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<document id="0AF0F0E4D25EB9327EE889E0C33FD66B" ID-CLB-Dataset="54188" ID-DOI="10.11646/zootaxa.5244.5.2" ID-GBIF-Dataset="77045783-e389-456c-8405-402d9a928da1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7663766" ID-ZooBank="B55396DA-15DB-4CDF-81FD-78686A0EC3A2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1677051835230" checkinUser="plazi" docAuthor="Ng, Peter K. L. &amp; Rahayu, Dwi Listyo" docDate="2023" docId="039087CD7E0B733847C6F942F59CFCCE" docLanguage="en" docName="zootaxa.5244.5.2.pdf" docOrigin="Zootaxa 5244 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5244.5.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lophoplax pannosa Ng &amp; Rahayu 2023, n. sp." docType="treatment" docVersion="6" lastPageNumber="444" masterDocId="FFA9FFB57E0573284751FF9DF768FFDA" masterDocTitle="Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura)" masterLastPageNumber="454" masterPageNumber="428" pageNumber="442" updateTime="1699475659855" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="14145A3FEC69DF096BFE8AAA4C432EC1">Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura)</mods:title>
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<mods:namePart id="F8D2C7B88E70B8EC9DE5C79D7E2DA305">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="F51989B5BC978B174B643BC055AC660D">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore. peterng @ nus. edu. sg; https: // orcid. org / 0000 - 0001 - 5946 - 0608</mods:affiliation>
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<mods:namePart id="DB9D90A8BA24CFFF7434C32DD95F4DF2">Rahayu, Dwi Listyo</mods:namePart>
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<mods:affiliation id="1B16F649F5250101903E3708F3C7241B">Research Center for Marine and Land Bioindustry, National Research and Innovation Agency, Dusun Teluk Kodek, Pemenang, Lombok Utara 83756, NTB, Indonesia. dwilistyo @ yahoo. com; https: // orcid. org / 0000 - 0003 - 4342 - 9045</mods:affiliation>
<mods:nameIdentifier id="6AC49A572775810FF4C3862A0AB47C6E" type="email">dwilistyo@yahoo.com</mods:nameIdentifier>
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<taxonomicName id="4C394D587E0B732647C6F942F6E0F920" ID-CoL="3bf981ac-668c-4b35-84b0-c4b774152b24" authority="Ng &amp; Rahayu, 2023" authorityName="Ng &amp; Rahayu" authorityYear="2023" box="[151,392,1759,1786]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="442" phylum="Arthropoda" rank="species" species="pannosa" status="sp. nov.">
<emphasis id="B94DEAC97E0B732647C6F942F6E0F920" bold="true" box="[151,392,1759,1786]" italics="true" pageId="14" pageNumber="442">Lophoplax pannosa</emphasis>
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<taxonomicNameLabel id="A27E57B27E0B732646DEF942F6B9F923" box="[399,465,1759,1785]" pageId="14" pageNumber="442" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8B8636DB7E0B732647C6F89EF54CF8C7" blockId="14.[151,548,1759,1822]" box="[151,548,1795,1822]" pageId="14" pageNumber="442">
(
<figureCitation id="13022A5E7E0B732647CEF89EF66BF8C7" box="[159,259,1795,1821]" captionStart="FIGURE 2" captionStartId="5.[151,250,1775,1799]" captionTargetBox="[151,1435,309,1752]" captionTargetId="figure-91@5.[151,1435,309,1752]" captionTargetPageId="5" captionText="FIGURE 2. Overall dorsal view. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; D, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; E, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; F, Lophoplax vermiculata, subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; H, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663772" httpUri="https://zenodo.org/record/7663772/files/figure.png" pageId="14" pageNumber="442">Figs. 2H</figureCitation>
,
<figureCitation id="13022A5E7E0B7326465EF89EF65CF8C7" box="[271,308,1795,1821]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="14" pageNumber="442">4D</figureCitation>
,
<figureCitation id="13022A5E7E0B7326466EF89EF614F8C4" box="[319,380,1795,1822]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. AF, frontal view of cephalothorax; GJ, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="14" pageNumber="442">5F, L</figureCitation>
,
