359 lines
35 KiB
XML
359 lines
35 KiB
XML
<document ID-DOI="10.1649/0010-066X-66.mo4.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="4907222" approvalRequired="4" approvalRequired_for_taxonomicNames="4" checkinTime="1623089673881" checkinUser="carolina" docAuthor="Orozco, Jesús" docDate="2012" docId="F449F723D56FB22485C344BCEAE0FB79" docLanguage="en" docName="TheColeopBull.66.mo11.1-182.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 11) 66" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Euphoria humilis Blanchard 1850" docType="treatment" docVersion="6" lastPageNumber="79" masterDocId="08708F5BD522B26A85194065E874FF87" masterDocTitle="Monographic Revision of the American Genus Euphoria Burmeister, 1842 (Coleoptera: Scarabaeidae: Cetoniinae)" masterLastPageNumber="182" masterPageNumber="1" pageNumber="78" updateTime="1663356291692" updateUser="tatiana" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Monographic Revision of the American Genus Euphoria Burmeister, 1842 (Coleoptera: Scarabaeidae: Cetoniinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Orozco, Jesús</mods:namePart>
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<mods:affiliation>Molecular Ecology and Systematics Group Department of Botany and Department of Zoology and Entomology Rhodes University Grahamstown, SOUTH AFRICA 6140</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="series">
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<mods:title>mo 11</mods:title>
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</mods:detail>
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<mods:detail type="pubDate">
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<mods:number>2012-12-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>66</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>182</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1649/0010-066X-66.mo4.1</mods:identifier>
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<mods:identifier type="ISSN">1938-4394</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4907222</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:F449F723D56FB22485C344BCEAE0FB79" httpUri="http://treatment.plazi.org/id/F449F723D56FB22485C344BCEAE0FB79" lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78">
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<subSubSection box="[218,571,1241,1262]" pageId="77" pageNumber="78" type="nomenclature">
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||
<paragraph blockId="77.[218,571,1241,1288]" box="[218,571,1241,1262]" pageId="77" pageNumber="78">
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<heading box="[218,571,1241,1262]" centered="true" fontSize="10" level="2" pageId="77" pageNumber="78" reason="2">
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<taxonomicName authority="Blanchard, 1850" authorityName="Blanchard" authorityYear="1850" box="[218,571,1241,1262]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="humilis">
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<emphasis bold="true" box="[218,571,1241,1262]" pageId="77" pageNumber="78">
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<emphasis bold="true" box="[218,392,1241,1262]" italics="true" pageId="77" pageNumber="78">Euphoria humilis</emphasis>
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<bibRefCitation author="Blanchard, C. E." box="[401,571,1241,1262]" editor="H. Milne-Edwards & C. E. Blanchard & H. Lucas" journalOrPublisher="Gide and Baudry, Paris, France" pageId="77" pageNumber="78" refId="ref121562" refString="Blanchard, C. E. 1850. Classe des Insectes. Order des Coleopteres. Volume I. In: Museum d' Histoire Naturelle de Paris. Catalogue de la Collection Entomologique (H. Milne-Edwards, C. E. Blanchard and H. Lucas, editors). Gide and Baudry, Paris, France." title="Classe des Insectes. Order des Coleopteres. Volume I." type="book chapter" volumeTitle="Museum d' Histoire Naturelle de Paris. Catalogue de la Collection Entomologique" year="1850">Blanchard, 1850</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[287,502,1267,1288]" pageId="77" pageNumber="78" type="description">
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<paragraph blockId="77.[218,571,1241,1288]" box="[287,502,1267,1288]" pageId="77" pageNumber="78">
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(Appendix 4:
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<figureCitation box="[425,495,1267,1288]" captionStart="Fig" captionStartId="162.[197,229,1686,1705]" captionTargetBox="[284,1148,183,1662]" captionTargetId="figure-0@162.[174,1192,183,1662]" captionTargetPageId="162" captionText="Fig. 43. Euphoria humilis. a) Habitus of male, b) Head, c) Parameres, d) Female, e) Temporal distribution, f) Geographic distribution in Mexico." figureDoi="http://doi.org/10.5281/zenodo.4907340" httpUri="https://zenodo.org/record/4907340/files/figure.png" pageId="77" pageNumber="78">Fig. 43</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="77" pageNumber="78" type="reference_group">
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<paragraph blockId="77.[151,638,1310,1570]" pageId="77" pageNumber="78">
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<treatmentCitationGroup pageId="77" pageNumber="78">
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<treatmentCitation author="Blanchard, C. E." box="[151,542,1310,1331]" page="13" pageId="77" pageNumber="78" year="1850">
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<taxonomicName authority="Blanchard 1850: 13" authorityName="Blanchard" authorityPageNumber="13" authorityYear="1850" box="[151,542,1310,1331]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="humilis">
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<emphasis box="[151,328,1310,1331]" italics="true" pageId="77" pageNumber="78">Euphoria humilis</emphasis>
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<bibRefCitation author="Blanchard, C. E." box="[337,542,1310,1331]" editor="H. Milne-Edwards & C. E. Blanchard & H. Lucas" journalOrPublisher="Gide and Baudry, Paris, France" pageId="77" pageNumber="78" refId="ref121562" refString="Blanchard, C. E. 1850. Classe des Insectes. Order des Coleopteres. Volume I. In: Museum d' Histoire Naturelle de Paris. Catalogue de la Collection Entomologique (H. Milne-Edwards, C. E. Blanchard and H. Lucas, editors). Gide and Baudry, Paris, France." title="Classe des Insectes. Order des Coleopteres. Volume I." type="book chapter" volumeTitle="Museum d' Histoire Naturelle de Paris. Catalogue de la Collection Entomologique" year="1850">Blanchard 1850: 13</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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. Original combination.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection box="[310,617,1337,1358]" pageId="77" pageNumber="78" type="materials_examined">
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<paragraph blockId="77.[151,638,1310,1570]" box="[310,617,1337,1358]" pageId="77" pageNumber="78">
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<typeStatus box="[310,398,1337,1358]" pageId="77" pageNumber="78">Holotype</typeStatus>
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at MNHN, examined.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="77" pageNumber="78" type="reference_group">
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<paragraph blockId="77.[151,638,1310,1570]" pageId="77" pageNumber="78">
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<treatmentCitationGroup pageId="77" pageNumber="78">
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<treatmentCitation author="Bates, W. D." box="[151,530,1363,1384]" page="371" pageId="77" pageNumber="78" year="1889">
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<taxonomicName authority="Bates 1889: 371" authorityName="Bates" authorityPageNumber="371" authorityYear="1889" box="[151,530,1363,1384]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="nigriventris" status="New synonymy">
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<emphasis box="[151,361,1363,1384]" italics="true" pageId="77" pageNumber="78">Euphoria nigriventris</emphasis>
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<bibRefCitation author="Bates, W. D." box="[370,530,1363,1384]" editor="F. D. Godman & O. Salvin" journalOrPublisher="London, UK" pageId="77" pageNumber="78" pagination="362 - 374" refId="ref121351" refString="Bates, W. D. 1889. Insecta. Coleoptera. Pectinicornia and Lamellicornia, [pp. 362 - 374]. In: Biologia Centrali-Americana. Vol II, Part 2 (F. D. Godman and O. Salvin, editors). London, UK." title="Insecta. Coleoptera. Pectinicornia and Lamellicornia" type="book chapter" volumeTitle="Biologia Centrali-Americana. Vol II, Part 2" year="1889">Bates 1889: 371</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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<emphasis bold="true" pageId="77" pageNumber="78">
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<taxonomicNameLabel pageId="77" pageNumber="78">New synonymy</taxonomicNameLabel>
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.
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</emphasis>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="77" pageNumber="78" type="materials_examined">
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<paragraph blockId="77.[151,638,1310,1570]" pageId="77" pageNumber="78">
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<typeStatus box="[266,368,1390,1411]" pageId="77" pageNumber="78">Lectotype</typeStatus>
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at BMNH designated by
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<bibRefCitation author="Hardy" box="[177,313,1416,1437]" firstAuthor="Hardy" pageId="77" pageNumber="78" pagination="127 - 143" refId="ref122900" refString="Hardy, A. R. 2001. Studies in the Euphoriina of the Americas (Coleoptera: Scarabaeidae) II. Status of names in Euphoria, types and synonymies, with notes on the South American species. Pan- Pacific Entomologist 77: 127 - 143." type="journal article" year="2001">Hardy (2001)</bibRefCitation>
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, not examined. Three
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<typeStatus pageId="77" pageNumber="78">paralectotypes</typeStatus>
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at BMNH, one examined.
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,637,1469,1491]" pageId="77" pageNumber="78" type="reference_group">
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||
<paragraph blockId="77.[151,638,1310,1570]" box="[151,637,1469,1491]" pageId="77" pageNumber="78">
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<treatmentCitationGroup box="[151,637,1469,1491]" pageId="77" pageNumber="78">
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<treatmentCitation author="Bates, W. D." box="[151,464,1469,1491]" page="371" pageId="77" pageNumber="78" year="1889">
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<taxonomicName authority="Bates 1889: 371" authorityName="Bates" authorityPageNumber="371" authorityYear="1889" box="[151,464,1469,1491]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="atra" status="New synonymy">
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<emphasis box="[151,292,1470,1491]" italics="true" pageId="77" pageNumber="78">Euphoria atra</emphasis>
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<bibRefCitation author="Bates, W. D." box="[301,464,1469,1490]" editor="F. D. Godman & O. Salvin" journalOrPublisher="London, UK" pageId="77" pageNumber="78" pagination="362 - 374" refId="ref121351" refString="Bates, W. D. 1889. Insecta. Coleoptera. Pectinicornia and Lamellicornia, [pp. 362 - 374]. In: Biologia Centrali-Americana. Vol II, Part 2 (F. D. Godman and O. Salvin, editors). London, UK." title="Insecta. Coleoptera. Pectinicornia and Lamellicornia" type="book chapter" volumeTitle="Biologia Centrali-Americana. Vol II, Part 2" year="1889">Bates 1889: 371</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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||
<emphasis bold="true" box="[474,637,1469,1490]" pageId="77" pageNumber="78">
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<taxonomicNameLabel box="[474,633,1469,1490]" pageId="77" pageNumber="78">New synonymy</taxonomicNameLabel>
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.
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</emphasis>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="77" pageNumber="78" type="materials_examined">
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<paragraph blockId="77.[151,638,1310,1570]" pageId="77" pageNumber="78">
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<typeStatus box="[177,283,1496,1517]" pageId="77" pageNumber="78">Lectotype</typeStatus>
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at BMNH designated by
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<bibRefCitation author="Hardy" firstAuthor="Hardy" pageId="77" pageNumber="78" pagination="127 - 143" refId="ref122900" refString="Hardy, A. R. 2001. Studies in the Euphoriina of the Americas (Coleoptera: Scarabaeidae) II. Status of names in Euphoria, types and synonymies, with notes on the South American species. Pan- Pacific Entomologist 77: 127 - 143." type="journal article" year="2001">Hardy (2001)</bibRefCitation>
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, examined. One
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<typeStatus box="[403,530,1523,1544]" pageId="77" pageNumber="78">paralectotype</typeStatus>
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at BMNH, not examined.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="77" pageNumber="78" type="description">
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<paragraph blockId="77.[151,638,1602,1729]" lastBlockId="77.[685,1173,194,1730]" pageId="77" pageNumber="78">
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<emphasis bold="true" box="[175,395,1602,1623]" pageId="77" pageNumber="78">
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Description (
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<emphasis bold="true" box="[310,323,1602,1623]" italics="true" pageId="77" pageNumber="78">n</emphasis>
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= 76).
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</emphasis>
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Length
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<quantity box="[486,634,1602,1624]" metricMagnitude="-2" metricUnit="m" metricValue="1.215" metricValueMax="1.38" metricValueMin="1.05" pageId="77" pageNumber="78" unit="mm" value="12.15" valueMax="13.8" valueMin="10.5">10.5–13.8 mm</quantity>
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; width
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<quantity box="[219,346,1628,1650]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="8.1" metricValueMin="6.2" pageId="77" pageNumber="78" unit="mm" value="7.15" valueMax="8.1" valueMin="6.2">6.2–8.1 mm</quantity>
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.
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<emphasis bold="true" box="[360,421,1629,1650]" pageId="77" pageNumber="78">Color</emphasis>
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: Dorsal surface yellowish brown, usually tomentous in males, shiny in females. Pronotum vittate in males, entirely black in females; males with 2 longitudinal, black vittae and 1 small, dark, semicircular, anterolateral mark at each side. Elytra yellowish brown with black markings; markings frequently covering elytra entirely in females. Ventral surface, including legs, light brown to black. Melanistic forms observed.
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<emphasis bold="true" box="[781,834,327,348]" pageId="77" pageNumber="78">Head</emphasis>
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: Frons frequently moderately raised at middle, densely punctate; punctures round, deep, moderate in size, occasionally confluent, moderately densely to densely setose; setae, yellowish, moderate to long. Clypeus subrectangular, surface as on frons, sides subparallel, weakly to moderately raised, apex vaguely to weakly reflexed, moderately sinuate. Antennal club slightly shorter than stem in males, much shorter in females.
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<emphasis bold="true" box="[988,1088,540,561]" pageId="77" pageNumber="78">Pronotum</emphasis>
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: Surface moderately densely punctate, anterolateral margins with sparse rugae; punctures round to lunulate, small to moderate, sparsely to moderately densely setose; setae short to moderate, yellowish. Sides weakly to moderately angulate. Base in front of scutellum weakly to moderately emarginate. Scutellum impunctate to sparsely punctate; punctures minute, bearing setae.
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<emphasis bold="true" box="[906,970,752,773]" pageId="77" pageNumber="78">Elytra</emphasis>
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: Surface moderately densely to densely punctate; striae bearing 2, densely punctate, lateral rows and 1 sparsely punctate medial row; punctures small to moderate, lunulate, geminate, and double grooves in lateral rows, only lunulate in medial row; apex and posterior end of sides rugose.
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<emphasis bold="true" box="[759,854,911,932]" pageId="77" pageNumber="78">Pygidium</emphasis>
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: Surface subconcentrically striate; striae dense, discontinuous, moderately to densely setose, setae short to moderate.
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<emphasis bold="true" box="[1018,1067,964,985]" pageId="77" pageNumber="78">Legs</emphasis>
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: Protibiae slender in males, robust in females, teeth equidistant or apical and medial teeth closer to each other than to basal tooth; teeth frequently strongly oblique, basal tooth frequently subobsolete. Meso- and metatibial carinae moderately developed in males, strongly developed in females. Metatibiae moderately to strongly expanded in females at apex, metatarsi longer than metatibiae in males, shorter in females.
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<emphasis bold="true" box="[879,944,1204,1225]" pageId="77" pageNumber="78">Venter</emphasis>
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: Mesometasternal process moderately compressed laterally, extending anteriorly to same level as mesocoxae or slightly beyond. Mesepimera, metasternum, and metacoxae setose, setae as on legs. Metasternum rugose, moderately to densely setose laterally, weakly to moderately punctate and setose at middle, median sulcus strongly impressed. Abdomen in lateral view flat in males, strongly convex in females.
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<emphasis bold="true" pageId="77" pageNumber="78">Male genitalia</emphasis>
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: Parameres as in
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<figureCitation box="[946,1028,1443,1464]" captionStart="Fig" captionStartId="162.[197,229,1686,1705]" captionTargetBox="[284,1148,183,1662]" captionTargetId="figure-0@162.[174,1192,183,1662]" captionTargetPageId="162" captionText="Fig. 43. Euphoria humilis. a) Habitus of male, b) Head, c) Parameres, d) Female, e) Temporal distribution, f) Geographic distribution in Mexico." figureDoi="http://doi.org/10.5281/zenodo.4907340" httpUri="https://zenodo.org/record/4907340/files/figure.png" pageId="77" pageNumber="78">Fig. 43c</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="77" pageNumber="78" type="diagnosis">
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<paragraph blockId="77.[685,1173,194,1730]" pageId="77" pageNumber="78">
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<emphasis bold="true" box="[709,814,1469,1490]" pageId="77" pageNumber="78">Diagnosis.</emphasis>
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This species is separated from other species in the group by the following combination of characters: subrectangular clypeus; clypeal sides weakly to moderately raised, clypeal apex vaguely to weakly reflexed; antennal club shorter than stem in both sexes (slightly shorter than stem in males, much shorter in females); double grooved elytral striae; oblique protibial teeth; and form of the parameres.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78" type="discussion">
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<paragraph blockId="77.[685,1173,194,1730]" lastBlockId="78.[173,660,194,1278]" lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78">
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<emphasis bold="true" box="[709,907,1682,1703]" pageId="77" pageNumber="78">Taxonomic History.</emphasis>
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<bibRefCitation author="Blanchard, C. E." box="[921,1080,1682,1703]" editor="H. Milne-Edwards & C. E. Blanchard & H. Lucas" journalOrPublisher="Gide and Baudry, Paris, France" pageId="77" pageNumber="78" refId="ref121562" refString="Blanchard, C. E. 1850. Classe des Insectes. Order des Coleopteres. Volume I. In: Museum d' Histoire Naturelle de Paris. Catalogue de la Collection Entomologique (H. Milne-Edwards, C. E. Blanchard and H. Lucas, editors). Gide and Baudry, Paris, France." title="Classe des Insectes. Order des Coleopteres. Volume I." type="book chapter" volumeTitle="Museum d' Histoire Naturelle de Paris. Catalogue de la Collection Entomologique" year="1850">Blanchard (1850)</bibRefCitation>
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described the species from a female specimen from “
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<collectingCountry box="[1086,1156,1708,1729]" name="Mexico" pageId="77" pageNumber="78">Mexico</collectingCountry>
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”.
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<bibRefCitation author="Bates, W. D." box="[173,301,194,215]" editor="F. D. Godman & O. Salvin" journalOrPublisher="London, UK" pageId="78" pageNumber="79" pagination="362 - 374" refId="ref121351" refString="Bates, W. D. 1889. Insecta. Coleoptera. Pectinicornia and Lamellicornia, [pp. 362 - 374]. In: Biologia Centrali-Americana. Vol II, Part 2 (F. D. Godman and O. Salvin, editors). London, UK." title="Insecta. Coleoptera. Pectinicornia and Lamellicornia" type="book chapter" volumeTitle="Biologia Centrali-Americana. Vol II, Part 2" year="1889">Bates (1889)</bibRefCitation>
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described
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<taxonomicName authorityName="Bates" authorityYear="1889" box="[415,559,194,215]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="nigriventris">
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<emphasis box="[415,429,194,215]" italics="true" pageId="78" pageNumber="79">E</emphasis>
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.
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<emphasis box="[445,559,194,215]" italics="true" pageId="78" pageNumber="79">nigriventris</emphasis>
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</taxonomicName>
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based on
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<specimenCount box="[173,295,221,242]" pageId="78" pageNumber="79" type="male">eight males</specimenCount>
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from Jalapa, Etla, and Peras, all
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<collectingCountry box="[173,250,247,268]" name="Mexico" pageId="78" pageNumber="79">Mexico</collectingCountry>
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, and
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<taxonomicName box="[308,378,247,268]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="atra">
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<emphasis box="[308,322,247,268]" italics="true" pageId="78" pageNumber="79">E</emphasis>
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.
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<emphasis box="[338,378,247,268]" italics="true" pageId="78" pageNumber="79">atra</emphasis>
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</taxonomicName>
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based on
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<specimenCount box="[488,602,247,268]" pageId="78" pageNumber="79" type="male">three males</specimenCount>
|
||
from Peras and
|
||
<collectingRegion box="[274,349,274,295]" country="Mexico" name="Oaxaca" pageId="78" pageNumber="79">Oaxaca</collectingRegion>
|
||
,
|
||
<collectingCountry box="[360,437,274,295]" name="Mexico" pageId="78" pageNumber="79">Mexico</collectingCountry>
|
||
.
|
||
<taxonomicName authorityName="Bates" authorityYear="1889" box="[447,659,274,295]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="nigriventris">
|
||
<emphasis box="[447,659,274,295]" italics="true" pageId="78" pageNumber="79">Euphoria nigriventris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[218,291,300,322]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="atra">
|
||
<emphasis box="[218,232,301,322]" italics="true" pageId="78" pageNumber="79">E</emphasis>
|
||
.
|
||
<emphasis box="[250,291,301,322]" italics="true" pageId="78" pageNumber="79">atra</emphasis>
|
||
</taxonomicName>
|
||
constitute intraspecific variation of
|
||
<taxonomicName authorityName="Blanchard" authorityYear="1850" box="[173,272,327,348]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="humilis">
|
||
<emphasis box="[173,187,327,348]" italics="true" pageId="78" pageNumber="79">E</emphasis>
|
||
.
|
||
<emphasis box="[201,272,327,348]" italics="true" pageId="78" pageNumber="79">humilis</emphasis>
|
||
</taxonomicName>
|
||
. Bates’ (1889) observations on the differences between these two species were all based on sexually dimorphic characters and color variation. Based on the examination of
|
||
<specimenCount box="[521,656,407,428]" pageId="78" pageNumber="79" type="generic">76 specimens</specimenCount>
|
||
, including the
|
||
<typeStatus box="[317,371,433,454]" pageId="78" pageNumber="79">types</typeStatus>
|
||
, I place
|
||
<taxonomicName authorityName="Bates" authorityYear="1889" box="[466,613,433,454]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="nigriventris">
|
||
<emphasis box="[466,480,433,454]" italics="true" pageId="78" pageNumber="79">E</emphasis>
|
||
.
|
||
<emphasis box="[498,613,433,454]" italics="true" pageId="78" pageNumber="79">nigriventris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[173,240,460,481]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="atra">
|
||
<emphasis box="[173,187,460,481]" italics="true" pageId="78" pageNumber="79">E</emphasis>
|
||
.
|
||
<emphasis box="[201,240,460,481]" italics="true" pageId="78" pageNumber="79">atra</emphasis>
|
||
</taxonomicName>
|
||
in synonymy with
|
||
<taxonomicName authorityName="Blanchard" authorityYear="1850" box="[432,529,460,481]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="humilis">
|
||
<emphasis box="[432,446,460,481]" italics="true" pageId="78" pageNumber="79">E</emphasis>
|
||
.
|
||
<emphasis box="[461,529,460,481]" italics="true" pageId="78" pageNumber="79">humilis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="78.[173,660,194,1278]" pageId="78" pageNumber="79">
|
||
<emphasis bold="true" box="[196,365,486,507]" pageId="78" pageNumber="79">Natural History.</emphasis>
|
||
Adults have been captured on
|
||
<taxonomicName box="[207,314,513,534]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="78" pageNumber="79" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis box="[207,275,513,534]" italics="true" pageId="78" pageNumber="79">Acacia</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
and are known to occur between
|
||
<quantity box="[173,318,539,561]" metricMagnitude="3" metricUnit="m" metricValue="2.0835" metricValueMax="2.4" metricValueMin="1.767" pageId="78" pageNumber="79" unit="m" value="2083.5" valueMax="2400.0" valueMin="1767.0">1,767–2,400 m</quantity>
|
||
elevation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="78" pageNumber="79" type="distribution">
|
||
<paragraph blockId="78.[173,660,194,1278]" pageId="78" pageNumber="79">
|
||
<emphasis bold="true" box="[196,431,566,587]" pageId="78" pageNumber="79">Temporal Distribution.</emphasis>
|
||
June (23), July (30), August (1) (
|
||
<figureCitation box="[292,375,593,614]" captionStart="Fig" captionStartId="162.[197,229,1686,1705]" captionTargetBox="[284,1148,183,1662]" captionTargetId="figure-0@162.[174,1192,183,1662]" captionTargetPageId="162" captionText="Fig. 43. Euphoria humilis. a) Habitus of male, b) Head, c) Parameres, d) Female, e) Temporal distribution, f) Geographic distribution in Mexico." figureDoi="http://doi.org/10.5281/zenodo.4907340" httpUri="https://zenodo.org/record/4907340/files/figure.png" pageId="78" pageNumber="79">Fig. 43e</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="78.[173,660,194,1278]" pageId="78" pageNumber="79">
|
||
<emphasis bold="true" box="[196,441,619,640]" pageId="78" pageNumber="79">Geographic Distribution.</emphasis>
|
||
Known from
|
||
<collectingRegion box="[578,655,619,640]" country="Mexico" name="Chiapas" pageId="78" pageNumber="79">Chiapas</collectingRegion>
|
||
,
|
||
<collectingRegion box="[173,245,646,667]" country="Mexico" name="Oaxaca" pageId="78" pageNumber="79">Oaxaca</collectingRegion>
|
||
, and
|
||
<collectingRegion box="[296,362,646,667]" country="Mexico" name="Puebla" pageId="78" pageNumber="79">Puebla</collectingRegion>
|
||
,
|
||
<collectingCountry box="[372,444,646,667]" name="Mexico" pageId="78" pageNumber="79">Mexico</collectingCountry>
|
||
(
|
||
<figureCitation box="[457,533,646,667]" captionStart="Fig" captionStartId="162.[197,229,1686,1705]" captionTargetBox="[284,1148,183,1662]" captionTargetId="figure-0@162.[174,1192,183,1662]" captionTargetPageId="162" captionText="Fig. 43. Euphoria humilis. a) Habitus of male, b) Head, c) Parameres, d) Female, e) Temporal distribution, f) Geographic distribution in Mexico." figureDoi="http://doi.org/10.5281/zenodo.4907340" httpUri="https://zenodo.org/record/4907340/files/figure.png" pageId="78" pageNumber="79">Fig. 43f</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="78" pageNumber="79" type="materials_examined">
|
||
<paragraph blockId="78.[173,660,194,1278]" pageId="78" pageNumber="79">
|
||
<emphasis bold="true" box="[196,473,672,693]" pageId="78" pageNumber="79">Specimens Examined (76).</emphasis>
|
||
Type material:
|
||
<materialsCitation collectionCode="MNHN" country="Mexico" location="Mexique" pageId="78" pageNumber="79" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||
<taxonomicName authority="Blanchard, 1850" authorityName="Blanchard" authorityYear="1850" box="[173,499,699,720]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="humilis">
|
||
<originalDetermination box="[173,499,699,720]" pageId="78" pageNumber="79">
|
||
<emphasis box="[173,335,699,720]" italics="true" pageId="78" pageNumber="79">Euphoria humilis</emphasis>
|
||
<bibRefCitation author="Blanchard, C. E." box="[342,499,699,720]" editor="H. Milne-Edwards & C. E. Blanchard & H. Lucas" journalOrPublisher="Gide and Baudry, Paris, France" pageId="78" pageNumber="79" refId="ref121562" refString="Blanchard, C. E. 1850. Classe des Insectes. Order des Coleopteres. Volume I. In: Museum d' Histoire Naturelle de Paris. Catalogue de la Collection Entomologique (H. Milne-Edwards, C. E. Blanchard and H. Lucas, editors). Gide and Baudry, Paris, France." title="Classe des Insectes. Order des Coleopteres. Volume I." type="book chapter" volumeTitle="Museum d' Histoire Naturelle de Paris. Catalogue de la Collection Entomologique" year="1850">Blanchard, 1850</bibRefCitation>
|
||
</originalDetermination>
|
||
</taxonomicName>
|
||
;
|
||
<typeStatus box="[508,590,699,720]" pageId="78" pageNumber="79">holotype</typeStatus>
|
||
<specimenCount box="[596,659,699,720]" pageId="78" pageNumber="79" type="female">female</specimenCount>
|
||
at
|
||
<collectionCode box="[200,274,725,746]" pageId="78" pageNumber="79">MNHN</collectionCode>
|
||
labeled “18/ 42//
|
||
<taxonomicName box="[469,560,725,746]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">Euphoria</taxonomicName>
|
||
/ humilis/ Blanch// MUSEUM PARIS / MEXIQUE/ GIESBREGHT 1842// TYPE//
|
||
<taxonomicName authorityName="Blanchard" authorityYear="1850" box="[525,646,779,800]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="humilis">C. humilis</taxonomicName>
|
||
,/ Blanch./
|
||
<collectingCountry box="[268,357,805,826]" pageId="78" pageNumber="79">Mexique</collectingCountry>
|
||
/ M. M. Giesbreght et Sallé” and my holotype label.
|
||
</materialsCitation>
|
||
<materialsCitation pageId="78" pageNumber="79">
|
||
<taxonomicName authority="Bates, 1889" authorityName="Bates" authorityYear="1889" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="nigriventris">
|
||
<emphasis box="[393,596,832,853]" italics="true" pageId="78" pageNumber="79">Euphoria nigriventris</emphasis>
|
||
<bibRefCitation author="Bates, W. D." editor="F. D. Godman & O. Salvin" journalOrPublisher="London, UK" pageId="78" pageNumber="79" pagination="362 - 374" refId="ref121351" refString="Bates, W. D. 1889. Insecta. Coleoptera. Pectinicornia and Lamellicornia, [pp. 362 - 374]. In: Biologia Centrali-Americana. Vol II, Part 2 (F. D. Godman and O. Salvin, editors). London, UK." title="Insecta. Coleoptera. Pectinicornia and Lamellicornia" type="book chapter" volumeTitle="Biologia Centrali-Americana. Vol II, Part 2" year="1889">Bates, 1889</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<typeStatus box="[230,355,858,879]" pageId="78" pageNumber="79">paralectotype</typeStatus>
|
||
male at BMNH labeled “PARA-/ LECTO-/TYPE// SYN-/ TYPE//Etla//
|
||
<collectingRegion box="[569,644,885,906]" country="Mexico" name="Mexico" pageId="78" pageNumber="79">Mexico</collectingRegion>
|
||
// Salle Coll//
|
||
<taxonomicName box="[291,376,911,932]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">Euphoria</taxonomicName>
|
||
/ nigriventris/ Bates.// B.C.A. Col., II(2)./
|
||
<taxonomicName box="[314,410,938,959]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">Euphoria</taxonomicName>
|
||
//
|
||
<typeStatus box="[439,651,938,959]" pageId="78" pageNumber="79">PARALECTOTYPE</typeStatus>
|
||
/
|
||
<taxonomicName box="[173,265,964,985]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">Euphoria</taxonomicName>
|
||
/ nigriventris/ Bates”.
|
||
</materialsCitation>
|
||
<materialsCitation pageId="78" pageNumber="79">
|
||
<taxonomicName authority="Bates, 1889" authorityName="Bates" authorityYear="1889" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="atra">
|
||
<emphasis box="[509,659,965,986]" italics="true" pageId="78" pageNumber="79">Euphoria atra</emphasis>
|
||
<bibRefCitation author="Bates, W. D." box="[173,297,991,1012]" editor="F. D. Godman & O. Salvin" journalOrPublisher="London, UK" pageId="78" pageNumber="79" pagination="362 - 374" refId="ref121351" refString="Bates, W. D. 1889. Insecta. Coleoptera. Pectinicornia and Lamellicornia, [pp. 362 - 374]. In: Biologia Centrali-Americana. Vol II, Part 2 (F. D. Godman and O. Salvin, editors). London, UK." title="Insecta. Coleoptera. Pectinicornia and Lamellicornia" type="book chapter" volumeTitle="Biologia Centrali-Americana. Vol II, Part 2" year="1889">Bates, 1889</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<typeStatus box="[313,403,991,1012]" pageId="78" pageNumber="79">lectotype</typeStatus>
|
||
male at BMNH labeled “LECTO-/TYPE/ SYN-/TYPE//
|
||
<collectingRegion box="[485,557,1018,1039]" country="Mexico" name="Oaxaca" pageId="78" pageNumber="79">Oaxaca</collectingRegion>
|
||
,/
|
||
<collectingRegion box="[575,649,1018,1039]" country="Mexico" name="Mexico" pageId="78" pageNumber="79">Mexico</collectingRegion>
|
||
./ Hoege// B.C.A. Col., II(2)./
|
||
<taxonomicName box="[445,530,1044,1065]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">Euphoria</taxonomicName>
|
||
/
|
||
<taxonomicName authority=", Bates" authorityName="Bates" box="[546,645,1044,1065]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="atra">atra, Bates</taxonomicName>
|
||
//
|
||
<typeStatus box="[173,311,1071,1092]" pageId="78" pageNumber="79">LECTOTYPE</typeStatus>
|
||
/
|
||
<taxonomicName box="[327,416,1071,1092]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">Euphoria</taxonomicName>
|
||
/ atra/ Bates = By/ A.R. Hardy, 78”.
|
||
</materialsCitation>
|
||
Other material:
|
||
<emphasis bold="true" box="[488,596,1097,1118]" pageId="78" pageNumber="79">
|
||
<collectingCountry box="[488,596,1097,1118]" name="Mexico" pageId="78" pageNumber="79">MEXICO</collectingCountry>
|
||
</emphasis>
|
||
(70):
|
||
<collectingRegion box="[173,273,1124,1145]" country="Mexico" name="Chiapas" pageId="78" pageNumber="79">CHIAPAS</collectingRegion>
|
||
: Arriaga (5), Navenchauc (3), Rincón las minas (1);
|
||
<collectingRegion box="[278,382,1151,1172]" country="Mexico" name="Oaxaca" pageId="78" pageNumber="79">OAXACA</collectingRegion>
|
||
: Díaz Ordaz (2), La Cumbre (4), Matatlán (1), Mitla (9), Monte Albán (16), Nejapa de Madero (1),
|
||
<collectingRegion box="[344,417,1204,1225]" country="Mexico" name="Oaxaca" pageId="78" pageNumber="79">Oaxaca</collectingRegion>
|
||
(12), Tapanatepec (1), Teotitlán (4), Tlacolula (6),
|
||
<taxonomicName box="[441,512,1230,1251]" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="78" pageNumber="103" phylum="Chordata" rank="species" species="data">no data</taxonomicName>
|
||
(1);
|
||
<collectingRegion box="[562,657,1230,1251]" country="Mexico" name="Puebla" pageId="78" pageNumber="79">PUEBLA</collectingRegion>
|
||
: Acatepec (3), Tlacotepec (1);
|
||
<taxonomicName box="[430,530,1257,1278]" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="78" pageNumber="103" phylum="Chordata" rank="species" species="data">NO DATA</taxonomicName>
|
||
: “
|
||
<collectingRegion box="[546,613,1257,1278]" country="Mexico" name="Mexico" pageId="78" pageNumber="79">Mexico</collectingRegion>
|
||
” (3).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |