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<document ID-DOI="10.1649/0010-066X-66.mo4.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="4907222" approvalRequired="4" approvalRequired_for_taxonomicNames="4" checkinTime="1623089673881" checkinUser="carolina" docAuthor="Orozco, Jesús" docDate="2012" docId="F449F723D516B25F85E245ADEB18FBD9" docLanguage="en" docName="TheColeopBull.66.mo11.1-182.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 11) 66" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Euphoria candezei Janson 1875" docType="treatment" docVersion="6" lastPageNumber="54" masterDocId="08708F5BD522B26A85194065E874FF87" masterDocTitle="Monographic Revision of the American Genus Euphoria Burmeister, 1842 (Coleoptera: Scarabaeidae: Cetoniinae)" masterLastPageNumber="182" masterPageNumber="1" pageNumber="53" updateTime="1663356291692" updateUser="tatiana" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Monographic Revision of the American Genus Euphoria Burmeister, 1842 (Coleoptera: Scarabaeidae: Cetoniinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Orozco, Jesús</mods:namePart>
<mods:affiliation>Molecular Ecology and Systematics Group Department of Botany and Department of Zoology and Entomology Rhodes University Grahamstown, SOUTH AFRICA 6140</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="series">
<mods:title>mo 11</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2012-12-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>66</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>182</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1649/0010-066X-66.mo4.1</mods:identifier>
<mods:identifier type="ISSN">1938-4394</mods:identifier>
<mods:identifier type="Zenodo-Dep">4907222</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:F449F723D516B25F85E245ADEB18FBD9" httpUri="http://treatment.plazi.org/id/F449F723D516B25F85E245ADEB18FBD9" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<subSubSection box="[251,580,1480,1501]" pageId="52" pageNumber="53" type="nomenclature">
<paragraph blockId="52.[251,580,1480,1528]" box="[251,580,1480,1501]" pageId="52" pageNumber="53">
<heading box="[251,580,1480,1501]" centered="true" fontSize="10" level="2" pageId="52" pageNumber="53" reason="2">
<taxonomicName authority="Janson, 1875" authorityName="Janson" authorityYear="1875" box="[251,580,1480,1501]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="candezei">
<emphasis bold="true" box="[251,580,1480,1501]" pageId="52" pageNumber="53">
<emphasis bold="true" box="[251,437,1480,1501]" italics="true" pageId="52" pageNumber="53">Euphoria candezei</emphasis>
<bibRefCitation author="Janson" box="[445,580,1480,1501]" firstAuthor="Janson" pageId="52" pageNumber="53" pagination="373 - 376" refId="ref123384" refString="Janson, O. E. 1875. Descriptions of three new species of American Cetoniidae. Cistula Entomologica 1: 373 - 376." type="journal article" year="1875">Janson, 1875</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[309,523,1507,1528]" pageId="52" pageNumber="53" type="description">
<paragraph blockId="52.[251,580,1480,1528]" box="[309,523,1507,1528]" pageId="52" pageNumber="53">
(Appendix 4:
<figureCitation box="[446,516,1507,1528]" captionStart="Fig" captionStartId="141.[175,207,1662,1681]" captionTargetBox="[262,1120,195,1633]" captionTargetId="figure-0@141.[161,1163,183,1639]" captionTargetPageId="141" captionText="Fig. 22. Euphoria candezei. a) Habitus, b) Head, c) Parameres, d) Color variation, e) Temporal distribution, f) Geographic distribution in Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, and Panama; open square indicates state record only." figureDoi="http://doi.org/10.5281/zenodo.4907282" httpUri="https://zenodo.org/record/4907282/files/figure.png" pageId="52" pageNumber="53">Fig. 22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="53" type="reference_group">
<paragraph blockId="52.[173,659,1549,1730]" pageId="52" pageNumber="53">
<treatmentCitationGroup pageId="52" pageNumber="53">
<treatmentCitation author="Janson" box="[173,562,1549,1570]" page="375" pageId="52" pageNumber="53" year="1875">
<taxonomicName authority="Janson 1875: 375" authorityName="Janson" authorityPageNumber="375" authorityYear="1875" box="[173,562,1549,1570]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="candezei">
<emphasis box="[173,366,1549,1570]" italics="true" pageId="52" pageNumber="53">Euphoria candezei</emphasis>
<bibRefCitation author="Janson" box="[376,562,1549,1570]" firstAuthor="Janson" page="375" pageId="52" pageNumber="53" pagination="373 - 376" refId="ref123384" refString="Janson, O. E. 1875. Descriptions of three new species of American Cetoniidae. Cistula Entomologica 1: 373 - 376." type="journal article" year="1875">Janson 1875: 375</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Original combination.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="53" type="materials_examined">
<paragraph blockId="52.[173,659,1549,1730]" pageId="52" pageNumber="53">
<typeStatus box="[333,428,1576,1597]" pageId="52" pageNumber="53">Lectotype</typeStatus>
at RMNH, here designated. One
<typeStatus box="[311,438,1602,1623]" pageId="52" pageNumber="53">paralectotype</typeStatus>
at RMNH, examined.