<figureCitation id="13022A5E7E0B732646D9F89EF6CEF8C7" box="[392,422,1795,1821]" captionStart="FIGURE 6" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,813,1703]" captionTargetId="figure-327@11.[151,1436,813,1703]" captionTargetPageId="11" captionText="FIGURE 6. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu.A, B, D, E, dorsal view of right cheliped; C, dorsal view of left cheliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663784" httpUri="https://zenodo.org/record/7663784/files/figure.png" pageId="14" pageNumber="442">6F</figureCitation>
,
<figureCitation id="13022A5E7E0B732646E0F899F685F8C4" box="[433,493,1796,1822]" captionStart="FIGURE 7" captionStartId="12.[151,250,1823,1847]" captionTargetBox="[229,1359,181,1799]" captionTargetId="figure-17@12.[229,1359,181,1799]" captionTargetPageId="12" captionText="FIGURE 7. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, B, right chela; CF, left chela; GL, right P4." figureDoi="http://doi.org/10.5281/zenodo.7663790" httpUri="https://zenodo.org/record/7663790/files/figure.png" pageId="14" pageNumber="442">7F, L</figureCitation>
,
<figureCitation id="13022A5E7E0B732646A8F89EF574F8C7" box="[505,540,1795,1821]" captionStart="FIGURE 8" captionStartId="13.[151,250,1105,1129]" captionTargetBox="[151,1435,181,1081]" captionTargetId="figure-25@13.[151,1435,181,1081]" captionTargetPageId="13" captionText="FIGURE 8. A, B, Lophoplax sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, female thoracic sternum and pleon; BD, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.7663793" httpUri="https://zenodo.org/record/7663793/files/figure.png" pageId="14" pageNumber="442">8D</figureCitation>
)
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<emphasis id="B94DEAC97E0B732647C6F8D6F656F8BF" bold="true" box="[151,318,1867,1893]" pageId="14" pageNumber="442">Type material</emphasis>
.
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<typeStatus id="548288797E0B7326461CF8D6F6DCF8BF" box="[333,436,1867,1893]" pageId="14" pageNumber="442" type="holotype">Holotype</typeStatus>
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: ovigerous female (7.4 ×
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) (
<specimenCode id="DB9F9EA07E0B73264417F8D6F369F8BF" box="[838,1025,1867,1893]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="14" pageNumber="442">ZRC 2019.1190</specimenCode>
), station LD07,
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, coll. SANTO 2006
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<collectingDate id="EFC3E9F37E0B7326441EF8F2F341F853" box="[847,1065,1903,1929]" pageId="14" pageNumber="442" value="2007-09-28">28 September 2007</collectingDate>
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<emphasis id="B94DEAC97E0B73264796F80EF6D9F877" bold="true" box="[199,433,1939,1965]" pageId="14" pageNumber="442">Diagnosis of female</emphasis>
. Carapace subquadrate (
<figureCitation id="13022A5E7E0B73264598F80EF44DF877" box="[713,805,1939,1965]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="14" pageNumber="442">Fig. 4D</figureCitation>
); epigastric, protogastric, hepatic, cardiac and intestinal areolets glabrous, prominent, smooth; epigastric areolets transversely narrow,relatively longer, not fused with protogastric areolets (
<figureCitation id="13022A5E7E0B732646C2F846F69AF82F" box="[403,498,2011,2037]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="14" pageNumber="442">Fig. 4D</figureCitation>
); protogastric areolet relatively transversely narrower, obliquely positioned (
<figureCitation id="13022A5E7E0B7327423EF846F7D6FF6B" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" lastPageId="15" lastPageNumber="443" pageId="14" pageNumber="442">Fig. 4D</figureCitation>
); cardiac areolet transverse in position (
<figureCitation id="13022A5E7E0A7327452FFF0AF5B1FF6B" box="[638,729,151,177]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="15" pageNumber="443">Fig. 4D</figureCitation>
); hepatic areolet prominent, more elongate, reaching to second anterolateral tooth (
<figureCitation id="13022A5E7E0A73274624FF26F6B9FF0F" box="[373,465,187,213]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="15" pageNumber="443">Fig. 4D</figureCitation>
); external orbital tooth triangular (
<figureCitation id="13022A5E7E0A73274408FF26F4DDFF0F" box="[857,949,187,213]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="15" pageNumber="443">Fig. 4D</figureCitation>
); tips of anterolateral teeth sharply pointed (
<figureCitation id="13022A5E7E0A732747CEFF42F792FF23" box="[159,250,223,249]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="15" pageNumber="443">Fig. 4D</figureCitation>
