treatments-xml/data/5F/06/51/5F0651448109FFB2FF1CFC7FFDD1F997.xml

<document id="BDD2650AF1E81F7AD8CCAE57620618BC" ID-DOI="10.11646/zootaxa.3945.1.1" ID-GBIF-Dataset="87f26c32-420d-4a40-8923-03b6cef48120" ID-ISSN="1175-5326" ID-Zenodo-Dep="288235" ID-ZooBank="2E17F87F-B07B-4394-A9C7-F288C456EAD4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461031140601" checkinUser="plazi" docAuthor="Totakura, Venkateswara Rao &amp; Reddy, Yenumula Ranga" docDate="2015" docId="5F0651448109FFB2FF1CFC7FFDD1F997" docLanguage="en" docName="zt03945p093.pdf" docOrigin="Zootaxa 3945 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Haplocyclops (Kiefercyclops) primitivus Totakura &amp; Reddy, 2015, n. sp." docType="treatment" docVersion="6" lastPageNumber="81" masterDocId="A33F293C814EFFE2FF8BFFB6FFA2FFC7" masterDocTitle="Groundwater cyclopoid copepods of peninsular India, with description of eight new species" masterLastPageNumber="93" masterPageNumber="1" pageNumber="72" updateTime="1698573267305" updateUser="plazi">
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<mods:title id="6E280181ED525FBC1B2CEC9ADF0D7265">Groundwater cyclopoid copepods of peninsular India, with description of eight new species</mods:title>
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<mods:namePart id="7D3339800D73AAB59EE5BE5E4C36650A">Totakura, Venkateswara Rao</mods:namePart>
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<mods:namePart id="55EDF42F7522070DFFF619CBB503475B">Reddy, Yenumula Ranga</mods:namePart>
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<treatment id="5F0651448109FFB2FF1CFC7FFDD1F997" ID-DOI="http://doi.org/10.5281/zenodo.6109720" ID-GBIF-Taxon="127690313" ID-Zenodo-Dep="6109720" LSID="urn:lsid:plazi:treatment:5F0651448109FFB2FF1CFC7FFDD1F997" httpUri="http://treatment.plazi.org/id/5F0651448109FFB2FF1CFC7FFDD1F997" lastPageId="80" lastPageNumber="81" pageId="71" pageNumber="72">
<subSubSection id="9FB5B3D98109FFA5FF1CFC7FFE8CFBC2" pageId="71" pageNumber="72" type="nomenclature">
<paragraph id="D710E0528109FFA5FF1CFC7FFD60FC24" blockId="71.[151,706,969,1029]" box="[151,706,969,995]" pageId="71" pageNumber="72">
<heading id="8C58573E8109FFA5FF1CFC7FFD60FC24" bold="true" box="[151,706,969,995]" fontSize="11" level="1" pageId="71" pageNumber="72" reason="1">
<emphasis id="E5DB3C408109FFA5FF1CFC7FFD60FC24" bold="true" box="[151,706,969,995]" pageId="71" pageNumber="72">
<taxonomicName id="10AF9BD18109FFA5FF1CFC7FFDDAFC24" box="[151,632,969,995]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="primitivus" status="sp. nov." subGenus="Kiefercyclops">
<emphasis id="E5DB3C408109FFA5FF1CFC7FFE98FC24" bold="true" box="[151,314,969,995]" italics="true" pageId="71" pageNumber="72">Haplocyclops</emphasis>
(
<emphasis id="E5DB3C408109FFA5FEC0FC7FFE4CFC24" bold="true" box="[331,494,969,995]" italics="true" pageId="71" pageNumber="72">Kiefercyclops</emphasis>
) 
<emphasis id="E5DB3C408109FFA5FE75FC7FFDDAFC24" bold="true" box="[510,632,969,995]" italics="true" pageId="71" pageNumber="72">primitivus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FEE8813B8109FFA5FD0BFC7FFD60FC24" box="[640,706,969,995]" pageId="71" pageNumber="72" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D710E0528109FFA5FF1CFC5AFE8CFBC2" blockId="71.[151,706,969,1029]" box="[151,302,1004,1029]" pageId="71" pageNumber="72">
(
<figureCitation id="4F94FCD78109FFA5FF14FC5AFE84FBC2" box="[159,294,1004,1029]" captionStart-0="FIGURE 45" captionStart-1="FIGURE 46" captionStart-2="FIGURE 47" captionStart-3="FIGURE 48" captionStart-4="FIGURE 49" captionStart-5="FIGURE 50" captionStartId-0="73.[151,250,1953,1975]" captionStartId-1="74.[151,250,1953,1975]" captionStartId-2="75.[151,250,1953,1975]" captionStartId-3="76.[151,250,1953,1975]" captionStartId-4="77.[151,250,1953,1975]" captionStartId-5="78.[151,250,1953,1975]" captionTargetBox-0="[216,1393,209,1914]" captionTargetBox-1="[174,1403,207,1926]" captionTargetBox-2="[192,1389,207,1915]" captionTargetBox-3="[182,1394,210,1921]" captionTargetBox-4="[213,1378,214,1927]" captionTargetBox-5="[197,1409,203,1905]" captionTargetId-0="figure@73.[178,1409,193,1932]" captionTargetId-1="figure@74.[163,1423,193,1932]" captionTargetId-2="figure@75.[178,1409,193,1932]" captionTargetId-3="figure@76.[178,1409,193,1932]" captionTargetId-4="figure@77.[178,1409,193,1932]" captionTargetId-5="figure@78.[178,1409,193,1932]" captionTargetPageId-0="73" captionTargetPageId-1="74" captionTargetPageId-2="75" captionTargetPageId-3="76" captionTargetPageId-4="77" captionTargetPageId-5="78" captionText-0="FIGURE 45. Haplocyclops primitivus n. sp. (a) paratype female, habitus, dorsal; (b) holotype female, antennule, lateral." captionText-1="FIGURE 46. Haplocyclops primitivus n. sp., holotype female (a), paratype female (b – e). (a) urosome, dorsal; (b) same, ventral; (c) same, lateral; (d) leg 5, latero-ventral; (e) leg 6, latero-ventral." captionText-2="FIGURE 47. Haplocyclops primitivus n. sp., holotype female. (a) rostrum, ventral; (b) antenna, lateral; (c) labrum, ventral; (d) mandible, lateral; (e) maxillule, lateral; (f) maxilla, lateral; (g) maxilliped, lateral." captionText-3="FIGURE 48. Haplocyclops primitivus n. sp., holotype female (a – d), paratype female (e), allotype male (f). (a) leg 1, anterior; (b) leg 2, anterior; (c) leg 3, anterior; (d) leg 4, posterior; (e) endopod of leg 3; (f) endopod of leg 4." captionText-4="FIGURE 49. Haplocyclops primitivus n. sp. (a) paratype male, habitus, dorsal; (b) allotype male, antennule, lateral; (c) same, ultimate and penultimate segments." captionText-5="FIGURE 50. Haplocyclops primitivus n. sp. (a) paratype male, urosome, dorsal; (b) allotype male, same, ventral." httpUri-0="https://zenodo.org/record/288280/files/figure.png" httpUri-1="https://zenodo.org/record/288281/files/figure.png" httpUri-2="https://zenodo.org/record/288282/files/figure.png" httpUri-3="https://zenodo.org/record/288283/files/figure.png" httpUri-4="https://zenodo.org/record/288284/files/figure.png" httpUri-5="https://zenodo.org/record/288285/files/figure.png" pageId="71" pageNumber="72">Figs. 45–50</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9FB5B3D98109FFA5FF1CFB82FAD7FAAA" pageId="71" pageNumber="72" type="materials_examined">
<paragraph id="D710E0528109FFA5FF1CFB82FB32FBB7" blockId="71.[151,1437,1076,2037]" pageId="71" pageNumber="72">
<emphasis id="E5DB3C408109FFA5FF1CFB82FE98FB8A" bold="true" box="[151,314,1076,1101]" pageId="71" pageNumber="72">
<typeStatus id="08145EF08109FFA5FF1CFB82FF70FB8A" box="[151,210,1076,1101]" pageId="71" pageNumber="72">Type</typeStatus>
locality.
</emphasis>
River 
<taxonomicName id="10AF9BD18109FFA5FE05FB82FE5BFB8A" box="[398,505,1076,1101]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="71" pageNumber="89" phylum="Arthropoda" rank="species" species="godavari" status="sp. nov.">Godavari</taxonomicName>
, Kapileswarapuram village (
<geoCoordinate id="B29B86958109FFA5FCC3FB82FC46FB8A" box="[840,996,1076,1101]" direction="north" orientation="latitude" pageId="71" pageNumber="72" precision="1" value="16.785833">16°47′09.0′′N</geoCoordinate>
<geoCoordinate id="B29B86958109FFA5FC7BFB82FB29FB8A" box="[1008,1163,1076,1101]" direction="east" orientation="longitude" pageId="71" pageNumber="72" precision="1" value="82.06092">82°03′39.3′′E</geoCoordinate>
; elevation 
<quantity id="10574DB78109FFA5FA86FB82FAEFFB8B" box="[1293,1357,1076,1100]" metricMagnitude="1" metricUnit="m" metricValue="2.3" pageId="71" pageNumber="72" unit="m" value="23.0">23 m</quantity>
; water temperature 28ºC, pH 7.0) in East 
<taxonomicName id="10AF9BD18109FFA5FD92FBE1FD23FBB7" box="[537,641,1111,1136]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="71" pageNumber="89" phylum="Arthropoda" rank="species" species="godavari" status="sp. nov.">Godavari</taxonomicName>
District, Andhra Pradesh, South 
<collectingCountry id="AFB8A0C28109FFA5FC7EFBE1FB8DFBB7" box="[1013,1071,1111,1136]" name="India" pageId="71" pageNumber="72">India</collectingCountry>
(
<figureCitation id="4F94FCD78109FFA5FBB5FBE1FBDDFBB7" box="[1086,1151,1111,1136]" captionStart="FIGURE 1" captionStartId="5.[151,250,1958,1980]" captionTargetBox="[185,1404,203,1932]" captionTargetId="figure@5.[178,1409,195,1934]" captionTargetPageId="5" captionText="FIGURE 1. Map showing the type localities of the cyclopoid species on the Indian peninsula." httpUri="https://zenodo.org/record/288236/files/figure.png" pageId="71" pageNumber="72">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="D710E0528109FFA5FF4CFBCAFAC9FAC7" blockId="71.[151,1437,1076,2037]" pageId="71" pageNumber="72">
<emphasis id="E5DB3C408109FFA5FF4CFBCAFE54FB52" bold="true" box="[199,502,1148,1173]" pageId="71" pageNumber="72">
<typeStatus id="08145EF08109FFA5FF4CFBCAFEA0FB52" box="[199,258,1148,1173]" pageId="71" pageNumber="72">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="08145EF08109FFA5FD8BFBCAFDCAFB52" box="[512,616,1148,1173]" pageId="71" pageNumber="72" type="holotype">Holotype</typeStatus>
female (MNHN-IU-2013-11878) and allotype male (MNHN-IU-2013- 11879), dissected on 3 slides each, and 6 
<typeStatus id="08145EF08109FFA5FDF3FB16FD4AFB7F" box="[632,744,1184,1208]" pageId="71" pageNumber="72" type="paratype">paratypes</typeStatus>
: 
<specimenCount id="C1A92BDB8109FFA5FD7EFB16FCF9FB7F" box="[757,859,1183,1208]" pageId="71" pageNumber="72" type="female">1 female</specimenCount>
, dissected on 3 slides (MNHN-IU-2013-11890); 
<specimenCount id="C1A92BDB8109FFA5FA05FB16FF4CFB1A" pageId="71" pageNumber="72" type="female">2 females</specimenCount>
whole-mounted on 1 slide each (MNHN-IU-2013-11891–11892), 
<specimenCount id="C1A92BDB8109FFA5FC57FB72FB8AFB1A" box="[988,1064,1220,1245]" pageId="71" pageNumber="72" type="male">1 male</specimenCount>
and 
<specimenCount id="C1A92BDB8109FFA5FBEBFB72FB63FB1A" box="[1120,1217,1220,1245]" pageId="71" pageNumber="72" type="female">1 female</specimenCount>
in alcohol in 1 vial (MNHN-IU-2013-11893); 
<specimenCount id="C1A92BDB8109FFA5FE4FFB5EFD92FAC7" box="[452,560,1255,1280]" pageId="71" pageNumber="72" type="female">2 females</specimenCount>
in senior author’s personal collection. 
<date id="A311C6928109FFA5FC6FFB5EFBDAFAC7" box="[996,1144,1255,1280]" pageId="71" pageNumber="72" value="2008-07-20">20 July 2008</date>
, Coll. V. R. Totakura.