</paragraph>
</subSubSection>
<subSubSection box="[173,591,1629,1650]" pageId="52" pageNumber="53" type="reference_group">
<paragraph blockId="52.[173,659,1549,1730]" box="[173,591,1629,1650]" pageId="52" pageNumber="53">
<treatmentCitationGroup box="[173,591,1629,1650]" pageId="52" pageNumber="53">
<treatmentCitation author="Sharp" box="[173,482,1629,1650]" page="137" pageId="52" pageNumber="53" year="1877">
<taxonomicName authority="Sharp 1877: 137" authorityName="Sharp" authorityPageNumber="137" authorityYear="1877" box="[173,482,1629,1650]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="belti">
<emphasis box="[173,312,1629,1650]" italics="true" pageId="52" pageNumber="53">Euphoria belti</emphasis>
<bibRefCitation author="Sharp" box="[319,482,1629,1650]" firstAuthor="Sharp" page="137" pageId="52" pageNumber="53" pagination="129 - 138" refId="ref125333" refString="Sharp, D. 1877. Description of some new species of beetles from Central America. Journal of the Linnean Society 13: 129 - 138." type="journal article" year="1877">Sharp 1877: 137</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Synonym.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="53" type="materials_examined">
<paragraph blockId="52.[173,659,1549,1730]" pageId="52" pageNumber="53">
<typeStatus pageId="52" pageNumber="53">Lectotype</typeStatus>
at BMNH designated by
<bibRefCitation author="Hardy" box="[514,654,1655,1676]" firstAuthor="Hardy" pageId="52" pageNumber="53" pagination="127 - 143" refId="ref122900" refString="Hardy, A. R. 2001. Studies in the Euphoriina of the Americas (Coleoptera: Scarabaeidae) II. Status of names in Euphoria, types and synonymies, with notes on the South American species. Pan- Pacific Entomologist 77: 127 - 143." type="journal article" year="2001">Hardy (2001)</bibRefCitation>
, examined. Two
<typeStatus box="[378,530,1682,1703]" pageId="52" pageNumber="53">paralectotypes</typeStatus>
at BMNH, not examined.
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="53" type="reference_group">
<paragraph blockId="52.[707,1193,194,348]" pageId="52" pageNumber="53">
<treatmentCitationGroup pageId="52" pageNumber="53">
<treatmentCitation author="Deloya, C. &amp; R. Woodruff" page="83" pageId="52" pageNumber="53" year="1995">
<taxonomicName authority="Deloya and Woodruff 1995: 83" authorityName="Deloya and Woodruff" authorityPageNumber="83" authorityYear="1995" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="vazquezae">
<emphasis box="[707,905,194,215]" italics="true" pageId="52" pageNumber="53">Euphoria vazquezae</emphasis>
<bibRefCitation author="Deloya, C. &amp; R. Woodruff" journalOrPublisher="Folia Entomologica Mexicana" pageId="52" pageNumber="53" pagination="81 - 86" part="93" refId="ref122352" refString="Deloya, C., and R. Woodruff. 1995. Descripcion de una nueva especie de Euphoria Burmeister de Honduras (Coleoptera: Melolonthidae, Cetoniinae). Folia Entomologica Mexicana 93: 81 - 86." title="Descripcion de una nueva especie de Euphoria Burmeister de Honduras (Coleoptera: Melolonthidae, Cetoniinae)" type="journal article" year="1995">Deloya and Woodruff 1995: 83</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<emphasis bold="true" box="[773,942,221,242]" pageId="52" pageNumber="53">New synonymy.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="53" type="materials_examined">
<paragraph blockId="52.[707,1193,194,348]" pageId="52" pageNumber="53">
<typeStatus box="[952,1047,221,242]" pageId="52" pageNumber="53">Holotype</typeStatus>
,
<typeStatus box="[1061,1144,221,242]" pageId="52" pageNumber="53">allotype</typeStatus>
, and
<specimenCount box="[732,855,247,268]" pageId="52" pageNumber="53" type="generic" typeStatus="paratype">one paratype</specimenCount>
at
<collectionCode box="[889,951,247,268]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="52" pageNumber="53" type="Museum">FSCA</collectionCode>
. One additional
<typeStatus box="[1112,1192,247,268]" pageId="52" pageNumber="53">paratype</typeStatus>
at
<collectionCode box="[756,822,274,295]" pageId="52" pageNumber="53">CCDX</collectionCode>
and another in
<collectorName box="[970,1192,274,295]" pageId="52" pageNumber="53">M.A. Morón Collection</collectorName>
(
<location LSID="urn:lsid:plazi:treatment:F449F723D516B25F85E245ADEB18FBD9:793F10EED516B25E87FA4149EB58FEC6" box="[739,812,300,321]" country="Mexico" county="Woodruff" municipality="Deloya" name="Xalapa" pageId="52" pageNumber="53" stateProvince="Veracruz de Ignacio de la Llave">Xalapa</location>
,
<collectingRegion box="[825,914,300,321]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="52" pageNumber="53">Veracruz</collectingRegion>
,
<collectingCountry box="[928,1004,300,321]" name="Mexico" pageId="52" pageNumber="53">Mexico</collectingCountry>
) (
<emphasis box="[1032,1068,301,322]" italics="true" pageId="52" pageNumber="53">fide</emphasis>
<bibRefCitation author="Deloya, C. &amp; R. Woodruff" journalOrPublisher="Folia Entomologica Mexicana" pageId="52" pageNumber="53" pagination="81 - 86" part="93" refId="ref122352" refString="Deloya, C., and R. Woodruff. 1995. Descripcion de una nueva especie de Euphoria Burmeister de Honduras (Coleoptera: Melolonthidae, Cetoniinae). Folia Entomologica Mexicana 93: 81 - 86." title="Descripcion de una nueva especie de Euphoria Burmeister de Honduras (Coleoptera: Melolonthidae, Cetoniinae)" type="journal article" year="1995">
<collectingMunicipality box="[1078,1148,300,321]" pageId="52" pageNumber="53">Deloya</collectingMunicipality>
and
<collectingCounty box="[732,822,327,348]" pageId="52" pageNumber="53">Woodruff</collectingCounty>
1995
</bibRefCitation>
).
<typeStatus box="[898,979,327,348]" pageId="52" pageNumber="53">Paratype</typeStatus>
at
<collectionCode box="[1011,1072,327,348]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="52" pageNumber="53" type="Museum">FSCA</collectionCode>
, examined.
</paragraph>
</subSubSection>
<subSubSection pageId="52" pageNumber="53" type="description">
<paragraph blockId="52.[707,1194,380,1729]" pageId="52" pageNumber="53">
<emphasis bold="true" box="[731,963,380,401]" pageId="52" pageNumber="53">
Description (
<emphasis bold="true" box="[865,878,380,401]" italics="true" pageId="52" pageNumber="53">n</emphasis>
= 310).
