365 lines
35 KiB
XML
365 lines
35 KiB
XML
<document ID-DOI="10.1649/0010-066X-66.mo4.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="4907222" approvalRequired="218" approvalRequired_for_taxonomicNames="4" approvalRequired_for_textStreams="213" approvalRequired_for_treatments="1" checkinTime="1623089673881" checkinUser="carolina" docAuthor="Orozco, Jesús" docDate="2012" docId="F449F723D538B27185F642A0EB71FD6D" docLanguage="en" docName="TheColeopBull.66.mo11.1-182.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 11) 66" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Euphoria subtomentosa" docType="treatment" docVersion="3" lastPageNumber="28" masterDocId="08708F5BD522B26A85194065E874FF87" masterDocTitle="Monographic Revision of the American Genus Euphoria Burmeister, 1842 (Coleoptera: Scarabaeidae: Cetoniinae)" masterLastPageNumber="182" masterPageNumber="1" pageNumber="27" updateTime="1663005054238" updateUser="tatiana" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Monographic Revision of the American Genus Euphoria Burmeister, 1842 (Coleoptera: Scarabaeidae: Cetoniinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Orozco, Jesús</mods:namePart>
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<mods:affiliation>Molecular Ecology and Systematics Group Department of Botany and Department of Zoology and Entomology Rhodes University Grahamstown, SOUTH AFRICA 6140</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="series">
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<mods:title>mo 11</mods:title>
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</mods:detail>
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<mods:detail type="pubDate">
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<mods:number>2012-12-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>66</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>182</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1649/0010-066X-66.mo4.1</mods:identifier>
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<mods:identifier type="ISSN">1938-4394</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4907222</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:F449F723D538B27185F642A0EB71FD6D" httpUri="http://treatment.plazi.org/id/F449F723D538B27185F642A0EB71FD6D" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
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<subSubSection pageId="26" pageNumber="27" type="nomenclature">
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<paragraph blockId="26.[239,592,709,784]" pageId="26" pageNumber="27">
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<heading centered="true" fontSize="10" level="2" pageId="26" pageNumber="27" reason="2">
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<taxonomicName authority="(Gory and Percheron, 1833)" baseAuthorityName="Gory and Percheron" baseAuthorityYear="1833" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="subtomentosa">
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<emphasis bold="true" pageId="26" pageNumber="27">
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<emphasis bold="true" box="[239,475,709,730]" italics="true" pageId="26" pageNumber="27">Euphoria subtomentosa</emphasis>
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(Gory and Percheron, 1833)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[314,517,763,784]" pageId="26" pageNumber="27" type="description">
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<paragraph blockId="26.[239,592,709,784]" box="[314,517,763,784]" pageId="26" pageNumber="27">
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(Appendix 4:
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<figureCitation box="[452,509,763,784]" captionStart="Fig" captionStartId="128.[197,229,1686,1705]" captionTargetBox="[256,1148,185,1662]" captionTargetId="figure-0@128.[174,1192,183,1662]" captionTargetPageId="128" captionText="Fig. 9. Euphoria subtomentosa. a) Habitus, b) Head, c) Parameres, d) Temporal distribution, e) Geographic distribution in Mexico." figureDoi="http://doi.org/10.5281/zenodo.4907252" httpUri="https://zenodo.org/record/4907252/files/figure.png" pageId="26" pageNumber="27">Fig. 9</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="27" type="reference_group">
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<paragraph blockId="26.[173,660,805,879]" pageId="26" pageNumber="27">
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<taxonomicName authority="Gory and Percheron 1833: 64" authorityName="Gory and Percheron" authorityPageNumber="64" authorityYear="1833" class="Insecta" family="Cetoniidae" genus="Cetonia" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="subtomentosa">
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<emphasis box="[173,387,805,826]" italics="true" pageId="26" pageNumber="27">Cetonia subtomentosa</emphasis>
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<materialsCitation collectionCode="MHNG" pageId="26" pageNumber="27" specimenCount="1" typeStatus="holotype">Gory and Percheron 1833: 64</materialsCitation>
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</taxonomicName>
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. Original combination.
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<typeStatus box="[457,544,832,853]" pageId="26" pageNumber="27">Holotype</typeStatus>
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at MHNG, examined.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="27" type="description">
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<paragraph blockId="26.[173,660,911,1729]" lastBlockId="26.[707,1194,194,1729]" pageId="26" pageNumber="27">
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<emphasis bold="true" box="[197,428,911,932]" pageId="26" pageNumber="27">
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Description (
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<emphasis bold="true" box="[331,344,911,932]" italics="true" pageId="26" pageNumber="27">n</emphasis>
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= 444).
