544 lines
56 KiB
XML
544 lines
56 KiB
XML
<document ID-DOI="10.3897/zookeys.28.325" ID-GBIF-Dataset="cac72c82-2c83-401f-b6d0-8afb5321fde3" ID-ISSN="1313–2970" ID-Zenodo-Dep="576570" ID-ZooBank="A979ECDE-871F-4AFC-9ABA-63A0FD6DC323" approvalRequired="46" approvalRequired_for_textStreams="46" checkinTime="1587979922717" checkinUser="plazi" docAuthor="Sereno, Paul & Larsson, Hans" docDate="2009" docId="039B2B68FFBF9B6BA5FAB1BFFBB250AE" docLanguage="en" docName="ZK_article_2104.pdf" docOrigin="ZooKeys 28 (28)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@fc5bfa5" docStyleName="zookeys.2008.journal_article" docTitle="Araripesuchus rattoides Sereno & Larsson 2009, sp. n." docType="treatment" docUuid="CF171699-D3FD-4B5C-909A-21C4412BCB0E" docUuidSource="ZooBank" docVersion="6" lastPageNumber="61" masterDocId="FFA25310FF879B57A576B119FF895175" masterDocTitle="Cretaceous Crocodyliforms from the Sahara" masterLastPageNumber="143" masterPageNumber="1" pageNumber="57" updateTime="1643567712823" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Cretaceous Crocodyliforms from the Sahara</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sereno, Paul</mods:namePart>
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<mods:affiliation>University of Chicago, Chicago, United States of America</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Larsson, Hans</mods:namePart>
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<mods:affiliation>Redpath Museum, McGill University, Montreal ,, Canada</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-11-19</mods:number>
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</mods:detail>
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||
<mods:detail type="volume">
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<mods:number>28</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>28</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>143</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.28.325</mods:identifier>
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<mods:identifier type="GBIF-Dataset">cac72c82-2c83-401f-b6d0-8afb5321fde3</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576570</mods:identifier>
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<mods:identifier type="ZooBank">A979ECDE-871F-4AFC-9ABA-63A0FD6DC323</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3790375" ID-GBIF-Taxon="163750071" ID-Zenodo-Dep="3790375" LSID="urn:lsid:zoobank.org:act:CF171699-D3FD-4B5C-909A-21C4412BCB0E" httpUri="http://treatment.plazi.org/id/039B2B68FFBF9B6BA5FAB1BFFBB250AE" lastPageId="60" lastPageNumber="61" pageId="56" pageNumber="57">
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<subSubSection pageId="56" pageNumber="57" type="nomenclature">
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||
<paragraph blockId="56.[140,985,166,298]" box="[140,491,166,193]" pageId="56" pageNumber="57">
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<heading box="[140,491,166,193]" fontSize="11" level="2" pageId="56" pageNumber="57" reason="3">
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<emphasis bold="true" box="[140,491,166,193]" pageId="56" pageNumber="57">
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<taxonomicName authority="Sereno & Larsson, 2009" authorityName="Sereno & Larsson" authorityYear="2009" box="[140,419,166,193]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="rattoides" status="sp. n.">
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<emphasis bold="true" box="[140,419,166,193]" italics="true" pageId="56" pageNumber="57">
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<emphasis box="[140,308,166,193]" inLineHeading="true" pageId="56" pageNumber="57" reason="1">Araripesuchus</emphasis>
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rattoides
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</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[426,491,167,193]" pageId="56" pageNumber="57" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph blockId="56.[140,985,166,298]" box="[140,985,202,229]" pageId="56" pageNumber="57">
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||
<uri box="[140,985,202,229]" pageId="56" pageNumber="57">
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||
urn:lsid:zoobank.org:act:
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<uuid box="[421,985,202,229]" pageId="56" pageNumber="57">CF171699-D3FD-4B5C-909A-21C4412BCB0E</uuid>
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||
</uri>
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection pageId="56" pageNumber="57" type="description">
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||
<paragraph blockId="56.[140,985,166,298]" box="[140,273,237,263]" pageId="56" pageNumber="57">
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<figureCitation box="[140,273,237,263]" captionStart-0="Figure 27" captionStart-1="Figure 28" captionStart-2="Figure 29" captionStart-3="Figure 30" captionStartId-0="57.[140,212,1526,1548]" captionStartId-1="58.[140,212,774,796]" captionStartId-2="58.[140,212,1494,1516]" captionStartId-3="59.[140,212,1462,1484]" captionTargetBox-0="[294,941,368,1483]" captionTargetBox-1="[150,1091,170,747]" captionTargetBox-2="[153,1091,955,1472]" captionTargetBox-3="[162,1079,1108,1429]" captionTargetId-0="figure@57.[268,980,351,1506]" captionTargetId-1="figure@58.[140,1108,162,757]" captionTargetId-2="figure@58.[145,1103,940,1477]" captionTargetId-3="figure@59.[149,1098,1093,1443]" captionTargetPageId-0="57" captionTargetPageId-1="58" captionTargetPageId-2="58" captionTargetPageId-3="59" captionText-0="Figure 27. Right dentary of the crocodyliform Araripesuchus rattoides sp. n. Isolated right dentary lacking teeth (CMN 41893). A Lateral view (reversed). B Medial view. C Dorsal view. D Anterior view. Scale bars equal 2 cm in A-C and 1 cm in D. Abbreviations: ad1, 3, 4, 8, 11, 13, 14, alveolus for dentary tooth 1, 3, 4, 8, 11, 13, 14; asp, articular surface for the splenial; dsym, dentary symphysis; fen, fenestra; fo, foramen; Mc, Meckel’s canal; rdsym, rough dentary symphysis; sdsym, smooth dentary symphysis." captionText-1="Figure 28. Left dentary of the crocodyliform Araripesuchus rattoides sp. n. Pencil drawing of isolated left dentary ramus lacking teeth (UCRC PV3). A Dorsal view. B Ventral view. C Medial view. Scale bar equals 1 cm. Parallel lines indicate broken bone surface. Abbreviations: ad1, 4, 5, 8, alveolus for dentary tooth 1, 4, 5, 8; asp, articular surface for the splenial; fen, fenestra; fo, foramen; sym, symphysis." captionText-2="Figure 29. Computed-tomographic scan of the crocodyliform Araripesuchus rattoides sp. n. Isolated left dentary ramus lacking teeth (UCRC PV3). A Drawing in dorsal view showing the location of cross-sections (B-D). B Parasagittal section showing the size and orientation of the alveolus for dentary tooth 1. C Cross-section through the alveolus of the fourth dentary tooth. D Cross-section through the alveolus of the sixth dentary tooth. Scale bar for B-D equals 1 cm. Abbreviations: ad1, 4, 6, alveolus for dentary tooth 1, 4, 6; sym, symphysis; vc, vascular cavity." captionText-3="Figure 30. Reconstruction of the dentition of the crocodyliform Araripesuchus rattoides sp. n. Anterior dentition restored based on the size and orientation of the alveoli in CMN 41893 and UCRC PV3. A Dorsal view. B Ventral view. C Anterolateral view with premaxillary and anterior maxillary dentition restored to match those in the dentary. Given the presence of large, adjacent first dentary teeth, there may have been a median diastema between the premaxillary teeth and one or two fewer teeth in each premaxillary tooth row. Scale bar equals 2 cm in A and B. Abbreviations: d1, 4, 8, dentary tooth 1, 4, 8; d, dentary; di, diastema; m1, maxillary tooth 1; pm1, 2, premaxillary tooth 1, 2." figureDoi-0="http://doi.org/10.5281/zenodo.3768353" figureDoi-1="http://doi.org/10.5281/zenodo.3768355" figureDoi-2="http://doi.org/10.5281/zenodo.3768357" figureDoi-3="http://doi.org/10.5281/zenodo.3768361" httpUri-0="https://zenodo.org/record/3768353/files/figure.png" httpUri-1="https://zenodo.org/record/3768355/files/figure.png" httpUri-2="https://zenodo.org/record/3768357/files/figure.png" httpUri-3="https://zenodo.org/record/3768361/files/figure.png" pageId="56" pageNumber="57">Figs. 27–30</figureCitation>
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</paragraph>
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<paragraph blockId="56.[140,985,166,298]" box="[140,224,272,298]" pageId="56" pageNumber="57">
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<tableCitation box="[140,224,272,298]" captionStart="Table 9" captionStartId="59.[140,200,329,351]" captionText="Table 9. Dimensions (mm) of the alveoli in the holotype right dentary of Araripesuchus rattoides (CMN 41893). Width is labiolingual; length is mesiodistal. Parentheses indicate estimated measurement; dash indicates partially preserved alveolus, the dimension for which cannot be determined." pageId="56" pageNumber="57">Table 9</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="56" pageNumber="57" type="etymology">
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<paragraph blockId="56.[140,1108,343,862]" pageId="56" pageNumber="57">
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<emphasis bold="true" box="[140,275,343,369]" pageId="56" pageNumber="57">
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<emphasis box="[140,270,343,369]" inLineHeading="true" pageId="56" pageNumber="57" reason="1">Etymology</emphasis>
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.