</emphasis>
Length
<quantity box="[1049,1189,380,402]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" metricValueMax="2.15" metricValueMin="1.71" pageId="52" pageNumber="53" unit="mm" value="19.3" valueMax="21.5" valueMin="17.1">17.121.5 mm</quantity>
; width
<quantity box="[767,895,406,428]" metricMagnitude="-2" metricUnit="m" metricValue="1.105" metricValueMax="1.22" metricValueMin="0.99" pageId="52" pageNumber="53" unit="mm" value="11.05" valueMax="12.2" valueMin="9.9">9.912.2 mm</quantity>
.
<emphasis bold="true" box="[906,963,407,428]" pageId="52" pageNumber="53">Color</emphasis>
: Dorsal and ventral surfaces shiny, bright to dark green, rarely with violaceous or red areas.
<emphasis bold="true" box="[914,967,460,481]" pageId="52" pageNumber="53">Head</emphasis>
: Frons depressed, with 1 central or 2 longitudinal depressions separated by ridge, densely punctate; punctures small to moderate, round, deeply impressed, at times coalescent; sparsely to moderately setose; setae short to moderate, yellowish. Clypeal surface moderately to densely punctate; punctures moderate in size, round; clypeus short, lateral margins strongly raised, sides moderately convergent at apex, lateral declivity moderately developed; apex strongly reflexed, moderately to strongly sinuate in frontal view, frequently forming 2 small denticles, glabrous to moderately densely setose; setae yellowish to whitish, short to moderate.
<emphasis bold="true" box="[810,912,805,826]" pageId="52" pageNumber="53">Pronotum</emphasis>
: Surface sparsely to moderately densely punctate; punctures minute to moderate, round to lunulate, denser and larger towards sides; glabrous. Lateral pronotal margins moderately to weakly angulate, females frequently with margins subparallel at base, males generally with margins convergent at base. Base in front of scutellum moderately to strongly emarginate. Scutellum longer than wide, impunctate.
<emphasis bold="true" box="[933,996,1018,1039]" pageId="52" pageNumber="53">Elytra</emphasis>
: Surface sparsely to moderately densely punctate; punctures round to lunulate, minute to moderate, larger towards apex and sides; costae obsolete to subobsolete. Posterior half of sutural costa strongly raised in lateral view. Surface glabrous. Apex occasionally with short sutural spine.
<emphasis bold="true" box="[841,936,1177,1198]" pageId="52" pageNumber="53">Pygidium</emphasis>
: Surface subconcentrically striate, striae discontinuous, moderately to strongly impressed, sparsely to moderately densely setose; setae short, yellowish.
<emphasis bold="true" box="[938,986,1257,1278]" pageId="52" pageNumber="53">Legs</emphasis>
: Protibial teeth well developed in both sexes, apical and medial teeth closer to each other than to basal tooth, basal tooth variably developed. Meso- and metatibial carinae moderately to well developed. Metatarsi compressed in females, much shorter than metatibiae, slightly shorter than metatibiae in males.
<emphasis bold="true" box="[1045,1115,1416,1437]" pageId="52" pageNumber="53">Venter</emphasis>
: Mesometasternal process extended anteriorly well beyond mesocoxae, glabrous on mesosternal lobe, apex variably rounded. Mesepimera, metasternum, and metacoxae setose, setae as on legs. Metasternum rugose, setose laterally, glabrous and impunctate at middle. Median sulcus well impressed. Abdominal sternites moderately densely setose laterally, glabrous to sparsely setose medially, sides rounded.
<emphasis bold="true" box="[707,848,1655,1676]" pageId="52" pageNumber="53">Male genitalia</emphasis>
: Parameres as in
<figureCitation box="[1017,1098,1655,1676]" captionStart="Fig" captionStartId="141.[175,207,1662,1681]" captionTargetBox="[262,1120,195,1633]" captionTargetId="figure-0@141.[161,1163,183,1639]" captionTargetPageId="141" captionText="Fig. 22. Euphoria candezei. a) Habitus, b) Head, c) Parameres, d) Color variation, e) Temporal distribution, f) Geographic distribution in Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, and Panama; open square indicates state record only." figureDoi="http://doi.org/10.5281/zenodo.4907282" httpUri="https://zenodo.org/record/4907282/files/figure.png" pageId="52" pageNumber="53">Fig. 22c</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="diagnosis">
<paragraph blockId="52.[707,1194,380,1729]" lastBlockId="53.[151,639,194,1730]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<emphasis bold="true" box="[731,836,1682,1703]" pageId="52" pageNumber="53">Diagnosis.</emphasis>
<taxonomicName box="[849,1038,1682,1703]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="candezei">
<emphasis box="[849,1038,1682,1703]" italics="true" pageId="52" pageNumber="53">Euphoria candezei</emphasis>
</taxonomicName>
is easily separated from other species in the group based on the large and shiny green body without markings, vertex without a protuberance, and form of the parameres. Dark purple specimens (
<figureCitation box="[546,630,247,268]" captionStart="Fig" captionStartId="141.[175,207,1662,1681]" captionTargetBox="[262,1120,195,1633]" captionTargetId="figure-0@141.[161,1163,183,1639]" captionTargetPageId="141" captionText="Fig. 22. Euphoria candezei. a) Habitus, b) Head, c) Parameres, d) Color variation, e) Temporal distribution, f) Geographic distribution in Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, and Panama; open square indicates state record only." figureDoi="http://doi.org/10.5281/zenodo.4907282" httpUri="https://zenodo.org/record/4907282/files/figure.png" pageId="53" pageNumber="54">Fig. 22d</figureCitation>
) show a green reflection when examined under a strong light.
</paragraph>
</subSubSection>
<subSubSection pageId="53" pageNumber="54" type="discussion">
<paragraph blockId="53.[151,639,194,1730]" pageId="53" pageNumber="54">
<emphasis bold="true" box="[175,374,327,348]" pageId="53" pageNumber="54">Taxonomic History.</emphasis>
<bibRefCitation author="Janson" box="[387,531,327,348]" firstAuthor="Janson" pageId="53" pageNumber="54" pagination="373 - 376" refId="ref123384" refString="Janson, O. E. 1875. Descriptions of three new species of American Cetoniidae. Cistula Entomologica 1: 373 - 376." type="journal article" year="1875">Janson (1875)</bibRefCitation>
described this species from specimens from
<collectingCountry box="[491,594,354,375]" name="Guatemala" pageId="53" pageNumber="54">Guatemala</collectingCountry>
and
<collectingCountry box="[151,251,380,401]" name="Nicaragua" pageId="53" pageNumber="54">Nicaragua</collectingCountry>
. The species was named after Dr. Ernest Candéze, who collected the first specimen.