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</emphasis>
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Length
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<quantity box="[519,655,911,933]" metricMagnitude="-2" metricUnit="m" metricValue="1.0100000000000002" metricValueMax="1.1800000000000002" metricValueMin="0.8400000000000001" pageId="26" pageNumber="27" unit="mm" value="10.1" valueMax="11.8" valueMin="8.4">8.4 –11.8 mm</quantity>
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; width
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<quantity box="[236,356,938,960]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="7.4" metricValueMin="5.1" pageId="26" pageNumber="27" unit="mm" value="6.25" valueMax="7.4" valueMin="5.1">5.1–7.4 mm</quantity>
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.
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<emphasis bold="true" box="[368,426,938,959]" pageId="26" pageNumber="27">Color</emphasis>
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: Head black, pronotum and elytra black or reddish brown. Elytra with bright yellow, thick, cretaceous markings distributed longitudinally on striae, covering elytral apex entirely. Pygidium almost entirely covered by cretaceous layer.
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<emphasis bold="true" box="[387,443,1071,1092]" pageId="26" pageNumber="27">Head</emphasis>
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: Frons flat, strongly strigose, densely setose; setae moderate to long, yellowish. Clypeus short, subtrapezoidal, moderately expanded anteriorly, strongly strigopunctate, sides not raised, apex strongly truncate, weakly reflexed. Antennal club shorter than stem, subequal in both sexes.
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<emphasis bold="true" box="[320,423,1230,1251]" pageId="26" pageNumber="27">Pronotum</emphasis>
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: Surface densely punctate, punctures small to moderate in size, confluent towards sides, densely setose, setae long, yellowish; sides weakly angulate to evenly rounded. Scutellum longer than wide, densely punctate, punctures lunulate, setigerous; setae moderate to long, yellowish. Base in front of scutellum weakly to strongly emarginate.
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<emphasis bold="true" box="[397,461,1416,1437]" pageId="26" pageNumber="27">Elytra</emphasis>
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: Surface moderately densely punctate, punctures lunulate, small to moderate in size, setigerous; setae short to long, yellowish. Posterior half of sutural costa raised in lateral view.
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<emphasis bold="true" box="[299,396,1523,1544]" pageId="26" pageNumber="27">Pygidium</emphasis>
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: Surface subconcentrically striate; striae discontinuous, weakly impressed, moderately densely setose; setae short, yellowish.
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<emphasis bold="true" box="[173,221,1602,1623]" pageId="26" pageNumber="27">Legs</emphasis>
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: Protibial teeth sharp, apical and medial teeth closer to each other than to basal tooth. Mesotibial carina well developed into 1 long and 1 short spine. Metatibiae with carina oblique, apex moderately expanded, spurs subequal in length both sexes.
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<emphasis bold="true" box="[776,843,194,215]" pageId="26" pageNumber="27">Venter</emphasis>
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: Mesometasternal process extending anteriorly to same level as mesocoxae or slightly beyond, apex truncate to rounded, densely setose. Mesepimera, metasternum, and metacoxae setose, setae as on legs. Metasternum rugose, setose laterally, glabrous and impunctate at middle. Median sulcus vaguely to weakly defined. Abdominal sternites densely setose, setae as on rest of body; abdomen in lateral view subequal in shape in both sexes.
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<emphasis bold="true" box="[861,1014,433,454]" pageId="26" pageNumber="27">Male genitalia</emphasis>
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: Parameres as in
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<figureCitation box="[707,778,460,481]" captionStart="Fig" captionStartId="128.[197,229,1686,1705]" captionTargetBox="[256,1148,185,1662]" captionTargetId="figure-0@128.[174,1192,183,1662]" captionTargetPageId="128" captionText="Fig. 9. Euphoria subtomentosa. a) Habitus, b) Head, c) Parameres, d) Temporal distribution, e) Geographic distribution in Mexico." figureDoi="http://doi.org/10.5281/zenodo.4907252" httpUri="https://zenodo.org/record/4907252/files/figure.png" pageId="26" pageNumber="27">Fig. 9c</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="27" type="diagnosis">
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<paragraph blockId="26.[707,1194,194,1729]" pageId="26" pageNumber="27">
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<emphasis bold="true" box="[731,836,486,507]" pageId="26" pageNumber="27">Diagnosis.</emphasis>
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<taxonomicName baseAuthorityName="Gory and Percheron" baseAuthorityYear="1833" box="[849,1076,486,507]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="subtomentosa">
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<emphasis box="[849,1076,486,507]" italics="true" pageId="26" pageNumber="27">Euphoria subtomentosa</emphasis>
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</taxonomicName>
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is separated from other species in the group by the subtrapezoidal clypeus and the cretaceous markings on the elytra that are distributed longitudinally on the elytral striae.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="discussion">
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<paragraph blockId="26.[707,1194,194,1729]" pageId="26" pageNumber="27">
