" type="journal article" year="2012">Zhao &amp; Ren, 2012a</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="49F6F8155A052B6AFE54FE85C462055A" box="[451,532,296,322]" country="China" name="Xizang" pageId="4" pageNumber="94">Xizang</collectingRegion>
(figures and illustrations of these species see Zhao 2012), all these species are found in high mountains far from
<collectingRegion id="49F6F8155A052B6AFD8AFEE1C4FF057E" box="[541,649,332,358]" country="China" name="Shanghai" pageId="4" pageNumber="94">Shanghai</collectingRegion>
. The punctation and shape of pronotum, elytra humer, aspect ratio of elytra and shape of aedeagus readily distinguish
<taxonomicName id="4C324D745A052B6AFD25FEDCC54F0592" box="[690,825,368,394]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="hongqiao">
<emphasis id="B946EAE55A052B6AFD25FEDCC54F0592" box="[690,825,368,394]" italics="true" pageId="4" pageNumber="94">L. hongqiao</emphasis>
</taxonomicName>
from
<taxonomicName id="4C324D745A052B6AFCEBFEDCC59C0592" authorityName="Schawaller" authorityYear="2008" box="[892,1002,369,394]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="moxica">
<emphasis id="B946EAE55A052B6AFCEBFEDCC59C0592" box="[892,1002,369,394]" italics="true" pageId="4" pageNumber="94">L. moxica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C324D745A052B6AFC60FEDCC2FF0592" authorityName="Schawaller" authorityYear="2008" box="[1015,1161,369,394]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="guangxica">
<emphasis id="B946EAE55A052B6AFC60FEDCC2FF0592" box="[1015,1161,369,394]" italics="true" pageId="4" pageNumber="94">L. guangxica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C324D745A052B6AFB01FEDCC3610592" authorityName="Masumoto &amp; Yin" authorityYear="1994" box="[1174,1303,368,394]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis id="B946EAE55A052B6AFB01FEDCC3610592" box="[1174,1303,368,394]" italics="true" pageId="4" pageNumber="94">L. daliensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C324D745A052B6AFAB3FEDCC6CF05B6" authorityName="Masumoto &amp; Yin" authorityYear="1993" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="watanabei">
<emphasis id="B946EAE55A052B6AFAB3FEDCC6CF05B6" italics="true" pageId="4" pageNumber="94">L. watanabei</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C324D745A052B6AFF60FE38C7F205B6" authorityName="Zhao &amp; Ren" authorityYear="2012" box="[247,388,405,430]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="motogana">
<emphasis id="B946EAE55A052B6AFF60FE38C7F205B6" box="[247,388,405,430]" italics="true" pageId="4" pageNumber="94">L. motogana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8D36F75A052B6AFF50FE14C7320627" blockId="4.[151,1437,152,863]" pageId="4" pageNumber="94">
<taxonomicName id="4C324D745A052B6AFF50FE14C70B05CA" box="[199,381,440,466]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="hongqiao">
<emphasis id="B946EAE55A052B6AFF50FE14C70B05CA" box="[199,381,440,466]" italics="true" pageId="4" pageNumber="94">Laena hongqiao</emphasis>
</taxonomicName>
closely resembles
<emphasis id="B946EAE55A052B6AFDD8FE14C50F05CA" box="[591,889,440,466]" italics="true" pageId="4" pageNumber="94">
<taxonomicName id="4C324D745A052B6AFDD8FE14C4A105CA" authorityName="Schawaller" authorityYear="2008" box="[591,727,441,466]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="jiangxica">L. jiangxica</taxonomicName>
,
<taxonomicName id="4C324D745A052B6AFD76FE14C50F05CA" authorityName="Schawaller" authorityYear="2008" box="[737,889,440,466]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="guizhouica">L. guizhouica</taxonomicName>
</emphasis>
and
<taxonomicName id="4C324D745A052B6AFC39FE14C26505CA" authorityName="Zhao &amp; Ren" authorityYear="2011" box="[942,1043,440,466]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="zhengi">