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</emphasis>
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<taxonomicName authority="(Latin)" baseAuthorityName="Latin" box="[283,442,343,370]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="56" pageNumber="57" phylum="Chordata" rank="genus">
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<emphasis box="[283,352,343,368]" italics="true" pageId="56" pageNumber="57">Rattus</emphasis>
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(Latin)
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</taxonomicName>
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;
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<emphasis box="[455,516,343,369]" italics="true" pageId="56" pageNumber="57">-oides</emphasis>
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, likeness (Latin). Named for the enlarged, procumbent first dentary tooth, which is reminiscent of the condition in many rodents.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="56" pageNumber="57" type="materials_examined">
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<paragraph blockId="56.[140,1108,343,862]" box="[185,902,413,439]" pageId="56" pageNumber="57">
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<materialsCitation ID-GBIF-Occurrence="2608868534" box="[185,902,413,439]" collectionCode="CMN" pageId="56" pageNumber="57" specimenCount="1" typeStatus="holotype">
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<emphasis bold="true" box="[185,306,413,439]" pageId="56" pageNumber="57">
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<emphasis box="[185,301,413,439]" inLineHeading="true" pageId="56" pageNumber="57" reason="1">
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<typeStatus box="[185,301,413,439]" pageId="56" pageNumber="57">Holotype</typeStatus>
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</emphasis>
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.
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</emphasis>
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<accessionNumber box="[313,466,413,439]" httpUri="http://www.ncbi.nlm.nih.gov/protein/CMN41893" pageId="56" pageNumber="57">
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<collectionCode box="[313,383,413,439]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="56" pageNumber="57" type="Museum">CMN</collectionCode>
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41893
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</accessionNumber>
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; right dentary preserving alveoli 1–14.
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</materialsCitation>
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</paragraph>
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<paragraph blockId="56.[140,1108,343,862]" box="[185,1107,448,474]" pageId="56" pageNumber="57">
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<emphasis bold="true" box="[185,389,448,474]" pageId="56" pageNumber="57">
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<emphasis box="[185,282,448,474]" inLineHeading="true" pageId="56" pageNumber="57" reason="1">Referred</emphasis>
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material.
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</emphasis>
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<collectionCode box="[394,472,448,474]" country="USA" httpUri="http://grbio.org/cool/m2jc-wmeq" name="University of California, Riverside" pageId="56" pageNumber="57" type="University or college">UCRC</collectionCode>
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PV3; anterior portion of left dentary preserving alveoli 1–8.
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</paragraph>
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<paragraph blockId="56.[140,1108,343,862]" pageId="56" pageNumber="57">
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<emphasis bold="true" box="[185,348,483,509]" pageId="56" pageNumber="57">
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<emphasis box="[185,243,483,509]" inLineHeading="true" pageId="56" pageNumber="57" reason="1">Type</emphasis>
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locality.
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</emphasis>
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Er Rachidia District (exact locality unknown), eastern
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<collectingCountry box="[1006,1108,484,510]" name="Morocco" pageId="56" pageNumber="57">Morocco</collectingCountry>
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(Fig. 1A, B). A referred specimen (
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<collectionCode box="[547,629,519,545]" country="USA" httpUri="http://grbio.org/cool/m2jc-wmeq" name="University of California, Riverside" pageId="56" pageNumber="57" type="University or college">UCRC</collectionCode>
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<collectionCode box="[640,680,519,545]" pageId="56" pageNumber="57">PV</collectionCode>
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3) was surface collected in
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<quantity box="[993,1085,519,545]" metricMagnitude="1" metricUnit="m" metricValue="5.0546" pageId="56" pageNumber="57" unit="in" value="1990.0">1990 in</quantity>
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a small wash at Darelkarib (south of Erfoud).
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</paragraph>
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<paragraph blockId="56.[140,1108,343,862]" pageId="56" pageNumber="57">
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<emphasis bold="true" box="[185,294,589,615]" pageId="56" pageNumber="57">
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<emphasis box="[185,288,589,615]" inLineHeading="true" pageId="56" pageNumber="57" reason="1">Horizon</emphasis>
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.
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</emphasis>
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Kem Kem Beds; Upper Cretaceous (Cenomanian), ca. 95 Mya (
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<bibRefCitation author="Sereno PC & Dutheil DB & Iarochene M & Larsson HCE & Lyon GH" pageId="56" pageNumber="57" pagination="986 - 991" refId="ref47291" refString="Sereno PC, Dutheil DB, Iarochene M, Larsson HCE, Lyon GH et al. (1996) Predatory dinosaurs from the Sahara and Late Cretaceous faunal differentiation. Science 272: 986 - 991." type="journal article" year="1996">Sereno et al. 1996</bibRefCitation>
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). Th e referred specimen (
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<collectionCode box="[543,625,624,650]" country="USA" httpUri="http://grbio.org/cool/m2jc-wmeq" name="University of California, Riverside" pageId="56" pageNumber="57" type="University or college">UCRC</collectionCode>
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<collectionCode box="[631,671,624,650]" pageId="56" pageNumber="57">PV</collectionCode>
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3) appears to have come from the lower member (pers. commun. D. Dutheil).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="60" lastPageNumber="61" pageId="56" pageNumber="57" type="diagnosis">
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<paragraph blockId="56.[140,1108,343,862]" pageId="56" pageNumber="57">
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<emphasis bold="true" box="[185,312,694,720]" pageId="56" pageNumber="57">
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<emphasis box="[185,307,694,720]" inLineHeading="true" pageId="56" pageNumber="57" reason="1">Diagnosis</emphasis>
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.
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</emphasis>
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Small-bodied metasuchian (<
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<quantity box="[658,705,695,721]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="56" pageNumber="57" unit="m" value="1.0">1 m</quantity>
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) with an enlarged procumbent first dentary tooth that is set immediately adjacent to the midline; smaller procumbent second dentary tooth; a caniniform fourth dentary tooth that is particularly large (twice the basal dimensions of adjacent crowns); and a smooth anterior surface on the dentary symphyses with an oval fenestra opening into the first alveolus.