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<emphasis bold="true" box="[731,929,593,614]" pageId="26" pageNumber="27">Taxonomic History.</emphasis>
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Gory and Percheron (1833) described
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<taxonomicName box="[805,985,619,640]" class="Insecta" family="Cetoniidae" genus="Cetonia" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="tomentosa">
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<emphasis box="[805,985,619,640]" italics="true" pageId="26" pageNumber="27">Cetonia tomentosa</emphasis>
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</taxonomicName>
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twice in their monograph under the same name from specimens labeled as being from
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<collectingCountry box="[847,921,672,693]" name="Mexico" pageId="26" pageNumber="27">Mexico</collectingCountry>
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. The first description (pages 63, 266), “
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<taxonomicName authority="Klug." authorityName="Klug." box="[813,1055,699,720]" class="Insecta" family="Cetoniidae" genus="Cetonia" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="tomentosa">Cetonia tomentosa Klug.</taxonomicName>
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”, corresponds to a mislabeled
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<taxonomicName box="[873,1135,726,747]" class="Insecta" family="Cetoniidae" genus="Anoplocheilus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="tomentosus">
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<emphasis box="[873,1135,726,747]" italics="true" pageId="26" pageNumber="27">Anoplocheilus tomentosus</emphasis>
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</taxonomicName>
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from
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<collectingCountry box="[707,842,752,773]" name="South Africa" pageId="26" pageNumber="27">South Africa</collectingCountry>
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. The second specimen (pages 64, 278) “
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<taxonomicName authority="Steven" authorityName="Steven" box="[774,1042,779,800]" class="Insecta" family="Cetoniidae" genus="Cetonia" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="tomentosa">Cetonia tomentosa Stéven</taxonomicName>
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” constitutes a Mexican specimen. The fact that the same name was used for two different species, and the use of other authors names after the species name, has caused considerable confusion regarding the valid name for the Mexican species and to whom authorship should be attributed to (
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<emphasis box="[1010,1016,938,959]" italics="true" pageId="26" pageNumber="27">i</emphasis>
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.
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<emphasis box="[1024,1034,938,959]" italics="true" pageId="26" pageNumber="27">e</emphasis>
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., Dejean 1836;
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<bibRefCitation author="Mannerheim" box="[707,882,964,985]" pageId="26" pageNumber="27" pagination="127 - 163" refId="ref123933" refString="Mannerheim, C. G. 1837. Observations critiques sur quelques ouvrages entomologiques. Bulletin de la Societe Imperiale des Naturalistes de Moscou 10: 127 - 163." type="journal article" year="1837">Mannerheim 1837</bibRefCitation>
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; Bates 1889;
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<bibRefCitation author="Blackwelder" box="[1016,1189,964,985]" pageId="26" pageNumber="27" pagination="189 - 341" refId="ref121444" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America, part 2. Bulletin of the United States National Museum 185: 189 - 341." type="journal article" year="1944">Blackwelder 1944</bibRefCitation>
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;
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<bibRefCitation author="Hardy" box="[707,823,991,1012]" pageId="26" pageNumber="27" pagination="127 - 143" refId="ref122900" refString="Hardy, A. R. 2001. Studies in the Euphoriina of the Americas (Coleoptera: Scarabaeidae) II. Status of names in Euphoria, types and synonymies, with notes on the South American species. Pan- Pacific Entomologist 77: 127 - 143." type="journal article" year="2001">Hardy 2001</bibRefCitation>
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).
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</paragraph>
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<paragraph blockId="26.[707,1194,194,1729]" pageId="26" pageNumber="27">
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Dejean’ s catalogue, considered the “bible” for coleopterists at the time (Barber and Bridwell 1940) indicated that, “Subtomentosa Hope”, “
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<taxonomicName authority="Sturm" authorityName="Sturm" box="[717,886,1097,1118]" class="Insecta" family="Cetoniidae" genus="Cetonia" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="28" phylum="Arthropoda" rank="species" species="tomentosa">Tomentosa Sturm</taxonomicName>
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,” “Multipunctata Höpfner”, and “Spardalina Dupont” all referred to the same species. “Multipunctata Höpfner”, and “Spardalina Dupont” were never described and are considered
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<emphasis box="[707,835,1204,1225]" italics="true" pageId="26" pageNumber="27">nomina nuda</emphasis>
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. In Gory and Percheron (1833) and Dejean (1836), the name following a species does not necessarily refer to the species’ author, but can also refer to the collection from where the specimen came or to the collection where the specimen was observed with that name.
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</paragraph>
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<paragraph blockId="26.[707,1194,194,1729]" pageId="26" pageNumber="27">
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<bibRefCitation author="Mannerheim" box="[731,925,1363,1384]" pageId="26" pageNumber="27" pagination="127 - 163" refId="ref123933" refString="Mannerheim, C. G. 1837. Observations critiques sur quelques ouvrages entomologiques. Bulletin de la Societe Imperiale des Naturalistes de Moscou 10: 127 - 163." type="journal article" year="1837">Mannerheim (1837)</bibRefCitation>
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, in his review of Gory and Percheron (1833), noted that
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<taxonomicName authority="Steven" authorityName="Steven" class="Insecta" family="Cetoniidae" genus="Cetonia" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="tomentosa">
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<emphasis box="[1005,1193,1390,1411]" italics="true" pageId="26" pageNumber="27">Cetonia tomentosa</emphasis>
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Stéven
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</taxonomicName>
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was not named by Stéven but by Sturm (origin of the name, not the description). He also noted that Dejean (1836) adopted the name “Subtomentosa Hope” for the Mexican species because of the two homonyms.
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</paragraph>
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<paragraph blockId="26.[707,1194,194,1729]" lastBlockId="27.[151,639,194,1729]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
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Burmeister (1842) used Gory and Percheron as the authors of the species (
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<taxonomicName box="[966,1179,1576,1597]" class="Insecta" family="Cetoniidae" genus="Cetonia" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="subtomentosa">
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<emphasis box="[966,1179,1576,1597]" italics="true" pageId="26" pageNumber="27">Cetonia subtomentosa</emphasis>
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</taxonomicName>
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). In addition, he agreed with
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<bibRefCitation author="Mannerheim" box="[993,1193,1602,1623]" pageId="26" pageNumber="27" pagination="127 - 163" refId="ref123933" refString="Mannerheim, C. G. 1837. Observations critiques sur quelques ouvrages entomologiques. Bulletin de la Societe Imperiale des Naturalistes de Moscou 10: 127 - 163." type="journal article" year="1837">Mannerheim (1837)</bibRefCitation>
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that Sturm was the originator of the name and not Stéven. Thomson (1878) also used Gory and Percheron as the authors of this species. Bates (1889),
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<bibRefCitation author="Schenkling" box="[781,955,1708,1729]" pageId="26" pageNumber="27" pagination="334 - 344" refId="ref125224" refString="Schenkling, S. 1921. Scarabaeidae: Cetoniinae [pp. 334 - 344]. In: Coleopterorum Catalogus, 21 (72): Scarabaeidae III. (S. Schenkling, editor). W. Junk, Berlin, Germany." type="journal article" year="1921">Schenkling (1921)</bibRefCitation>
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, and
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<bibRefCitation author="Blackwelder" box="[1008,1193,1708,1729]" pageId="26" pageNumber="27" pagination="189 - 341" refId="ref121444" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America, part 2. Bulletin of the United States National Museum 185: 189 - 341." type="journal article" year="1944">Blackwelder (1944)</bibRefCitation>
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used Mannerheim as the author of the species. None of these three authors provided a rationale for this decision.
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<bibRefCitation author="Hardy" box="[335,472,247,268]" pageId="27" pageNumber="28" pagination="127 - 143" refId="ref122900" refString="Hardy, A. R. 2001. Studies in the Euphoriina of the Americas (Coleoptera: Scarabaeidae) II. Status of names in Euphoria, types and synonymies, with notes on the South American species. Pan- Pacific Entomologist 77: 127 - 143." type="journal article" year="2001">Hardy (2001)</bibRefCitation>
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used Dejean as the author of the species arguing Dejean used
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<taxonomicName box="[151,329,301,322]" class="Insecta" family="Cetoniidae" genus="Cetonia" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="subtomentosa">
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<emphasis box="[151,329,301,322]" italics="true" pageId="27" pageNumber="28">C. subtomentosa</emphasis>
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</taxonomicName>
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as a replacement name for “
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<taxonomicName authority="Sturm" authorityName="Sturm" box="[162,325,327,348]" class="Insecta" family="Cetoniidae" genus="Cetonia" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="tomentosa">tomentosa Sturm</taxonomicName>
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”.
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</paragraph>
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<paragraph blockId="27.[151,639,194,1729]" pageId="27" pageNumber="28">
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Gory and Percheron (1833) provided a short description of the species in Latin at the beginning of their book (
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<emphasis box="[351,566,407,428]" italics="true" pageId="27" pageNumber="28">Tableau Diagnostique</emphasis>
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) and a longer one in French later on. Both descriptions of
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<taxonomicName box="[180,310,460,481]" class="Insecta" family="Cetoniidae" genus="Cetonia" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="tomentosa">
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<emphasis box="[180,196,460,481]" italics="true" pageId="27" pageNumber="28">C</emphasis>
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.