<emphasis id="B946EAE55A052B6AFC39FE14C26505CA" box="[942,1043,440,466]" italics="true" pageId="4" pageNumber="94">L. zhengi</emphasis>
</taxonomicName>
in general appearance, however,
<taxonomicName id="4C324D745A052B6AFA11FE14C69605EE" authorityName="Zhao &amp; Ren" authorityYear="2011" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="zhengi">
<emphasis id="B946EAE55A052B6AFA11FE14C69605EE" italics="true" pageId="4" pageNumber="94">L. zhengi</emphasis>
</taxonomicName>
possesses quinquangular aedeagal parameres which distinctly differ from that of
<taxonomicName id="4C324D745A052B6AFBEAFE70C37005EE" box="[1149,1286,476,502]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="hongqiao">
<emphasis id="B946EAE55A052B6AFBEAFE70C37005EE" box="[1149,1286,476,502]" italics="true" pageId="4" pageNumber="94">L. hongqiao</emphasis>
</taxonomicName>
;
<taxonomicName id="4C324D745A052B6AFA82FE70C3EA05EE" authorityName="Schawaller" authorityYear="2008" box="[1301,1436,477,502]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="jiangxica">
<emphasis id="B946EAE55A052B6AFA82FE70C3EA05EE" box="[1301,1436,477,502]" italics="true" pageId="4" pageNumber="94">L. jiangxica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C324D745A052B6AFF51FDACC7280602" authorityName="Schawaller" authorityYear="2008" box="[198,350,512,538]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="guizhouica">
<emphasis id="B946EAE55A052B6AFF51FDACC7280602" box="[198,350,512,538]" italics="true" pageId="4" pageNumber="94">L. guizhouica</emphasis>
</taxonomicName>
bears much shorter parameres with obtuse apex, which is different from the longer and acute one of
<taxonomicName id="4C324D745A052B6AFF21FD88C74B0626" box="[182,317,548,574]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="hongqiao">
<emphasis id="B946EAE55A052B6AFF21FD88C74B0626" box="[182,317,548,574]" italics="true" pageId="4" pageNumber="94">L. hongqiao</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8D36F75A052B6AFF50FDE4C55706B3" blockId="4.[151,1437,152,863]" pageId="4" pageNumber="94">
<taxonomicName id="4C324D745A052B6AFF50FDE4C4F8067B" authority="Lobl &amp; Schawaller, 2008" authorityName="Lobl &amp; Schawaller" authorityYear="2008" box="[199,654,584,611]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="confusa">
<emphasis id="B946EAE55A052B6AFF50FDE4C71A067B" box="[199,364,585,611]" italics="true" pageId="4" pageNumber="94">Laena confusa</emphasis>
<bibRefCitation id="EFA34B065A052B6AFEE4FDE5C4F8067B" author="Lobl, I. &amp; Merkl, K. &amp; Ando, K. &amp; Bouchard, P. &amp; Lillig, M. &amp; Masumoto, K. &amp; Schawaller, W." box="[371,654,584,611]" pageId="4" pageNumber="94" pagination="105 - 352" refId="ref19275" refString="Lobl, I., Merkl, K., Ando, K., Bouchard, P., Lillig, M., Masumoto, K. &amp; Schawaller, W. (2008) Tenebrionidae Latreille, 1802. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera 5. Tenebrionoidea. Apollo Books, Stenstrup, pp. 105 - 352. https: // doi. org / 10.1163 / 9789004260900 _ 011" type="book chapter" year="2008">Löbl &amp; Schawaller, 2008</bibRefCitation>
</taxonomicName>
is a replacement name for
<taxonomicName id="4C324D745A052B6AFC57FDE4C34A067B" authority="Li &amp; Chen, 1993" authorityName="Li &amp; Chen" authorityYear="1993" box="[960,1340,584,611]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B946EAE55A052B6AFC57FDE4C200067A" box="[960,1142,584,610]" italics="true" pageId="4" pageNumber="94">Laena chinensis</emphasis>
Li &amp; Chen, 1993
</taxonomicName>
(a homonym of
<taxonomicName id="4C324D745A052B6AFF69FDC0C45C069F" authority="Kaszab, 1965" authorityName="Kaszab" authorityYear="1965" box="[254,554,620,647]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B946EAE55A052B6AFF69FDC0C778069E" box="[254,270,621,646]" italics="true" pageId="4" pageNumber="94">L</emphasis>
.