</paragraph>
<paragraph id="D710E0528109FFA5FF4CFABAFAD7FAAA" blockId="71.[151,1437,1076,2037]" pageId="71" pageNumber="72">
<emphasis id="E5DB3C408109FFA5FF4CFABAFDA7FAE2" bold="true" box="[199,517,1292,1317]" pageId="71" pageNumber="72">Other material examined.</emphasis>
South 
<collectingCountry id="AFB8A0C28109FFA5FDD6FABAFD3BFAE2" box="[605,665,1292,1317]" name="India" pageId="71" pageNumber="72">India</collectingCountry>
, Andhra Pradesh, East 
<taxonomicName id="10AF9BD18109FFA5FC3BFABAFBB9FAE2" box="[944,1051,1292,1317]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="71" pageNumber="89" phylum="Arthropoda" rank="species" species="godavari" status="sp. nov.">Godavari</taxonomicName>
, River 
<taxonomicName id="10AF9BD18109FFA5FBFFFABAFB7EFAE2" box="[1140,1244,1292,1317]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="71" pageNumber="89" phylum="Arthropoda" rank="species" species="godavari" status="sp. nov.">Godavari</taxonomicName>
at Atreyapuram village (water temperature 28ºC, pH 7.0), 
<date id="A311C6928109FFA5FDFCFA86FCADFA8F" box="[631,783,1327,1352]" pageId="71" pageNumber="72" value="2008-07-20">20 July 2008</date>
: 
<specimenCount id="C1A92BDB8109FFA5FC97FA86FCCAFA8F" box="[796,872,1327,1352]" pageId="71" pageNumber="72" type="male">1 male</specimenCount>
and 
<specimenCount id="C1A92BDB8109FFA5FC29FA86FBB1FA8F" box="[930,1043,1327,1352]" pageId="71" pageNumber="72" type="female">3 females</specimenCount>
; at Dhawaleswaram village (water temperature 27ºC, pH 7.5), 
<date id="A311C6928109FFA5FE46FAE2FDC3FAAB" box="[461,609,1364,1389]" pageId="71" pageNumber="72" value="2008-07-20">20 July 2008</date>
: 
<specimenCount id="C1A92BDB8109FFA5FDE7FAE2FD7AFAAA" box="[620,728,1364,1389]" pageId="71" pageNumber="72" type="female">2 females</specimenCount>
in senior author’s personal collection; Coll. V. R. Totakura.
</paragraph>
</subSubSection>
<subSubSection id="9FB5B3D98109FFA5FF4CFAC1FD28F8A3" pageId="71" pageNumber="72" type="diagnosis">
<paragraph id="D710E0528109FFA5FF4CFAC1FD28F8A3" blockId="71.[151,1437,1076,2037]" pageId="71" pageNumber="72">
<emphasis id="E5DB3C408109FFA5FF4CFAC1FEE3FA57" bold="true" box="[199,321,1399,1424]" pageId="71" pageNumber="72">Diagnosis.</emphasis>
Very small cyclopinae, cyclopiform and integument smooth. Genital double-somite large and of characteristic shape, swollen both laterally and ventrally, with genital field situated at 3/4 of genital doublesomite’s length; anal operculum bowl-shaped, extending up to or beyond the somite’s limit; caudal rami about 1.5–1.7 times as long as wide, with lateral seta inserted in proximal half almost dorsally; ornamented with transverse row fine spinules at the base of lateral seta and dorsal seta; principal inner apical seta 1.3 times as long as principal outer apical seta; female antennule 11-segmented, lateral seta present on ultimate segment, and with only 2 setae on seventh segment; male antennule 16-segmented, digeniculate; antenna 4-segmented, with setal formula 2.1.5.7 and without exopodal seta; mandibular palp lacking; maxillulary palp 2-segmented; maxilla 4- segmented, with small lateral seta on basis; and maxilliped 4-segmented, setal formula 1.2.1.2; 2-segmented exopod and endopod on legs 1–4, without any coxal armature; leg 1 basis without inner spine; spine formula of second exopodal segment of legs 1–4: 2.2.2.2 and setal formula: 5.4.4.4; endopod of leg 4 with 1 spine and 1 seta apically; leg 5 represented by 3 setae, of which 2 setae on small protuberance and 1 basal plumose seta; female leg 6 as distinct cuticular plate, armed with 2 unequal setae and 1 tiny spinous process; the same leg in male with 2 minute spinous processes and 1 slender seta.
</paragraph>
</subSubSection>
<subSubSection id="9FB5B3D98109FFADFF4CF8D9FD1AF972" lastPageId="79" lastPageNumber="80" pageId="71" pageNumber="72" type="description">
<paragraph id="D710E0528109FFA5FF4CF8D9FA3FF832" blockId="71.[151,1437,1076,2037]" pageId="71" pageNumber="72">
<emphasis id="E5DB3C408109FFA5FF4CF8D9FDB5F84F" bold="true" box="[199,535,1903,1928]" pageId="71" pageNumber="72">Description of adult female.</emphasis>
Total body length, measured from tip of rostrum to posterior margin of caudal rami (excluding caudal setae), 318–347µm (
<typeStatus id="08145EF08109FFA5FD00F822FD4DF86A" box="[651,751,1940,1965]" pageId="71" pageNumber="72" type="holotype">holotype</typeStatus>
338 µm). Habitus (
<figureCitation id="4F94FCD78109FFA5FC44F822FB83F86A" box="[975,1057,1940,1965]" captionStart="FIGURE 45" captionStartId="73.[151,250,1953,1975]" captionTargetBox="[216,1393,209,1914]" captionTargetId="figure@73.[178,1409,193,1932]" captionTargetPageId="73" captionText="FIGURE 45. Haplocyclops primitivus n. sp. (a) paratype female, habitus, dorsal; (b) holotype female, antennule, lateral." httpUri="https://zenodo.org/record/288280/files/figure.png" pageId="71" pageNumber="72">Fig. 45</figureCitation>
a) robust, cyclopiform and dorsoventrally compressed, with prosome/urosome ratio 1.5 and greatest width (102 Μm) at distal end of cephalothorax. Body length/width ratio about 3.4 (dorsal view); cephalothorax almost 1.5 times as wide as genital double-somite.
</paragraph>
<paragraph id="D710E0528106FFAAFF1CFF2EFB18FD6F" blockId="72.[151,1437,151,2013]" pageId="72" pageNumber="73">
Free pedigerous somites without particular expansions, lateral margins somewhat curved. Preserved specimens colourless. Naupliar eye absent. Rostral projection (
<figureCitation id="4F94FCD78106FFAAFD6BFF0AFC90FF13" box="[736,818,188,213]" captionStart="FIGURE 47" captionStartId="75.[151,250,1953,1975]" captionTargetBox="[192,1389,207,1915]" captionTargetId="figure@75.[178,1409,193,1932]" captionTargetPageId="75" captionText="FIGURE 47. Haplocyclops primitivus n. sp., holotype female. (a) rostrum, ventral; (b) antenna, lateral; (c) labrum, ventral; (d) mandible, lateral; (e) maxillule, lateral; (f) maxilla, lateral; (g) maxilliped, lateral." httpUri="https://zenodo.org/record/288282/files/figure.png" pageId="72" pageNumber="73">Fig. 47</figureCitation>
a) well developed, membranous, broadly triangular, tip rounded and furnished with 2 large sensilla. Cephalothorax (
<figureCitation id="4F94FCD78106FFAAFCC1FF69FC3FFF3F" box="[842,925,223,248]" captionStart="FIGURE 45" captionStartId="73.[151,250,1953,1975]" captionTargetBox="[216,1393,209,1914]" captionTargetId="figure@73.[178,1409,193,1932]" captionTargetPageId="73" captionText="FIGURE 45. Haplocyclops primitivus n. sp. (a) paratype female, habitus, dorsal; (b) holotype female, antennule, lateral." httpUri="https://zenodo.org/record/288280/files/figure.png" pageId="72" pageNumber="73">Fig. 45</figureCitation>
a) 1.7 times as long as greatest width (dorsal view) and 35.7% of total body length; pleurotergites of free pedigerous somites ornamented with small sensilla; no other ornamentation visible. Hyaline fringes of prosomites narrow and smooth. Arthrodial membranes well developed. Third pediger longer than fourth one, fourth pediger unornamented. Fifth pediger (
<figureCitation id="4F94FCD78106FFAAFAA7FEFAFA25FEA2" box="[1324,1415,332,357]" captionStart="FIGURE 46" captionStartId="74.[151,250,1953,1975]" captionTargetBox="[174,1403,207,1926]" captionTargetId="figure@74.[163,1423,193,1932]" captionTargetPageId="74" captionText="FIGURE 46. Haplocyclops primitivus n. sp., holotype female (a), paratype female (b – e). (a) urosome, dorsal; (b) same, ventral; (c) same, lateral; (d) leg 5, latero-ventral; (e) leg 6, latero-ventral." httpUri="https://zenodo.org/record/288281/files/figure.png" pageId="72" pageNumber="73">Fig. 46</figureCitation>
a) ornamented with 4 small dorsal sensilla; hyaline fringe smooth dorso-ventrally. Genital double-somite (
<figureCitation id="4F94FCD78106FFAAFA97FED9FACFFE4F" box="[1308,1389,367,392]" captionStart="FIGURE 46" captionStartId="74.[151,250,1953,1975]" captionTargetBox="[174,1403,207,1926]" captionTargetId="figure@74.[163,1423,193,1932]" captionTargetPageId="74" captionText="FIGURE 46. Haplocyclops primitivus n. sp., holotype female (a), paratype female (b – e). (a) urosome, dorsal; (b) same, ventral; (c) same, lateral; (d) leg 5, latero-ventral; (e) leg 6, latero-ventral." httpUri="https://zenodo.org/record/288281/files/figure.png" pageId="72" pageNumber="73">Fig. 46</figureCitation>
a–c) proximally expanded, large, about as wide as long (dorsal view) and ornamented with 2 large dorsal sensilla; hyaline fringe of genital double-somite as well as next 2 urosomites smooth. Copulatory pore (
<figureCitation id="4F94FCD78106FFAAFB6FFE01FA9AFE17" box="[1252,1336,439,464]" captionStart="FIGURE 46" captionStartId="74.[151,250,1953,1975]" captionTargetBox="[174,1403,207,1926]" captionTargetId="figure@74.[163,1423,193,1932]" captionTargetPageId="74" captionText="FIGURE 46. Haplocyclops primitivus n. sp., holotype female (a), paratype female (b – e). (a) urosome, dorsal; (b) same, ventral; (c) same, lateral; (d) leg 5, latero-ventral; (e) leg 6, latero-ventral." httpUri="https://zenodo.org/record/288281/files/figure.png" pageId="72" pageNumber="73">Fig. 46</figureCitation>
b) small, ovoid, situated at 3/4 of somite’s length; copulatory duct narrow, well sclerotized. Seminal receptacle with large anterior expansion but somewhat narrow behind, representing 61.5% of double-somite’s length; oviducts broad and weakly sclerotized. Ovipores situated dorso-laterally, covered with reduced sixth legs. Anal somite ornamented with 2 large sensilla dorsally and 1 transverse row of minute spinules along posterior margin ventrally and laterally. Anal operculum large, bowl-shaped/sublinguiform, extending slightly beyond somite’s limit, smooth, with almost straight caudal margin, representing 65.3% of anal somite’s width. Anal sinus somewhat wide.
</paragraph>
<paragraph id="D710E0528106FFAAFF4CFD02FB14FC62" blockId="72.[151,1437,151,2013]" pageId="72" pageNumber="73">
Caudal rami (
<figureCitation id="4F94FCD78106FFAAFEE1FD02FE61FD0A" box="[362,451,692,717]" captionStart="FIGURE 46" captionStartId="74.[151,250,1953,1975]" captionTargetBox="[174,1403,207,1926]" captionTargetId="figure@74.[163,1423,193,1932]" captionTargetPageId="74" captionText="FIGURE 46. Haplocyclops primitivus n. sp., holotype female (a), paratype female (b – e). (a) urosome, dorsal; (b) same, ventral; (c) same, lateral; (d) leg 5, latero-ventral; (e) leg 6, latero-ventral." httpUri="https://zenodo.org/record/288281/files/figure.png" pageId="72" pageNumber="73">Fig. 46</figureCitation>
a–c): subcylindrical, parallel, 1.5 times as long as wide; ornamented with a row of spinules on subdistal dorsal surface close to dorsal seta. Dorsal seta about 1.9 times as long as ramus, inserted at 4/ 5 of ramus and just above the level of outermost apical seta, biarticulate at base (
<figureCitation id="4F94FCD78106FFAAFBCAFD4AFB03FCD2" box="[1089,1185,764,789]" captionStart="FIGURE 45" captionStartId="73.[151,250,1953,1975]" captionTargetBox="[216,1393,209,1914]" captionTargetId="figure@73.[178,1409,193,1932]" captionTargetPageId="73" captionText="FIGURE 45. Haplocyclops primitivus n. sp. (a) paratype female, habitus, dorsal; (b) holotype female, antennule, lateral." httpUri="https://zenodo.org/record/288280/files/figure.png" pageId="72" pageNumber="73">Figs. 45</figureCitation>
a, 46a–c). Lateral seta arising somewhat dorso-laterally at 3/5 of ramus length, 0.7 times as long as ramus width; 1 short transverse row of fine spinules lying beside its base. Outermost apical seta slightly longer than ramus. Innermost apical seta small and slender, 0.4 times as long as outermost apical seta. Principal apical setae without breaking planes. Inner apical seta 3.3 times as long as caudal rami, 1.3 times as long as outer apical seta. All setae plumose.