<bibRefCitation author="Sharp" firstAuthor="Sharp" pageId="53" pageNumber="54" pagination="129 - 138" refId="ref125333" refString="Sharp, D. 1877. Description of some new species of beetles from Central America. Journal of the Linnean Society 13: 129 - 138." type="journal article" year="1877">Sharp (1877)</bibRefCitation>
described
<taxonomicName authorityName="Sharp" authorityYear="1877" box="[327,472,433,454]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="belti">
<emphasis box="[327,472,433,454]" italics="true" pageId="53" pageNumber="54">Euphoria belti</emphasis>
</taxonomicName>
from specimens collected by Mr. Thomas Belt in
<collectingCountry box="[474,574,460,481]" name="Nicaragua" pageId="53" pageNumber="54">Nicaragua</collectingCountry>
.
<bibRefCitation author="Bates, W. D." editor="F. D. Godman &amp; O. Salvin" journalOrPublisher="London, UK" pageId="53" pageNumber="54" pagination="362 - 374" refId="ref121351" refString="Bates, W. D. 1889. Insecta. Coleoptera. Pectinicornia and Lamellicornia, [pp. 362 - 374]. In: Biologia Centrali-Americana. Vol II, Part 2 (F. D. Godman and O. Salvin, editors). London, UK." title="Insecta. Coleoptera. Pectinicornia and Lamellicornia" type="book chapter" volumeTitle="Biologia Centrali-Americana. Vol II, Part 2" year="1889">Bates (1889)</bibRefCitation>
treated
<taxonomicName authorityName="Sharp" authorityYear="1877" box="[289,356,486,508]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="belti">
<emphasis box="[289,303,487,508]" italics="true" pageId="53" pageNumber="54">E</emphasis>
.
<emphasis box="[315,356,487,508]" italics="true" pageId="53" pageNumber="54">belti</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName box="[524,632,486,508]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="candezei">
<emphasis box="[524,538,487,508]" italics="true" pageId="53" pageNumber="54">E</emphasis>
.
<emphasis box="[551,632,487,508]" italics="true" pageId="53" pageNumber="54">candezei</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="53.[151,639,194,1730]" pageId="53" pageNumber="54">
<bibRefCitation author="Deloya, C. &amp; R. Woodruff" box="[175,430,513,534]" journalOrPublisher="Folia Entomologica Mexicana" pageId="53" pageNumber="54" pagination="81 - 86" part="93" refId="ref122352" refString="Deloya, C., and R. Woodruff. 1995. Descripcion de una nueva especie de Euphoria Burmeister de Honduras (Coleoptera: Melolonthidae, Cetoniinae). Folia Entomologica Mexicana 93: 81 - 86." title="Descripcion de una nueva especie de Euphoria Burmeister de Honduras (Coleoptera: Melolonthidae, Cetoniinae)" type="journal article" year="1995">Deloya and Woodruff (1995)</bibRefCitation>
described
<taxonomicName authorityName="Deloya and Woodruff" authorityYear="1995" box="[522,637,513,534]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="vazquezae">
<emphasis box="[522,536,513,534]" italics="true" pageId="53" pageNumber="54">E</emphasis>
.
<emphasis box="[545,637,513,534]" italics="true" pageId="53" pageNumber="54">vazquezae</emphasis>
</taxonomicName>
from
<specimenCount box="[208,360,540,561]" pageId="53" pageNumber="54" type="generic">five specimens</specimenCount>
collected on Cerro Uyuca,
<collectingCountry box="[151,256,566,587]" name="Honduras" pageId="53" pageNumber="54">Honduras</collectingCountry>
. The characters listed to separate
<taxonomicName authorityName="Deloya and Woodruff" authorityYear="1995" box="[151,280,593,614]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="vazquezae">
<emphasis box="[151,165,593,614]" italics="true" pageId="53" pageNumber="54">E</emphasis>
.
<emphasis box="[181,280,593,614]" italics="true" pageId="53" pageNumber="54">vazquezae</emphasis>
</taxonomicName>
from
<taxonomicName box="[346,460,593,614]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="candezei">
<emphasis box="[346,360,593,614]" italics="true" pageId="53" pageNumber="54">E</emphasis>
.
<emphasis box="[375,460,593,614]" italics="true" pageId="53" pageNumber="54">candezei</emphasis>
</taxonomicName>
(color reflections of the dorsum, development of the basal protibial tooth, development of teeth on mesotibial carina, and angle of the parameres apex) all vary intraspecifically. Based on the study of the intraspecific variation of
<specimenCount box="[267,408,725,746]" pageId="53" pageNumber="54" type="generic">310 specimens</specimenCount>
, I place
<taxonomicName authorityName="Deloya and Woodruff" authorityYear="1995" box="[489,613,725,747]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="vazquezae">
<emphasis box="[489,503,726,747]" italics="true" pageId="53" pageNumber="54">E</emphasis>
.
<emphasis box="[517,613,726,747]" italics="true" pageId="53" pageNumber="54">vazquezae</emphasis>
</taxonomicName>
in synonymy with
<taxonomicName box="[303,410,752,773]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="candezei">
<emphasis box="[303,317,752,773]" italics="true" pageId="53" pageNumber="54">E</emphasis>
.
<emphasis box="[329,410,752,773]" italics="true" pageId="53" pageNumber="54">candezei</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="53.[151,639,194,1730]" pageId="53" pageNumber="54">
<emphasis bold="true" box="[175,349,779,800]" pageId="53" pageNumber="54">Natural History.</emphasis>
Adults have been found between
<quantity box="[239,355,805,827]" metricMagnitude="3" metricUnit="m" metricValue="1.0455" metricValueMax="2.0" metricValueMin="0.091" pageId="53" pageNumber="54" unit="m" value="1045.5" valueMax="2000.0" valueMin="91.0">912,000 m</quantity>
elevation in mesophytic and cloud forests. Adult
<taxonomicName box="[341,448,832,853]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="candezei">
<emphasis box="[341,448,832,853]" italics="true" pageId="53" pageNumber="54">E. candezei</emphasis>
</taxonomicName>
have been observed swarming at ground level over a newly cut area or digging in the soil. This behavior suggests copula probably takes place in the soil.
<bibRefCitation author="Solis" box="[507,637,911,932]" firstAuthor="Solis" pageId="53" pageNumber="54" refId="ref125513" refString="Solis, A. 2004. Escarabajos Fruteros de Costa Rica. INBio, Santo Domingo Heredia, Costa Rica." type="book" year="2004">Solís (2004)</bibRefCitation>
observed the species feeding on
<taxonomicName box="[455,549,938,959]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="53" pageNumber="54" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[455,513,938,959]" italics="true" pageId="53" pageNumber="54">Rubus</emphasis>
sp.
</taxonomicName>
, mangos, banana, and fermented honey in
<collectingCountry box="[485,597,964,985]" name="Costa Rica" pageId="53" pageNumber="54">Costa Rica</collectingCountry>
. He also observed adults in large numbers perching on trees at the forest edge.
<bibRefCitation author="Moron, M. A. &amp; B. C. Ratcliffe &amp; C. Deloya." box="[369,551,1018,1039]" journalOrPublisher="CONABIO and Sociedad Mexicana de Entomologia, Mexico" pageId="53" pageNumber="54" refId="ref124137" refString="Moron, M. A., B. C. Ratcliffe, and C. Deloya. 1997. Atlas de los Escarabajos de Mexico (Coleoptera: Lamellicornia). Vol. 1. Familia Melolonthidae. CONABIO and Sociedad Mexicana de Entomologia, Mexico." title="Atlas de los Escarabajos de Mexico (Coleoptera: Lamellicornia). Vol. 1. Familia Melolonthidae" type="book" year="1997">
Morón
<emphasis box="[438,478,1018,1039]" italics="true" pageId="53" pageNumber="54">et al</emphasis>
. (1997)
</bibRefCitation>
recorded the species from a variety of habitats in
<collectingCountry box="[558,634,1044,1065]" name="Mexico" pageId="53" pageNumber="54">Mexico</collectingCountry>
, from undisturbed to highly disturbed forests.
</paragraph>
</subSubSection>
<subSubSection pageId="53" pageNumber="54" type="distribution">
<paragraph blockId="53.[151,639,194,1730]" pageId="53" pageNumber="54">
<emphasis bold="true" box="[175,409,1097,1118]" pageId="53" pageNumber="54">Temporal Distribution.</emphasis>
January (2), February (2), March (1), April (2), May (78), June (29), July (23), August (25), September (25), October (23), November (46), December (4) (
<figureCitation box="[514,597,1177,1198]" captionStart="Fig" captionStartId="141.[175,207,1662,1681]" captionTargetBox="[262,1120,195,1633]" captionTargetId="figure-0@141.[161,1163,183,1639]" captionTargetPageId="141" captionText="Fig. 22. Euphoria candezei. a) Habitus, b) Head, c) Parameres, d) Color variation, e) Temporal distribution, f) Geographic distribution in Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, and Panama; open square indicates state record only." figureDoi="http://doi.org/10.5281/zenodo.4907282" httpUri="https://zenodo.org/record/4907282/files/figure.png" pageId="53" pageNumber="54">Fig. 22e</figureCitation>
).
</paragraph>
<paragraph blockId="53.[151,639,194,1730]" pageId="53" pageNumber="54">
<emphasis bold="true" box="[175,432,1204,1225]" pageId="53" pageNumber="54">Geographic Distribution.</emphasis>
Known from southern
<collectingCountry box="[189,261,1230,1251]" name="Mexico" pageId="53" pageNumber="54">Mexico</collectingCountry>
to northern
<collectingCountry box="[384,459,1230,1251]" name="Panama" pageId="53" pageNumber="54">Panama</collectingCountry>
(
<figureCitation box="[474,553,1230,1251]" captionStart="Fig" captionStartId="141.[175,207,1662,1681]" captionTargetBox="[262,1120,195,1633]" captionTargetId="figure-0@141.[161,1163,183,1639]" captionTargetPageId="141" captionText="Fig. 22. Euphoria candezei. a) Habitus, b) Head, c) Parameres, d) Color variation, e) Temporal distribution, f) Geographic distribution in Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, and Panama; open square indicates state record only." figureDoi="http://doi.org/10.5281/zenodo.4907282" httpUri="https://zenodo.org/record/4907282/files/figure.png" pageId="53" pageNumber="54">Fig. 22f</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="53" pageNumber="54" type="materials_examined">
<paragraph blockId="53.[151,639,194,1730]" lastBlockId="53.[685,1173,194,1118]" pageId="53" pageNumber="54">
<emphasis bold="true" box="[175,464,1257,1278]" pageId="53" pageNumber="54">Specimens Examined (310).</emphasis>
Type material:
<materialsCitation collectionCode="RMNH" collectorName="O. E. Janson &amp; O. Jans." county="Type." location="Chont." municipality="Lucassen" pageId="53" pageNumber="54" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<taxonomicName authority="Janson, 1875" authorityName="Janson" authorityYear="1875" box="[151,464,1283,1305]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="candezei">
<emphasis box="[151,329,1284,1305]" italics="true" pageId="53" pageNumber="54">Euphoria candezei</emphasis>
<bibRefCitation author="Janson" box="[337,464,1283,1304]" firstAuthor="Janson" pageId="53" pageNumber="54" pagination="373 - 376" refId="ref123384" refString="Janson, O. E. 1875. Descriptions of three new species of American Cetoniidae. Cistula Entomologica 1: 373 - 376." type="journal article" year="1875">Janson, 1875</bibRefCitation>
</taxonomicName>
;
<typeStatus box="[475,560,1283,1304]" pageId="53" pageNumber="54">lectotype</typeStatus>
male at
<collectionCode box="[151,222,1310,1331]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="53" pageNumber="54" type="Museum">RMNH</collectionCode>
<emphasis bold="true" box="[229,385,1310,1331]" pageId="53" pageNumber="54">here designated</emphasis>
labeled “
<location LSID="urn:lsid:plazi:treatment:F449F723D516B25F85E245ADEB18FBD9:793F10EED517B25F84C5457BEA6FFAB4" box="[476,539,1310,1331]" county="Type." municipality="Lucassen" name="Chont." pageId="53" pageNumber="54">Chont.</location>
/
<location LSID="urn:lsid:plazi:treatment:F449F723D516B25F85E245ADEB18FBD9:793F10EED517B25F8731457BEA04FAB4" box="[552,624,1310,1331]" county="Type." municipality="Lucassen" name="Nicarag" pageId="53" pageNumber="54">Nicarag</location>
//
<location LSID="urn:lsid:plazi:treatment:F449F723D516B25F85E245ADEB18FBD9:793F10EED517B25F8581455CE946FAC9" box="[152,306,1337,1358]" county="Type." municipality="Lucassen" name="Museum Leiden" pageId="53" pageNumber="54">Museum Leiden</location>
/ verz. F.