<emphasis id="B946EAE55A052B6AFE8AFDC1C7F3069E" box="[285,389,620,646]" italics="true" pageId="4" pageNumber="94">chinensis</emphasis>
Kaszab, 1965
</taxonomicName>
). According to the original description,
<taxonomicName id="4C324D745A052B6AFC79FDC0C215069F" authorityName="Lobl &amp; Schawaller" authorityYear="2008" box="[1006,1123,621,647]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="confusa">
<emphasis id="B946EAE55A052B6AFC79FDC0C215069F" box="[1006,1123,621,647]" italics="true" pageId="4" pageNumber="94">L. confusa</emphasis>
</taxonomicName>
has armed anterior femora, which can be used to clearly separate it from
<taxonomicName id="4C324D745A052B6AFD04FD3CC56C06B2" box="[659,794,656,682]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="hongqiao">
<emphasis id="B946EAE55A052B6AFD04FD3CC56C06B2" box="[659,794,656,682]" italics="true" pageId="4" pageNumber="94">L. hongqiao</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8D36F75A052B6AFF50FD19C52106EB" blockId="4.[151,1437,152,863]" pageId="4" pageNumber="94">
<emphasis id="B946EAE55A052B6AFF50FD19C73D06D7" bold="true" box="[199,331,692,719]" pageId="4" pageNumber="94">Bionomics.</emphasis>
<typeStatus id="548988555A052B6AFEC3FD18C7FA06D7" box="[340,396,693,719]" pageId="4" pageNumber="94">Type</typeStatus>
specimens of
<taxonomicName id="4C324D745A052B6AFDA2FD18C4C906D6" box="[565,703,692,718]" class="Insecta" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="94" phylum="Arthropoda" rank="species" species="hongqiao">
<emphasis id="B946EAE55A052B6AFDA2FD18C4C906D6" box="[565,703,692,718]" italics="true" pageId="4" pageNumber="94">L. hongqiao</emphasis>
</taxonomicName>
were collected by sifting leaf litter in an artificial mixed forest mainly consists of
<taxonomicName id="4C324D745A052B6AFEFDFD74C4F606EA" box="[362,640,728,755]" class="Magnoliopsida" family="Lauraceae" genus="Cinnamomum" kingdom="Plantae" order="Laurales" pageId="4" pageNumber="94" phylum="Tracheophyta" rank="species" species="camphora">
<emphasis id="B946EAE55A052B6AFEFDFD74C4F606EA" box="[362,640,728,755]" italics="true" pageId="4" pageNumber="94">Cinnamomum camphora</emphasis>
</taxonomicName>
(L.) J. Presl, 1825.
</paragraph>
</subSubSection>
<subSubSection id="C328657C5A052B6AFF50FD51C416070E" box="[199,608,764,791]" pageId="4" pageNumber="94" type="distribution">
<paragraph id="8B8D36F75A052B6AFF50FD51C416070E" blockId="4.[151,1437,152,863]" box="[199,608,764,791]" pageId="4" pageNumber="94">
<emphasis id="B946EAE55A052B6AFF50FD51C717070F" bold="true" box="[199,353,764,791]" pageId="4" pageNumber="94">Distribution.</emphasis>
East
<collectingCountry id="F32576675A052B6AFE08FD51C790070E" box="[415,486,764,790]" name="China" pageId="4" pageNumber="94">China</collectingCountry>
:
<collectingRegion id="49F6F8155A052B6AFE67FD51C42A070E" box="[496,604,764,790]" country="China" name="Shanghai" pageId="4" pageNumber="94">Shanghai</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C328657C5A052B6AFF50FC8CC42A0746" pageId="4" pageNumber="94" type="etymology">
<paragraph id="8B8D36F75A052B6AFF50FC8CC42A0746" blockId="4.[151,1437,152,863]" pageId="4" pageNumber="94">
<emphasis id="B946EAE55A052B6AFF50FC8CC7380723" bold="true" box="[199,334,801,827]" pageId="4" pageNumber="94">Etymology.</emphasis>
Named after Hong Qiao Economic and Technological Development Zone (ṘŘḠaeḃẆHȐ ³), where the
<typeStatus id="548988555A052B6AFEA8FCE8C7190747" box="[319,367,837,863]" pageId="4" pageNumber="94">type</typeStatus>
series was collected.
</paragraph>
</subSubSection>
</treatment>
</document>
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