</paragraph>
<paragraph id="D710E0528106FFAAFF4CFC19FC55FBBB" blockId="72.[151,1437,151,2013]" pageId="72" pageNumber="73">
Antennule (
<figureCitation id="4F94FCD78106FFAAFED9FC19FE0EFC0F" box="[338,428,943,968]" captionStart="FIGURE 45" captionStartId="73.[151,250,1953,1975]" captionTargetBox="[216,1393,209,1914]" captionTargetId="figure@73.[178,1409,193,1932]" captionTargetPageId="73" captionText="FIGURE 45. Haplocyclops primitivus n. sp. (a) paratype female, habitus, dorsal; (b) holotype female, antennule, lateral." httpUri="https://zenodo.org/record/288280/files/figure.png" pageId="72" pageNumber="73">Fig. 45</figureCitation>
b): 11-segmented, 73.6% of the length of cephalothoracic shield and unornamented; armature formula: 7.2.5.2.1.2.2.3.3.3.7+aes. Setae articulating at basal part on segments 7, 8, and 11, and segments 1, 3, 7, 8, 10, and 11 with 4, 1, 2, 1, 1, and 3 long bipinnate setae, respectively; all other setae smooth. Length ratios of antennular segments from proximal end and along caudal margins 1.0: 0.2: 0.5: 0.2: 0.1: 0.3: 0.5: 0.5: 0.3: 0.4: 0.5. Probable segmental homology: 1 (I–V), 2 (VI–VII), 3 (VIII–XI), 4 (XII–XIII), 5 (XIV), 6 (XV–XVI), 7 (XVII–XX), 8 (XXI–XXIII), 9 (XXIV), 10 (XXV), and 11 (XXVI–XXVIII).
</paragraph>
<paragraph id="D710E0528106FFAAFF4CFB3EFB4BFA92" blockId="72.[151,1437,151,2013]" pageId="72" pageNumber="73">
Antenna (
<figureCitation id="4F94FCD78106FFAAFECBFB31FE3CFB67" box="[320,414,1159,1184]" captionStart="FIGURE 47" captionStartId="75.[151,250,1953,1975]" captionTargetBox="[192,1389,207,1915]" captionTargetId="figure@75.[178,1409,193,1932]" captionTargetPageId="75" captionText="FIGURE 47. Haplocyclops primitivus n. sp., holotype female. (a) rostrum, ventral; (b) antenna, lateral; (c) labrum, ventral; (d) mandible, lateral; (e) maxillule, lateral; (f) maxilla, lateral; (g) maxilliped, lateral." httpUri="https://zenodo.org/record/288282/files/figure.png" pageId="72" pageNumber="73">Fig. 47</figureCitation>
b): 4-segmented, comprising coxobasis and 3-segmented endopod. Coxobasis large, somewhat cylindrical, 2.5 times as long as wide, armed with 2 short, smooth seta at inner distal corner; exopodal seta absent. First endopodal segment stout, 1.3 times as long as wide, with inner smooth seta, and 1 arched row of spinules on outer margin. Second endopodal segment about twice as long as wide, bearing 5 setae and ornamented with 1 spinular row on outer distal margin. Third endopodal segment cylindrical, 2.6 times as long as wide, armed with 7 smooth, prehensile apical setae, and ornamented with 1 spinular row on outer distal margin.
</paragraph>
<paragraph id="D710E0528106FFAAFF4CFAE9FAFEFA5A" blockId="72.[151,1437,151,2013]" pageId="72" pageNumber="73">
Labrum (
<figureCitation id="4F94FCD78106FFAAFEA5FAE9FE20FABF" box="[302,386,1375,1400]" captionStart="FIGURE 47" captionStartId="75.[151,250,1953,1975]" captionTargetBox="[192,1389,207,1915]" captionTargetId="figure@75.[178,1409,193,1932]" captionTargetPageId="75" captionText="FIGURE 47. Haplocyclops primitivus n. sp., holotype female. (a) rostrum, ventral; (b) antenna, lateral; (c) labrum, ventral; (d) mandible, lateral; (e) maxillule, lateral; (f) maxilla, lateral; (g) maxilliped, lateral." httpUri="https://zenodo.org/record/288282/files/figure.png" pageId="72" pageNumber="73">Fig. 47</figureCitation>
c): relatively small, trapezoidal plate; anterior edge slightly concave, with about 20 small blunt teeth between rounded lateral corners; ornamented with 2 groups of long hair-like spinules on dorsal surface.
</paragraph>
<paragraph id="D710E0528106FFAAFF4CFA11FBA6F9CF" blockId="72.[151,1437,151,2013]" pageId="72" pageNumber="73">
Mandible (
<figureCitation id="4F94FCD78106FFAAFEC8FA11FE35FA07" box="[323,407,1447,1472]" captionStart="FIGURE 47" captionStartId="75.[151,250,1953,1975]" captionTargetBox="[192,1389,207,1915]" captionTargetId="figure@75.[178,1409,193,1932]" captionTargetPageId="75" captionText="FIGURE 47. Haplocyclops primitivus n. sp., holotype female. (a) rostrum, ventral; (b) antenna, lateral; (c) labrum, ventral; (d) mandible, lateral; (e) maxillule, lateral; (f) maxilla, lateral; (g) maxilliped, lateral." httpUri="https://zenodo.org/record/288282/files/figure.png" pageId="72" pageNumber="73">Fig. 47</figureCitation>
d): coxal gnathobase roughly divided into 3 groups of teeth; inner group of 3 large unequal teeth, innermost one being larger; middle group of 3 moderate teeth and 1 small sub-distal spinular row; distal group with 2 long spinules on ventral surface and 1 unipinnate outermost seta.
</paragraph>
<paragraph id="D710E0528106FFAAFF4CF9A2FA2EF95F" blockId="72.[151,1437,151,2013]" pageId="72" pageNumber="73">
Maxillule (
<figureCitation id="4F94FCD78106FFAAFECDF9A2FE3EF9EB" box="[326,412,1556,1581]" captionStart="FIGURE 47" captionStartId="75.[151,250,1953,1975]" captionTargetBox="[192,1389,207,1915]" captionTargetId="figure@75.[178,1409,193,1932]" captionTargetPageId="75" captionText="FIGURE 47. Haplocyclops primitivus n. sp., holotype female. (a) rostrum, ventral; (b) antenna, lateral; (c) labrum, ventral; (d) mandible, lateral; (e) maxillule, lateral; (f) maxilla, lateral; (g) maxilliped, lateral." httpUri="https://zenodo.org/record/288282/files/figure.png" pageId="72" pageNumber="73">Fig. 47</figureCitation>
e): composed of praecoxa and 2-segmented palp. Praecoxal arthrite bearing 3 very strong claw-like spinous processes and 1 stout pinnate seta and 7 medial elements, of which second proximal one largest and pinnate. Palp composed of coxobasis and endopod. Coxobasis with smooth proximal (exopodal) seta and 3 medial setae (2 slender, smooth and 1 strong, bipinnate); endopod with 3 smooth (2 apical and 1 subapical) setae.
</paragraph>
<paragraph id="D710E0528106FFAAFF4CF912FAF1F852" blockId="72.[151,1437,151,2013]" pageId="72" pageNumber="73">
Maxilla (
<figureCitation id="4F94FCD78106FFAAFEBBF912FE24F97A" box="[304,390,1700,1725]" captionStart="FIGURE 47" captionStartId="75.[151,250,1953,1975]" captionTargetBox="[192,1389,207,1915]" captionTargetId="figure@75.[178,1409,193,1932]" captionTargetPageId="75" captionText="FIGURE 47. Haplocyclops primitivus n. sp., holotype female. (a) rostrum, ventral; (b) antenna, lateral; (c) labrum, ventral; (d) mandible, lateral; (e) maxillule, lateral; (f) maxilla, lateral; (g) maxilliped, lateral." httpUri="https://zenodo.org/record/288282/files/figure.png" pageId="72" pageNumber="73">Fig. 47</figureCitation>
f): 4-segmented, consisting of syncoxa, basis and 2-segmented endopod; praecoxa fused to coxa; proximal endite of praecoxa robust, armed with 2 setae; distal endite small, unarmed. Proximal endite of coxa with 1 bipinnate seta; distal endite fused at base, moderate in size, elongate and armed apically with 2 unequal pinnate setae, proximal seta being 1.2 times longer; syncoxa unornamented. Basis expanded into robust claw, armed with 2 setae; 1 small seta at base and 1 strong pinnate proximal seta about as long as claw. Endopod 2- segmented; proximal segment armed with 2 robust, unequal, bipinnate setae; distal segment small, with 1 robust, unipinnate apical claw and 2 slender and smooth subapical setae. All strong setae and basal claw prehensile.
</paragraph>
<paragraph id="D710E0528106FFAAFF4CF816FB6AF81A" blockId="72.[151,1437,151,2013]" pageId="72" pageNumber="73">
Maxilliped (
<figureCitation id="4F94FCD78106FFAAFED2F816FE17F87F" box="[345,437,1952,1977]" captionStart="FIGURE 47" captionStartId="75.[151,250,1953,1975]" captionTargetBox="[192,1389,207,1915]" captionTargetId="figure@75.[178,1409,193,1932]" captionTargetPageId="75" captionText="FIGURE 47. Haplocyclops primitivus n. sp., holotype female. (a) rostrum, ventral; (b) antenna, lateral; (c) labrum, ventral; (d) mandible, lateral; (e) maxillule, lateral; (f) maxilla, lateral; (g) maxilliped, lateral." httpUri="https://zenodo.org/record/288282/files/figure.png" pageId="72" pageNumber="73">
Fig. 
<quantity id="10574DB78106FFAAFE1EF816FE17F87F" box="[405,437,1952,1976]" metricMagnitude="-2" metricUnit="kg" metricValue="4.7" pageId="72" pageNumber="73" unit="g" value="47.0">47</quantity>
</figureCitation>
g): 4-segmented, composed of syncoxa, basis and 2-segmented endopod. Armature formula: 1.2.1.2. Ornamentation consisting of longitudinal rows of spinules on first 3 segments.
</paragraph>
<caption id="83D0B0DA8107FFABFF1CF817FAFDF870" box="[151,1375,1953,1975]" httpUri="https://zenodo.org/record/288280/files/figure.png" pageId="73" pageNumber="74" targetBox="[216,1393,209,1914]" targetPageId="73">
<paragraph id="D710E0528107FFABFF1CF817FAFDF870" blockId="73.[151,1375,1953,1975]" box="[151,1375,1953,1975]" pageId="73" pageNumber="74">
<emphasis id="E5DB3C408107FFABFF1CF817FEBAF871" bold="true" box="[151,280,1953,1975]" pageId="73" pageNumber="74">FIGURE 45</emphasis>
. 
<taxonomicName id="10AF9BD18107FFABFEAEF817FDBCF870" box="[293,542,1953,1975]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="primitivus" status="sp. nov.">
<emphasis id="E5DB3C408107FFABFEAEF817FDBCF870" box="[293,542,1953,1975]" italics="true" pageId="73" pageNumber="74">Haplocyclops primitivus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FEE8813B8107FFABFDAFF817FDFFF871" box="[548,605,1953,1974]" pageId="73" pageNumber="74" rank="species">
<emphasis id="E5DB3C408107FFABFDAFF817FDFFF871" bold="true" box="[548,605,1953,1974]" pageId="73" pageNumber="74">n. sp.</emphasis>
</taxonomicNameLabel>
(a) paratype female, habitus, dorsal; (b) holotype female, antennule, lateral.
</paragraph>
</caption>
<caption id="83D0B0DA8104FFA8FF1CF817FC21F811" httpUri="https://zenodo.org/record/288281/files/figure.png" pageId="74" pageNumber="75" targetBox="[174,1403,207,1926]" targetPageId="74">
<paragraph id="D710E0528104FFA8FF1CF817FC21F811" blockId="74.[151,1436,1953,2006]" pageId="74" pageNumber="75">
<emphasis id="E5DB3C408104FFA8FF1CF817FEBFF871" bold="true" box="[151,285,1953,1975]" pageId="74" pageNumber="75">FIGURE 46</emphasis>
. 