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<emphasis box="[211,310,460,481]" italics="true" pageId="27" pageNumber="28">tomentosa</emphasis>
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</taxonomicName>
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use this name in all parts of the book. In the case of the Mexican specimen, an additional name is given in the
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<emphasis box="[399,599,513,534]" italics="true" pageId="27" pageNumber="28">Tableau Diagnostique</emphasis>
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followed by the description (page 64): “
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<emphasis italics="true" pageId="27" pageNumber="28">TOMENTOSA, Steven</emphasis>
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.,
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<emphasis box="[238,384,566,587]" italics="true" pageId="27" pageNumber="28">Sub-Tomentosa</emphasis>
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.
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<emphasis box="[401,453,566,587]" italics="true" pageId="27" pageNumber="28">Hope</emphasis>
|
||
.
|
||
<emphasis box="[471,554,566,587]" italics="true" pageId="27" pageNumber="28">
|
||
<collectingCountry box="[471,554,566,587]" name="Mexico" pageId="27" pageNumber="28">Mexique</collectingCountry>
|
||
</emphasis>
|
||
.
|
||
<emphasis italics="true" pageId="27" pageNumber="28">
|
||
<taxonomicName authorityName="Scott" authorityYear="1963" box="[571,634,566,587]" class="Collembola" family="Entomobryidae" genus="Rubra" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">Rubra</taxonomicName>
|
||
, thorace corporeque obscurioribus vittis duabus longitudinabilus griseis in elytris, prima exteriore
|
||
<taxonomicName box="[151,225,646,667]" class="Magnoliopsida" family="Asteraceae" genus="Viguiera" kingdom="Plantae" order="Asterales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="dentata">dentata</taxonomicName>
|
||
, secunda posticali”
|
||
</emphasis>
|
||
. This constitutes the description of
|
||
<taxonomicName box="[298,461,672,693]" class="Insecta" family="Cetoniidae" genus="Cetonia" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="subtomentosa">
|
||
<emphasis box="[298,312,672,693]" italics="true" pageId="27" pageNumber="28">E</emphasis>
|
||
.
|
||
<emphasis box="[329,461,672,693]" italics="true" pageId="27" pageNumber="28">subtomentosa</emphasis>
|
||
</taxonomicName>
|
||
even in this rare two-names scenario. Gory and Percheron (1833) were probably illustrating the fact that there were specimens of this species in the Stéven collection under the name “Tomentosa” and some others in the Hope collection under the name “Sub-Tomentosa”. The fact that there is a description attached to the name of the species (“Sub-tomentosa”) and that the other name used in the description (“Tomentosa”) is currently a senior homonym for a South African specimen justifies the use of Gory and Percheron as the authors of this species.
|
||
</paragraph>
|
||
<paragraph blockId="27.[151,639,194,1729]" pageId="27" pageNumber="28">
|
||
<emphasis bold="true" box="[175,334,991,1012]" pageId="27" pageNumber="28">Natural History.</emphasis>
|
||
Adults of
|
||
<taxonomicName box="[437,589,991,1012]" class="Insecta" family="Cetoniidae" genus="Cetonia" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="subtomentosa">
|
||
<emphasis box="[437,451,991,1012]" italics="true" pageId="27" pageNumber="28">E</emphasis>
|
||
.
|
||
<emphasis box="[463,589,991,1012]" italics="true" pageId="27" pageNumber="28">subtomentosa</emphasis>
|
||
</taxonomicName>
|
||
have been observed on various species of
|
||
<taxonomicName baseAuthorityName="Deloya" baseAuthorityYear="1988" box="[524,635,1018,1039]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
|
||
:
|
||
<taxonomicName authority="Cav." authorityName="Cav." box="[151,367,1044,1066]" class="Magnoliopsida" family="Asteraceae" genus="Bidens" kingdom="Plantae" order="Asterales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="odorata">
|
||
<emphasis box="[151,309,1045,1066]" italics="true" pageId="27" pageNumber="28">Bidens odorata</emphasis>
|
||
Cav.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="(Cav.)" baseAuthorityName="Cav." box="[379,634,1044,1066]" class="Magnoliopsida" family="Asteraceae" genus="Dyssodia" kingdom="Plantae" order="Asterales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="pinnata">
|
||
<emphasis box="[379,559,1045,1066]" italics="true" pageId="27" pageNumber="28">Dyssodia pinnata</emphasis>
|
||
(Cav.)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName box="[151,335,1071,1092]" class="Magnoliopsida" family="Asteraceae" genus="Helianthus" kingdom="Plantae" order="Asterales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="annus">
|
||
<emphasis box="[151,335,1071,1092]" italics="true" pageId="27" pageNumber="28">Helianthus annus</emphasis>
|
||
</taxonomicName>
|
||
L. flowers, and
|
||
<taxonomicName box="[525,631,1071,1092]" class="Magnoliopsida" family="Asteraceae" genus="Viguiera" kingdom="Plantae" order="Asterales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="dentata">
|
||
<emphasis box="[525,631,1071,1092]" italics="true" pageId="27" pageNumber="28">V. dentata</emphasis>
|
||
</taxonomicName>
|
||
. Adults have also been collected in
|
||
<taxonomicName box="[514,637,1098,1119]" class="Insecta" family="Formicidae" genus="Atta" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mexicana">
|
||
<emphasis box="[514,637,1098,1119]" italics="true" pageId="27" pageNumber="28">A. mexicana</emphasis>
|
||
</taxonomicName>
|
||
debris piles (
|
||
<bibRefCitation author="Deloya" box="[285,417,1124,1145]" pageId="27" pageNumber="28" pagination="77 - 91" refId="ref122112" refString="Deloya, C. 1988. Coleopteros lamelicornios asociados a depositos de detritos de Atta mexicana (Smith) (Hymenoptera: Formicidae) en el sur del estado de Morelos, Mexico. Folia Entomologica Mexicana 75: 77 - 91." type="journal article" year="1988">Deloya 1988</bibRefCitation>
|
||
) and at elevations of
|
||
<quantity box="[151,305,1150,1172]" metricMagnitude="3" metricUnit="m" metricValue="1.8" metricValueMax="2.2" metricValueMin="1.4" pageId="27" pageNumber="28" unit="m" value="1800.0" valueMax="2200.0" valueMin="1400.0">
|
||
<geoCoordinate box="[151,206,1151,1172]" degrees="1,400" direction="north" orientation="latitude" pageId="27" pageNumber="28" precision="55" value="1.4">1,400</geoCoordinate>
|
||
<geoCoordinate box="[206,274,1150,1172]" degrees="2,200" direction="west" orientation="longitude" pageId="27" pageNumber="28" precision="55" value="-2.2">–2,200</geoCoordinate>
|
||
m
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="27" pageNumber="28" type="distribution">
|
||
<paragraph blockId="27.[151,639,194,1729]" pageId="27" pageNumber="28">
|
||
<emphasis bold="true" box="[175,409,1177,1198]" pageId="27" pageNumber="28">Temporal Distribution.</emphasis>
|
||
April (1), May (1), June (17), July (4), August (4), September (148), October (172), November (27) (
|
||
<figureCitation box="[463,536,1230,1251]" captionStart="Fig" captionStartId="128.[197,229,1686,1705]" captionTargetBox="[256,1148,185,1662]" captionTargetId="figure-0@128.[174,1192,183,1662]" captionTargetPageId="128" captionText="Fig. 9. Euphoria subtomentosa. a) Habitus, b) Head, c) Parameres, d) Temporal distribution, e) Geographic distribution in Mexico." figureDoi="http://doi.org/10.5281/zenodo.4907252" httpUri="https://zenodo.org/record/4907252/files/figure.png" pageId="27" pageNumber="28">Fig. 9d</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="27.[151,639,194,1729]" pageId="27" pageNumber="28">
|
||
<emphasis bold="true" box="[175,455,1257,1278]" pageId="27" pageNumber="28">Geographic Distribution.</emphasis>
|
||
K n o w n f r o m
|
||
<materialsCitation country="Mexico" location="Aguascalientes" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Aguascalientes">
|
||
<collectingRegion box="[151,314,1283,1304]" country="Mexico" name="Aguascalientes" pageId="27" pageNumber="28">Aguascalientes</collectingRegion>
|
||
,
|
||
<collectingRegion box="[331,419,1283,1304]" country="Mexico" name="Chiapas" pageId="27" pageNumber="28">Chiapas</collectingRegion>
|
||
,
|
||
<collectingRegion box="[435,515,1283,1304]" country="Mexico" name="Colima" pageId="27" pageNumber="28">Colima</collectingRegion>
|
||
, Estado de
|
||
<collectingCountry box="[151,223,1310,1331]" name="Mexico" pageId="27" pageNumber="28">México</collectingCountry>
|
||
</materialsCitation>
|
||
,
|
||
<collectingRegion box="[235,319,1310,1331]" country="Mexico" name="Guerrero" pageId="27" pageNumber="28">Guerrero</collectingRegion>
|
||
,
|
||
<collectingRegion box="[330,435,1310,1331]" country="Mexico" name="Guanajuato" pageId="27" pageNumber="28">Guanajuato</collectingRegion>
|
||
(
|
||
<bibRefCitation author="Deloya" box="[449,627,1310,1331]" pageId="27" pageNumber="28" pagination="15 - 55" refId="ref122155" refString="Deloya, C., A. Burgos, J. Blackaller, and J. M. Lobo. 1993. Los coleopteros lamelicornios de Cuernavaca, Morelos, Mexico (Passalidae, Trogidae, Scarabaeidae y Melolonthidae). Boletin de la Sociedad Veracruzana de Zoologia 3: 15 - 55." type="journal article" year="1993">
|
||
Deloya
|
||
<emphasis box="[523,565,1310,1331]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
|
||
. 1993
|
||
</bibRefCitation>
|
||
),
|
||
<collectingRegion box="[151,233,1337,1358]" country="Mexico" name="Hidalgo" pageId="27" pageNumber="28">Hidalgo</collectingRegion>
|
||
,
|
||
<collectingRegion box="[248,318,1337,1358]" country="Mexico" name="Jalisco" pageId="27" pageNumber="28">Jalisco</collectingRegion>
|
||
,
|
||
<collectingRegion box="[332,444,1337,1358]" country="Mexico" name="Michoacan de Ocampo" pageId="27" pageNumber="28">Michoacán</collectingRegion>
|
||
,
|
||
<collectingRegion box="[459,544,1337,1358]" country="Mexico" name="Morelos" pageId="27" pageNumber="28">Morelos</collectingRegion>
|
||
,
|
||
<collectingRegion box="[558,634,1337,1358]" country="Mexico" name="Nayarit" pageId="27" pageNumber="28">Nayarit</collectingRegion>
|
||
,
|
||
<collectingRegion box="[151,223,1363,1384]" country="Mexico" name="Oaxaca" pageId="27" pageNumber="28">Oaxaca</collectingRegion>
|
||
,
|
||
<collectingRegion box="[235,301,1363,1384]" country="Mexico" name="Puebla" pageId="27" pageNumber="28">Puebla</collectingRegion>
|
||
, and
|
||
<materialsCitation box="[355,616,1363,1384]" country="Mexico" location="Veracruz" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
|
||
<collectingRegion box="[355,440,1363,1384]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="27" pageNumber="28">Veracruz</collectingRegion>
|
||
,
|
||
<collectingCountry box="[451,522,1363,1384]" name="Mexico" pageId="27" pageNumber="28">Mexico</collectingCountry>
|
||
(
|
||
<figureCitation box="[538,608,1363,1384]" captionStart="Fig" captionStartId="128.[197,229,1686,1705]" captionTargetBox="[256,1148,185,1662]" captionTargetId="figure-0@128.[174,1192,183,1662]" captionTargetPageId="128" captionText="Fig. 9. Euphoria subtomentosa. a) Habitus, b) Head, c) Parameres, d) Temporal distribution, e) Geographic distribution in Mexico." figureDoi="http://doi.org/10.5281/zenodo.4907252" httpUri="https://zenodo.org/record/4907252/files/figure.png" pageId="27" pageNumber="28">Fig. 9e</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="27" pageNumber="28" type="materials_examined">
|
||
<paragraph blockId="27.[151,639,194,1729]" lastBlockId="27.[685,1172,194,746]" pageId="27" pageNumber="28">
|
||
<emphasis bold="true" box="[175,464,1390,1411]" pageId="27" pageNumber="28">Specimens Examined (444).</emphasis>
|
||
Type material:
|
||
<typeStatus box="[151,239,1416,1437]" pageId="27" pageNumber="28">Holotype</typeStatus>
|
||
female at MHNG labeled “Gory/ TYPE// subtomentosa/ B/ tomentosa G.P./ Mexico” and my
|
||
<typeStatus box="[151,236,1469,1490]" pageId="27" pageNumber="28">holotype</typeStatus>
|
||
label. Other material:
|
||
<emphasis bold="true" box="[467,569,1469,1490]" pageId="27" pageNumber="28">
|
||
<collectingCountry box="[467,569,1469,1490]" name="Mexico" pageId="27" pageNumber="28">MEXICO</collectingCountry>
|
||
</emphasis>
|
||
(443):
|
||
<collectingRegion box="[151,373,1496,1517]" country="Mexico" name="Aguascalientes" pageId="27" pageNumber="28">AGUASCALIENTES</collectingRegion>
|
||
: Pabellón (1).