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</paragraph>
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<paragraph blockId="56.[140,1108,906,1672]" pageId="56" pageNumber="57">
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<emphasis bold="true" box="[140,244,906,932]" pageId="56" pageNumber="57">
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<emphasis box="[140,239,906,932]" inLineHeading="true" pageId="56" pageNumber="57" reason="1">Dentary</emphasis>
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.
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</emphasis>
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The dentary of
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<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[423,546,906,932]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="rattoides">
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<emphasis box="[423,546,906,932]" italics="true" pageId="56" pageNumber="57">A. rattoides</emphasis>
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</taxonomicName>
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show a series of features that distinguishes it from the previously named species
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<taxonomicName authorityName="Buffetaut & Taquet" authorityYear="1979" box="[489,618,941,967]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="wegeneri">
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<emphasis box="[489,618,941,967]" italics="true" pageId="56" pageNumber="57">A. wegeneri</emphasis>
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</taxonomicName>
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and from a contemporary unnamed species from Cenomanian beds in
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<collectingCountry box="[494,558,976,1002]" name="Niger" pageId="56" pageNumber="57">Niger</collectingCountry>
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that closely resembles
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<taxonomicName authority="(Turner 2006)" baseAuthorityName="Turner" baseAuthorityYear="2006" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="tsangatsangana">
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<emphasis box="[817,1013,977,1003]" italics="true" pageId="56" pageNumber="57">A. tsangatsangana</emphasis>
|
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(
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<bibRefCitation author="Turner AH" pageId="56" pageNumber="57" pagination="255 - 369" refId="ref47827" refString="Turner AH (2006) Osteology and phylogeny of a new species of Araripesuchus (Crocodyliformes: Mesoeucrocodylia) from the Late Cretaceous of Madagascar. Historical Biology 18: 255 - 369." type="journal article" year="2006">Turner 2006</bibRefCitation>
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||
)
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</taxonomicName>
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. Th e skull in
|
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<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[367,491,1011,1038]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="rattoides">
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<emphasis box="[367,491,1011,1038]" italics="true" pageId="56" pageNumber="57">A. rattoides</emphasis>
|
||
</taxonomicName>
|
||
appears to be proportionately narrower than in
|
||
<taxonomicName authorityName="Buffetaut & Taquet" authorityYear="1979" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="wegeneri">
|
||
<emphasis italics="true" pageId="56" pageNumber="57">A wegeneri</emphasis>
|
||
</taxonomicName>
|
||
, based on the angle of divergence of the dentary tooth row from the midline. In
|
||
<taxonomicName authorityName="Buffetaut & Taquet" authorityYear="1979" box="[140,266,1082,1108]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="wegeneri">
|
||
<emphasis box="[140,266,1082,1108]" italics="true" pageId="56" pageNumber="57">A. wegeneri</emphasis>
|
||
</taxonomicName>
|
||
, the tooth row diverges at an angle between 20 and 25° from the midline (Fig. 18C), an angle matching the divergence of the upper tooth row (Figs. 14C, 15C). In
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[175,293,1152,1179]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="rattoides">
|
||
<emphasis box="[175,293,1152,1179]" italics="true" pageId="56" pageNumber="57">A rattoides</emphasis>
|
||
</taxonomicName>
|
||
, by contrast, the angle of divergence is approximately 10° (
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<figureCitation box="[987,1094,1152,1178]" captionStart="Figure 27" captionStartId="57.[140,212,1526,1548]" captionTargetBox="[294,941,368,1483]" captionTargetId="figure@57.[268,980,351,1506]" captionTargetPageId="57" captionText="Figure 27. Right dentary of the crocodyliform Araripesuchus rattoides sp. n. Isolated right dentary lacking teeth (CMN 41893). A Lateral view (reversed). B Medial view. C Dorsal view. D Anterior view. Scale bars equal 2 cm in A-C and 1 cm in D. Abbreviations: ad1, 3, 4, 8, 11, 13, 14, alveolus for dentary tooth 1, 3, 4, 8, 11, 13, 14; asp, articular surface for the splenial; dsym, dentary symphysis; fen, fenestra; fo, foramen; Mc, Meckel’s canal; rdsym, rough dentary symphysis; sdsym, smooth dentary symphysis." figureDoi="http://doi.org/10.5281/zenodo.3768353" httpUri="https://zenodo.org/record/3768353/files/figure.png" pageId="56" pageNumber="57">Fig. 27C</figureCitation>
|
||
), or less than half that in
|
||
<taxonomicName authorityName="Buffetaut & Taquet" authorityYear="1979" box="[412,538,1188,1214]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="wegeneri">
|
||
<emphasis box="[412,538,1188,1214]" italics="true" pageId="56" pageNumber="57">A. wegeneri</emphasis>
|
||
</taxonomicName>
|
||
. Th e anterior end of the dentary in
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[951,1082,1187,1214]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="rattoides">A. rattoides</taxonomicName>
|
||
is proportionately deeper than in
|
||
<taxonomicName authorityName="Buffetaut & Taquet" authorityYear="1979" box="[491,615,1223,1249]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="wegeneri">
|
||
<emphasis box="[491,615,1223,1249]" italics="true" pageId="56" pageNumber="57">A. wegeneri</emphasis>
|
||
</taxonomicName>
|
||
and other species of
|
||
<taxonomicName baseAuthorityName="Pol and Apesteguia" baseAuthorityYear="2005" box="[849,998,1223,1249]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="genus">
|
||
<emphasis box="[849,998,1223,1249]" italics="true" pageId="56" pageNumber="57">Araripesuchus</emphasis>
|
||
</taxonomicName>
|
||
. This difference is visible in both anterior and lateral view (Figs. 18A, D, 27A, D).