); junction between antero- and posterolateral margins with 2 low lobes, not swollen or tuberculiform (
<figureCitation id="13022A5E7E0A7327423EFF42F7D6FEC7" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="15" pageNumber="443">Fig. 4D</figureCitation>
); epistome relatively wider longitudinally (
<figureCitation id="13022A5E7E0A732745F2FE9EF591FEC7" box="[675,761,258,285]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. AF, frontal view of cephalothorax; GJ, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="15" pageNumber="443">Fig. 5F</figureCitation>
); median lobe of posterior margin of the epistome relatively low with lateral margins gently concave (
<figureCitation id="13022A5E7E0A7327453FFEBAF5ADFE9B" box="[622,709,294,321]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. AF, frontal view of cephalothorax; GJ, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="15" pageNumber="443">Fig. 5F</figureCitation>
). Third maxilliped with merus quadrate, ischium subrectangular (
<figureCitation id="13022A5E7E0A732747CEFED6F79EFEBF" box="[159,246,330,357]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. AF, frontal view of cephalothorax; GJ, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="15" pageNumber="443">Fig. 5L</figureCitation>
). Dorsal and lateral surfaces of carpus of cheliped with 5 weakly defined areolets, deeply eroded (
<figureCitation id="13022A5E7E0A7327426BFED6F2E6FEBF" box="[1338,1422,331,357]" captionStart="FIGURE 6" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,813,1703]" captionTargetId="figure-327@11.[151,1436,813,1703]" captionTargetPageId="11" captionText="FIGURE 6. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu.A, B, D, E, dorsal view of right cheliped; C, dorsal view of left cheliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663784" httpUri="https://zenodo.org/record/7663784/files/figure.png" pageId="15" pageNumber="443">Fig. 6F</figureCitation>
); cheliped fingers not distinctly bent, pollex subparallel with the ventral margin of palm (
<figureCitation id="13022A5E7E0A73274327FEF2F3A4FE53" box="[1142,1228,367,393]" captionStart="FIGURE 7" captionStartId="12.[151,250,1823,1847]" captionTargetBox="[229,1359,181,1799]" captionTargetId="figure-17@12.[229,1359,181,1799]" captionTargetPageId="12" captionText="FIGURE 7. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, B, right chela; CF, left chela; GL, right P4." figureDoi="http://doi.org/10.5281/zenodo.7663790" httpUri="https://zenodo.org/record/7663790/files/figure.png" pageId="15" pageNumber="443">Fig. 7F</figureCitation>
). P2P5 relatively not elongate (
<figureCitation id="13022A5E7E0A73274661FE0EF6FCFE77" box="[304,404,403,429]" captionStart="FIGURE 2" captionStartId="5.[151,250,1775,1799]" captionTargetBox="[151,1435,309,1752]" captionTargetId="figure-91@5.[151,1435,309,1752]" captionTargetPageId="5" captionText="FIGURE 2. Overall dorsal view. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; D, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; E, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; F, Lophoplax vermiculata, subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; H, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663772" httpUri="https://zenodo.org/record/7663772/files/figure.png" pageId="15" pageNumber="443">Figs. 2H</figureCitation>
,
<figureCitation id="13022A5E7E0A732746F1FE0EF6AAFE77" box="[416,450,403,429]" captionStart="FIGURE 7" captionStartId="12.[151,250,1823,1847]" captionTargetBox="[229,1359,181,1799]" captionTargetId="figure-17@12.[229,1359,181,1799]" captionTargetPageId="12" captionText="FIGURE 7. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, B, right chela; CF, left chela; GL, right P4." figureDoi="http://doi.org/10.5281/zenodo.7663790" httpUri="https://zenodo.org/record/7663790/files/figure.png" pageId="15" pageNumber="443">7L</figureCitation>
); outer surface of P3P5 merus, carpus and propodus gently rugose (
<figureCitation id="13022A5E7E0A73274397FE0EF275FE77" box="[1222,1309,403,429]" captionStart="FIGURE 7" captionStartId="12.[151,250,1823,1847]" captionTargetBox="[229,1359,181,1799]" captionTargetId="figure-17@12.[229,1359,181,1799]" captionTargetPageId="12" captionText="FIGURE 7. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, B, right chela; CF, left chela; GL, right P4." figureDoi="http://doi.org/10.5281/zenodo.7663790" httpUri="https://zenodo.org/record/7663790/files/figure.png" pageId="15" pageNumber="443">Fig. 7L</figureCitation>
).
</paragraph>
<caption id="DF4666537E0A732747C6F813F201F834" ID-DOI="http://doi.org/10.5281/zenodo.7663799" ID-Zenodo-Dep="7663799" httpUri="https://zenodo.org/record/7663799/files/figure.png" pageId="15" pageNumber="443" startId="15.[151,247,1934,1958]" targetBox="[227,1360,463,1902]" targetPageId="15">
<paragraph id="8B8636DB7E0A732747C6F813F201F834" blockId="15.[151,1436,1934,2031]" pageId="15" pageNumber="443">
<emphasis id="B94DEAC97E0A732747C6F813F673F87C" bold="true" box="[151,283,1934,1958]" pageId="15" pageNumber="443">FIGURE 10.</emphasis>
AE,
<taxonomicName id="4C394D587E0A73274606F813F5D4F87C" authority="(Stimpson, 1858)" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[343,700,1934,1958]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="443" phylum="Arthropoda" rank="species" species="sculpta">
<emphasis id="B94DEAC97E0A73274606F813F564F87C" box="[343,524,1934,1958]" italics="true" pageId="15" pageNumber="443">Lophoplax sculpta</emphasis>
(
<bibRefCitation id="EFA84B2A7E0A73274549F813F5DDF87C" author="Stimpson, W." box="[536,693,1934,1958]" pageId="15" pageNumber="443" pagination="93 - 110" refId="ref18446" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 (39 - 56). https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
)
</taxonomicName>
, neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; FJ,
<taxonomicName id="4C394D587E0A73274268F812F2F4F87C" authority="Ng &amp; Rahayu, 2023" authorityName="Ng &amp; Rahayu" authorityYear="2023" box="[1337,1436,1934,1958]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="443" phylum="Arthropoda" rank="species" species="sordida" status="sp. nov.">
<emphasis id="B94DEAC97E0A73274268F812F2F4F87C" box="[1337,1436,1934,1958]" italics="true" pageId="15" pageNumber="443">L. sordida</emphasis>
</taxonomicName>
<emphasis id="B94DEAC97E0A732747C6F82FF7A7F810" bold="true" box="[151,207,1970,1994]" pageId="15" pageNumber="443">
<taxonomicNameLabel id="A27E57B27E0A732747C6F82FF7A7F810" box="[151,207,1970,1994]" pageId="15" pageNumber="443" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok. A, B, pleon; B, G, left G1 (ventral view); C, H, distal part of left G1 (ventral view); D, I, left G1 (dorsal view); E, J, left G2. Scales: A, F = 1.0 mm; B, E, G, J = 0.5 mm; C, D, H, I = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B8636DB7E1573384796FF0BF690FF6A" blockId="16.[151,1437,150,789]" box="[199,504,150,177]" pageId="16" pageNumber="444">
<emphasis id="B94DEAC97E1573384796FF0BF60DFF6A" bold="true" box="[199,357,150,176]" pageId="16" pageNumber="444">Colour in life</emphasis>
. Not known.
</paragraph>
</subSubSection>
<subSubSection id="C32365507E1573384796FF27F5F7FF22" pageId="16" pageNumber="444" type="etymology">
<paragraph id="8B8636DB7E1573384796FF27F5F7FF22" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="444">
<emphasis id="B94DEAC97E1573384796FF27F620FF0E" bold="true" box="[199,328,186,212]" pageId="16" pageNumber="444">Etymology</emphasis>
. The name is derived from the Latin “pannosus” for scruffy or unkempt, alluding to the general appearance of the species, even after cleaning.
</paragraph>
</subSubSection>
<subSubSection id="C32365507E1573384796FE9FF64CFD2A" pageId="16" pageNumber="444" type="discussion">
<paragraph id="8B8636DB7E1573384796FE9FF2E3FD72" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="444">
<emphasis id="B94DEAC97E1573384796FE9FF65CFEC6" bold="true" box="[199,308,258,284]" pageId="16" pageNumber="444">Remarks</emphasis>
.
<taxonomicName id="4C394D587E1573384616FE9FF542FEC6" authority="Ng &amp; Rahayu, 2023" authorityName="Ng &amp; Rahayu" authorityYear="2023" box="[327,554,258,284]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="444" phylum="Arthropoda" rank="species" species="pannosa" status="sp. nov.">
<emphasis id="B94DEAC97E1573384616FE9FF542FEC6" box="[327,554,258,284]" italics="true" pageId="16" pageNumber="444">Lophoplax pannosa</emphasis>
</taxonomicName>
<emphasis id="B94DEAC97E1573384567FE9FF512FEC6" bold="true" box="[566,634,258,284]" pageId="16" pageNumber="444">
<taxonomicNameLabel id="A27E57B27E1573384567FE9FF512FEC6" box="[566,634,258,284]" pageId="16" pageNumber="444" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C394D587E157338444FFE9EF4FAFEC6" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[798,914,258,284]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="444" phylum="Arthropoda" rank="species" species="scuplta">
<emphasis id="B94DEAC97E157338444FFE9EF4FAFEC6" box="[798,914,258,284]" italics="true" pageId="16" pageNumber="444">L. scuplta</emphasis>
</taxonomicName>
in that the epigastric areolet is transversely narrower and longer (
<figureCitation id="13022A5E7E15733846D9FEBBF6B7FE9A" box="[392,479,294,320]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="16" pageNumber="444">Fig. 4D</figureCitation>
) (versus relatively wider and shorter;
<figureCitation id="13022A5E7E15733844D4FEBBF497FE9A" box="[901,1023,294,320]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="16" pageNumber="444">Fig. 3AC</figureCitation>
); the protogastric areolet is relatively transversely narrower and obliquely positioned (
<figureCitation id="13022A5E7E15733845E1FED7F46EFEBE" box="[688,774,330,356]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="16" pageNumber="444">Fig. 4D</figureCitation>
) (versus transversely wider and subparallel in position;
<figureCitation id="13022A5E7E1573384221FED7F7B7FE52" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="16" pageNumber="444">Fig. 3AC</figureCitation>
); the cardiac areolet is obliquely positioned (
<figureCitation id="13022A5E7E1573384589FEF3F458FE52" box="[728,816,366,392]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="16" pageNumber="444">Fig. 4D</figureCitation>
) (versus clearly transverse in position;
<figureCitation id="13022A5E7E15733843BAFEF3F20EFE52" box="[1259,1382,366,392]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="16" pageNumber="444">Fig. 3AC</figureCitation>
); the hepatic areolet is proportionately longer, reaching to the beginning of the second anterolateral tooth (
<figureCitation id="13022A5E7E15733843B9FE0FF256FE76" box="[1256,1342,402,428]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="16" pageNumber="444">Fig. 4D</figureCitation>
) (versus shorter, reaching only to the base of the first tooth;
<figureCitation id="13022A5E7E1573384587FE2BF438FE0A" box="[726,848,438,464]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="16" pageNumber="444">Fig. 3AC</figureCitation>
); the junction between the antero- and posterolateral margins is demarcated by two low lobes but neither swollen or tuberculiform (
<figureCitation id="13022A5E7E1573384369FE47F3FCFE2E" box="[1080,1172,474,500]" captionStart="FIGURE 4" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[151,1436,417,1430]" captionTargetId="figure-139@7.[151,1436,417,1430]" captionTargetPageId="7" captionText="FIGURE 4. Dorsal view of carapace. A, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="16" pageNumber="444">Fig. 4D</figureCitation>
) (versus junction with two large rounded to subtruncate tubercles with the tips rounded to flattened, often with median depression;
<figureCitation id="13022A5E7E1573384234FE63F7B4FDE6" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="16" pageNumber="444">Figs. 3AC</figureCitation>