<taxonomicName id="10AF9BD18104FFA8FEA5F817FD89F870" box="[302,555,1953,1975]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="primitivus" status="sp. nov.">
<emphasis id="E5DB3C408104FFA8FEA5F817FD89F870" box="[302,555,1953,1975]" italics="true" pageId="74" pageNumber="75">Haplocyclops primitivus</emphasis>
</taxonomicName>
<emphasis id="E5DB3C408104FFA8FDBEF817FDDAF871" bold="true" box="[565,632,1953,1974]" pageId="74" pageNumber="75">
<taxonomicNameLabel id="FEE8813B8104FFA8FDBEF817FDD6F871" box="[565,628,1953,1974]" pageId="74" pageNumber="75" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
holotype female (a), paratype female (b–e). (a) urosome, dorsal; (b) same, ventral; (c) same, lateral; (d) leg 5, latero-ventral; (e) leg 6, latero-ventral.
</paragraph>
</caption>
<caption id="83D0B0DA8105FFA9FF1CF817FC70F811" httpUri="https://zenodo.org/record/288282/files/figure.png" pageId="75" pageNumber="76" targetBox="[192,1389,207,1915]" targetPageId="75">
<paragraph id="D710E0528105FFA9FF1CF817FC70F811" blockId="75.[151,1436,1953,2006]" pageId="75" pageNumber="76">
<emphasis id="E5DB3C408105FFA9FF1CF817FEBAF871" bold="true" box="[151,280,1953,1975]" pageId="75" pageNumber="76">FIGURE 47</emphasis>
. 
<taxonomicName id="10AF9BD18105FFA9FEAEF817FDBFF870" box="[293,541,1953,1975]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="primitivus" status="sp. nov.">
<emphasis id="E5DB3C408105FFA9FEAEF817FDBFF870" box="[293,541,1953,1975]" italics="true" pageId="75" pageNumber="76">Haplocyclops primitivus</emphasis>
</taxonomicName>
<emphasis id="E5DB3C408105FFA9FDA8F817FDC0F871" bold="true" box="[547,610,1953,1974]" pageId="75" pageNumber="76">
<taxonomicNameLabel id="FEE8813B8105FFA9FDA8F817FDFFF871" box="[547,605,1953,1974]" pageId="75" pageNumber="76" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
holotype female. (a) rostrum, ventral; (b) antenna, lateral; (c) labrum, ventral; (d) mandible, lateral; (e) maxillule, lateral; (f) maxilla, lateral; (g) maxilliped, lateral.
</paragraph>
</caption>
<caption id="83D0B0DA8102FFAEFF1CF817FB2AF811" httpUri="https://zenodo.org/record/288283/files/figure.png" pageId="76" pageNumber="77" targetBox="[182,1394,210,1921]" targetPageId="76">
<paragraph id="D710E0528102FFAEFF1CF817FB2AF811" blockId="76.[151,1436,1953,2006]" pageId="76" pageNumber="77">
<emphasis id="E5DB3C408102FFAEFF1CF817FEBBF871" bold="true" box="[151,281,1953,1975]" pageId="76" pageNumber="77">FIGURE 48</emphasis>
. 
<taxonomicName id="10AF9BD18102FFAEFEADF817FDBDF870" box="[294,543,1953,1975]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="primitivus" status="sp. nov.">
<emphasis id="E5DB3C408102FFAEFEADF817FDBDF870" box="[294,543,1953,1975]" italics="true" pageId="76" pageNumber="77">Haplocyclops primitivus</emphasis>
</taxonomicName>
<emphasis id="E5DB3C408102FFAEFDADF817FDC7F871" bold="true" box="[550,613,1953,1974]" pageId="76" pageNumber="77">
<taxonomicNameLabel id="FEE8813B8102FFAEFDADF817FDC2F871" box="[550,608,1953,1974]" pageId="76" pageNumber="77" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
holotype female (a–d), paratype female (e), allotype male (f). (a) leg 1, anterior; (b) leg 2, anterior; (c) leg 3, anterior; (d) leg 4, posterior; (e) endopod of leg 3; (f) endopod of leg 4.
</paragraph>
</caption>
<caption id="83D0B0DA8103FFAFFF1CF817FE5CF811" httpUri="https://zenodo.org/record/288284/files/figure.png" pageId="77" pageNumber="78" targetBox="[213,1378,214,1927]" targetPageId="77">
<paragraph id="D710E0528103FFAFFF1CF817FE5CF811" blockId="77.[151,1436,1953,2006]" pageId="77" pageNumber="78">
<emphasis id="E5DB3C408103FFAFFF1CF817FEBBF871" bold="true" box="[151,281,1953,1975]" pageId="77" pageNumber="78">FIGURE 49</emphasis>
. 
<taxonomicName id="10AF9BD18103FFAFFEADF817FDBDF870" box="[294,543,1953,1975]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="primitivus" status="sp. nov.">
<emphasis id="E5DB3C408103FFAFFEADF817FDBDF870" box="[294,543,1953,1975]" italics="true" pageId="77" pageNumber="78">Haplocyclops primitivus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FEE8813B8103FFAFFDADF817FDC2F871" box="[550,608,1953,1974]" pageId="77" pageNumber="78" rank="species">
<emphasis id="E5DB3C408103FFAFFDADF817FDC2F871" bold="true" box="[550,608,1953,1974]" pageId="77" pageNumber="78">n. sp.</emphasis>
</taxonomicNameLabel>
(a) paratype male, habitus, dorsal; (b) allotype male, antennule, lateral; (c) same, ultimate and penultimate segments.
</paragraph>
</caption>
<caption id="83D0B0DA8100FFACFF1CF817FABAF870" box="[151,1304,1953,1975]" httpUri="https://zenodo.org/record/288285/files/figure.png" pageId="78" pageNumber="79" targetBox="[197,1409,203,1905]" targetPageId="78">
<paragraph id="D710E0528100FFACFF1CF817FABAF870" blockId="78.[151,1304,1953,1975]" box="[151,1304,1953,1975]" pageId="78" pageNumber="79">
<emphasis id="E5DB3C408100FFACFF1CF817FEBAF871" bold="true" box="[151,280,1953,1975]" pageId="78" pageNumber="79">FIGURE 50</emphasis>
. 
<taxonomicName id="10AF9BD18100FFACFEAEF817FDBCF870" box="[293,542,1953,1975]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="primitivus" status="sp. nov.">
<emphasis id="E5DB3C408100FFACFEAEF817FDBCF870" box="[293,542,1953,1975]" italics="true" pageId="78" pageNumber="79">Haplocyclops primitivus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FEE8813B8100FFACFDAFF817FDFFF871" box="[548,605,1953,1974]" pageId="78" pageNumber="79" rank="species">
<emphasis id="E5DB3C408100FFACFDAFF817FDFFF871" bold="true" box="[548,605,1953,1974]" pageId="78" pageNumber="79">n. sp.</emphasis>
</taxonomicNameLabel>
(a) paratype male, urosome, dorsal; (b) allotype male, same, ventral.
</paragraph>
</caption>
<paragraph id="D710E0528101FFADFF4CFF2EFE7FFF3F" blockId="79.[151,1436,151,248]" pageId="79" pageNumber="80">
Legs 1–4 (
<figureCitation id="4F94FCD78101FFADFEB5FF21FE33FF77" box="[318,401,151,176]" captionStart="FIGURE 48" captionStartId="76.[151,250,1953,1975]" captionTargetBox="[182,1394,210,1921]" captionTargetId="figure@76.[178,1409,193,1932]" captionTargetPageId="76" captionText="FIGURE 48. Haplocyclops primitivus n. sp., holotype female (a – d), paratype female (e), allotype male (f). (a) leg 1, anterior; (b) leg 2, anterior; (c) leg 3, anterior; (d) leg 4, posterior; (e) endopod of leg 3; (f) endopod of leg 4." httpUri="https://zenodo.org/record/288283/files/figure.png" pageId="79" pageNumber="80">Fig. 48</figureCitation>
a–d): relatively short, with 2-segmented exopod and endopod. Endopod only slightly shorter than exopod on legs 1–3, but distinctly reduced in size on leg 4. Armature formula of legs 1–4 (legend: same as that of 
<taxonomicName id="10AF9BD18101FFADFF3EFF69FE6EFF3F" box="[181,460,223,248]" class="Maxillopoda" family="Cyclopinidae" genus="Paracyclopina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="E5DB3C408101FFADFF3EFF69FE6EFF3F" box="[181,460,223,248]" italics="true" pageId="79" pageNumber="80">Paracyclopina orientalis</emphasis>
</taxonomicName>
):
</paragraph>
<paragraph id="D710E0528101FFADFEDCFEA2FBD4FEED" blockId="79.[159,1285,276,504]" box="[343,1142,276,298]" pageId="79" pageNumber="80">Coxa Basis Exopod Endopod</paragraph>
<paragraph id="D710E0528101FFADFDF9FE8BFB19FE94" blockId="79.[159,1285,276,504]" box="[626,1211,317,339]" pageId="79" pageNumber="80">1 2 1 2</paragraph>
<paragraph id="D710E0528101FFADFF14FED1FC18FCF7" blockId="79.[159,1285,276,504]" lastBlockId="79.[151,1437,575,2005]" pageId="79" pageNumber="80">
Leg 1 0-0 0-1 0- 
<date id="A311C6928101FFADFD0DFED0FCB3FEBB" box="[646,785,358,380]" pageId="79" pageNumber="80">I 4</date>
, 1+I, 
<date id="A311C6928101FFADFCC7FED1FB86FEBB" box="[844,1060,358,380]" pageId="79" pageNumber="80">I 1</date>
-0 1, 1+I, 1 Leg 2 0-0 0-1 0- 
<date id="A311C6928101FFADFD0DFE26FCB3FE61" box="[646,785,400,422]" pageId="79" pageNumber="80">I 3</date>
, 1+I, 
<date id="A311C6928101FFADFCC7FE27FB86FE61" box="[844,1060,400,422]" pageId="79" pageNumber="80">I 1</date>
-0 1, 1+I, 1 Leg 3 0-0 0-1 0- 
<date id="A311C6928101FFADFD0DFE0FFCB3FE08" box="[646,785,441,463]" pageId="79" pageNumber="80">I 3</date>
, 1+I, 
<date id="A311C6928101FFADFCC7FE0CFB86FE08" box="[844,1060,441,463]" pageId="79" pageNumber="80">I 1</date>
-0 2, 1+I, 1 Leg 4 0-0 0-1 0- 
<date id="A311C6928101FFADFD0DFE54FCB3FE3F" box="[646,785,482,504]" pageId="79" pageNumber="80">I 3</date>
, 1+I, 
<date id="A311C6928101FFADFCC7FE55FB86FE3F" box="[844,1060,482,504]" pageId="79" pageNumber="80">I 1</date>
-0 1+I Second exopodal segment spine formula 2.2.2.2 and setal formula 5.4.4.4. Apical spine on second endopodal segment of legs 1–3 about as long as segment, but distinctly longer on leg 4. Intercoxal sclerites of all swimming legs with concave distal margins and rounded lateral prominences fringed with long hairs. Praecoxae short, narrow and ornamented with 1 row of spinules each at outer distal margin. All coxae ornamented with 1 row of spinules each at outer distal margin. Basis with round inner distal corners, ornamented with long hairs. Inner spine on basis of leg 1 absent. Second endopodal segment of leg 4 as long as wide, armed with 1 spine and 1 seta apically; first segment ornamented with a row of spinules on distal margin (
<figureCitation id="4F94FCD78101FFADFCC0FCA1FC3CFCF7" box="[843,926,791,816]" captionStart="FIGURE 47" captionStartId="75.[151,250,1953,1975]" captionTargetBox="[192,1389,207,1915]" captionTargetId="figure@75.[178,1409,193,1932]" captionTargetPageId="75" captionText="FIGURE 47. Haplocyclops primitivus n. sp., holotype female. (a) rostrum, ventral; (b) antenna, lateral; (c) labrum, ventral; (d) mandible, lateral; (e) maxillule, lateral; (f) maxilla, lateral; (g) maxilliped, lateral." httpUri="https://zenodo.org/record/288282/files/figure.png" pageId="79" pageNumber="80">Fig. 47</figureCitation>
d).