|
||
</paragraph>
|
||
<paragraph blockId="56.[140,1108,906,1672]" pageId="56" pageNumber="57">
|
||
The orientation of the alveoli for teeth d1–11 is more procumbent in
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[977,1100,1293,1319]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="rattoides">
|
||
<emphasis box="[977,1100,1293,1319]" italics="true" pageId="56" pageNumber="57">A. rattoides</emphasis>
|
||
</taxonomicName>
|
||
. The first and second alveoli project more strongly anteriorly than dorsally, a difference best appreciated in anterior view (Figs. 18D, 27D). Succeeding alveoli, including the caniniform (d4) and d5–11, are visible in lateral view (
|
||
<figureCitation box="[750,852,1399,1425]" captionStart="Figure 27" captionStartId="57.[140,212,1526,1548]" captionTargetBox="[294,941,368,1483]" captionTargetId="figure@57.[268,980,351,1506]" captionTargetPageId="57" captionText="Figure 27. Right dentary of the crocodyliform Araripesuchus rattoides sp. n. Isolated right dentary lacking teeth (CMN 41893). A Lateral view (reversed). B Medial view. C Dorsal view. D Anterior view. Scale bars equal 2 cm in A-C and 1 cm in D. Abbreviations: ad1, 3, 4, 8, 11, 13, 14, alveolus for dentary tooth 1, 3, 4, 8, 11, 13, 14; asp, articular surface for the splenial; dsym, dentary symphysis; fen, fenestra; fo, foramen; Mc, Meckel’s canal; rdsym, rough dentary symphysis; sdsym, smooth dentary symphysis." figureDoi="http://doi.org/10.5281/zenodo.3768353" httpUri="https://zenodo.org/record/3768353/files/figure.png" pageId="56" pageNumber="57">Fig. 27A</figureCitation>
|
||
), whereas they are hidden by the dorsal edge of the alveolar margin in
|
||
<taxonomicName box="[704,824,1434,1460]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="wegener">
|
||
<emphasis box="[704,824,1434,1460]" italics="true" pageId="56" pageNumber="57">A. wegener</emphasis>
|
||
</taxonomicName>
|
||
i (Fig. 18A). Despite the more pronounced anterior projection of the anteriormost pair of teeth, the symphyseal region below these teeth (
|
||
<figureCitation box="[433,535,1504,1530]" captionStart="Figure 27" captionStartId="57.[140,212,1526,1548]" captionTargetBox="[294,941,368,1483]" captionTargetId="figure@57.[268,980,351,1506]" captionTargetPageId="57" captionText="Figure 27. Right dentary of the crocodyliform Araripesuchus rattoides sp. n. Isolated right dentary lacking teeth (CMN 41893). A Lateral view (reversed). B Medial view. C Dorsal view. D Anterior view. Scale bars equal 2 cm in A-C and 1 cm in D. Abbreviations: ad1, 3, 4, 8, 11, 13, 14, alveolus for dentary tooth 1, 3, 4, 8, 11, 13, 14; asp, articular surface for the splenial; dsym, dentary symphysis; fen, fenestra; fo, foramen; Mc, Meckel’s canal; rdsym, rough dentary symphysis; sdsym, smooth dentary symphysis." figureDoi="http://doi.org/10.5281/zenodo.3768353" httpUri="https://zenodo.org/record/3768353/files/figure.png" pageId="56" pageNumber="57">Fig. 27A</figureCitation>
|
||
) is deeper than in
|
||
<taxonomicName authorityName="Buffetaut & Taquet" authorityYear="1979" box="[752,878,1505,1531]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="wegeneri">
|
||
<emphasis box="[752,878,1505,1531]" italics="true" pageId="56" pageNumber="57">A. wegeneri</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 18A) and in a larger contemporary of
|
||
<taxonomicName authorityName="Buffetaut & Taquet" authorityYear="1979" box="[413,540,1540,1566]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="wegeneri">
|
||
<emphasis box="[413,540,1540,1566]" italics="true" pageId="56" pageNumber="57">A. wegeneri</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 31A). Moreover, unlike these other species, the symphyseal articular surface of the dentary is not uniformly rugose in
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[959,1080,1575,1601]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="rattoides">
|
||
<emphasis box="[959,1080,1575,1601]" italics="true" pageId="56" pageNumber="57">A. rattoides</emphasis>
|
||
</taxonomicName>
|
||
as it is in
|
||
<taxonomicName authorityName="Buffetaut & Taquet" authorityYear="1979" box="[219,344,1610,1636]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="wegeneri">
|
||
<emphasis box="[219,344,1610,1636]" italics="true" pageId="56" pageNumber="57">A. wegeneri</emphasis>
|
||
</taxonomicName>
|
||
and its larger contemporary (Figs. 18B, 31B). Th e anterior portion is smooth and fenestrated, as seen in two specimens (
|
||
<figureCitation box="[738,849,1645,1671]" captionStart="Figure 27" captionStartId="57.[140,212,1526,1548]" captionTargetBox="[294,941,368,1483]" captionTargetId="figure@57.[268,980,351,1506]" captionTargetPageId="57" captionText="Figure 27. Right dentary of the crocodyliform Araripesuchus rattoides sp. n. Isolated right dentary lacking teeth (CMN 41893). A Lateral view (reversed). B Medial view. C Dorsal view. D Anterior view. Scale bars equal 2 cm in A-C and 1 cm in D. Abbreviations: ad1, 3, 4, 8, 11, 13, 14, alveolus for dentary tooth 1, 3, 4, 8, 11, 13, 14; asp, articular surface for the splenial; dsym, dentary symphysis; fen, fenestra; fo, foramen; Mc, Meckel’s canal; rdsym, rough dentary symphysis; sdsym, smooth dentary symphysis." figureDoi="http://doi.org/10.5281/zenodo.3768353" httpUri="https://zenodo.org/record/3768353/files/figure.png" pageId="56" pageNumber="57">Figs. 27B</figureCitation>
|
||
,
|
||
<figureCitation box="[861,914,1645,1671]" captionStart="Figure 28" captionStartId="58.[140,212,774,796]" captionTargetBox="[150,1091,170,747]" captionTargetId="figure@58.[140,1108,162,757]" captionTargetPageId="58" captionText="Figure 28. Left dentary of the crocodyliform Araripesuchus rattoides sp. n. Pencil drawing of isolated left dentary ramus lacking teeth (UCRC PV3). A Dorsal view. B Ventral view. C Medial view. Scale bar equals 1 cm. Parallel lines indicate broken bone surface. Abbreviations: ad1, 4, 5, 8, alveolus for dentary tooth 1, 4, 5, 8; asp, articular surface for the splenial; fen, fenestra; fo, foramen; sym, symphysis." figureDoi="http://doi.org/10.5281/zenodo.3768355" httpUri="https://zenodo.org/record/3768355/files/figure.png" pageId="56" pageNumber="57">28C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="57.[140,1108,166,299]" pageId="57" pageNumber="58">
|
||
Tooth size is also distinctive in
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[549,674,167,193]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="rattoides">
|
||
<emphasis box="[549,674,167,193]" italics="true" pageId="57" pageNumber="58">A. rattoides</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation box="[691,781,167,193]" captionStart="Table 9" captionStartId="59.[140,200,329,351]" captionText="Table 9. Dimensions (mm) of the alveoli in the holotype right dentary of Araripesuchus rattoides (CMN 41893). Width is labiolingual; length is mesiodistal. Parentheses indicate estimated measurement; dash indicates partially preserved alveolus, the dimension for which cannot be determined." pageId="57" pageNumber="58">Table 9</tableCitation>
|
||
). The first tooth is 75% the average diameter of the caniniform tooth (d4), which is already twice the diameter of adjacent crowns. In
|
||
<taxonomicName authorityName="Buffetaut & Taquet" authorityYear="1979" box="[401,527,237,263]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="wegeneri">
|
||
<emphasis box="[401,527,237,263]" italics="true" pageId="57" pageNumber="58">A. wegeneri</emphasis>
|
||
</taxonomicName>
|
||
the first dentary tooth is small (
|
||
<figureCitation box="[895,1000,237,263]" captionStart="Figure 27" captionStartId="57.[140,212,1526,1548]" captionTargetBox="[294,941,368,1483]" captionTargetId="figure@57.[268,980,351,1506]" captionTargetPageId="57" captionText="Figure 27. Right dentary of the crocodyliform Araripesuchus rattoides sp. n. Isolated right dentary lacking teeth (CMN 41893). A Lateral view (reversed). B Medial view. C Dorsal view. D Anterior view. Scale bars equal 2 cm in A-C and 1 cm in D. Abbreviations: ad1, 3, 4, 8, 11, 13, 14, alveolus for dentary tooth 1, 3, 4, 8, 11, 13, 14; asp, articular surface for the splenial; dsym, dentary symphysis; fen, fenestra; fo, foramen; Mc, Meckel’s canal; rdsym, rough dentary symphysis; sdsym, smooth dentary symphysis." figureDoi="http://doi.org/10.5281/zenodo.3768353" httpUri="https://zenodo.org/record/3768353/files/figure.png" pageId="57" pageNumber="58">Fig. 27C</figureCitation>
|
||
), and the caniniform is considerably less than twice as large as adjacent crowns (
|
||
<figureCitation box="[930,1032,272,298]" captionStart="Figure 20" captionStartId="49.[140,212,1462,1484]" captionTargetBox="[180,1068,388,1428]" captionTargetId="figure@49.[165,1082,363,1453]" captionTargetPageId="49" captionText="Figure 20. Unworn dentition of the crocodyliform Araripesuchus wegeneri. Detailed views of the anterior and middle portions of the tooth rows (MNN GAD20). A Left tooth rows in medial view. B Anterior portionof left tooth rows in medial view. C Middle portion of left tooth rows in medial view. Scale bar equals 1 cm in A and 5 mm in B and C. Abbreviations: ca, carina; d2, 4, 5, 7, 9, 11, 14, 16, dentary tooth 2, 4, 5, 7, 9, 11, 14, 16; m1, 3, 5, 7, 9, 13, maxillary tooth 1, 3, 5, 7, 9, 13; ne, neck; pm2–5, premaxillary tooth 2–5; rt, root." figureDoi="http://doi.org/10.5281/zenodo.3768341" httpUri="https://zenodo.org/record/3768341/files/figure.png" pageId="57" pageNumber="58">Fig. 20B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3768353" ID-Zenodo-Dep="3768353" httpUri="https://zenodo.org/record/3768353/files/figure.png" pageId="57" pageNumber="58" startId="57.[140,212,1526,1548]" targetBox="[294,941,368,1483]" targetPageId="57">
|
||
<paragraph blockId="57.[140,1109,1526,1676]" pageId="57" pageNumber="58">
|
||
<emphasis bold="true" box="[140,901,1526,1548]" pageId="57" pageNumber="58">
|
||
Figure 27. Right dentary of the crocodyliform
|
||
<taxonomicName authority="Sereno & Larsson, 2009" authorityName="Sereno & Larsson" authorityYear="2009" box="[611,840,1526,1548]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="rattoides" status="sp. n.">
|
||
<emphasis bold="true" box="[611,840,1526,1548]" italics="true" pageId="57" pageNumber="58">Araripesuchus rattoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[847,901,1526,1547]" pageId="57" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Isolated right dentary lacking teeth (CMN 41893).