<collectionCode box="[392,407,1337,1358]" name="Tavera, Department of Geology and Geophysics" pageId="53" pageNumber="54">T</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:F449F723D516B25F85E245ADEB18FBD9:793F10EED517B25F84BB455CE9A3FAC9" box="[418,471,1337,1358]" county="Type." municipality="Lucassen" name="Valck" pageId="53" pageNumber="54">Valck</location>
/
<collectingMunicipality box="[486,572,1337,1358]" pageId="53" pageNumber="54">Lucassen</collectingMunicipality>
/ (
<collectorName pageId="53" pageNumber="54">O.E. Janson</collectorName>
)// type//
<specimenCount box="[309,327,1362,1384]" pageId="53" pageNumber="54" type="male"></specimenCount>
/
<collectionCode box="[343,401,1363,1384]" pageId="53" pageNumber="54">TYPE</collectionCode>
//
<taxonomicName authority=", O. Jns" authorityName="O. Jns" class="Insecta" family="Cetoniidae" genus="Erirhipis" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="candezei">Erirhipis candezei, O. Jns</taxonomicName>
/
<collectingCounty box="[194,246,1390,1411]" pageId="53" pageNumber="54">Type.</collectingCounty>
// type// candezei/
<collectorName box="[417,493,1390,1411]" pageId="53" pageNumber="54">O. Jans.</collectorName>
” and my
<typeStatus pageId="53" pageNumber="54">lectotype</typeStatus>
label;
</materialsCitation>
<materialsCitation collectionCode="RMNH" collectorName="O. E. Janson" county="Lucassen" location="Chontal" municipality="Valck" pageId="53" pageNumber="54" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<typeStatus box="[260,385,1416,1437]" pageId="53" pageNumber="54">paralectotype</typeStatus>
female at
<collectionCode box="[491,562,1416,1437]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="53" pageNumber="54" type="Museum">RMNH</collectionCode>
labeled “
<location LSID="urn:lsid:plazi:treatment:F449F723D516B25F85E245ADEB18FBD9:793F10EED517B25F85BB45C6E89BFA3F" box="[162,239,1443,1464]" county="Lucassen" municipality="Valck" name="Chontal" pageId="53" pageNumber="54">Chontal</location>
/ Nicarag.// Museum Leiden/ verz. F.
<collectionCode box="[618,634,1443,1464]" name="Tavera, Department of Geology and Geophysics" pageId="53" pageNumber="54">T</collectionCode>
.
<collectingMunicipality box="[151,204,1469,1490]" pageId="53" pageNumber="54">Valck</collectingMunicipality>
/
<collectingCounty box="[220,306,1469,1490]" pageId="53" pageNumber="54">Lucassen</collectingCounty>
/ (
<collectorName box="[328,443,1469,1490]" pageId="53" pageNumber="54">O.E. Janson</collectorName>
)// type//
<specimenCount box="[534,552,1469,1491]" pageId="53" pageNumber="54" type="female"></specimenCount>
/
<collectionCode box="[566,624,1469,1490]" pageId="53" pageNumber="54">TYPE</collectionCode>
// type// type” and my
<typeStatus box="[355,481,1496,1517]" pageId="53" pageNumber="54">paralectotype</typeStatus>
label.
</materialsCitation>
<materialsCitation collectionCode="BMNH" collectorName="B. C. A. Col." county="Hardy" location="Sharp" municipality="By" pageId="53" pageNumber="54" specimenCount="2" typeStatus="lectotype">
<taxonomicName authority="Sharp, 1877" authorityName="Sharp" authorityYear="1877" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="belti">
<emphasis italics="true" pageId="53" pageNumber="54">Euphoria belti</emphasis>
<bibRefCitation author="Sharp" box="[210,346,1522,1543]" firstAuthor="Sharp" pageId="53" pageNumber="54" pagination="129 - 138" refId="ref125333" refString="Sharp, D. 1877. Description of some new species of beetles from Central America. Journal of the Linnean Society 13: 129 - 138." type="journal article" year="1877">Sharp, 1877</bibRefCitation>
</taxonomicName>
;
<typeStatus box="[361,459,1522,1543]" pageId="53" pageNumber="54">lectotype</typeStatus>
male at
<collectionCode box="[562,638,1522,1543]" pageId="53" pageNumber="54">BMNH</collectionCode>
labeled “LECTO-/
<collectionCode box="[326,384,1549,1570]" pageId="53" pageNumber="54">TYPE</collectionCode>
/ SYN-/
<collectionCode box="[458,516,1549,1570]" pageId="53" pageNumber="54">TYPE</collectionCode>
// Chontales/ Nicaragua/
<collectionCode box="[271,287,1576,1597]" name="Tavera, Department of Geology and Geophysics" pageId="53" pageNumber="54">T</collectionCode>
. Belt.//
<taxonomicName box="[375,465,1576,1597]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">Euphoria</taxonomicName>
/belti/ Ind. Typ./ D.S.//
<collectorName box="[221,346,1602,1623]" pageId="53" pageNumber="54">B.C.A. Col.</collectorName>
, II(2)./
<taxonomicName box="[431,520,1602,1623]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">Euphoria</taxonomicName>
/ candezei//
<typeStatus box="[151,288,1629,1650]" pageId="53" pageNumber="54">LECTOTYPE</typeStatus>
/
<taxonomicName box="[303,391,1629,1650]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">Euphoria</taxonomicName>
/ belti/
<location LSID="urn:lsid:plazi:treatment:F449F723D516B25F85E245ADEB18FBD9:793F10EED517B25F84C94638EA7CF9F5" box="[464,520,1629,1650]" county="Hardy" municipality="By" name="Sharp" pageId="53" pageNumber="54">Sharp</location>
=
<collectingMunicipality box="[549,576,1629,1650]" pageId="53" pageNumber="54">By</collectingMunicipality>
/ A.
<collectionCode box="[615,634,1629,1650]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="53" pageNumber="54">R</collectionCode>
.
<collectingCounty box="[151,217,1655,1676]" pageId="53" pageNumber="54">Hardy</collectingCounty>
, 78”.
</materialsCitation>
<materialsCitation collectionCode="FSCA" country="Honduras" county="Hardy" location="Sharp" municipality="By" pageId="53" pageNumber="54" specimenCount="2" typeStatus="lectotype">
<taxonomicName authority="Deloya and Woodruff, 1995" authorityName="Deloya and Woodruff" authorityYear="1995" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="vazquezae">
<emphasis box="[288,503,1655,1676]" italics="true" pageId="53" pageNumber="54">Euphoria vazquezae</emphasis>
<bibRefCitation author="Deloya, C. &amp; R. Woodruff" journalOrPublisher="Folia Entomologica Mexicana" pageId="53" pageNumber="54" pagination="81 - 86" part="93" refId="ref122352" refString="Deloya, C., and R. Woodruff. 1995. Descripcion de una nueva especie de Euphoria Burmeister de Honduras (Coleoptera: Melolonthidae, Cetoniinae). Folia Entomologica Mexicana 93: 81 - 86." title="Descripcion de una nueva especie de Euphoria Burmeister de Honduras (Coleoptera: Melolonthidae, Cetoniinae)" type="journal article" year="1995">Deloya and Woodruff, 1995</bibRefCitation>
</taxonomicName>
;
<typeStatus box="[324,407,1682,1703]" pageId="53" pageNumber="54">paratype</typeStatus>
male at
<collectionCode box="[499,559,1682,1703]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="53" pageNumber="54" type="Museum">FSCA</collectionCode>
labeled “
<collectingCountry box="[162,299,1708,1729]" name="Honduras" pageId="53" pageNumber="54">HONDURAS</collectingCountry>
:/
<location LSID="urn:lsid:plazi:treatment:F449F723D516B25F85E245ADEB18FBD9:793F10EED517B25F842646C9E9AFF946" box="[319,475,1708,1729]" country="Honduras" municipality="Morazon" name="Dept. Francisco" pageId="53" pageNumber="54">Dept. Francisco</location>
/
<collectingMunicipality box="[492,579,1708,1729]" pageId="53" pageNumber="54">Morazon</collectingMunicipality>
[
<emphasis box="[597,624,1709,1730]" italics="true" pageId="53" pageNumber="54">sic</emphasis>
], Cerro Uyuca/
<quantity box="[840,922,194,215]" metricMagnitude="3" metricUnit="m" metricValue="1.9812" pageId="53" pageNumber="54" unit="ft" value="6500.0">6,500ft.</quantity>
,
<date box="[938,1042,194,215]" pageId="53" pageNumber="54" value="1993-06-06">6-VI-1993</date>
/ coll. M.C. Thomas//
<taxonomicName box="[785,903,221,242]" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">EUPHORIA</taxonomicName>
/ VAZQUEZAE/ DELOYA Y WOODRUFF/ A. C. Deloya, det.//
<taxonomicName authorityName="Deloya and Woodruff" authorityYear="1995" class="Insecta" family="Cetoniidae" genus="Euphoria" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="vazquezae">Euphoria vazquezae</taxonomicName>
/ PARATIPO/ Deloya &amp; Woodruff/ 1995”.
</materialsCitation>
Other material:
<emphasis bold="true" box="[945,1108,300,321]" pageId="53" pageNumber="54">
<collectingCountry box="[945,1108,300,321]" name="Costa Rica" pageId="53" pageNumber="54">COSTA RICA</collectingCountry>
</emphasis>
(74):
<collectingRegion box="[685,814,327,348]" country="Costa Rica" name="Alajuela" pageId="53" pageNumber="54">ALAJUELA</collectingRegion>
: Cariblanco (1), Parque Nacional Peñas Blancas (4) Upala (2);
<collectingRegion box="[973,1087,354,375]" country="Costa Rica" name="Cartago" pageId="53" pageNumber="54">CARTAGO</collectingRegion>
: Cartago (5), Orosí (2), Navarro (1), Pavones (1), Río Macho (4), Tres Ríos (3), Turrialba (2), Volcán Irazú (4);
<collectingRegion box="[685,852,433,454]" country="Costa Rica" name="Guanacaste" pageId="53" pageNumber="54">GUANACASTE</collectingRegion>
: Bebedero (1), Parque Nacional Guanacaste (5), La Palma (1);
<collectingRegion box="[980,1086,460,481]" country="Costa Rica" name="Heredia" pageId="53" pageNumber="54">HEREDIA</collectingRegion>
: La Caja (1);
<collectingRegion box="[725,835,486,507]" country="Costa Rica" name="SAN JOSE" pageId="53" pageNumber="54">SAN JOSÉ</collectingRegion>
: Coronado (5), San José (18), Tibas (2);
<collectingRegion box="[729,891,513,534]" country="Costa Rica" name="Puntarenas" pageId="53" pageNumber="54">PUNTARENAS</collectingRegion>
: San Vito (3), Surubres (2);
<collectingRegion box="[685,847,540,561]" country="Costa Rica" name="SAN RAFAEL" pageId="53" pageNumber="54">SAN RAFAEL</collectingRegion>
: Parque Nacional La Amistad (1);
<taxonomicName box="[726,833,566,587]" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="103" phylum="Chordata" rank="species" species="data">NO DATA</taxonomicName>
: “Costa Rica” (6).