|
||
<collectingRegion box="[528,635,1496,1517]" country="Mexico" name="Chiapas" pageId="27" pageNumber="28">CHIAPAS</collectingRegion>
|
||
: Parque Nacional Cañon del Sumidero (6), Tuxtla Gutiérrez (16),
|
||
<taxonomicName box="[296,365,1549,1570]" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="103" phylum="Chordata" rank="species" species="data">no data</taxonomicName>
|
||
(5);
|
||
<collectingRegion box="[413,508,1549,1570]" country="Mexico" name="Colima" pageId="27" pageNumber="28">COLIMA</collectingRegion>
|
||
: Colima (10);
|
||
<collectingRegion box="[151,375,1576,1597]" country="Mexico" name="Mexico" pageId="27" pageNumber="28">ESTADO DE MÉXICO</collectingRegion>
|
||
: Coatlinchan (1), Xochimilco (1);
|
||
<collectingRegion box="[196,333,1602,1623]" country="Mexico" name="Guerrero" pageId="27" pageNumber="28">GUERRERO</collectingRegion>
|
||
: Chichihualco (
|
||
<quantity box="[495,561,1602,1623]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="27" pageNumber="28" unit="km" value="22.0">22 km</quantity>
|
||
E) (2), Chilpancingo (7), Iguala (3), Mochitlán (3), Taxco (58), Teloloapan (2), Tixtla (2);
|
||
<collectingRegion box="[517,635,1655,1676]" country="Mexico" name="Hidalgo" pageId="27" pageNumber="28">HIDALGO</collectingRegion>
|
||
: Zimapán (1);
|
||
<collectingRegion box="[292,394,1682,1703]" country="Mexico" name="Jalisco" pageId="27" pageNumber="28">JALISCO</collectingRegion>
|
||
: Ajijíc (45), Autlán (3), Barra de Navidad (1), Chapala (1), Cocula (1), Guadalajara (7), Huentitán (2), Jocotepec (8), Lago de Chapala (1), Melaque (1), San Fandia (3), San Gabriel (5), Sayula (2), Tecolotlán (1), Teocaltriche (1), Tequila (1), Tesistán (1), Tizapán (2), Tlaquepaque (2), Tuxcacuesco (1), Volcán Colima (28), Zapopan (3);
|
||
<collectingRegion box="[732,893,327,348]" country="Mexico" name="Michoacan de Ocampo" pageId="27" pageNumber="28">MICHOACÁN</collectingRegion>
|
||
: Acahuato (1), Cotija (2);
|
||
<collectingRegion box="[685,809,354,375]" country="Mexico" name="Morelos" pageId="27" pageNumber="28">MORELOS</collectingRegion>
|
||
: Amacuzac (5), Cuernavaca (34), Tepoztlán (3), Tizapán (3), Xochicalco (15), Xochitepec (6), Yautepec (1), Zacatepec de Hidalgo (1);
|
||
<collectingRegion box="[814,920,433,454]" country="Mexico" name="Nayarit" pageId="27" pageNumber="28">NAYARIT</collectingRegion>
|
||
: Volcán El Ceboruco (1);
|
||
<collectingRegion box="[685,797,460,481]" country="Mexico" name="Oaxaca" pageId="27" pageNumber="28">OAXACA</collectingRegion>
|
||
: Ocotepec (5), Huajuapan (9), San Francisco Huapanapa (1), Monte Albán (3), Oaxaca (11), Mitla (1), Nochixtlán (1), Puebla (1), San Juan Baustista Huicatlán (2), San Sebastian Tutla (2), Santiago Cacaloxtepec (13), Santiago Chazumba (37),
|
||
<taxonomicName box="[842,910,593,614]" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="103" phylum="Chordata" rank="species" species="data">no data</taxonomicName>
|
||
(1);
|
||
<collectingRegion box="[957,1051,593,614]" country="Mexico" name="Puebla" pageId="27" pageNumber="28">PUEBLA</collectingRegion>
|
||
: Acatlán (4), Calipán (2), Coxcatlán (1), Izúcar de Matamoros (14), Petlalcingo (6), Piaxtla (1), Tehuacán (1), Tepexco (4), Zapotitlán (6);
|
||
<collectingRegion box="[953,1081,672,693]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="27" pageNumber="28">VERACRUZ</collectingRegion>
|
||
: Córdoba (1), Xalapa (1);
|
||
<taxonomicName box="[871,990,699,720]" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="103" phylum="Chordata" rank="species" species="data">NO DATA</taxonomicName>
|
||
: Mexico (1),
|
||
<taxonomicName class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="103" phylum="Chordata" rank="species" species="data">no data</taxonomicName>
|
||
(4).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |