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<document ID-DOI="10.1649/0010-066X-66.mo4.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="4907222" approvalRequired="4" approvalRequired_for_taxonomicNames="4" checkinTime="1623089673881" checkinUser="carolina" docAuthor="Orozco, Jesús" docDate="2012" docId="F449F723D560B22985EC4496E9FCFEF0" docLanguage="en" docName="TheColeopBull.66.mo11.1-182.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 11) 66" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Euphoria mystica Thomson 1878" docType="treatment" docVersion="7" lastPageNumber="67" masterDocId="08708F5BD522B26A85194065E874FF87" masterDocTitle="Monographic Revision of the American Genus Euphoria Burmeister, 1842 (Coleoptera: Scarabaeidae: Cetoniinae)" masterLastPageNumber="182" masterPageNumber="1" pageNumber="67" updateTime="1663356291692" updateUser="tatiana" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Monographic Revision of the American Genus Euphoria Burmeister, 1842 (Coleoptera: Scarabaeidae: Cetoniinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Orozco, Jesús</mods:namePart>
<mods:affiliation>Molecular Ecology and Systematics Group Department of Botany and Department of Zoology and Entomology Rhodes University Grahamstown, SOUTH AFRICA 6140</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="series">
<mods:title>mo 11</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2012-12-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>66</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>182</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1649/0010-066X-66.mo4.1</mods:identifier>
<mods:identifier type="ISSN">1938-4394</mods:identifier>
<mods:identifier type="Zenodo-Dep">4907222</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7083413" ID-Zenodo-Dep="7083413" LSID="urn:lsid:plazi:treatment:F449F723D560B22985EC4496E9FCFEF0" httpUri="http://treatment.plazi.org/id/F449F723D560B22985EC4496E9FCFEF0" lastPageId="67" lastPageNumber="67" pageId="66" pageNumber="67">
<subSubSection box="[245,587,1267,1288]" pageId="66" pageNumber="67" type="nomenclature">
<paragraph blockId="66.[245,587,1267,1315]" box="[245,587,1267,1288]" pageId="66" pageNumber="67">
<heading box="[245,587,1267,1288]" centered="true" fontSize="10" level="2" pageId="66" pageNumber="67" reason="2">
<taxonomicName authority="Thomson, 1878" authorityName="Thomson" authorityYear="1878" box="[245,587,1267,1288]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="mystica">
<emphasis bold="true" box="[245,587,1267,1288]" pageId="66" pageNumber="67">
<emphasis bold="true" box="[245,420,1267,1288]" italics="true" pageId="66" pageNumber="67">Euphoria mystica</emphasis>
<bibRefCitation author="Thomson, J." box="[428,587,1267,1288]" journalOrPublisher="E. Deyrolle, Paris, France" pageId="66" pageNumber="67" refId="ref125576" refString="Thomson, J. 1878. Typi Cetonidarum, Suivis de Typi Molommidarum et de Typi Nilionidarum Musaei Thomsoniani. E. Deyrolle, Paris, France." title="Typi Cetonidarum, Suivis de Typi Molommidarum et de Typi Nilionidarum Musaei Thomsoniani." type="book" year="1878">Thomson, 1878</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[309,523,1294,1315]" pageId="66" pageNumber="67" type="description">
<paragraph blockId="66.[245,587,1267,1315]" box="[309,523,1294,1315]" pageId="66" pageNumber="67">
(Appendix 4:
<figureCitation box="[446,516,1294,1315]" captionStart="Fig" captionStartId="152.[197,229,1686,1705]" captionTargetBox="[303,1131,187,1662]" captionTargetId="figure-0@152.[174,1192,183,1662]" captionTargetPageId="152" captionText="Fig. 33. Euphoria mystica. a) Habitus of male, b) Head, c) Parameres, d) Female, e) Temporal distribution, f) Geographic distribution in Guatemala and Mexico." figureDoi="http://doi.org/10.5281/zenodo.4907312" httpUri="https://zenodo.org/record/4907312/files/figure.png" pageId="66" pageNumber="67">Fig. 33</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="66" pageNumber="67" type="reference_group">
<paragraph blockId="66.[173,659,1337,1384]" pageId="66" pageNumber="67">
<treatmentCitationGroup pageId="66" pageNumber="67">
<treatmentCitation author="Thomson, J." box="[173,562,1337,1358]" page="27" pageId="66" pageNumber="67" year="1878">
<taxonomicName authority="Thomson 1878: 27" authorityName="Thomson" authorityPageNumber="27" authorityYear="1878" box="[173,562,1337,1358]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="mystica">
<emphasis box="[173,353,1337,1358]" italics="true" pageId="66" pageNumber="67">Euphoria mystica</emphasis>
<bibRefCitation author="Thomson, J." box="[363,562,1337,1358]" journalOrPublisher="E. Deyrolle, Paris, France" pageId="66" pageNumber="67" refId="ref125576" refString="Thomson, J. 1878. Typi Cetonidarum, Suivis de Typi Molommidarum et de Typi Nilionidarum Musaei Thomsoniani. E. Deyrolle, Paris, France." title="Typi Cetonidarum, Suivis de Typi Molommidarum et de Typi Nilionidarum Musaei Thomsoniani." type="book" year="1878">Thomson 1878: 27</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Original combination.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection box="[331,638,1363,1384]" pageId="66" pageNumber="67" type="materials_examined">
<paragraph blockId="66.[173,659,1337,1384]" box="[331,638,1363,1384]" pageId="66" pageNumber="67">
<typeStatus box="[331,419,1363,1384]" pageId="66" pageNumber="67">Holotype</typeStatus>
at MNHN, examined.
</paragraph>
</subSubSection>
<subSubSection pageId="66" pageNumber="67" type="description">
<paragraph blockId="66.[173,659,1416,1729]" lastBlockId="66.[707,1194,194,1729]" pageId="66" pageNumber="67">
<emphasis bold="true" box="[196,416,1416,1437]" pageId="66" pageNumber="67">
Description (
<emphasis bold="true" box="[331,344,1416,1437]" italics="true" pageId="66" pageNumber="67">n</emphasis>
= 21).
</emphasis>
Length
<quantity box="[507,655,1416,1438]" metricMagnitude="-2" metricUnit="m" metricValue="1.405" metricValueMax="1.52" metricValueMin="1.29" pageId="66" pageNumber="67" unit="mm" value="14.05" valueMax="15.2" valueMin="12.9">12.915.2 mm</quantity>
; width
<quantity box="[240,368,1442,1464]" metricMagnitude="-3" metricUnit="m" metricValue="8.15" metricValueMax="9.4" metricValueMin="6.9" pageId="66" pageNumber="67" unit="mm" value="8.15" valueMax="9.4" valueMin="6.9">6.99.4 mm</quantity>
.
<emphasis bold="true" box="[381,442,1443,1464]" pageId="66" pageNumber="67">Color</emphasis>
: Dorsal surface yellowish brown to dark brown, anterior half shiny, posterior half tomentous. Pronotum shiny, unicolored in males, with 4 longitudinal, irregular, dark vittae in females. Elytra tomentous, with black to dark brown markings, markings frequently covering most of surface. Ventral surface, including legs, light brown, reddish brown, or a combination of these colors.
<emphasis bold="true" box="[424,479,1655,1676]" pageId="66" pageNumber="67">Head</emphasis>
: Frons frequently with strongly raised, longitudinal ridge at middle, densely punctate; punctures round, deep, moderate in size, confluent, moderately densely to densely setose; setae, yellowish, moderate to long. Clypeus subrectangular, sides moderately raised in males, weakly raised in females; apex truncate in dorsal view, sinuate in frontal view, strongly reflexed in males, weakly reflexed in females. Antennal club longer than stem in males, shorter in females.
<emphasis bold="true" box="[707,812,380,401]" pageId="66" pageNumber="67">Pronotum</emphasis>
: Surface densely punctate; punctures small, round to lunulate, moderately densely to densely setose; setae short, yellowish. Sides at base weakly convergent anteriorly, anterior corners with few rugae. Base in front of scutellum strongly emarginate. Scutellum densely punctate; punctures round to lunulate, moderate in size, some bearing short, yellowish setae.
<emphasis bold="true" box="[943,1010,566,587]" pageId="66" pageNumber="67">Elytra</emphasis>
: Surface sparsely to moderately densely punctate, striae bearing 3 irregular rows of small, lunulate or irregular punctures, glabrous to sparsely setose; setae short to moderate, yellowish. Costae sometimes weak. Apical declivity and posterior sides weakly rugose.
<emphasis bold="true" box="[707,804,725,746]" pageId="66" pageNumber="67">Pygidium</emphasis>
: Surface subconcentrically striate; striae dense, discontinuous, moderately densely to densely setose, setae short to moderate in size.
<emphasis bold="true" box="[707,754,805,826]" pageId="66" pageNumber="67">Legs</emphasis>
: Protibiae slender in males, robust in females, apical and medial teeth closer to each other than to basal tooth in males, frequently equidistant in females, basal tooth frequently obsolete to subobsolete in both sexes. Metatibiae not apically expanded in both sexes, metatarsi shorter than metatibiae in females, unknown in males (tarsi missing, presumably as long or longer).
<emphasis bold="true" box="[1042,1111,991,1012]" pageId="66" pageNumber="67">Venter</emphasis>
: Mesometasternal process weakly compressed laterally, extending anteriorly well beyond mesocoxae, apex variably rounded. Metasternum rugose, moderately to densely setose laterally, weakly to moderately punctate and setose at middle, median sulcus weakly to strongly impressed. Abdomen in lateral view flat to weakly concave in males, flat to weakly convex in females.
<emphasis bold="true" box="[791,933,1204,1225]" pageId="66" pageNumber="67">Male genitalia</emphasis>
: Parameres as in
<figureCitation box="[1101,1182,1204,1225]" captionStart="Fig" captionStartId="152.[197,229,1686,1705]" captionTargetBox="[303,1131,187,1662]" captionTargetId="figure-0@152.[174,1192,183,1662]" captionTargetPageId="152" captionText="Fig. 33. Euphoria mystica. a) Habitus of male, b) Head, c) Parameres, d) Female, e) Temporal distribution, f) Geographic distribution in Guatemala and Mexico." figureDoi="http://doi.org/10.5281/zenodo.4907312" httpUri="https://zenodo.org/record/4907312/files/figure.png" pageId="66" pageNumber="67">Fig. 33c</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="66" pageNumber="67" type="diagnosis">
<paragraph blockId="66.[707,1194,194,1729]" pageId="66" pageNumber="67">
<emphasis bold="true" box="[731,836,1230,1251]" pageId="66" pageNumber="67">Diagnosis.</emphasis>
<taxonomicName authorityName="Thomson" authorityYear="1878" box="[849,1019,1231,1252]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="mystica">
<emphasis box="[849,1019,1231,1252]" italics="true" pageId="66" pageNumber="67">Euphoria mystica</emphasis>
</taxonomicName>
is separated from other species in the group by the shiny pronotum, tomentous elytra, densely punctate scutellum, and form of the parameres.
<bibRefCitation author="Sakai, K. &amp; S. Nagai" box="[925,1146,1310,1331]" journalOrPublisher="Mushi-Sha, Tokyo, Japan" pageId="66" pageNumber="67" refId="ref124976" refString="Sakai, K., and S. Nagai. 1998. The Cetoniine Beetles of the World. Mushi-Sha, Tokyo, Japan." title="The Cetoniine Beetles of the World." type="book" year="1998">Sakai and Nagai (1998)</bibRefCitation>
misidentified this species as
<taxonomicName authorityName="Burmeister" authorityYear="1842" box="[935,1056,1337,1358]" class="Insecta" family="Cetoniidae" genus="Erirhipis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="subguttata">
<emphasis box="[935,1056,1337,1358]" italics="true" pageId="66" pageNumber="67">E. subguttata</emphasis>
</taxonomicName>
(Appendix 5).
</paragraph>
</subSubSection>
<subSubSection pageId="66" pageNumber="67" type="discussion">
<paragraph blockId="66.[707,1194,194,1729]" pageId="66" pageNumber="67">
<emphasis bold="true" box="[731,922,1363,1384]" pageId="66" pageNumber="67">Taxonomic History.</emphasis>
<bibRefCitation author="Thomson, J." box="[934,1095,1363,1384]" journalOrPublisher="E. Deyrolle, Paris, France" pageId="66" pageNumber="67" refId="ref125576" refString="Thomson, J. 1878. Typi Cetonidarum, Suivis de Typi Molommidarum et de Typi Nilionidarum Musaei Thomsoniani. E. Deyrolle, Paris, France." title="Typi Cetonidarum, Suivis de Typi Molommidarum et de Typi Nilionidarum Musaei Thomsoniani." type="book" year="1878">Thomson (1878)</bibRefCitation>
described this species based on a male specimen from “
<collectingCountry box="[1109,1177,1390,1411]" name="Mexico" pageId="66" pageNumber="67">Mexico</collectingCountry>
”. Additional specimens in the MNHN are labeled “
<taxonomicName authority="Guerin" authorityName="Guerin" box="[717,994,1443,1464]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="segmentata">
<emphasis box="[717,921,1443,1464]" italics="true" pageId="66" pageNumber="67">Euphoria segmentata</emphasis>
Guerin
</taxonomicName>
”, but this name was never published and is therefore unavailable.
</paragraph>
<paragraph blockId="66.[707,1194,194,1729]" box="[731,1193,1496,1517]" pageId="66" pageNumber="67">
<emphasis bold="true" box="[731,889,1496,1517]" pageId="66" pageNumber="67">Natural History.</emphasis>
Nothing is known of its biology.
</paragraph>
</subSubSection>
<subSubSection pageId="66" pageNumber="67" type="distribution">
<paragraph blockId="66.[707,1194,194,1729]" pageId="66" pageNumber="67">
<emphasis bold="true" box="[731,965,1523,1544]" pageId="66" pageNumber="67">Temporal Distribution.</emphasis>
March (2), May (1), August (2), September (1) (
<figureCitation box="[976,1060,1549,1570]" captionStart="Fig" captionStartId="152.[197,229,1686,1705]" captionTargetBox="[303,1131,187,1662]" captionTargetId="figure-0@152.[174,1192,183,1662]" captionTargetPageId="152" captionText="Fig. 33. Euphoria mystica. a) Habitus of male, b) Head, c) Parameres, d) Female, e) Temporal distribution, f) Geographic distribution in Guatemala and Mexico." figureDoi="http://doi.org/10.5281/zenodo.4907312" httpUri="https://zenodo.org/record/4907312/files/figure.png" pageId="66" pageNumber="67">Fig. 33e</figureCitation>
).
</paragraph>
<paragraph blockId="66.[707,1194,194,1729]" pageId="66" pageNumber="67">
<emphasis bold="true" box="[731,975,1576,1597]" pageId="66" pageNumber="67">Geographic Distribution.</emphasis>
Known from Veracruz and Chiapas,
<collectingCountry box="[837,908,1602,1623]" name="Mexico" pageId="66" pageNumber="67">Mexico</collectingCountry>
and
<collectingRegion box="[957,1082,1602,1623]" country="Guatemala" name="Alta Verapaz" pageId="66" pageNumber="67">Alta Verapaz</collectingRegion>
,
<collectingCountry box="[1093,1193,1602,1623]" name="Guatemala" pageId="66" pageNumber="67">Guatemala</collectingCountry>
(
<figureCitation box="[713,788,1629,1650]" captionStart="Fig" captionStartId="152.[197,229,1686,1705]" captionTargetBox="[303,1131,187,1662]" captionTargetId="figure-0@152.[174,1192,183,1662]" captionTargetPageId="152" captionText="Fig. 33. Euphoria mystica. a) Habitus of male, b) Head, c) Parameres, d) Female, e) Temporal distribution, f) Geographic distribution in Guatemala and Mexico." figureDoi="http://doi.org/10.5281/zenodo.4907312" httpUri="https://zenodo.org/record/4907312/files/figure.png" pageId="66" pageNumber="67">Fig. 33f</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67" type="materials_examined">
<paragraph blockId="66.[707,1194,194,1729]" lastBlockId="67.[151,639,194,375]" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67">
<emphasis bold="true" box="[731,1006,1655,1676]" pageId="66" pageNumber="67">Specimens Examined (21).</emphasis>
Type material:
<materialsCitation collectionCode="MNHN" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[707,792,1682,1703]" pageId="66" pageNumber="67">Holotype</typeStatus>
<specimenCount box="[798,842,1682,1703]" pageId="66" pageNumber="67" type="male">male</specimenCount>
at
<collectionCode box="[871,942,1682,1703]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="66" pageNumber="67">MNHN</collectionCode>
labeled “mex// Ex-musaeo/ VAN LANSBERGE// MUSÉUM PARIS/ COLL. R. OBERTHUR” and my holotype label.
</materialsCitation>
Other material:
<emphasis bold="true" box="[245,404,221,242]" pageId="67" pageNumber="68">
<collectingCountry box="[245,404,221,242]" name="Guatemala" pageId="67" pageNumber="68">GUATEMALA</collectingCountry>
</emphasis>
(3):
<collectingRegion box="[457,635,221,242]" country="Guatemala" name="Alta Verapaz" pageId="67" pageNumber="68">ALTA VERAPAZ</collectingRegion>
: Baleu (1), Panzós (1);
<taxonomicName box="[375,483,247,268]" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="67" pageNumber="103" phylum="Chordata" rank="species" species="data">NO DATA</taxonomicName>
:
<collectingRegion box="[494,595,247,268]" country="Guatemala" name="Guatemala" pageId="67" pageNumber="68">Guatemala</collectingRegion>
(1).
<emphasis bold="true" box="[152,251,274,295]" pageId="67" pageNumber="68">
<collectingCountry box="[152,251,274,295]" name="Mexico" pageId="67" pageNumber="68">MEXICO</collectingCountry>
</emphasis>
(17):
<collectingRegion box="[307,409,274,295]" country="Mexico" name="Chiapas" pageId="67" pageNumber="68">CHIAPAS</collectingRegion>
: Jetja (1), Pueblo Nuevo (1), Rancho Santa
<taxonomicName box="[352,402,300,321]" class="Magnoliopsida" family="Rosaceae" genus="Rosa" kingdom="Plantae" order="Rosales" pageId="67" pageNumber="103" phylum="Tracheophyta" rank="genus">Rosa</taxonomicName>
(2), San Antonio (3).
<collectingRegion box="[152,283,327,348]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="67" pageNumber="68">VERACRUZ</collectingRegion>
. Córdoba (1), Orizaba (3), Soteapan (2), Xalapa (3), Xico (1).
</paragraph>
</subSubSection>
</treatment>
</document>