,
<figureCitation id="13022A5E7E15733847B6FDBFF665FDE6" box="[231,269,546,572]" captionStart="FIGURE 9" captionStartId="14.[151,250,1509,1533]" captionTargetBox="[176,1410,181,1485]" captionTargetId="figure-17@14.[176,1410,181,1485]" captionTargetPageId="14" captionText="FIGURE 9. AD, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia. A, right lateral view of posterior part of sub-branchial region; B, EG, anterior male thoracic sternum and pleon; C, male pleonal somites 14; D, male pleonal somites 13." figureDoi="http://doi.org/10.5281/zenodo.7663797" httpUri="https://zenodo.org/record/7663797/files/figure.png" pageId="16" pageNumber="444">9A</figureCitation>
); the median lobe of the posterior margin of the epistome is relatively low with the lateral margins gently concave (
<figureCitation id="13022A5E7E1573384655FDDBF631FDBB" box="[260,345,582,609]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. AF, frontal view of cephalothorax; GJ, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="16" pageNumber="444">Fig. 5F</figureCitation>
) (versus median lobe is more salient with the lateral margins deeply concave;
<figureCitation id="13022A5E7E15733843B1FDDBF234FDBA" box="[1248,1372,582,609]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. AF, frontal view of cephalothorax; GJ, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="16" pageNumber="444">Fig. 5A, B</figureCitation>
); and the dorsal and outer surfaces of the carpus of the cheliped are covered with five weakly defined areolets which are deeply eroded (
<figureCitation id="13022A5E7E1573384615FD13F6FFFD72" box="[324,407,654,680]" captionStart="FIGURE 6" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,813,1703]" captionTargetId="figure-327@11.[151,1436,813,1703]" captionTargetPageId="11" captionText="FIGURE 6. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu.A, B, D, E, dorsal view of right cheliped; C, dorsal view of left cheliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663784" httpUri="https://zenodo.org/record/7663784/files/figure.png" pageId="16" pageNumber="444">Fig. 6F</figureCitation>
) (versus with four distinct smooth areolets;
<figureCitation id="13022A5E7E15733844D7FD13F496FD72" box="[902,1022,654,680]" captionStart="FIGURE 6" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,813,1703]" captionTargetId="figure-327@11.[151,1436,813,1703]" captionTargetPageId="11" captionText="FIGURE 6. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu.A, B, D, E, dorsal view of right cheliped; C, dorsal view of left cheliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663784" httpUri="https://zenodo.org/record/7663784/files/figure.png" pageId="16" pageNumber="444">Fig. 6A, B</figureCitation>
). No male specimens are available.
</paragraph>
<paragraph id="8B8636DB7E1573384796FD2FF64CFD2A" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="444">
<emphasis id="B94DEAC97E1573384796FD2FF64BFD16" bold="true" box="[199,291,690,716]" pageId="16" pageNumber="444">Habitat</emphasis>
. The
<typeStatus id="548288797E1573384633FD2EF6FAFD17" box="[354,402,691,717]" pageId="16" pageNumber="444">type</typeStatus>
was collected from shallow waters by diving (
<quantity id="4CC19B3E7E15733844C6FD2FF4B5FD17" box="[919,989,690,717]" metricMagnitude="0" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="16" pageNumber="444" unit="m" value="2.0" valueMax="3.0" valueMin="1.0">13 m</quantity>
depth) in a reef area with some seagrass beds nearby.
</paragraph>
</subSubSection>
<subSubSection id="C32365507E1573384796FD67F59CFCCE" box="[199,756,762,789]" pageId="16" pageNumber="444" type="distribution">
<paragraph id="8B8636DB7E1573384796FD67F59CFCCE" blockId="16.[151,1437,150,789]" box="[199,756,762,789]" pageId="16" pageNumber="444">
<emphasis id="B94DEAC97E1573384796FD67F631FCCE" bold="true" box="[199,345,762,788]" pageId="16" pageNumber="444">Distribution</emphasis>
. Known only from
<collectingCountry id="F32E764B7E1573384566FD66F5FBFCCF" box="[567,659,763,789]" name="Vanuatu" pageId="16" pageNumber="444">Vanuatu</collectingCountry>
thus far.
</paragraph>
</subSubSection>
</treatment>
</document>