<emphasis bold="true" box="[1015,1171,566,587]" pageId="53" pageNumber="54">
<collectingCountry box="[1015,1171,566,587]" name="Guatemala" pageId="53" pageNumber="54">GUATEMALA</collectingCountry>
</emphasis>
(5):
<collectingRegion box="[731,917,593,614]" country="Guatemala" name="Baja Verapaz" pageId="53" pageNumber="54">BAJA VERAPAZ</collectingRegion>
: Cobán (“Road to”) (2);
<collectingRegion box="[685,857,619,640]" country="Guatemala" name="Alta Verapaz" pageId="53" pageNumber="54">ALTA VERAPAZ</collectingRegion>
: Baleu (1), Cobán (1), Santa Cruz El Chol (1).
<emphasis bold="true" box="[818,959,646,667]" pageId="53" pageNumber="54">
<collectingCountry box="[818,959,646,667]" name="Honduras" pageId="53" pageNumber="54">HONDURAS</collectingCountry>
</emphasis>
(76):
<collectingRegion box="[1026,1169,646,667]" country="Honduras" name="Atlantida" pageId="53" pageNumber="54">ATLÁNTIDA</collectingRegion>
: La Ceiba (2);
<collectingRegion box="[833,995,672,693]" country="Honduras" name="Comayagua" pageId="53" pageNumber="54">COMAYAGUA</collectingRegion>
: Comayagua (2), Forestry School (3), Siguatepeque (21), Taulabé (1);
<collectingRegion box="[727,811,725,746]" country="Honduras" name="Copan" pageId="53" pageNumber="54">COPÁN</collectingRegion>
:
<taxonomicName box="[821,897,725,746]" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="data">No data</taxonomicName>
(1);
<collectingRegion box="[946,1039,725,746]" country="Honduras" name="Cortes" pageId="53" pageNumber="54">CORTÉS</collectingRegion>
: La Lima (1), Omoa (1), Río Lindo (1), Yojoa (10);
<collectingRegion box="[1033,1169,752,773]" country="Honduras" name="El Paraiso" pageId="53" pageNumber="54">EL PARAÍSO</collectingRegion>
: Yuscarán (1);
<collectingRegion box="[830,1103,779,800]" country="Honduras" name="Francisco Morazan" pageId="53" pageNumber="54">FRANCISCO MORAZÁN</collectingRegion>
: Cerro Uyuca (29), Valle de Ángeles (1);
<collectingRegion box="[1008,1078,805,826]" country="Honduras" name="Yoro" pageId="53" pageNumber="54">YORO</collectingRegion>
: Yoro (2).
<emphasis bold="true" box="[685,789,832,853]" pageId="53" pageNumber="54">
<collectingCountry box="[685,789,832,853]" name="Mexico" pageId="53" pageNumber="54">MEXICO</collectingCountry>
</emphasis>
(105):
<collectingRegion box="[871,980,832,853]" country="Mexico" name="Chiapas" pageId="53" pageNumber="54">CHIAPAS</collectingRegion>
: La Revancha (5), Ocozocoautla de Espinosa (12), Oxchuc (10), Parque Laguna Bélgica (1), Pueblo Nuevo (8), Rancho Santa
<taxonomicName box="[750,799,911,932]" class="Magnoliopsida" family="Rosaceae" genus="Rosa" kingdom="Plantae" order="Rosales" pageId="53" pageNumber="103" phylum="Tracheophyta" rank="genus">Rosa</taxonomicName>
(29), San Antonio Buenavista (30), San Cristóbal de las Casas (4), Santa
<taxonomicName box="[1078,1126,938,959]" class="Magnoliopsida" family="Rosaceae" genus="Rosa" kingdom="Plantae" order="Rosales" pageId="53" pageNumber="103" phylum="Tracheophyta" rank="genus">Rosa</taxonomicName>
(3),
<taxonomicName box="[685,764,964,985]" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="103" phylum="Chordata" rank="species" species="data">no data</taxonomicName>
(1);
<collectingRegion box="[825,1023,964,985]" country="Mexico" name="Quintana Roo" pageId="53" pageNumber="54">QUINTANA ROO</collectingRegion>
:
<taxonomicName box="[1039,1123,964,985]" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="data">No data</taxonomicName>
(1);
<collectingRegion box="[685,816,991,1012]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="53" pageNumber="54">VERACRUZ</collectingRegion>
: Catemaco (1).
<emphasis bold="true" box="[976,1128,991,1012]" pageId="53" pageNumber="54">
<collectingCountry box="[976,1128,991,1012]" name="Nicaragua" pageId="53" pageNumber="54">NICARAGUA</collectingCountry>
</emphasis>
(7):
<collectingRegion box="[685,824,1018,1039]" country="Nicaragua" name="Chontales" pageId="53" pageNumber="54">CHONTALES</collectingRegion>
: Chontales (1);
<collectingRegion box="[972,1087,1018,1039]" country="Nicaragua" name="Jinotega" pageId="53" pageNumber="54">JINOTEGA</collectingRegion>
: Jinotega (1);
<taxonomicName box="[729,834,1044,1065]" class="Mammalia" family="Muridae" genus="Neotoma" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="103" phylum="Chordata" rank="species" species="data">NO DATA</taxonomicName>
(5).
<emphasis bold="true" box="[888,994,1044,1065]" pageId="53" pageNumber="54">
<collectingCountry box="[888,994,1044,1065]" name="Panama" pageId="53" pageNumber="54">PANAMA</collectingCountry>
</emphasis>
(39):
<collectingRegion box="[1060,1170,1044,1065]" country="Panama" name="Chiriqui" pageId="53" pageNumber="54">CHIRIQUÍ</collectingRegion>
: Finca La Suiza (4), Los Planes (21), Valle Hornito (11), Potrerillos (3).
</paragraph>
</subSubSection>
</treatment>
</document>