|
||
<emphasis bold="true" box="[416,435,1558,1580]" pageId="57" pageNumber="58">A</emphasis>
|
||
Lateral view (reversed).
|
||
<emphasis bold="true" box="[661,678,1558,1580]" pageId="57" pageNumber="58">B</emphasis>
|
||
Medial view.
|
||
<emphasis bold="true" box="[807,826,1558,1580]" pageId="57" pageNumber="58">C</emphasis>
|
||
Dorsal view.
|
||
<emphasis bold="true" box="[952,971,1558,1580]" pageId="57" pageNumber="58">D</emphasis>
|
||
Anterior view. Scale bars equal 2 cm in A-C and 1 cm in D. Abbreviations:
|
||
<emphasis box="[700,914,1590,1612]" italics="true" pageId="57" pageNumber="58">ad1, 3, 4, 8, 11, 13, 14</emphasis>
|
||
, alveolus for dentary tooth 1, 3, 4, 8, 11, 13, 14;
|
||
<emphasis box="[393,421,1623,1644]" italics="true" pageId="57" pageNumber="58">asp</emphasis>
|
||
, articular surface for the splenial;
|
||
<emphasis box="[730,774,1622,1643]" italics="true" pageId="57" pageNumber="58">dsym</emphasis>
|
||
, dentary symphysis;
|
||
<emphasis box="[965,992,1622,1643]" italics="true" pageId="57" pageNumber="58">fen</emphasis>
|
||
, fenestra;
|
||
<emphasis box="[1085,1101,1622,1643]" italics="true" pageId="57" pageNumber="58">fo</emphasis>
|
||
, foramen;
|
||
<emphasis box="[229,258,1655,1676]" italics="true" pageId="57" pageNumber="58">Mc</emphasis>
|
||
, Meckel’s canal;
|
||
<emphasis box="[414,466,1654,1675]" italics="true" pageId="57" pageNumber="58">rdsym</emphasis>
|
||
, rough dentary symphysis;
|
||
<emphasis box="[721,771,1654,1675]" italics="true" pageId="57" pageNumber="58">sdsym</emphasis>
|
||
, smooth dentary symphysis.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3768355" ID-Zenodo-Dep="3768355" httpUri="https://zenodo.org/record/3768355/files/figure.png" pageId="58" pageNumber="59" startId="58.[140,212,774,796]" targetBox="[150,1091,170,747]" targetPageId="58">
|
||
<paragraph blockId="58.[140,1108,774,892]" pageId="58" pageNumber="59">
|
||
<emphasis bold="true" box="[140,868,774,796]" pageId="58" pageNumber="59">
|
||
Figure 28. Left dentary of the crocodyliform
|
||
<taxonomicName authority="Sereno & Larsson, 2009" authorityName="Sereno & Larsson" authorityYear="2009" box="[583,810,774,796]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="rattoides" status="sp. n.">
|
||
<emphasis bold="true" box="[583,810,774,796]" italics="true" pageId="58" pageNumber="59">Araripesuchus rattoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[815,868,774,795]" pageId="58" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Pencil drawing of isolated left dentary ramus lacking teeth (UCRC PV3).
|
||
<emphasis bold="true" box="[582,601,806,828]" pageId="58" pageNumber="59">A</emphasis>
|
||
Dorsal view.
|
||
<emphasis bold="true" box="[728,745,806,828]" pageId="58" pageNumber="59">B</emphasis>
|
||
Ventral view.
|
||
<emphasis bold="true" box="[876,895,806,828]" pageId="58" pageNumber="59">C</emphasis>
|
||
Medial view. Scale bar equals 1 cm. Parallel lines indicate broken bone surface. Abbreviations:
|
||
<emphasis box="[805,912,839,860]" italics="true" pageId="58" pageNumber="59">ad1, 4, 5, 8</emphasis>
|
||
, alveolus for dentary tooth 1, 4, 5, 8;
|
||
<emphasis box="[292,320,871,892]" italics="true" pageId="58" pageNumber="59">asp</emphasis>
|
||
, articular surface for the splenial;
|
||
<emphasis box="[634,661,871,892]" italics="true" pageId="58" pageNumber="59">fen</emphasis>
|
||
, fenestra;
|
||
<emphasis box="[756,772,871,892]" italics="true" pageId="58" pageNumber="59">fo</emphasis>
|
||
, foramen;
|
||
<emphasis box="[874,907,871,892]" italics="true" pageId="58" pageNumber="59">sym</emphasis>
|
||
, symphysis.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3768357" ID-Zenodo-Dep="3768357" httpUri="https://zenodo.org/record/3768357/files/figure.png" pageId="58" pageNumber="59" startId="58.[140,212,1494,1516]" targetBox="[153,1091,955,1472]" targetPageId="58">
|
||
<paragraph blockId="58.[140,1108,1494,1676]" pageId="58" pageNumber="59">
|
||
<emphasis bold="true" box="[140,1065,1494,1516]" pageId="58" pageNumber="59">
|
||
Figure 29. Computed-tomographic scan of the crocodyliform
|
||
<taxonomicName authority="Sereno & Larsson, 2009" authorityName="Sereno & Larsson" authorityYear="2009" box="[771,1001,1494,1516]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="rattoides" status="sp. n.">
|
||
<emphasis bold="true" box="[771,1001,1494,1516]" italics="true" pageId="58" pageNumber="59">Araripesuchus rattoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1010,1065,1494,1515]" pageId="58" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Isolated left dentary ramus lacking teeth (UCRC PV3).