</paragraph>
<paragraph id="D710E0528101FFADFF4CFC8BFDB7FC5A" blockId="79.[151,1437,575,2005]" pageId="79" pageNumber="80">
Leg 5 (
<figureCitation id="4F94FCD78101FFADFE9CFC8AFED0FC92" box="[279,370,828,853]" captionStart="FIGURE 45" captionStartId="73.[151,250,1953,1975]" captionTargetBox="[216,1393,209,1914]" captionTargetId="figure@73.[178,1409,193,1932]" captionTargetPageId="73" captionText="FIGURE 45. Haplocyclops primitivus n. sp. (a) paratype female, habitus, dorsal; (b) holotype female, antennule, lateral." httpUri="https://zenodo.org/record/288280/files/figure.png" pageId="79" pageNumber="80">Figs. 45</figureCitation>
a, 46a–d): with 3 setae, basipodal seta strong, visible clearly in dorsal and lateral views, arising from prominent protuberance; other 2 setae unequal, smooth and located on small protuberance; 1 row of spinules occurring at base of protuberance.
</paragraph>
<paragraph id="D710E0528101FFADFF4CFC1EFC7AFC23" blockId="79.[151,1437,575,2005]" pageId="79" pageNumber="80">
Leg 6 (
<figureCitation id="4F94FCD78101FFADFE9EFC11FECBFC07" box="[277,361,935,960]" captionStart="FIGURE 46" captionStartId="74.[151,250,1953,1975]" captionTargetBox="[174,1403,207,1926]" captionTargetId="figure@74.[163,1423,193,1932]" captionTargetPageId="74" captionText="FIGURE 46. Haplocyclops primitivus n. sp., holotype female (a), paratype female (b – e). (a) urosome, dorsal; (b) same, ventral; (c) same, lateral; (d) leg 5, latero-ventral; (e) leg 6, latero-ventral." httpUri="https://zenodo.org/record/288281/files/figure.png" pageId="79" pageNumber="80">Fig. 46</figureCitation>
e): distinct at base, small, semicircular cuticular plate, armed with 2 unequal, spiniform setae and 1 small spinous projection; outer seta about 1.5 times as long as inner one.
</paragraph>
<paragraph id="D710E0528101FFADFF4CFC59FAD6FAC2" blockId="79.[151,1437,575,2005]" pageId="79" pageNumber="80">
<emphasis id="E5DB3C408101FFADFF4CFC59FDA1FBCF" bold="true" box="[199,515,1007,1032]" pageId="79" pageNumber="80">Description of adult male.</emphasis>
Total body length 338 µm. Habitus (
<figureCitation id="4F94FCD78101FFADFC3AFC59FBA7FBCF" box="[945,1029,1007,1032]" captionStart="FIGURE 49" captionStartId="77.[151,250,1953,1975]" captionTargetBox="[213,1378,214,1927]" captionTargetId="figure@77.[178,1409,193,1932]" captionTargetPageId="77" captionText="FIGURE 49. Haplocyclops primitivus n. sp. (a) paratype male, habitus, dorsal; (b) allotype male, antennule, lateral; (c) same, ultimate and penultimate segments." httpUri="https://zenodo.org/record/288284/files/figure.png" pageId="79" pageNumber="80">Fig. 49</figureCitation>
a) slightly slenderer than in female, prosome/urosome ratio about 1.5 and greatest width (96 Μm) at posterior end of cephalothorax. Body length/width ratio 3.6; cephalothorax about 1.7 times as wide as genital somite. Cephalothorax 1.2 times as long as wide (dorsal view) and 35.2% of total body length. Ornamentation of prosomites as illustrated. Hyaline fringe of fifth pedigerous somite smooth both ventrally and dorsally. Genital somite (
<figureCitation id="4F94FCD78101FFADFC40FBC9FB82FB5F" box="[971,1056,1151,1176]" captionStart="FIGURE 50" captionStartId="78.[151,250,1953,1975]" captionTargetBox="[197,1409,203,1905]" captionTargetId="figure@78.[178,1409,193,1932]" captionTargetPageId="78" captionText="FIGURE 50. Haplocyclops primitivus n. sp. (a) paratype male, urosome, dorsal; (b) allotype male, same, ventral." httpUri="https://zenodo.org/record/288285/files/figure.png" pageId="79" pageNumber="80">Fig. 50</figureCitation>
a, b) about as long as wide, with smooth hyaline fringe dorsally, ornamented with 2 sensilla dorsally; 2 relatively small, ovoid spermatophores visible from inside; next 3 urosomites without any sensilla, integument ornamented only with smooth hyaline frill. Anal somite with transverse row of spinules all along caudal margin. Anal operculum almost similar to female.
</paragraph>
<paragraph id="D710E0528101FFADFF4CFAB9FD11FA1B" blockId="79.[151,1437,575,2005]" pageId="79" pageNumber="80">
Antennule (
<figureCitation id="4F94FCD78101FFADFEDAFAB9FE09FAEF" box="[337,427,1295,1320]" captionStart="FIGURE 49" captionStartId="77.[151,250,1953,1975]" captionTargetBox="[213,1378,214,1927]" captionTargetId="figure@77.[178,1409,193,1932]" captionTargetPageId="77" captionText="FIGURE 49. Haplocyclops primitivus n. sp. (a) paratype male, habitus, dorsal; (b) allotype male, antennule, lateral; (c) same, ultimate and penultimate segments." httpUri="https://zenodo.org/record/288284/files/figure.png" pageId="79" pageNumber="80">Fig. 49</figureCitation>
b, c): 16-segmented, digeniculate, proximal geniculation between 8 and 9 and distal geniculation between 14 and 15 segments; ornamented with spinules only on first segment. Armature formula: 10.3.3.1.0.1.0.0.1+aes.2.1.1.1.0.1.10. Short unipinnate seta present on segment 13; segments 1, 6 and 16 with 1, 1 and 4 long pinnate seta(e), respectively. Aesthetascs on segments 4 and 9 smooth and of moderate length; 4 setae on segment 16 with articulate base. Length ratios of antennular segments along medial axis 1.0: 0.3: 0.2: 0.1: 0.1: 0.1: 0.2: 0.3: 0.3: 0.2: 0.3: 0.2: 0.3: 0.7: 0.7: 1.2.
</paragraph>
<paragraph id="D710E0528101FFADFF4CFA5EFF49F9E2" blockId="79.[151,1437,575,2005]" pageId="79" pageNumber="80">Antenna, labrum, mandible, maxillule, maxilla, maxilliped, legs 1–3 and 5, and caudal rami similar to those of female.</paragraph>
<paragraph id="D710E0528101FFADFF4CF987FDB7F9AA" blockId="79.[151,1437,575,2005]" pageId="79" pageNumber="80">
Leg 4 (
<figureCitation id="4F94FCD78101FFADFE97F999FED2F98F" box="[284,368,1583,1608]" captionStart="FIGURE 48" captionStartId="76.[151,250,1953,1975]" captionTargetBox="[182,1394,210,1921]" captionTargetId="figure@76.[178,1409,193,1932]" captionTargetPageId="76" captionText="FIGURE 48. Haplocyclops primitivus n. sp., holotype female (a – d), paratype female (e), allotype male (f). (a) leg 1, anterior; (b) leg 2, anterior; (c) leg 3, anterior; (d) leg 4, posterior; (e) endopod of leg 3; (f) endopod of leg 4." httpUri="https://zenodo.org/record/288283/files/figure.png" pageId="79" pageNumber="80">Fig. 48</figureCitation>
f): second endopodal segment about as long as wide, with 1 apical seta, 1 apical spine and 1 inner seta; otherwise as in female.
</paragraph>
<paragraph id="D710E0528101FFADFF4CF9CFFD1AF972" blockId="79.[151,1437,575,2005]" pageId="79" pageNumber="80">
Leg 6 (
<figureCitation id="4F94FCD78101FFADFEABF9C1FEDAF957" box="[288,376,1655,1680]" captionStart="FIGURE 50" captionStartId="78.[151,250,1953,1975]" captionTargetBox="[197,1409,203,1905]" captionTargetId="figure@78.[178,1409,193,1932]" captionTargetPageId="78" captionText="FIGURE 50. Haplocyclops primitivus n. sp. (a) paratype male, urosome, dorsal; (b) allotype male, same, ventral." httpUri="https://zenodo.org/record/288285/files/figure.png" pageId="79" pageNumber="80">Fig. 50</figureCitation>
b): large, unornamented cuticular plate fused at base; armed with 1 seta and 2 diminutive prominences on outer distal corner, as in female.
</paragraph>
</subSubSection>
<subSubSection id="9FB5B3D98101FFADFF4CF909FD68F93A" pageId="79" pageNumber="80" type="etymology">
<paragraph id="D710E0528101FFADFF4CF909FD68F93A" blockId="79.[151,1437,575,2005]" pageId="79" pageNumber="80">
<emphasis id="E5DB3C408101FFADFF4CF909FEECF91F" bold="true" box="[199,334,1727,1752]" pageId="79" pageNumber="80">Etymology.</emphasis>
The specific epithet is derived from the Latin adjective, 
<taxonomicName id="10AF9BD18101FFADFBA4F977FB03F91F" box="[1071,1185,1729,1752]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="primitivus">
<emphasis id="E5DB3C408101FFADFBA4F977FB03F91F" box="[1071,1185,1729,1752]" italics="true" pageId="79" pageNumber="80">primitivus</emphasis>
</taxonomicName>
, alluding to several plesiomorphic character states. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="9FB5B3D98101FFB2FF4CF8B1FDD1FF12" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80" type="description">
<paragraph id="D710E0528101FFADFF4CF8B1FD2BF882" blockId="79.[151,1437,575,2005]" pageId="79" pageNumber="80">
<emphasis id="E5DB3C408101FFADFF4CF8B1FE99F8E7" bold="true" box="[199,315,1799,1824]" pageId="79" pageNumber="80">Variation.</emphasis>
Second exopodal segment of leg 3 with additional inner seta (
<figureCitation id="4F94FCD78101FFADFC71F8B1FBEFF8E7" box="[1018,1101,1799,1824]" captionStart="FIGURE 48" captionStartId="76.[151,250,1953,1975]" captionTargetBox="[182,1394,210,1921]" captionTargetId="figure@76.[178,1409,193,1932]" captionTargetPageId="76" captionText="FIGURE 48. Haplocyclops primitivus n. sp., holotype female (a – d), paratype female (e), allotype male (f). (a) leg 1, anterior; (b) leg 2, anterior; (c) leg 3, anterior; (d) leg 4, posterior; (e) endopod of leg 3; (f) endopod of leg 4." httpUri="https://zenodo.org/record/288283/files/figure.png" pageId="79" pageNumber="80">Fig. 48</figureCitation>
e), second endopodal segment of leg 4 with additional inner seta (
<figureCitation id="4F94FCD78101FFADFD94F89AFDD3F882" box="[543,625,1836,1861]" captionStart="FIGURE 48" captionStartId="76.[151,250,1953,1975]" captionTargetBox="[182,1394,210,1921]" captionTargetId="figure@76.[178,1409,193,1932]" captionTargetPageId="76" captionText="FIGURE 48. Haplocyclops primitivus n. sp., holotype female (a – d), paratype female (e), allotype male (f). (a) leg 1, anterior; (b) leg 2, anterior; (c) leg 3, anterior; (d) leg 4, posterior; (e) endopod of leg 3; (f) endopod of leg 4." httpUri="https://zenodo.org/record/288283/files/figure.png" pageId="79" pageNumber="80">Fig. 48</figureCitation>
f).
</paragraph>
<paragraph id="D710E0528101FFB2FF4CF8F9FDD1FF12" blockId="79.[151,1437,575,2005]" lastBlockId="80.[151,1437,151,1616]" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
<emphasis id="E5DB3C408101FFADFF4CF8F9FE89F8AF" bold="true" box="[199,299,1871,1896]" pageId="79" pageNumber="80">Ecology.</emphasis>
The new species was found in very small numbers in the present samples. It was, however, accompanied by various taxa at the 
<typeStatus id="08145EF08101FFADFDC9F8C3FDD0F84A" box="[578,626,1909,1933]" pageId="79" pageNumber="80">type</typeStatus>
locality: 
<emphasis id="E5DB3C408101FFADFD6FF8C2FB83F84A" box="[740,1057,1908,1933]" italics="true" pageId="79" pageNumber="80">Parvulobathynella distincta</emphasis>
Ranga Reddy, Elia &amp; Totakura, 
<taxonomicName id="10AF9BD18101FFADFF1CF821FD54F877" authority="Ranga Reddy &amp; Schminke" authorityName="Ranga Reddy &amp; Schminke" box="[151,758,1943,1969]" class="Malacostraca" family="Parabathynellidae" genus="Habrobathynella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Bathynellacea" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="plenituda">
<emphasis id="E5DB3C408101FFADFF1CF821FE64F877" box="[151,454,1943,1968]" italics="true" pageId="79" pageNumber="80">Habrobathynella plenituda</emphasis>
Ranga Reddy &amp; Schminke
</taxonomicName>
, and 
<taxonomicName id="10AF9BD18101FFADFCB8F82FFAECF877" authority="Ranga Reddy &amp; Schminke" authorityName="Ranga Reddy &amp; Schminke" box="[819,1358,1943,1969]" class="Maxillopoda" family="Parastenocarididae" genus="Kinnecaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="godavari">
<emphasis id="E5DB3C408101FFADFCB8F82FFBB8F877" box="[819,1050,1943,1968]" italics="true" pageId="79" pageNumber="80">Kinnecaris godavari</emphasis>
Ranga Reddy &amp; Schminke
</taxonomicName>
; and at Atreyapuram, it co-existed with 
<emphasis id="E5DB3C408101FFADFD8EF80AFC94F812" box="[517,822,1980,2005]" italics="true" pageId="79" pageNumber="80">Serbanibathynella secunda</emphasis>
Totakura &amp; Ranga Reddy, 
<taxonomicName id="10AF9BD18101FFB2FBE7F80AFE60FF77" authority="Schminke &amp; Ranga Reddy" authorityName="Schminke &amp; Ranga Reddy" class="Malacostraca" family="Parabathynellidae" genus="Habrobathynella" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="80" lastPageNumber="81" order="Bathynellacea" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="plenituda">
<emphasis id="E5DB3C408101FFADFBE7F80AFA3EF812" box="[1132,1436,1980,2005]" italics="true" pageId="79" pageNumber="80">Habrobathynella plenituda</emphasis>
Schminke &amp; Ranga Reddy
</taxonomicName>
, 
<taxonomicName id="10AF9BD1811EFFB2FE46FF21FC59FF77" authority="Ranga Reddy &amp; Totakura" authorityName="Ranga Reddy &amp; Totakura" box="[461,1019,151,176]" class="Malacostraca" family="Parabathynellidae" genus="Habrobathynella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Bathynellacea" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="savita">
<emphasis id="E5DB3C40811EFFB2FE46FF21FD77FF77" box="[461,725,151,176]" italics="true" pageId="80" pageNumber="81">Habrobathynella savita</emphasis>
Ranga Reddy &amp; Totakura
</taxonomicName>
, 
<taxonomicName id="10AF9BD1811EFFB2FB8DFF2FFA36FF77" authority="Enckell" authorityName="Enckell" box="[1030,1428,151,176]" class="Maxillopoda" family="Parastenocarididae" genus="Parastenocaris" kingdom="Animalia" order="Harpacticoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="curvispinus">
<emphasis id="E5DB3C40811EFFB2FB8DFF2FFA9BFF77" box="[1030,1337,153,176]" italics="true" pageId="80" pageNumber="81">Parastenocaris curvispinus</emphasis>
Enckell
</taxonomicName>
, oligochaetes, nematodes, and insect larvae
</paragraph>
</subSubSection>
<subSubSection id="9FB5B3D9811EFFB2FF4CFF69FDD1F997" pageId="80" pageNumber="81" type="discussion">
<paragraph id="D710E052811EFFB2FF4CFF69FCE1FD6F" blockId="80.[151,1437,151,1616]" pageId="80" pageNumber="81">
<emphasis id="E5DB3C40811EFFB2FF4CFF69FE99FF3F" bold="true" box="[199,315,223,248]" pageId="80" pageNumber="81">Remarks.</emphasis>
<bibRefCitation id="B33E9DA3811EFFB2FECCFF69FE44FF3F" author="Kiefer" box="[327,486,223,248]" pageId="80" pageNumber="81" refString="Kiefer, F. (1952) Haplocyclops Gudrunae n. g. et n. sp. ein neuer Ruderfuβkrebs (Crustacea Copepoda) aus Madagascar. Zoologischer Anzeiger, 149, 240 - 243." type="journal article" year="1952">Kiefer (1952)</bibRefCitation>
erected the genus 
<taxonomicName id="10AF9BD1811EFFB2FD47FF69FBA5FF3F" authority="Kiefer, 1952" authorityName="Kiefer" authorityYear="1952" box="[716,1031,223,248]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E5DB3C40811EFFB2FD47FF69FCC4FF3F" box="[716,870,223,248]" italics="true" pageId="80" pageNumber="81">Haplocyclops</emphasis>
<bibRefCitation id="B33E9DA3811EFFB2FCF9FF69FBA5FF3F" author="Kiefer" box="[882,1031,223,248]" pageId="80" pageNumber="81" refString="Kiefer, F. (1952) Haplocyclops Gudrunae n. g. et n. sp. ein neuer Ruderfuβkrebs (Crustacea Copepoda) aus Madagascar. Zoologischer Anzeiger, 149, 240 - 243." type="journal article" year="1952">Kiefer, 1952</bibRefCitation>
</taxonomicName>
, to accommodate the Madagascan 
<taxonomicName id="10AF9BD1811EFFB2FF1CFEB2FDF0FEDA" authority="Kiefer, 1952" authorityName="Kiefer" authorityYear="1952" box="[151,594,260,285]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="gudrunae">
<emphasis id="E5DB3C40811EFFB2FF1CFEB2FE09FEDA" box="[151,427,260,285]" italics="true" pageId="80" pageNumber="81">Haplocyclops gudrunae</emphasis>
<bibRefCitation id="B33E9DA3811EFFB2FE31FEB2FDF0FEDA" author="Kiefer" box="[442,594,260,285]" pageId="80" pageNumber="81" refString="Kiefer, F. (1952) Haplocyclops Gudrunae n. g. et n. sp. ein neuer Ruderfuβkrebs (Crustacea Copepoda) aus Madagascar. Zoologischer Anzeiger, 149, 240 - 243." type="journal article" year="1952">Kiefer, 1952</bibRefCitation>
</taxonomicName>
. Later, 
<bibRefCitation id="B33E9DA3811EFFB2FD38FEB2FCF6FEDA" author="Kiefer" box="[691,852,260,285]" pageId="80" pageNumber="81" refString="Kiefer, F. (1955) Neue Cyclopoida Gnathostoma (Crustacea Copepoda) aus Madagascar, II. Cyclopinae. Zoologischer Anzeiger, 154, 222 - 232." type="journal article" year="1955">Kiefer (1955)</bibRefCitation>
added 
<taxonomicName id="10AF9BD1811EFFB2FC3FFEB2FAC2FEDA" authority="Kiefer, 1955" authorityName="Kiefer" authorityYear="1955" box="[948,1376,260,285]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="E5DB3C40811EFFB2FC3FFEB2FB18FEDA" box="[948,1210,260,285]" italics="true" pageId="80" pageNumber="81">Haplocyclops pauliani</emphasis>
<bibRefCitation id="B33E9DA3811EFFB2FB42FEB2FAC2FEDA" author="Kiefer" box="[1225,1376,260,285]" pageId="80" pageNumber="81" refString="Kiefer, F. (1955) Neue Cyclopoida Gnathostoma (Crustacea Copepoda) aus Madagascar, II. Cyclopinae. Zoologischer Anzeiger, 154, 222 - 232." type="journal article" year="1955">Kiefer, 1955</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="10AF9BD1811EFFB2FF1CFE91FDBFFE87" authority="Kiefer, 1955" authorityName="Kiefer" authorityYear="1955" box="[151,541,295,320]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="neuter">
<emphasis id="E5DB3C40811EFFB2FF1CFE91FE20FE87" box="[151,386,295,320]" italics="true" pageId="80" pageNumber="81">Haplocyclops neuter</emphasis>
<bibRefCitation id="B33E9DA3811EFFB2FE00FE91FDBFFE87" author="Kiefer" box="[395,541,295,320]" pageId="80" pageNumber="81" refString="Kiefer, F. (1955) Neue Cyclopoida Gnathostoma (Crustacea Copepoda) aus Madagascar, II. Cyclopinae. Zoologischer Anzeiger, 154, 222 - 232." type="journal article" year="1955">Kiefer, 1955</bibRefCitation>
</taxonomicName>
, from the same region. The description of both these species was repeated by Kiefer himself (1956), but only with some additional drawings. 
<bibRefCitation id="B33E9DA3811EFFB2FC05FEFAFBE8FEA2" author="Lindberg" box="[910,1098,332,357]" pageId="80" pageNumber="81" refString="Lindberg, K. (1956) Les Cyclopides (Crustaces Copepodes) tres evolues en tant qu'habitants des eaux souterraines; Revue des travaux recents concernant les Bryocyclops Kiefer et Speocyclops Kiefer. Actes prem. Congres International de Speleologie, 1953, 71 - 83. [Paris]" type="journal article" year="1956">Lindberg (1956)</bibRefCitation>
relegated 
<taxonomicName id="10AF9BD1811EFFB2FB43FEFAFAC0FEA2" box="[1224,1378,332,357]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E5DB3C40811EFFB2FB43FEFAFAC0FEA2" box="[1224,1378,332,357]" italics="true" pageId="80" pageNumber="81">Haplocyclops</emphasis>
</taxonomicName>
to a species group in the genus 
<taxonomicName id="10AF9BD1811EFFB2FE50FED9FCA7FE4F" authority="Kiefer, 1927" authorityName="Kiefer" authorityYear="1927" box="[475,773,367,392]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E5DB3C40811EFFB2FE50FED9FDC4FE4F" box="[475,614,367,392]" italics="true" pageId="80" pageNumber="81">Bryocyclops</emphasis>
Kiefer, 1927
</taxonomicName>
. 
<bibRefCitation id="B33E9DA3811EFFB2FC9FFED9FC15FE4F" author="Kiefer" box="[788,951,367,392]" pageId="80" pageNumber="81" refString="Kiefer, F. (1960 a) Zwei neue Cyclopiden (Crust. Cop.) als Bewohner des Sandluckensystems im Niger (Franzosischer Sudan). Bulletin de l'Institut Francais d'Afrique Noire, 22 (A 2), 395 - 401." type="journal article" year="1960" yearSuffix="a">Kiefer (1960a</bibRefCitation>
, 
<bibRefCitation id="B33E9DA3811EFFB2FC4CFED9FC74FE4F" author="Kiefer" box="[967,982,367,392]" pageId="80" pageNumber="81" refString="Kiefer, F. (1960 b) Beitrage zur Copepodenkunde (XX). Zoologischer Anzeiger, 165, 37 - 45." type="journal article" year="1960" yearSuffix="b">b</bibRefCitation>
) did not completely accept Lindberg’s (1956) decision, but treated 
<taxonomicName id="10AF9BD1811EFFB2FE7FFE22FD2CFE6A" box="[500,654,404,429]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E5DB3C40811EFFB2FE7FFE22FD2CFE6A" box="[500,654,404,429]" italics="true" pageId="80" pageNumber="81">Haplocyclops</emphasis>
</taxonomicName>
as a subgenus of 
<taxonomicName id="10AF9BD1811EFFB2FCF4FE22FBABFE6A" box="[895,1033,404,429]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E5DB3C40811EFFB2FCF4FE22FBABFE6A" box="[895,1033,404,429]" italics="true" pageId="80" pageNumber="81">Bryocyclops</emphasis>
</taxonomicName>
, when he described 
<taxonomicName id="10AF9BD1811EFFB2FA9AFE22FE06FE17" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="monodi" subGenus="Haplocyclops">
<emphasis id="E5DB3C40811EFFB2FA9AFE22FA3EFE6A" box="[1297,1436,404,429]" italics="true" pageId="80" pageNumber="81">Bryocyclops</emphasis>
(
<emphasis id="E5DB3C40811EFFB2FF2BFE01FE98FE17" box="[160,314,439,464]" italics="true" pageId="80" pageNumber="81">Haplocyclops</emphasis>
) 
<emphasis id="E5DB3C40811EFFB2FEDBFE01FE06FE17" box="[336,420,439,464]" italics="true" pageId="80" pageNumber="81">monodi</emphasis>
</taxonomicName>
from the continental Africa. 
<taxonomicName id="10AF9BD1811EFFB2FC83FE01FC00FE17" box="[776,930,439,464]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E5DB3C40811EFFB2FC83FE01FC00FE17" box="[776,930,439,464]" italics="true" pageId="80" pageNumber="81">Haplocyclops</emphasis>
</taxonomicName>
was considered as subgenus of the genus 
<taxonomicName id="10AF9BD1811EFFB2FF1CFE6AFE80FE32" box="[151,290,476,501]" class="Maxillopoda" family="Cyclopidae" genus="Bryocyclops" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E5DB3C40811EFFB2FF1CFE6AFE80FE32" box="[151,290,476,501]" italics="true" pageId="80" pageNumber="81">Bryocyclops</emphasis>
</taxonomicName>
by several authors, e. g. 
<bibRefCitation id="B33E9DA3811EFFB2FDCEFE6BFD4AFE33" author="Dussart" box="[581,744,476,500]" pageId="80" pageNumber="81" refString="Dussart, B. H. (1981) Contribution a l'etude des Copepodes d'Iran. Crustaceana, 41, 162 - 166. http: // dx. doi. org / 10.1163 / 156854081 X 00200" type="journal article" year="1981">Dussart (1981</bibRefCitation>
, 
<bibRefCitation id="B33E9DA3811EFFB2FD7DFE6AFC90FE33" author="Dussart" box="[758,818,476,500]" pageId="80" pageNumber="81" refString="Dussart, B. H. (1982) Crustaces Copepodes des eaux interieures. Faune Madagascar, ORSTOM-CNRS, 58, 1 - 146. [Paris]" type="journal article" year="1982">1982</bibRefCitation>
), 
<bibRefCitation id="B33E9DA3811EFFB2FCCCFE6BFBCEFE32" author="Dussart" box="[839,1132,476,501]" pageId="80" pageNumber="81" refString="Dussart, B. H. &amp; Defaye, D. (1985) Repertoire mondial des Copepodes Cyclopoides. C. N. R. S. Centre Regional de Publications de Bordeaux, 1985, 1 - 236." type="journal article" year="1985">Dussart &amp; Defaye (1985)</bibRefCitation>
, 
<bibRefCitation id="B33E9DA3811EFFB2FBF1FE6AFA35FE32" author="Lescher-Moutoue" box="[1146,1431,476,501]" pageId="80" pageNumber="81" refString="Lescher-Moutoue. F. (1986) Copepoda Cyclopoida Cyclopidae des eaux douces souterraines continentales. In: Botosaneanu, L. (Ed.), Stygofauna Mundi. E. J. Brill / Backhuys, Leiden, pp. 299 - 312." type="book chapter" year="1986">Lescher-Moutoué (1986)</bibRefCitation>
, 
<bibRefCitation id="B33E9DA3811EFFB2FF1CFE49FDF1FDDF" box="[151,595,511,536]" pageId="80" pageNumber="81" refString="Rocha, C. E. F. da &amp; Carvalho Bjornbberg, M. H. G. (1987) Copepods of the Jureia Ecological Reserve, State of Sao Paulo, Brazil, II. The genera Hesperocyclops, Muscocyclops, and Bryocyclops (Copepoda, Cyclopidae). Hydrobiologia, 153, 97 - 107. http: // dx. doi. org / 10.1007 / BF 00006642" type="journal article">Rocha &amp; Carvalho Bjornbberg (1987)</bibRefCitation>
, and 
<bibRefCitation id="B33E9DA3811EFFB2FD1DFDB6FC8DFDDF" author="Pesce" box="[662,815,511,536]" pageId="80" pageNumber="81" refString="Pesce, G. L. (1996) Towards a revision of Cyclopinae copepods (Crustacea, Cyclopidae). Fragment a Entomologica Roma, 28, 189 - 200." type="journal article" year="1996">Pesce (1996)</bibRefCitation>
. The validity of 
<taxonomicName id="10AF9BD1811EFFB2FC7CFE49FB33FDDF" box="[1015,1169,511,536]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E5DB3C40811EFFB2FC7CFE49FB33FDDF" box="[1015,1169,511,536]" italics="true" pageId="80" pageNumber="81">Haplocyclops</emphasis>
</taxonomicName>
was accepted by 
<bibRefCitation id="B33E9DA3811EFFB2FAECFE49FF41FDFB" author="Reid" pageId="80" pageNumber="81" refString="Reid, J. W. (1999) New records of Bryocyclops from the continental U. S. A. Puerto Rico, and Brazil (Copepoda: Cyclopoida). Journal of Crustacean Biology, 19, 84 - 92. http: // dx. doi. org / 10.2307 / 1549550" type="journal article" year="1999">Reid (1999)</bibRefCitation>
, 
<bibRefCitation id="B33E9DA3811EFFB2FF7AFD93FDB0FDFB" author="Dussart" box="[241,530,548,573]" pageId="80" pageNumber="81" refString="Dussart, B. H. &amp; Defaye, D. (2001) Introduction to Copepoda. 2 nd Edition. Vol. 16. Guides to the Identification of the Microinvertebrates of the Continental Waters of the World, 344 pp." type="book" year="2001">Dussart &amp; Defaye (2001)</bibRefCitation>
and 
<bibRefCitation id="B33E9DA3811EFFB2FDC7FD92FD75FDFB" author="Fiers" box="[588,727,548,573]" pageId="80" pageNumber="81" refString="Fiers, F. (2002) The genus Haplocyclops Kiefer, 1952 (Copepoda, Cyclopoida, Cyclopidae): Redescription of the type-species, H. gudrunae, and its congeners. Hydrobiologia, 474, 155 - 169. http: // dx. doi. org / 10.1023 / A: 1016579619924" type="journal article" year="2002">Fiers (2002)</bibRefCitation>
after it was critically examined and redefined by 
<bibRefCitation id="B33E9DA3811EFFB2FA99FD92FF47FDA7" pageId="80" pageNumber="81" refString="Rocha, C. E. F. da, Torres, I. C. &amp; Maia-Barbosa, P. (1998) Haplocyclops torresi n. sp. and Potamocaris estevesi Reid, 1991 from Brazil, with a proposal for revalidation of the genus Haplocyclops Kiefer, 1952 (Copepoda). Beaufortia, 48, 1 - 15." type="journal article">
Rocha 
<emphasis id="E5DB3C40811EFFB2FAE9FD93FA37FDFA" box="[1378,1429,548,573]" italics="true" pageId="80" pageNumber="81">et al</emphasis>
. (1998)
</bibRefCitation>
. Finally, six species have so far been recognized under 
<taxonomicName id="10AF9BD1811EFFB2FCDBFDF1FC48FDA7" box="[848,1002,583,608]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E5DB3C40811EFFB2FCDBFDF1FC48FDA7" box="[848,1002,583,608]" italics="true" pageId="80" pageNumber="81">Haplocyclops</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="B33E9DA3811EFFB2FBA7FDF1FB17FDA7" author="Kiefer" box="[1068,1205,583,608]" pageId="80" pageNumber="81" refString="Kiefer, F. (1955) Neue Cyclopoida Gnathostoma (Crustacea Copepoda) aus Madagascar, II. Cyclopinae. Zoologischer Anzeiger, 154, 222 - 232." type="journal article" year="1955">Kiefer 1955</bibRefCitation>
, 
<bibRefCitation id="B33E9DA3811EFFB2FB4AFDF1FAA8FDA7" author="Kiefer" box="[1217,1290,583,608]" pageId="80" pageNumber="81" refString="Kiefer, F. (1960 b) Beitrage zur Copepodenkunde (XX). Zoologischer Anzeiger, 165, 37 - 45." type="journal article" year="1960" yearSuffix="b">1960b</bibRefCitation>
; 
<bibRefCitation id="B33E9DA3811EFFB2FA9DFDF1FF72FD43" author="Rocha" pageId="80" pageNumber="81" refString="Rocha, C. E. F. da, Torres, I. C. &amp; Maia-Barbosa, P. (1998) Haplocyclops torresi n. sp. and Potamocaris estevesi Reid, 1991 from Brazil, with a proposal for revalidation of the genus Haplocyclops Kiefer, 1952 (Copepoda). Beaufortia, 48, 1 - 15." type="journal article" year="1998">
Rocha 
<emphasis id="E5DB3C40811EFFB2FAEFFDFFFA3EFDA7" box="[1380,1436,583,608]" italics="true" pageId="80" pageNumber="81">et al.</emphasis>
1998
</bibRefCitation>
) including the Indian 
<taxonomicName id="10AF9BD1811EFFB2FE43FDDBFC77FD42" authority="Karanovic &amp; Ranga Reddy, 2005" authorityName="Karanovic &amp; Ranga Reddy" authorityYear="2005" box="[456,981,620,645]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="fiersi" subGenus="Kiefercyclops">
<emphasis id="E5DB3C40811EFFB2FE43FDDBFDF5FD43" box="[456,599,620,644]" italics="true" pageId="80" pageNumber="81">H. (K.) fiersi</emphasis>
<bibRefCitation id="B33E9DA3811EFFB2FDD5FDDAFC77FD42" author="Karanovic" box="[606,981,620,645]" pageId="80" pageNumber="81" refString="Karanovic, T. &amp; Ranga Reddy, Y. (2005) First Haplocyclops Kiefer (Crustacea, Copepoda) from Indian subterranean waters: the most reduced free-living cyclopoid. Annales de Limnologie, 41, 83 - 92. http: // dx. doi. org / 10.1051 / limn / 2005011" type="journal article" year="2005">Karanovic &amp; Ranga Reddy, 2005</bibRefCitation>
</taxonomicName>
, which is, incidentally, the most reduced free-living cyclopoid (see 
<bibRefCitation id="B33E9DA3811EFFB2FE36FD39FC95FD6F" author="Karanovic" box="[445,823,655,680]" pageId="80" pageNumber="81" refString="Karanovic, T. &amp; Ranga Reddy, Y. (2005) First Haplocyclops Kiefer (Crustacea, Copepoda) from Indian subterranean waters: the most reduced free-living cyclopoid. Annales de Limnologie, 41, 83 - 92. http: // dx. doi. org / 10.1051 / limn / 2005011" type="journal article" year="2005">Karanovic &amp; Ranga Reddy, 2005</bibRefCitation>
).
</paragraph>
<paragraph id="D710E052811EFFB2FF4CFD03FF49FC2A" blockId="80.[151,1437,151,1616]" pageId="80" pageNumber="81">
Now, the rationale behind assigning 
<taxonomicName id="10AF9BD1811EFFB2FDF8FD03FCA9FD0B" box="[627,779,693,716]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="primitivus" status="sp. nov.">
<emphasis id="E5DB3C40811EFFB2FDF8FD03FCA9FD0B" box="[627,779,693,716]" italics="true" pageId="80" pageNumber="81">H. primitivus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FEE8813B811EFFB2FC9EFD02FCF5FD0A" box="[789,855,692,717]" pageId="80" pageNumber="81" rank="species">
<emphasis id="E5DB3C40811EFFB2FC9EFD02FCF5FD0A" bold="true" box="[789,855,692,717]" pageId="80" pageNumber="81">n. sp.</emphasis>
</taxonomicNameLabel>
to 
<taxonomicName id="10AF9BD1811EFFB2FC08FD02FBBFFD0A" box="[899,1053,692,717]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E5DB3C40811EFFB2FC08FD02FBBFFD0A" box="[899,1053,692,717]" italics="true" pageId="80" pageNumber="81">Haplocyclops</emphasis>
</taxonomicName>
needs to be explained. The new species has the following characters, which are considered by 
<bibRefCitation id="B33E9DA3811EFFB2FCEEFD61FB53FD37" box="[869,1265,727,752]" pageId="80" pageNumber="81" refString="Karanovic, T. &amp; Ranga Reddy, Y. (2005) First Haplocyclops Kiefer (Crustacea, Copepoda) from Indian subterranean waters: the most reduced free-living cyclopoid. Annales de Limnologie, 41, 83 - 92. http: // dx. doi. org / 10.1051 / limn / 2005011" type="journal article">Karanovic &amp; Ranga Reddy (2005)</bibRefCitation>
the significant ‘autapomorphies’ of 
<taxonomicName id="10AF9BD1811EFFB2FE0EFD4AFDBDFCD2" box="[389,543,764,789]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E5DB3C40811EFFB2FE0EFD4AFDBDFCD2" box="[389,543,764,789]" italics="true" pageId="80" pageNumber="81">Haplocyclops</emphasis>
</taxonomicName>
: the genital double-somite with ovipores situated in the posterior half and the caudal rami with lateral seta inserted in the proximal half. The other supportive characters include: large anal operculum; principal apical setae without breaking planes; seventh antennular segment armed with 2 setae; fifth legs completely fused to somite and represented by 3 slender setae in both sexes. It must, however, be clarified that 
<taxonomicName id="10AF9BD1811EFFB2FF1CFC3BFE90FC63" box="[151,306,909,932]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="primitivus" status="sp. nov.">
<emphasis id="E5DB3C40811EFFB2FF1CFC3BFE90FC63" box="[151,306,909,932]" italics="true" pageId="80" pageNumber="81">H. primitivus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FEE8813B811EFFB2FEB4FC3AFE27FC62" box="[319,389,908,933]" pageId="80" pageNumber="81" rank="species">
<emphasis id="E5DB3C40811EFFB2FEB4FC3AFE27FC62" bold="true" box="[319,389,908,933]" pageId="80" pageNumber="81">n. sp.</emphasis>
</taxonomicNameLabel>
has at least two interesting plesiomorphic character states that are as yet unknown in 
<taxonomicName id="10AF9BD1811EFFB2FF1CFC19FE93FC0F" box="[151,305,943,968]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E5DB3C40811EFFB2FF1CFC19FE93FC0F" box="[151,305,943,968]" italics="true" pageId="80" pageNumber="81">Haplocyclops</emphasis>
</taxonomicName>
: both rami of legs 1–4 are 2-segmented, and the second exopodal segment has a setal formula of 5.4.4.4.
</paragraph>
<paragraph id="D710E052811EFFB2FF5BFC41FED1FABF" blockId="80.[151,1437,151,1616]" pageId="80" pageNumber="81">
<bibRefCitation id="B33E9DA3811EFFB2FF5BFC41FDFBFBD7" box="[208,601,1015,1040]" pageId="80" pageNumber="81" refString="Karanovic, T. &amp; Ranga Reddy, Y. (2005) First Haplocyclops Kiefer (Crustacea, Copepoda) from Indian subterranean waters: the most reduced free-living cyclopoid. Annales de Limnologie, 41, 83 - 92. http: // dx. doi. org / 10.1051 / limn / 2005011" type="journal article">Karanovic &amp; Ranga Reddy (2005)</bibRefCitation>
subdivided the genus 
<taxonomicName id="10AF9BD1811EFFB2FCD0FC41FC57FBD7" box="[859,1013,1015,1040]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="E5DB3C40811EFFB2FCD0FC41FC57FBD7" box="[859,1013,1015,1040]" italics="true" pageId="80" pageNumber="81">Haplocyclops</emphasis>
</taxonomicName>
into two subgenera: 
<emphasis id="E5DB3C40811EFFB2FB6CFC41FF11FBF3" italics="true" pageId="80" pageNumber="81">
<taxonomicName id="10AF9BD1811EFFB2FB6CFC41FA23FBD7" box="[1255,1409,1015,1040]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus" sensu="stricto">Haplocyclops</taxonomicName>
<taxonomicNameLabel id="FEE8813B811EFFB2FA01FC4FFF11FBF3" pageId="80" pageNumber="81" sensu="stricto">s. str</taxonomicNameLabel>
</emphasis>
. 
<bibRefCitation id="B33E9DA3811EFFB2FF48FBAAFEF3FBF2" author="Kiefer" box="[195,337,1052,1077]" pageId="80" pageNumber="81" refString="Kiefer, F. (1952) Haplocyclops Gudrunae n. g. et n. sp. ein neuer Ruderfuβkrebs (Crustacea Copepoda) aus Madagascar. Zoologischer Anzeiger, 149, 240 - 243." type="journal article" year="1952">Kiefer, 1952</bibRefCitation>
and 
<emphasis id="E5DB3C40811EFFB2FE00FBAAFD86FBF2" box="[395,548,1052,1077]" italics="true" pageId="80" pageNumber="81">Kiefercyclops</emphasis>
<bibRefCitation id="B33E9DA3811EFFB2FDA7FBAAFC09FBF2" author="Karanovic" box="[556,939,1052,1077]" pageId="80" pageNumber="81" refString="Karanovic, T. &amp; Ranga Reddy, Y. (2005) First Haplocyclops Kiefer (Crustacea, Copepoda) from Indian subterranean waters: the most reduced free-living cyclopoid. Annales de Limnologie, 41, 83 - 92. http: // dx. doi. org / 10.1051 / limn / 2005011" type="journal article" year="2005">Karanovic &amp; Ranga Reddy, 2005</bibRefCitation>
. The former contains seven species, mostly from the Gondwanan landmasses of 
<collectingCountry id="AFB8A0C2811EFFB2FDC0FB89FD76FB9F" box="[587,724,1087,1112]" name="Madagascar" pageId="80" pageNumber="81">Madagascar</collectingCountry>
, Africa, and South 
<collectingCountry id="AFB8A0C2811EFFB2FC4AFB89FB80FB9F" box="[961,1058,1087,1112]" name="United States of America" pageId="80" pageNumber="81">America</collectingCountry>
whereas the latter heretofore is monotypic with the Indian 
<taxonomicName id="10AF9BD1811EFFB2FE4FFBD3FDF0FBBB" box="[452,594,1124,1148]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="fiersi" subGenus="Kiefercyclops">
<emphasis id="E5DB3C40811EFFB2FE4FFBD3FDF0FBBB" box="[452,594,1124,1148]" italics="true" pageId="80" pageNumber="81">H. (K.) fiersi</emphasis>
</taxonomicName>
. Both 
<taxonomicName id="10AF9BD1811EFFB2FD16FBD3FCC6FBBB" box="[669,868,1124,1148]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="primitivus" status="sp. nov." subGenus="Kiefercyclops">
<emphasis id="E5DB3C40811EFFB2FD16FBD3FCC6FBBB" box="[669,868,1124,1148]" italics="true" pageId="80" pageNumber="81">H. (K.) primitivus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FEE8813B811EFFB2FCE1FBD2FC0BFBBB" box="[874,937,1124,1149]" pageId="80" pageNumber="81" rank="species">
<emphasis id="E5DB3C40811EFFB2FCE1FBD2FC00FBBA" bold="true" box="[874,930,1124,1149]" pageId="80" pageNumber="81">n. sp</emphasis>
.
</taxonomicNameLabel>
and 
<taxonomicName id="10AF9BD1811EFFB2FC55FBD3FB3AFBBA" box="[990,1176,1124,1149]" class="Maxillopoda" family="Parastenocarididae" genus="Kinnecaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="godavari" status="sp. nov." subGenus="Kiefercyclops">
<emphasis id="E5DB3C40811EFFB2FC55FBD3FB3AFBBA" box="[990,1176,1124,1149]" italics="true" pageId="80" pageNumber="81">H. (K.) godavari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FEE8813B811EFFB2FB14FBD2FB7FFBBB" box="[1183,1245,1124,1149]" pageId="80" pageNumber="81" rank="species">
<emphasis id="E5DB3C40811EFFB2FB14FBD2FB75FBBA" bold="true" box="[1183,1239,1124,1149]" pageId="80" pageNumber="81">n. sp</emphasis>
.
</taxonomicNameLabel>
described herein belong to the latter subgenus mainly because the mandibular palp is completely absent and the legs 1–4 have no coxal armature. 
<taxonomicName id="10AF9BD1811EFFB2FEC0FB1BFE7DFB03" box="[331,479,1197,1220]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="primitivus" status="sp. nov.">
<emphasis id="E5DB3C40811EFFB2FEC0FB1BFE7DFB03" box="[331,479,1197,1220]" italics="true" pageId="80" pageNumber="81">H. primitivus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FEE8813B811EFFB2FE6EFB1AFD81FB03" box="[485,547,1196,1221]" pageId="80" pageNumber="81" rank="species">
<emphasis id="E5DB3C40811EFFB2FE6EFB1AFDBFFB02" bold="true" box="[485,541,1196,1221]" pageId="80" pageNumber="81">n. sp</emphasis>
.
</taxonomicNameLabel>
has a set of primitive characters, and is thus at variance with 
<emphasis id="E5DB3C40811EFFB2FB45FB1BFB49FB03" box="[1230,1259,1197,1220]" italics="true" pageId="80" pageNumber="81">H.</emphasis>
(
<emphasis id="E5DB3C40811EFFB2FB71FB1AFA3EFB02" box="[1274,1436,1196,1221]" italics="true" pageId="80" pageNumber="81">Kiefercyclops)</emphasis>
diagnosis in the following features: coxobasis of antenna with 2 setae vs. 1 seta; maxillulary palp 2-segmented vs. 1-segmented; maxilla with vs. without lateral small seta on basis; leg 1 first exopodal segment armed vs. unarmed; exopod of leg 3, as well as endopod of legs 2 and 3, 2-segmented vs. 1-segmented; and leg 6 with 2 setae vs. 1 seta. Despite these differences, we think it advisable to allocate 
<taxonomicName id="10AF9BD1811EFFB2FCC0FA8BFC41FA93" box="[843,995,1341,1364]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="primitivus" status="sp. nov.">
<emphasis id="E5DB3C40811EFFB2FCC0FA8BFC41FA93" box="[843,995,1341,1364]" italics="true" pageId="80" pageNumber="81">H. primitivus</emphasis>
</taxonomicName>
<emphasis id="E5DB3C40811EFFB2FC66FA8AFB8DFA92" bold="true" box="[1005,1071,1340,1365]" pageId="80" pageNumber="81">
<taxonomicNameLabel id="FEE8813B811EFFB2FC66FA8AFB8DFA92" box="[1005,1071,1340,1365]" pageId="80" pageNumber="81" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
to 
<emphasis id="E5DB3C40811EFFB2FBD0FA8AFB56FA92" box="[1115,1268,1340,1365]" italics="true" pageId="80" pageNumber="81">Kiefercyclops</emphasis>
rather than to 
<emphasis id="E5DB3C40811EFFB2FF1CFAE9FED1FABF" box="[151,371,1375,1400]" italics="true" pageId="80" pageNumber="81">
<taxonomicName id="10AF9BD1811EFFB2FF1CFAE9FE93FABF" box="[151,305,1375,1400]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="genus" sensu="stricto">Haplocyclops</taxonomicName>
<taxonomicNameLabel id="FEE8813B811EFFB2FEB3FAD7FED1FABF" box="[312,371,1377,1400]" pageId="80" pageNumber="81" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="D710E052811EFFB2FF4CFA33FDD1F997" blockId="80.[151,1437,151,1616]" pageId="80" pageNumber="81">
<taxonomicName id="10AF9BD1811EFFB2FF4CFA33FEFDFA5B" box="[199,351,1413,1436]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="primitivus" status="sp. nov.">
<emphasis id="E5DB3C40811EFFB2FF4CFA33FEFDFA5B" box="[199,351,1413,1436]" italics="true" pageId="80" pageNumber="81">H. primitivus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FEE8813B811EFFB2FEE1FA32FE0FFA5A" box="[362,429,1412,1437]" pageId="80" pageNumber="81" rank="species">
<emphasis id="E5DB3C40811EFFB2FEE1FA32FE0FFA5A" bold="true" box="[362,429,1412,1437]" pageId="80" pageNumber="81">n. sp.</emphasis>
</taxonomicNameLabel>
can be readily separated from both 
<taxonomicName id="10AF9BD1811EFFB2FCD5FA33FC54FA5B" box="[862,1014,1412,1436]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="fiersi" subGenus="Kiefercyclops">
<emphasis id="E5DB3C40811EFFB2FCD5FA33FC54FA5B" box="[862,1014,1412,1436]" italics="true" pageId="80" pageNumber="81">H. (K.) fiersi</emphasis>
</taxonomicName>
and 
<taxonomicName id="10AF9BD1811EFFB2FBBEFA33FB5AFA5A" box="[1077,1272,1412,1437]" class="Maxillopoda" family="Parastenocarididae" genus="Kinnecaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="godavari" status="sp. nov." subGenus="Kiefercyclops">
<emphasis id="E5DB3C40811EFFB2FBBEFA33FB5AFA5A" box="[1077,1272,1412,1437]" italics="true" pageId="80" pageNumber="81">H. (K.) godavari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FEE8813B811EFFB2FA8FFA32FAE5FA5B" box="[1284,1351,1412,1437]" pageId="80" pageNumber="81" rank="species">
<emphasis id="E5DB3C40811EFFB2FA8FFA32FAE2FA5A" bold="true" box="[1284,1344,1412,1437]" pageId="80" pageNumber="81">n. sp</emphasis>
.
</taxonomicNameLabel>
by the following features: cyclopiform habitus; dorsal caudal seta 1.9 times as long as ramus length; presence of a row of transverse row of fine spinules at base of dorsal seta and lateral seta; lateral seta present on ultimate segment of antennule; leg 1 without spine on basis; legs 1–4 with 2-segmented exopod and endopod; and leg 4 second endopodal segment armed with one spine and one seta (the 1-segmented endopod in 
<taxonomicName id="10AF9BD1811EFFB2FBFCF9A3FAACF9EB" box="[1143,1294,1556,1580]" class="Maxillopoda" family="Cyclopidae" genus="Haplocyclops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="fiersi" subGenus="Kiefercyclops">
<emphasis id="E5DB3C40811EFFB2FBFCF9A3FAACF9EB" box="[1143,1294,1556,1580]" italics="true" pageId="80" pageNumber="81">H. (K.) fiersi</emphasis>
</taxonomicName>
and 
<taxonomicName id="10AF9BD1811EFFB2FAC1F9A3FF5EF997" class="Maxillopoda" family="Parastenocarididae" genus="Kinnecaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="godavari" status="sp. nov." subGenus="Kiefercyclops">
<emphasis id="E5DB3C40811EFFB2FAC1F9A3FF5EF997" italics="true" pageId="80" pageNumber="81">H. (K.) godavari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FEE8813B811EFFB2FE88F981FEE0F988" box="[259,322,1591,1616]" pageId="80" pageNumber="81" rank="species">
<emphasis id="E5DB3C40811EFFB2FE88F981FE99F997" bold="true" box="[259,315,1591,1616]" pageId="80" pageNumber="81">n. sp</emphasis>
.
</taxonomicNameLabel>
also has similar armature).
</paragraph>
</subSubSection>
</treatment>
</document>