|
||
<emphasis bold="true" box="[637,656,1526,1548]" pageId="58" pageNumber="59">A</emphasis>
|
||
Drawing in dorsal view showing the location of cross-sections (B-D).
|
||
<emphasis bold="true" box="[341,358,1558,1580]" pageId="58" pageNumber="59">B</emphasis>
|
||
Parasagittal section showing the size and orientation of the alveolus for dentary tooth 1.
|
||
<emphasis bold="true" box="[222,241,1590,1612]" pageId="58" pageNumber="59">C</emphasis>
|
||
Cross-section through the alveolus of the fourth dentary tooth.
|
||
<emphasis bold="true" box="[841,860,1590,1612]" pageId="58" pageNumber="59">D</emphasis>
|
||
Cross-section through the alveolus of the sixth dentary tooth. Scale bar for B-D equals 1 cm. Abbreviations:
|
||
<emphasis box="[905,995,1622,1644]" italics="true" pageId="58" pageNumber="59">ad1, 4, 6,</emphasis>
|
||
alveolus for dentary tooth 1, 4, 6;
|
||
<emphasis box="[345,378,1655,1676]" italics="true" pageId="58" pageNumber="59">sym</emphasis>
|
||
, symphysis;
|
||
<emphasis box="[495,513,1655,1676]" italics="true" pageId="58" pageNumber="59">vc</emphasis>
|
||
, vascular cavity.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="59.[140,1108,167,264]" pageId="59" pageNumber="60">
|
||
Tooth number may have been slightly greater in
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[751,877,167,193]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="59" pageNumber="60" phylum="Chordata" rank="species" species="rattoides">
|
||
<emphasis box="[751,877,167,193]" italics="true" pageId="59" pageNumber="60">A. rattoides</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName authorityName="Buffetaut & Taquet" authorityYear="1979" box="[927,1055,167,193]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="59" pageNumber="60" phylum="Chordata" rank="species" species="wegeneri">
|
||
<emphasis box="[927,1055,167,193]" italics="true" pageId="59" pageNumber="60">A. wegeneri</emphasis>
|
||
</taxonomicName>
|
||
, the largest postcaniniform teeth are d11 and d12 (Figs. 18B, 20A). In
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[885,1007,202,228]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="59" pageNumber="60" phylum="Chordata" rank="species" species="rattoides">
|
||
<emphasis box="[885,1007,202,228]" italics="true" pageId="59" pageNumber="60">A. rattoides</emphasis>
|
||
</taxonomicName>
|
||
the largest postcaniniform dentary teeth are d12 and d13 (
|
||
<figureCitation box="[715,799,237,263]" captionStart="Figure 27" captionStartId="57.[140,212,1526,1548]" captionTargetBox="[294,941,368,1483]" captionTargetId="figure@57.[268,980,351,1506]" captionTargetPageId="57" captionText="Figure 27. Right dentary of the crocodyliform Araripesuchus rattoides sp. n. Isolated right dentary lacking teeth (CMN 41893). A Lateral view (reversed). B Medial view. C Dorsal view. D Anterior view. Scale bars equal 2 cm in A-C and 1 cm in D. Abbreviations: ad1, 3, 4, 8, 11, 13, 14, alveolus for dentary tooth 1, 3, 4, 8, 11, 13, 14; asp, articular surface for the splenial; dsym, dentary symphysis; fen, fenestra; fo, foramen; Mc, Meckel’s canal; rdsym, rough dentary symphysis; sdsym, smooth dentary symphysis." figureDoi="http://doi.org/10.5281/zenodo.3768353" httpUri="https://zenodo.org/record/3768353/files/figure.png" pageId="59" pageNumber="60">Fig. 27</figureCitation>
|
||
,
|
||
<tableCitation box="[809,895,237,263]" captionStart="Table 9" captionStartId="59.[140,200,329,351]" captionText="Table 9. Dimensions (mm) of the alveoli in the holotype right dentary of Araripesuchus rattoides (CMN 41893). Width is labiolingual; length is mesiodistal. Parentheses indicate estimated measurement; dash indicates partially preserved alveolus, the dimension for which cannot be determined." pageId="59" pageNumber="60">Table 9</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<caption pageId="59" pageNumber="60">
|
||
<paragraph blockId="59.[140,1107,329,415]" pageId="59" pageNumber="60">
|
||
<emphasis bold="true" box="[140,227,329,351]" pageId="59" pageNumber="60">Table 9.</emphasis>
|
||
Dimensions (mm) of the alveoli in the holotype right dentary of
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[834,1036,329,350]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="59" pageNumber="60" phylum="Chordata" rank="species" species="rattoides">
|
||
<emphasis box="[834,1036,329,350]" italics="true" pageId="59" pageNumber="60">Araripesuchus rattoides</emphasis>
|
||
</taxonomicName>
|
||
(CMN 41893). Width is labiolingual; length is mesiodistal. Parentheses indicate estimated measurement; dash indicates partially preserved alveolus, the dimension for which cannot be determined.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="59" pageNumber="60">
|
||
<table box="[175,1042,444,985]" gridcols="4" gridrows="15" pageId="59" pageNumber="60">
|
||
<tr box="[175,1042,444,503]" gridrow="0" pageId="59" pageNumber="60">
|
||
<th box="[175,267,444,503]" gridcol="0" gridrow="0" pageId="59" pageNumber="60">
|
||
<emphasis bold="true" box="[175,267,444,468]" pageId="59" pageNumber="60">Alveolus</emphasis>
|
||
1
|
||
</th>
|
||
<th box="[326,413,444,503]" gridcol="1" gridrow="0" pageId="59" pageNumber="60">
|
||
<emphasis bold="true" box="[326,396,444,468]" pageId="59" pageNumber="60">Width</emphasis>
|
||
4.0
|
||
</th>
|
||
<th box="[449,548,444,503]" gridcol="2" gridrow="0" pageId="59" pageNumber="60">
|
||
<emphasis bold="true" box="[449,526,445,469]" pageId="59" pageNumber="60">Length</emphasis>
|
||
4.0
|
||
</th>
|
||
<th box="[563,1042,444,503]" gridcol="3" gridrow="0" pageId="59" pageNumber="60">
|
||
<emphasis bold="true" box="[772,891,444,468]" pageId="59" pageNumber="60">Comments</emphasis>
|
||
Enlarged incisiform tooth, subcircular alveolus
|
||
</th>
|
||
</tr>
|
||
<tr box="[175,1042,514,538]" gridrow="1" pageId="59" pageNumber="60" rowspan-3="1">
|
||
<th box="[175,267,514,538]" gridcol="0" gridrow="1" pageId="59" pageNumber="60">2</th>
|
||
<td box="[326,413,514,538]" gridcol="1" gridrow="1" pageId="59" pageNumber="60">2.6</td>
|
||
<td box="[449,548,514,538]" gridcol="2" gridrow="1" pageId="59" pageNumber="60">2.6</td>
|
||
</tr>
|
||
<tr box="[175,1042,548,572]" gridrow="2" pageId="59" pageNumber="60">
|
||
<th box="[175,267,548,572]" gridcol="0" gridrow="2" pageId="59" pageNumber="60">3</th>
|
||
<td box="[326,413,548,572]" gridcol="1" gridrow="2" pageId="59" pageNumber="60">2.4</td>
|
||
<td box="[449,548,548,572]" gridcol="2" gridrow="2" pageId="59" pageNumber="60">2.4</td>
|
||
<td box="[563,1042,548,572]" gridcol="3" gridrow="2" pageId="59" pageNumber="60">Smallest incisiform tooth, subcircular alveolus</td>
|
||
</tr>
|
||
<tr box="[175,1042,582,607]" gridrow="3" pageId="59" pageNumber="60">
|
||
<th box="[175,267,582,607]" gridcol="0" gridrow="3" pageId="59" pageNumber="60">4</th>
|
||
<td box="[326,413,582,607]" gridcol="1" gridrow="3" pageId="59" pageNumber="60">5.1</td>
|
||
<td box="[449,548,582,607]" gridcol="2" gridrow="3" pageId="59" pageNumber="60">6.3</td>
|
||
<td box="[563,1042,582,607]" gridcol="3" gridrow="3" pageId="59" pageNumber="60">Caniniform tooth</td>
|
||
</tr>
|
||
<tr box="[175,1042,617,641]" gridrow="4" pageId="59" pageNumber="60" rowspan-3="1">
|
||
<th box="[175,267,617,641]" gridcol="0" gridrow="4" pageId="59" pageNumber="60">5</th>
|
||
<td box="[326,413,617,641]" gridcol="1" gridrow="4" pageId="59" pageNumber="60">3.0</td>
|
||
<td box="[449,548,617,641]" gridcol="2" gridrow="4" pageId="59" pageNumber="60">3.5</td>
|
||
</tr>
|
||
<tr box="[175,1042,651,676]" gridrow="5" pageId="59" pageNumber="60" rowspan-3="1">
|
||
<th box="[175,267,651,676]" gridcol="0" gridrow="5" pageId="59" pageNumber="60">6</th>
|
||
<td box="[326,413,651,676]" gridcol="1" gridrow="5" pageId="59" pageNumber="60">2.4</td>
|
||
<td box="[449,548,651,676]" gridcol="2" gridrow="5" pageId="59" pageNumber="60">2.4</td>
|
||
</tr>
|
||
<tr box="[175,1042,686,710]" gridrow="6" pageId="59" pageNumber="60">
|
||
<th box="[175,267,686,710]" gridcol="0" gridrow="6" pageId="59" pageNumber="60">7</th>
|
||
<td box="[326,413,686,710]" gridcol="1" gridrow="6" pageId="59" pageNumber="60">1.9</td>
|
||
<td box="[449,548,686,710]" gridcol="2" gridrow="6" pageId="59" pageNumber="60">2.0</td>
|
||
<td box="[563,1042,686,710]" gridcol="3" gridrow="6" pageId="59" pageNumber="60">Smallest tooth, subcircular alveolus</td>
|
||
</tr>
|
||
<tr box="[175,1042,720,745]" gridrow="7" pageId="59" pageNumber="60">
|
||
<th box="[175,267,720,745]" gridcol="0" gridrow="7" pageId="59" pageNumber="60">8</th>
|
||
<td box="[326,413,720,745]" gridcol="1" gridrow="7" pageId="59" pageNumber="60">2.0</td>
|
||
<td box="[449,548,720,745]" gridcol="2" gridrow="7" pageId="59" pageNumber="60">2.3</td>
|
||
<td box="[563,1042,720,745]" gridcol="3" gridrow="7" pageId="59" pageNumber="60">Second smallest tooth, subcircular alveolus</td>
|
||
</tr>
|
||
<tr box="[175,1042,755,779]" gridrow="8" pageId="59" pageNumber="60" rowspan-3="1">
|
||
<th box="[175,267,755,779]" gridcol="0" gridrow="8" pageId="59" pageNumber="60">9</th>
|
||
<td box="[326,413,755,779]" gridcol="1" gridrow="8" pageId="59" pageNumber="60">2.1</td>
|
||
<td box="[449,548,755,779]" gridcol="2" gridrow="8" pageId="59" pageNumber="60">2.9</td>
|
||
</tr>
|
||
<tr box="[175,1042,790,814]" gridrow="9" pageId="59" pageNumber="60" rowspan-3="1">
|
||
<th box="[175,267,790,814]" gridcol="0" gridrow="9" pageId="59" pageNumber="60">10</th>
|
||
<td box="[326,413,790,814]" gridcol="1" gridrow="9" pageId="59" pageNumber="60">2.3</td>
|
||
<td box="[449,548,790,814]" gridcol="2" gridrow="9" pageId="59" pageNumber="60">3.3</td>
|
||
</tr>
|
||
<tr box="[175,1042,824,848]" gridrow="10" pageId="59" pageNumber="60" rowspan-3="1">
|
||
<th box="[175,267,824,848]" gridcol="0" gridrow="10" pageId="59" pageNumber="60">11</th>
|
||
<td box="[326,413,824,848]" gridcol="1" gridrow="10" pageId="59" pageNumber="60">3.2</td>
|
||
<td box="[449,548,824,848]" gridcol="2" gridrow="10" pageId="59" pageNumber="60">4.0</td>
|
||
</tr>
|
||
<tr box="[175,1042,858,882]" gridrow="11" pageId="59" pageNumber="60" rowspan-3="1">
|
||
<th box="[175,267,858,882]" gridcol="0" gridrow="11" pageId="59" pageNumber="60">12</th>
|
||
<td box="[326,413,858,882]" gridcol="1" gridrow="11" pageId="59" pageNumber="60">(4.0)</td>
|
||
<td box="[449,548,858,882]" gridcol="2" gridrow="11" pageId="59" pageNumber="60">5.0</td>
|
||
</tr>
|
||
<tr box="[175,1042,893,917]" gridrow="12" pageId="59" pageNumber="60">
|
||
<th box="[175,267,893,917]" gridcol="0" gridrow="12" pageId="59" pageNumber="60">13</th>
|
||
<td box="[326,413,893,917]" gridcol="1" gridrow="12" pageId="59" pageNumber="60">—</td>
|
||
<td box="[449,548,893,917]" gridcol="2" gridrow="12" pageId="59" pageNumber="60">5.3</td>
|
||
<td box="[563,1042,893,917]" gridcol="3" gridrow="12" pageId="59" pageNumber="60">Largest tooth</td>
|
||
</tr>
|
||
<tr box="[175,1042,927,951]" gridrow="13" pageId="59" pageNumber="60" rowspan-3="1">
|
||
<th box="[175,267,927,951]" gridcol="0" gridrow="13" pageId="59" pageNumber="60">14</th>
|
||
<td box="[326,413,927,951]" gridcol="1" gridrow="13" pageId="59" pageNumber="60">—</td>
|
||
<td box="[449,548,927,951]" gridcol="2" gridrow="13" pageId="59" pageNumber="60">3.8</td>
|
||
</tr>
|
||
<tr box="[175,1042,961,985]" gridrow="14" pageId="59" pageNumber="60" rowspan-3="1">
|
||
<th box="[175,267,961,985]" gridcol="0" gridrow="14" pageId="59" pageNumber="60">15</th>
|
||
<td box="[326,413,961,985]" gridcol="1" gridrow="14" pageId="59" pageNumber="60">—</td>
|
||
<td box="[449,548,961,985]" gridcol="2" gridrow="14" pageId="59" pageNumber="60">—</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3768361" ID-Zenodo-Dep="3768361" httpUri="https://zenodo.org/record/3768361/files/figure.png" pageId="59" pageNumber="60" startId="59.[140,212,1462,1484]" targetBox="[162,1079,1108,1429]" targetPageId="59">
|
||
<paragraph blockId="59.[140,1108,1462,1676]" pageId="59" pageNumber="60">
|
||
<emphasis bold="true" box="[140,1108,1462,1484]" pageId="59" pageNumber="60">
|
||
Figure 30. Reconstruction of the dentition of the crocodyliform
|
||
<taxonomicName authority="Sereno & Larsson, 2009" authorityName="Sereno & Larsson" authorityYear="2009" box="[809,1043,1462,1484]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="59" pageNumber="60" phylum="Chordata" rank="species" species="rattoides" status="sp. n.">
|
||
<emphasis bold="true" box="[809,1043,1462,1484]" italics="true" pageId="59" pageNumber="60">Araripesuchus rattoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1051,1108,1462,1483]" pageId="59" pageNumber="60" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Anterior dentition restored based on the size and orientation of the alveoli in CMN 41893 and UCRC PV3.
|
||
<emphasis bold="true" box="[197,216,1526,1548]" pageId="59" pageNumber="60">A</emphasis>
|
||
Dorsal view.
|
||
<emphasis bold="true" box="[351,368,1526,1548]" pageId="59" pageNumber="60">B</emphasis>
|
||
Ventral view.
|
||
<emphasis bold="true" box="[509,528,1526,1548]" pageId="59" pageNumber="60">C</emphasis>
|
||
Anterolateral view with premaxillary and anterior maxillary dentition restored to match those in the dentary. Given the presence of large, adjacent first dentary teeth, there may have been a median diastema between the premaxillary teeth and one or two fewer teeth in each premaxillary tooth row. Scale bar equals 2 cm in A and B. Abbreviations:
|
||
<emphasis box="[872,943,1622,1644]" italics="true" pageId="59" pageNumber="60">d1, 4, 8</emphasis>
|
||
, dentary tooth 1, 4, 8;
|
||
<emphasis box="[189,200,1654,1675]" italics="true" pageId="59" pageNumber="60">d</emphasis>
|
||
, dentary;
|
||
<emphasis box="[296,314,1654,1675]" italics="true" pageId="59" pageNumber="60">di</emphasis>
|
||
, diastema;
|
||
<emphasis box="[421,450,1655,1676]" italics="true" pageId="59" pageNumber="60">m1</emphasis>
|
||
, maxillary tooth 1;
|
||
<emphasis box="[637,677,1655,1676]" italics="true" pageId="59" pageNumber="60">pm1</emphasis>
|
||
,
|
||
<emphasis box="[690,702,1655,1676]" italics="true" pageId="59" pageNumber="60">2</emphasis>
|
||
, premaxillary tooth 1, 2.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="60.[140,1108,167,475]" pageId="60" pageNumber="61">
|
||
Other features in
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" box="[380,500,167,193]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="rattoides">
|
||
<emphasis box="[380,500,167,193]" italics="true" pageId="60" pageNumber="61">A. rattoides</emphasis>
|
||
</taxonomicName>
|
||
confirm its status as a species of
|
||
<taxonomicName baseAuthorityName="Pol and Apesteguia" baseAuthorityYear="2005" box="[860,1007,167,193]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="60" pageNumber="61" phylum="Chordata" rank="genus">
|
||
<emphasis box="[860,1007,167,193]" italics="true" pageId="60" pageNumber="61">Araripesuchus</emphasis>
|
||
</taxonomicName>
|
||
. Both
|
||
<taxonomicName authorityName="Sereno & Larsson" authorityYear="2009" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="rattoides">
|
||
<emphasis italics="true" pageId="60" pageNumber="61">A. rattoides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Buffetaut & Taquet" authorityYear="1979" box="[285,409,202,228]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="wegeneri">
|
||
<emphasis box="[285,409,202,228]" italics="true" pageId="60" pageNumber="61">A. wegeneri</emphasis>
|
||
</taxonomicName>
|
||
have an unusual anterior extension of the articular scar for the splenial located dorsal to the symphysis on the subhorizontal palatal surface. This articular extension of the splenial, which is located medial to the alveoli for d4–6 (
|
||
<figureCitation box="[991,1094,272,298]" captionStart="Figure 27" captionStartId="57.[140,212,1526,1548]" captionTargetBox="[294,941,368,1483]" captionTargetId="figure@57.[268,980,351,1506]" captionTargetPageId="57" captionText="Figure 27. Right dentary of the crocodyliform Araripesuchus rattoides sp. n. Isolated right dentary lacking teeth (CMN 41893). A Lateral view (reversed). B Medial view. C Dorsal view. D Anterior view. Scale bars equal 2 cm in A-C and 1 cm in D. Abbreviations: ad1, 3, 4, 8, 11, 13, 14, alveolus for dentary tooth 1, 3, 4, 8, 11, 13, 14; asp, articular surface for the splenial; dsym, dentary symphysis; fen, fenestra; fo, foramen; Mc, Meckel’s canal; rdsym, rough dentary symphysis; sdsym, smooth dentary symphysis." figureDoi="http://doi.org/10.5281/zenodo.3768353" httpUri="https://zenodo.org/record/3768353/files/figure.png" pageId="60" pageNumber="61">Fig. 27C</figureCitation>
|
||
), is continuous posteriorly with the more typical vertical splenial attachment scar dorsal to Meckel’s canal.
|
||
<taxonomicName authorityName="Buffetaut & Taquet" authorityYear="1979" box="[351,477,343,369]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="wegeneri">
|
||
<emphasis box="[351,477,343,369]" italics="true" pageId="60" pageNumber="61">A. wegeneri</emphasis>
|
||
</taxonomicName>
|
||
shows a similar articular extension of the splenial (Fig. 18C). The alveoli posterior to d11, in addition, are open medially with alveolar septa poorly developed as low rounded ridges (
|
||
<figureCitation box="[595,730,413,439]" captionStart="Figure 27" captionStartId="57.[140,212,1526,1548]" captionTargetBox="[294,941,368,1483]" captionTargetId="figure@57.[268,980,351,1506]" captionTargetPageId="57" captionText="Figure 27. Right dentary of the crocodyliform Araripesuchus rattoides sp. n. Isolated right dentary lacking teeth (CMN 41893). A Lateral view (reversed). B Medial view. C Dorsal view. D Anterior view. Scale bars equal 2 cm in A-C and 1 cm in D. Abbreviations: ad1, 3, 4, 8, 11, 13, 14, alveolus for dentary tooth 1, 3, 4, 8, 11, 13, 14; asp, articular surface for the splenial; dsym, dentary symphysis; fen, fenestra; fo, foramen; Mc, Meckel’s canal; rdsym, rough dentary symphysis; sdsym, smooth dentary symphysis." figureDoi="http://doi.org/10.5281/zenodo.3768353" httpUri="https://zenodo.org/record/3768353/files/figure.png" pageId="60" pageNumber="61">Fig. 27B, C</figureCitation>
|
||
). A similar condition is present in
|
||
<taxonomicName authorityName="Buffetaut & Taquet" authorityYear="1979" box="[140,264,449,475]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="wegeneri">
|
||
<emphasis box="[140,264,449,475]" italics="true" pageId="60" pageNumber="61">A. wegeneri</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 18B) and some other species (Fig. 31C) (
|
||
<bibRefCitation author="Pol D & Apesteguia S" box="[787,1070,448,475]" pageId="60" pageNumber="61" pagination="1 - 38" refId="ref46568" refString="Pol D, Apesteguia S (2005) New Araripesuchus remains from the early Late Cretaceous (Cenomanian-Turonian) of Patagonia. American Museum Novitates 3490: 1 - 38." type="journal article" year="2005">Pol and Apesteguia 2005</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |